taxonID	type	description	language	source
03DCBC11C25BFFBD0ED21862FD9A60C9.taxon	diagnosis	Diagnosis. Shell small (to 0.54 – 3.1 mm); without nacre; usually with slit or foramen (absent in Coronadoa); slit well above periphery; radula with serrated rachidian tooth, lateral teeth 1 – 3 similar, lateral tooth 4 reduced, hook-shaped, lateral tooth 5 enlarged by broadening, marginal teeth spoon shaped. Operculum thin, round, multispiral, with central nucleus.	en	Geiger, Daniel L., Mclean, James H. (2010): New species and records of Scissurellidae and Anatomidae from the Americas (Mollusca: Gastropoda: Vetigastropoda). Zootaxa 2356: 1-35, DOI: 10.5281/zenodo.275645
03DCBC11C25AFFBC0ED21FD5FE9C66EE.taxon	materials_examined	Type species. Schismope brevis Hedley, 1904 (OD).	en	Geiger, Daniel L., Mclean, James H. (2010): New species and records of Scissurellidae and Anatomidae from the Americas (Mollusca: Gastropoda: Vetigastropoda). Zootaxa 2356: 1-35, DOI: 10.5281/zenodo.275645
03DCBC11C25AFFBC0ED21FD5FE9C66EE.taxon	diagnosis	Diagnosis. Scissurellidae with slit closed to foramen at maturity, keels of foramen not elevated to form chimney.	en	Geiger, Daniel L., Mclean, James H. (2010): New species and records of Scissurellidae and Anatomidae from the Americas (Mollusca: Gastropoda: Vetigastropoda). Zootaxa 2356: 1-35, DOI: 10.5281/zenodo.275645
03DCBC11C25AFFBC0ED21FD5FE9C66EE.taxon	discussion	Remarks. In juveniles, the selenizone may be shorter and the slit is still not closed to a foramen. Juvenile Sinezona and adult Scissurella both have an open slit; mature specimens can be recognized by the descending last quarter whorl.	en	Geiger, Daniel L., Mclean, James H. (2010): New species and records of Scissurellidae and Anatomidae from the Americas (Mollusca: Gastropoda: Vetigastropoda). Zootaxa 2356: 1-35, DOI: 10.5281/zenodo.275645
03DCBC11C25AFFB90ED2199DFED165AD.taxon	materials_examined	Type material. Holotype (SBMNH 83542). Paratypes: from type locality (SBMNH 83543, 5); 0 m, NE coast, Oahu, Hawaii, USA, 21.620 ˚ N, 157.910 ˚ W (UF 350462, 1); 0 m, off Kala Place, Diamond Head, Oahu, Hawaii, USA, 21.250 ˚ N, 157.800 ˚ W (SBMNH 351023, 5: formalin fixed and ethanol preserved; SBMNH 351024, 4: formalin fixed and dried. SBMNH 351025, 1: ethanol fixed and preserved); Hakipu’u Reef, Kaneohe Bay, Oahu, Hawaii, USA (BPBM 261475, 1: complete). Moku Manu Island, Kaneohe Bay, Oahu, Hawaii, USA (BPBM 263630, 1: complete). Type locality. 0 m, Haleiwa, Oahu, Hawaii, USA, 21.590 ˚ N, 158.113 ˚ W.	en	Geiger, Daniel L., Mclean, James H. (2010): New species and records of Scissurellidae and Anatomidae from the Americas (Mollusca: Gastropoda: Vetigastropoda). Zootaxa 2356: 1-35, DOI: 10.5281/zenodo.275645
03DCBC11C25AFFB90ED2199DFED165AD.taxon	etymology	Etymology. Named in honor of the late Alison E. Kay of the University of Hawaii for her many accomplishments with respect to the Hawaiian malacofauna.	en	Geiger, Daniel L., Mclean, James H. (2010): New species and records of Scissurellidae and Anatomidae from the Americas (Mollusca: Gastropoda: Vetigastropoda). Zootaxa 2356: 1-35, DOI: 10.5281/zenodo.275645
03DCBC11C25AFFB90ED2199DFED165AD.taxon	description	Description. Shell small (to 0.65 mm), depressed ovoid. Protoconch 0.75 whorls, with 19 – 20 strong axials, no apertural varix, apertural margin slightly sinusoid. TI of 0.75 whorls, only axial sculpture, approximately 13 fine, distinct ribs, interstices with fine irregular growth lines. TII slightly> 0.5 whorl, on shoulder, axial sculpture slightly diminished in strength towards selenizone / foramen, approximately five indistinct axials appear towards aperture; base with fine, distinct axials only, base forming angle with walls of umbilicus. Selenizone approximately 0.25 whorl, slit closed to form foramen. Aperture ovoid, roof strongly protracted, adumbilical margin straight. Umbilicus open, quite wide, border to base with spiral cord (funiculus). Operculum thin, round, multispiral, with central nucleus. Radula (Fig. 3 A – B). Rachidian triangular, cusp with 5 denticles, central one largest. Lateral teeth 3 – 5 similar, 3 – 4 denticles on outer margin. Lateral tooth 4 reduced, pointed. Lateral tooth 5 broadened with approximately 6 – 8 denticles on inner margin. Radular interlock of central field moderate. Inner marginal teeth with two small denticles on inner margin of cusp, terminal denticle largest, approximately five denticles on outer margin. Outer marginal teeth spoon shaped, many fine bristles on each side of cusp. Differential diagnosis. Sinezona kayae has more distinct and fewer axials and is more depressed than Sin. insignis from South Africa. Sinezona costulata Geiger & Sasaki, 2009, from temperate Japan is overall wider, lacks the strong spiral cord bordering the umbilicus, and the axial lamellae on the base do not reach the selenizone. Sinezona danieldreieri Geiger, 2008, from the Indo-Pacific has an overall broader shell, more and finer axial cords on the teleoconch, axial cords on the shoulder reaching from the suture to the selenizone, and a protoconch with fewer (9 – 15 vs. 19 – 20) broader strong axial cords.	en	Geiger, Daniel L., Mclean, James H. (2010): New species and records of Scissurellidae and Anatomidae from the Americas (Mollusca: Gastropoda: Vetigastropoda). Zootaxa 2356: 1-35, DOI: 10.5281/zenodo.275645
03DCBC11C25AFFB90ED2199DFED165AD.taxon	distribution	Distribution. Hawaii, USA. Specimen records. 30 m, off Waikiki, Oahu, 21.270 ˚ N, 157.840 ˚ W (LACM 75 - 96, 1). 30 m, Koloa, Kauai, 21.883 ˚ N, 159.475 ˚ W (LACM 75 - 97, 1). 0 m, off Kala Place, Diamond Head, Oahu. 15 / X / 2001 (DLG 223, 12 formalin fixed; DLG 231, 11 ethanol fixed). 0 m, Ka’awa, Oahu, Hawaii, USA, 21.555 ˚ N, 157.851 ˚ W (DLG 473, 2). 0 m, Kona Village Resort, Big Island, Hawaii, USA, 19.831 ˚ N, 155.988 ˚ W (DLG 485, 3: complete). 0 m, Kala, Oahu, Hawaii, USA, 21.261 ˚ N, 157.787 ˚ W (DLG 495, 7: complete). 0 m, Kona Village Resort, Big Island, Hawaii, USA, 19.831 ˚ N, 155.988 ˚ W (DLG 751, 6). 0 m, off Kala Place, Diamond Head, Oahu, Hawaii, USA (DLG 752, 1). 0 m, off Kala Place, Diamond Head, Oahu, Hawaii, USA (DLG 753, 4). 0 m, Makai Pier, Makapu Beach, Oahu, Hawaii, USA (UF 350463, 1). 10 m, Maile, Oahu, Hawaii, USA (UF 350460, 1). 60 m, Kepuhi Point, Oahu, Hawaii, USA (UF 350461, 2).	en	Geiger, Daniel L., Mclean, James H. (2010): New species and records of Scissurellidae and Anatomidae from the Americas (Mollusca: Gastropoda: Vetigastropoda). Zootaxa 2356: 1-35, DOI: 10.5281/zenodo.275645
03DCBC11C25AFFB90ED2199DFED165AD.taxon	discussion	Remarks. Herbert (1986) included Kay’s (1979) listing and illustrations of this new species in the synonymy of the South African species, although noting that (p. 628) “ the identity of her specimens requires confirmation as the occurrence of S. insignis, an essentially warm temperate species, in the tropical Pacific is surprising. ”	en	Geiger, Daniel L., Mclean, James H. (2010): New species and records of Scissurellidae and Anatomidae from the Americas (Mollusca: Gastropoda: Vetigastropoda). Zootaxa 2356: 1-35, DOI: 10.5281/zenodo.275645
03DCBC11C25EFFB80ED21FD5FD7C63D0.taxon	materials_examined	Type material. Holotype (LACM 2466), leg. Twila Bratcher. Type locality. 30 m, off Waikiki, Oahu, HI, USA, 21.270 ˚ N, 157.840 ˚ W.	en	Geiger, Daniel L., Mclean, James H. (2010): New species and records of Scissurellidae and Anatomidae from the Americas (Mollusca: Gastropoda: Vetigastropoda). Zootaxa 2356: 1-35, DOI: 10.5281/zenodo.275645
03DCBC11C25EFFB80ED21FD5FD7C63D0.taxon	etymology	Etymology. Named after its provenance from Hawaii, USA.	en	Geiger, Daniel L., Mclean, James H. (2010): New species and records of Scissurellidae and Anatomidae from the Americas (Mollusca: Gastropoda: Vetigastropoda). Zootaxa 2356: 1-35, DOI: 10.5281/zenodo.275645
03DCBC11C25EFFB80ED21FD5FD7C63D0.taxon	description	Description. Shell small (0.5 mm), trochiform, globular. Protoconch of 1 whorl, with fine axial cords, no apertural varix. apertural margin prosocline. TI of 1.25 whorls. Shoulder with broad indistinct axial cords, approximately 30 on first whorl, without discernible spiral sculpture. Base with axials becoming more distinct towards umbilicus, approximately 13 low spiral cords between periphery and umbilicus, barely noticeable at periphery, becoming as strong as axials towards umbilicus. TII of 0.125 whorls, same sculpture as on TI. Umbilicus very narrow, bordered by strong cord, walls with distinct growth marks, no funiculus. Selenizone absent, foramen oval, keels low, strong. Aperture rounded, roof overhanging. Animal unknown. Differential diagnosis. Sinezona haliotimorpha (Bandel, 1998) from the Oligocene of France has a protoconch with fewer and stronger axials, and a teleococh with stronger axials and consistently finer spiral threads on the base, as well as a foramen with more elevated keels. Sinezona brevis Hedley, 1904 [= Ariella campbelli Bandel, 1998] from New Zealand has a protoconch like S. haliotimorpha, has a greater expansion rate of the shell giving it a more oval appearance, and when fully grown has a short selenizone.	en	Geiger, Daniel L., Mclean, James H. (2010): New species and records of Scissurellidae and Anatomidae from the Americas (Mollusca: Gastropoda: Vetigastropoda). Zootaxa 2356: 1-35, DOI: 10.5281/zenodo.275645
03DCBC11C25EFFB80ED21FD5FD7C63D0.taxon	distribution	Distribution. Known only from type locality.	en	Geiger, Daniel L., Mclean, James H. (2010): New species and records of Scissurellidae and Anatomidae from the Americas (Mollusca: Gastropoda: Vetigastropoda). Zootaxa 2356: 1-35, DOI: 10.5281/zenodo.275645
03DCBC11C25EFFB80ED21FD5FD7C63D0.taxon	discussion	Remarks. The absence of a selenizone has been taken as genus-level character by Bandel (1998) to establish Ariella Bandel, 1998. Marshall (2002), however, pointed out that there is significant intraspecific variability in the length of selenizone, with some species [e. g., Sinezona brevis (Hedley, 1904)] showing a distinct to no selenizone at all. Accordingly, the absence of a selenizone is not considered significant for generic classification. Equally, the presence and absence of a protoconch varix and details of teleoconch ornamentation vary extensively between species in the same genus and is also insignificant for beta taxonomy (Geiger 2003). The shell appears mature due to the descending apertural margin on the last portion of the whorl. The species is known from a single, but well-preserved specimen. The difference in protoconch sculpture, and the absence of any sign of a selenizone despite the sloping aperture indicating maturity, refutes the possibility of a mislocalized specimen from New Zealand.	en	Geiger, Daniel L., Mclean, James H. (2010): New species and records of Scissurellidae and Anatomidae from the Americas (Mollusca: Gastropoda: Vetigastropoda). Zootaxa 2356: 1-35, DOI: 10.5281/zenodo.275645
03DCBC11C25EFFB50ED21A78FB7960E5.taxon	description	Sinezona n. sp. Geiger, ms: Kaiser, 2009: 221, 228, fig. 237 I – K [paratype Fig. 7 B].	en	Geiger, Daniel L., Mclean, James H. (2010): New species and records of Scissurellidae and Anatomidae from the Americas (Mollusca: Gastropoda: Vetigastropoda). Zootaxa 2356: 1-35, DOI: 10.5281/zenodo.275645
03DCBC11C25EFFB50ED21A78FB7960E5.taxon	materials_examined	Type material. Holotype (SBMNH 83703). Paratypes: type locality (SBMNH 83704, 2); 16 – 23 m, Isla Maria Cleofa, Islas Tres Marias, Mexico, 21.300 ˚ N, 106.276 ˚ W (SBMNH 83705, 4, mounted radula); 23 – 30 m, Sharkfin Rock, SSW Punta Rodriguez, Isla del Coco, Costa Rica, 5.533 ˚ N, 86.983 ˚ W (SBMNH 83706, 3); 12 – 15 m, Clipperton Island, France, 10.305 ˚ N, 109.198 ˚ W (KKC 210001, 8); 12 – 15 m, 10.305 ˚ N, 109.198 ˚ W (LACM 3116, 5; MNHN 21265, 5; SDMNH, 5; WAM S 41348, 5; USNM 1117767, 5; all ex KKC 200518). 10 – 38 m, Clipperton Island, France, 10.320 ˚ N, 109.223 ˚ W (SBMNH 83707, ex KKC 210111, 2). Type locality. 15 – 30 m, East side, Clipperton Island, France, 10.295 ˚ N, 109.201 ˚ W, leg. Kirstie Kaiser. Additional material examined. France, Clipperton Island. 12 – 15 m, 10.305 ˚ N, 109.198 ˚ W (KKC 200518, 75). 11 – 22 m, 10.316 ˚ N, 109.214 ˚ W (KKC 210048, 5). 17 m, NW side, 10.322 ˚ N, 109.223 ˚ W (KKC 210108, 1). 15 – 30 m, East side, 10.295 ˚ N, 109.201 ˚ W (KKC 200516, 51). 12 – 43 m, 10.291 ˚ N, 109.225 ˚ W (KKC 200517, 80). 10 – 38 m, 10.320 ˚ N, 109.223 ˚ W (KKC 210047, 8). 15 – 20 m, 10.314 ˚ N, 109.238 ˚ W (KKC 210000, 12). 69 m, 10.320 ˚ N, 109.235 ˚ W (KKC 200515, 47). 15 m, 10.288 ˚ N, 109.350 ˚ W (KKC 200519, 40). Costa Rica. 23 – 30 m, Sharkfin Rock, SSW Punta Rodriguez, Isla del Coco, 5.533 ˚ N, 86.983 ˚ W (KKC, 5). Mexico. 16 – 23 m, Isla Maria Cleofa, Islas Tres Marias, 21.300 ˚ N, 106.276 ˚ W (KKC, 7).	en	Geiger, Daniel L., Mclean, James H. (2010): New species and records of Scissurellidae and Anatomidae from the Americas (Mollusca: Gastropoda: Vetigastropoda). Zootaxa 2356: 1-35, DOI: 10.5281/zenodo.275645
03DCBC11C25EFFB50ED21A78FB7960E5.taxon	etymology	Etymology. Named after Carole Hertz, editor of The Festivus, and Carol Skoglund, long time collector and student of the Panamic molluscan fauna, for their significant body of work and often collaborative efforts to better understand the taxonomy of the Panamic fauna. Genitive feminine plural of their common first name.	en	Geiger, Daniel L., Mclean, James H. (2010): New species and records of Scissurellidae and Anatomidae from the Americas (Mollusca: Gastropoda: Vetigastropoda). Zootaxa 2356: 1-35, DOI: 10.5281/zenodo.275645
03DCBC11C25EFFB50ED21A78FB7960E5.taxon	description	Description. Shell small (to 0.54 mm), trochiform. Protoconch of 0.875 whorls, fine axials on outer portion of whorl, weak apertural varix not connected to embryonic cap, apertural margin straight. TI of 0.8 – 1.1 whorls, with 15 – 38 axial cords, interstices with fine irregular growth marks, first spiral thread no earlier than after 0.66 whorls, increasing to 0 – 2 on shoulder, approximately six between position of selenizone and periphery. TII of 0.33 whorls. Shoulder almost flat, axial cords of same density as on TI, two to five indistinct spiral lines. Base with moderate constriction below selenizone, axials as on shoulder, approximately 12 spirals, increasing in strength from lines at periphery to low cords near umbilicus, sometimes forming diffuse nodules at intersection with axials. Umbilicus open, bordered by moderately strong spiral cord, walls straight, with fine axial growth marks. Selenizone slightly above periphery, keels low, strong, growth marks distinct, selenizone approximately twice as long as teardrop shaped foramen. Aperture subquadratic, roof overhanging. Radula, rachidian triangular with five denticles of cusp, central one largest, denticles arranged in V. Lateral teeth 1 – 3 similar, three to four denticles on outer edge of cusp. Lateral tooth 4 reduced, hook-shaped. Lateral tooth 5 enlarged by broadening, six denticles on inner margin of cusp. Radular interlock of central field moderate. Inner marginal teeth with terminal denticle largest, one on inner edge, four on outer edge. Outer marginal teeth spoon-shaped with many denticles on either side of cusp. Differential diagnosis. Sinezona rimuloides (Carpenter, 1856) from the Panamic province to central California is more elevated in overall shape, has a narrower umbilicus, and the selenizone is approximately three times as long as the foramen.	en	Geiger, Daniel L., Mclean, James H. (2010): New species and records of Scissurellidae and Anatomidae from the Americas (Mollusca: Gastropoda: Vetigastropoda). Zootaxa 2356: 1-35, DOI: 10.5281/zenodo.275645
03DCBC11C25EFFB50ED21A78FB7960E5.taxon	distribution	Distribution. Mexico (offshore Isla Maria Cleofa), Clipperton and Cocos Islands.	en	Geiger, Daniel L., Mclean, James H. (2010): New species and records of Scissurellidae and Anatomidae from the Americas (Mollusca: Gastropoda: Vetigastropoda). Zootaxa 2356: 1-35, DOI: 10.5281/zenodo.275645
03DCBC11C252FFB40ED21BFBFC72625B.taxon	materials_examined	Type species. Coronadoa simonsae Bartsch, 1915 (OD).	en	Geiger, Daniel L., Mclean, James H. (2010): New species and records of Scissurellidae and Anatomidae from the Americas (Mollusca: Gastropoda: Vetigastropoda). Zootaxa 2356: 1-35, DOI: 10.5281/zenodo.275645
03DCBC11C252FFB40ED21BFBFC72625B.taxon	diagnosis	Diagnosis. Scissurellidae without a selenizone, slit, or foramen.	en	Geiger, Daniel L., Mclean, James H. (2010): New species and records of Scissurellidae and Anatomidae from the Americas (Mollusca: Gastropoda: Vetigastropoda). Zootaxa 2356: 1-35, DOI: 10.5281/zenodo.275645
03DCBC11C252FFB10ED21AF5FDA0667D.taxon	materials_examined	Type material. Holotype (LACM 3113: Fig. 9 C). Paratypes: 26 – 40 m, Farnsworth Bank, off S side of Catalina Island, California, USA, 33.350 ˚ N, 118.517 ˚ W (LACM 3114); 13 – 30 m, Cortez Bank, 0.5 mile south of Bishop Rock, California, USA, 32.433 ˚ N, 119.125 ˚ W (LACM 3115: Fig. 9 A – B). All LACM type material leg. J. McLean. Type locality. 26 – 40 m, Farnsworth Bank, off S side of Catalina Island, California, USA, 33.350 ˚ N, 118.517 ˚ W. Additional material examined. 10 – 23 m, NW side of Guadalupe Island, (Pilot Rock and 5 miles S), Baja California, Mexico, 29.188 ˚ N, 118.253 ˚ W (LACM 72 - 121, 1). 20 – 26 m, SW side Natividad Island, Baja California, Mexico, 27.888 ˚ N, 115.212 ˚ W (LACM 72 - 117, 1). 13 – 17 m, 5 fathom Bank about 4 miles SW of SE tip of Cedros Island, outer coast Baja California, Mexico, 27.983 ˚ N, 115.212 ˚ W (LACM 71 - 93, 2). 26 m, San Miguel Island, California, USA, 34.020 ˚ N, 120.402 ˚ W (DLG 1131, 1). 46 m, off Santa Cruz Island, California, USA, 34.056 ˚ N, 119.961 ˚ W (DLG 1263, 3).	en	Geiger, Daniel L., Mclean, James H. (2010): New species and records of Scissurellidae and Anatomidae from the Americas (Mollusca: Gastropoda: Vetigastropoda). Zootaxa 2356: 1-35, DOI: 10.5281/zenodo.275645
03DCBC11C252FFB10ED21AF5FDA0667D.taxon	etymology	Etymology. demisi-: Latin for sunken; - spira: Latin for spire. Referring to the flat spire of the species.	en	Geiger, Daniel L., Mclean, James H. (2010): New species and records of Scissurellidae and Anatomidae from the Americas (Mollusca: Gastropoda: Vetigastropoda). Zootaxa 2356: 1-35, DOI: 10.5281/zenodo.275645
03DCBC11C252FFB10ED21AF5FDA0667D.taxon	description	Description. Shell small (to 0.75 mm), trochiform depressed. Protoconch of 0.875 whorls, embryonic cap without sculpture, remainder with strong axial ribs, no apertural varix, apertural margin prosocline. Teleoconch of 1.5 whorls, 11 – 13 raised axial cords, indistinct at suture and towards umbilicus, most pronounced at periphery, interstices with irregular axial growth marks; no spiral sculpture. Umbilicus open, occasionally undersurface of protoconch visible, bordered by knobby carina towards base. Aperture subquadratic D-shaped, lower adumbilical corner pronounced, forming knobby carina of umbilicus. Operculum thin, round, multispiral, nucleus central. Differential diagnosis. The sympatric C. simonsae Bartsch, 1946 (Fig. 9 A – B and Fig. 10 A, C; Fig. 9 C and Fig. 10 B) is proportionally taller (width to height: simonsae mean = 1.28, SD = 0.047, n = 4; demisispira mean = 1.79, SD = 0.22, n = 4; t-test t = - 4.36, p = 0.022). As a corollary, its umbilicus is wider, and tends to have fewer axial cords per unit whorl. The degree of axial sculpturing is variable in both species, from distinct cords to raised lamellae. Coronadoa hasegawai Geiger & Sasaki, 2009, from temperate Japan is as tall as C. simonsae.	en	Geiger, Daniel L., Mclean, James H. (2010): New species and records of Scissurellidae and Anatomidae from the Americas (Mollusca: Gastropoda: Vetigastropoda). Zootaxa 2356: 1-35, DOI: 10.5281/zenodo.275645
03DCBC11C252FFB10ED21AF5FDA0667D.taxon	discussion	Remarks. Coronadoa demisispira is much less common than C. simonsae; if the two species are present in the same lot, C. demisispira comprises at most 10 % of C. simonsae specimens. This skewed ratio suggests that the two are not dimorphic males and females of the same species. In general, Vetigastropoda show a male: female ratio of close to 1: 1 (Bretos et al. 1985, Hooker & Creese 1995, Wells & Malay 1995), although there are no data on Scissurellidae s. l.	en	Geiger, Daniel L., Mclean, James H. (2010): New species and records of Scissurellidae and Anatomidae from the Americas (Mollusca: Gastropoda: Vetigastropoda). Zootaxa 2356: 1-35, DOI: 10.5281/zenodo.275645
03DCBC11C257FFB10ED21C10FADB6060.taxon	materials_examined	Type material. Holotype USNM 1132066. 9 paratypes from type locality (USNM 1132067). 15 paratypes 137 – 146 m, off Pelican Island, Barbados, 13.167 ˚ N, 59.567 ˚ W (USNM 500678). One paratype 165 – 183 m, off Lazaretto, Barbados, 13.133 ˚ N, 59.617 ˚ W (USNM 1132065, 1). Type locality. 137 m, Off Pelican Island, Barbados, 13.167 ˚ N, 59.567 ˚ W.	en	Geiger, Daniel L., Mclean, James H. (2010): New species and records of Scissurellidae and Anatomidae from the Americas (Mollusca: Gastropoda: Vetigastropoda). Zootaxa 2356: 1-35, DOI: 10.5281/zenodo.275645
03DCBC11C257FFB10ED21C10FADB6060.taxon	etymology	Etymology. Alternati-, Latin for differing on two sides, - sculpta, Latin for sculpture, referring to the predominant axial sculpture on shoulder, while the base has predominantly spiral sculpture. Adjective.	en	Geiger, Daniel L., Mclean, James H. (2010): New species and records of Scissurellidae and Anatomidae from the Americas (Mollusca: Gastropoda: Vetigastropoda). Zootaxa 2356: 1-35, DOI: 10.5281/zenodo.275645
03DCBC11C257FFB10ED21C10FADB6060.taxon	description	Description. Shell medium size (to 2.09 × 2.15 mm W × H), trochiform globular. Protoconch of 0.875 whorls, 175 µm long in holotype, embryonic cap with some flocculent sculpture, remainder smooth, weak apertural varix not connected to embryonic cap, apertural margin slightly sinusoid. Teleoconch I of 0.66 – 0.75 whorls, 14 – 19 axial cords, some with a spiral cord in position of selenizone. Teleoconch II of 2.66 whorls, suture moderately impressed. Shoulder strongly convex, approximately 34 distinct equally spaced axial cords; first spiral line shortly after onset of selenizone, 3 – 4 thick spiral lines at apertural margin concentrated in middle of shoulder, running over axial cords forming minute thickenings. Base with weak constriction below selenizone, evenly curving into umbilicus, approximately 20 axial cords of decreasing strength from selenizone towards mid-base, then increasing in strength towards umbilicus; about twice as many thick spiral threads as on shoulder, weak below selenizone, increasing in strength to cords towards umbilicus. Umbilicus open, moderately wide. Selenizone at periphery, keels moderately strong, strongly elevated, lunules distinct; slit open, with parallel margins. Aperture round, flared, roof overhanging. Animal unknown. Differential diagnosis. Anatoma proxima from the SE United States to Brazil has similar sculpture on shoulder and base, a less convex shoulder, and a protoconch with flocculent sculpture. Anatoma atlantica (Bandel, 1998) from the eastern US is proportionally wider (width to height ratio 1.219 in holotype of A. atlantica, 0.973 in holotype of A. alternatisculpta), has finer sculpture and a much larger protoconch. Anatoma plicatazona n. sp. from the Caribbean has a more turreted shell form, and the keels of the selenizone are strongly curved towards the apex. Anatoma alta (Watson, 1886) from the NE Atlantic is more globular in shape, the selenizone distinctly above the periphery, and has similar sculpture on shoulder and base.	en	Geiger, Daniel L., Mclean, James H. (2010): New species and records of Scissurellidae and Anatomidae from the Americas (Mollusca: Gastropoda: Vetigastropoda). Zootaxa 2356: 1-35, DOI: 10.5281/zenodo.275645
03DCBC11C257FFB10ED21D17FB6765C6.taxon	materials_examined	Type species. Scissurella crispata Fleming, 1828 (M).	en	Geiger, Daniel L., Mclean, James H. (2010): New species and records of Scissurellidae and Anatomidae from the Americas (Mollusca: Gastropoda: Vetigastropoda). Zootaxa 2356: 1-35, DOI: 10.5281/zenodo.275645
03DCBC11C257FFB10ED21D17FB6765C6.taxon	diagnosis	Diagnosis. Anatomidae with open slit, protoconch smooth or with flocculent sculpture.	en	Geiger, Daniel L., Mclean, James H. (2010): New species and records of Scissurellidae and Anatomidae from the Americas (Mollusca: Gastropoda: Vetigastropoda). Zootaxa 2356: 1-35, DOI: 10.5281/zenodo.275645
03DCBC11C257FFB10ED21EDFFB1D66C5.taxon	diagnosis	Diagnosis. Shell small (0.9 – 11 mm), without nacre, with slit or foramen at periphery of whorl.	en	Geiger, Daniel L., Mclean, James H. (2010): New species and records of Scissurellidae and Anatomidae from the Americas (Mollusca: Gastropoda: Vetigastropoda). Zootaxa 2356: 1-35, DOI: 10.5281/zenodo.275645
03DCBC11C256FFAC0ED21FD5FF026266.taxon	description	Synonymy	en	Geiger, Daniel L., Mclean, James H. (2010): New species and records of Scissurellidae and Anatomidae from the Americas (Mollusca: Gastropoda: Vetigastropoda). Zootaxa 2356: 1-35, DOI: 10.5281/zenodo.275645
03DCBC11C256FFAC0ED21FD5FF026266.taxon	materials_examined	Type material. Holotype (ZIN 32372), 1.32 × 1.05 mm (W × H). One to four paratypes (repository uncertain: ZIN?). Type locality. Near Moneron Island, [Russia, northern Sea of Japan, 46.250 ˚ N, 141.250 ˚ E] 50 – 70 m. Leg. B. I. Sirenko. (OD).	en	Geiger, Daniel L., Mclean, James H. (2010): New species and records of Scissurellidae and Anatomidae from the Americas (Mollusca: Gastropoda: Vetigastropoda). Zootaxa 2356: 1-35, DOI: 10.5281/zenodo.275645
03DCBC11C256FFAC0ED21FD5FF026266.taxon	etymology	Etymology. More disk-like than other species (crispata, japonica) (OD).	en	Geiger, Daniel L., Mclean, James H. (2010): New species and records of Scissurellidae and Anatomidae from the Americas (Mollusca: Gastropoda: Vetigastropoda). Zootaxa 2356: 1-35, DOI: 10.5281/zenodo.275645
03DCBC11C256FFAC0ED21FD5FF026266.taxon	description	Description. Shell small (to 1.3 mm), translation of Golikov & Sirenko (1980: 107 – 108), the translation of the Russian original description was kindly provided by Vyacheslav Palchevskiy (University of Southern California, Los Angeles, California, USA): “ The shell is small, almost disk-like with thin walls and with 2.5 slightly convex whorls separated by a deeply impressed suture. The last whorl makes a small concave platform near the suture. The upper part of the whorl has a weakly convex, smooth slope towards the periphery, restricted by two raised keels, which border the slit near the outer edge of the aperture. The base of the shell under the keels is moderately convex but forms a deep depression in the region of the wide umbilicus. The diameter of the last whorl is 1.45 × as wide as high and 1.3 × greater than the height of the shell. The surface of the shell is milky-white though in one of the paratypes, the color of the shell is slightly yellow. The axial sculpture of the shell consists only of thin, non-contrasted growth lines. Only in the region of the umbilicus, there are slightly separated, barely visible (only under magnification) wrinkles. The spiral sculpture is absent except for separate, non-distinct, unshaped lines. The aperture is round-oval, essentially shifted in respect to the axis of the shell; its height is 3 / 4 of the shell’s height. The outer apertural lip has a slight angle in the upper part near the parietal wall of the whorl and slopes smoothly towards a deep and narrow slit; the region where the keels frame the slit is slightly raised. The inner lip near the columella is strongly curved outside making the narrow fold, and during the transition towards the base of the shell, forms a smooth, circular and dull angle. ” Redescription based on material from Alaska collected by the late Rae Baxter. Shell medium size, to 2.3 mm, trochiform biconical. Protoconch of 0.75 whorls, coarse flocculent sculpture, no apertural varix, apertural margin straight. Teleoconch I of 0.45 – 0.6 whorls, irregular axial thickenings sometimes as approximately 20 low, broad axial cords; no spiral sculpture. Teleoconch II up to 2.1 whorls. Suture below selenizone, deeply impressed in early whorls; descending by approximately 1.5 × width of selenizone on base on largest known specimen, weakly impressed. Shoulder convex, early sculpture from almost smooth to low axial cords only; subsequently coarsely to finely clathrate sculpture of variable density and distinctness. Base with similar sculpture as shoulder. Umbilicus open, distinct funiculus. Selenizone slightly above periphery, keels low, often broken, moderate strength; slit open, margins slightly converging. Aperture rounded Dshaped, roof overhanging. Animal unknown. Differential diagnosis. Anatoma obtusata (Golikov & Gublin, 1978) from the boreal north Pacific is more globular in overall shell shape, has a teleoconch I of 0.875 – 1 whorls, and a very narrow umbilicus. Thieleella baxteri (McLean, 1984) from the Northeastern Pacific shares the discoidal shell shape in smaller specimens, but has strong and sharp axial lamellae on shoulder and base, crossed by much finer spiral lines. Thieleella kelseyi (Dall, 1905) from the northeastern Pacific has an overall more turreted shell shape, a protoconch with reticulate sculpture and fine axial cords on shoulder and base crossed by finer spiral lines.	en	Geiger, Daniel L., Mclean, James H. (2010): New species and records of Scissurellidae and Anatomidae from the Americas (Mollusca: Gastropoda: Vetigastropoda). Zootaxa 2356: 1-35, DOI: 10.5281/zenodo.275645
03DCBC11C256FFAC0ED21FD5FF026266.taxon	distribution	Distribution. North Pacific from Moneron Island (46.250 ˚ N, 141.250 ˚ E) to Alaska (52.300 ˚ N, 170.040 ˚ W), 7 – 210 m (empty shells only). Specimen records. 210 m, S of Attu Island, W of Agattu Island, Near Islands, Aleutian Islands, Alaska, USA, 52.496 ˚ N, 172.908 ˚ W (LACM 86 - 310, 2). 152 m, S of Semichi Island, E of Cape Sabak, Agattu Island, Near Islands, Alaska, USA, 52.379 ˚ N, 174.121 ˚ W (LACM 86 - 356, 1). 121 m, N of Carlisle Island, Islands of Four Mountains, Alaska, USA, 52.976 ˚ N, 170.043 ˚ W (LACM 86 - 333, 1). 82 m, SE of Red bluff, NE end Kiska Island, Rat Islands, Alaska, USA, 52.074 ˚ N, 177.696 ˚ W (LACM 86 - 315, 3). 121 m, off SE side Semisopochoi Island, Rat Islands, Alaska, USA, 51.889 ˚ N, 179.760 ˚ W (LACM 86 - 322, 2). 7 m, Middle Rocks, Chicagof Harbor, Bering Sea side of Attu Island, Near Islands, Aleutian Islands, Alaska, USA, 52.940 ˚ N, 173.265 ˚ W (LACM 79 - 71, 4). All LACM material leg. Rae Baxter.	en	Geiger, Daniel L., Mclean, James H. (2010): New species and records of Scissurellidae and Anatomidae from the Americas (Mollusca: Gastropoda: Vetigastropoda). Zootaxa 2356: 1-35, DOI: 10.5281/zenodo.275645
03DCBC11C256FFAC0ED21FD5FF026266.taxon	discussion	Remarks. Five specimens were collected, and reference is made to at least one paratype, but it is unclear whether only one or more paratypes were designated. The holotype was illustrated by Kantor & Syosev (2006) and is strongly decalcified. Multiple attempts at obtaining the specimen on loan failed. The specimens from Alaska are considered conspecific as they agree as well as possible with the original illustration, the subsequent photograph of the holotype as well as with the original description. The overall shape of the specimens, the strong funiculus, the very weak sculpture in some of the examples examined, and the distribution in the boreal Pacific all support the identification of the Alaskan specimens as A. disciformis. The original illustration shows a specimen with a very rounded aperture, whereas some of the specimens shown here have a rather more angular D-shaped aperture. The latter condition is encountered in specimens with a broken lower adumbilical portion of the peristome, hence, is an artifact of no taxonomic significance. As in Anatomidae, juvenile specimens are more depressed than older ones as shown by the ontogenetic series in Figure 12 [see also Zelaya & Geiger 2007, figs 18 – 40, for ontogenetic series of A. euglypta (Pelseneer, 1903)]. The shared protoconch and early teleoconch features support that the specimens belong to the same species.	en	Geiger, Daniel L., Mclean, James H. (2010): New species and records of Scissurellidae and Anatomidae from the Americas (Mollusca: Gastropoda: Vetigastropoda). Zootaxa 2356: 1-35, DOI: 10.5281/zenodo.275645
03DCBC11C24AFFAA0ED21AF0FD2D6586.taxon	description	Anatoma n. sp.: Geiger, 2008: fig. 1.	en	Geiger, Daniel L., Mclean, James H. (2010): New species and records of Scissurellidae and Anatomidae from the Americas (Mollusca: Gastropoda: Vetigastropoda). Zootaxa 2356: 1-35, DOI: 10.5281/zenodo.275645
03DCBC11C24AFFAA0ED21AF0FD2D6586.taxon	materials_examined	Type material. Holotype (SBMNH 83518, ex. CRC 12843). Paratype (BMSM 17981, ex CRC 12842) from type locality. Type localit y. 7 m, Sandy Cay, Abaco, Bahamas, 26.399 ˚ N, 76.988 ˚ W. August 13, 2005, leg. Colin Redfern.	en	Geiger, Daniel L., Mclean, James H. (2010): New species and records of Scissurellidae and Anatomidae from the Americas (Mollusca: Gastropoda: Vetigastropoda). Zootaxa 2356: 1-35, DOI: 10.5281/zenodo.275645
03DCBC11C24AFFAA0ED21AF0FD2D6586.taxon	etymology	Etymology. Plicata, Latin for folded, - zona referring to the selenizone. Describing the diagnosing feature of the strongly upward folded selenizone.	en	Geiger, Daniel L., Mclean, James H. (2010): New species and records of Scissurellidae and Anatomidae from the Americas (Mollusca: Gastropoda: Vetigastropoda). Zootaxa 2356: 1-35, DOI: 10.5281/zenodo.275645
03DCBC11C24AFFAA0ED21AF0FD2D6586.taxon	description	Description. Shell to 1.5 mm, trochiform, turreted. Protoconch 0.75 whorls, flocculent sculpture, no apertural varix, apertural margin straight. TI of 0.25 – 0.33 whorls, 11 – 12 axial cords, no spiral sculpture, interstices with irregularities. TII> 2 whorls. Shoulder convex, approximately 50 distinct axial cords on first whorl, becoming gradually less distinct to diffuse towards apertural margin; first spiral after 0.125 – 0.25 TII whorls, half to third strength of axials, increasing to 5 – 7 after 1 TII whorl, of equal strength as axials, forming raised points at intersection with axials; raised points best developed after 1.25 TII whorls; sculpture becoming less distinct with growth, forming subdued cancellate pattern. Suture little impressed, position descending with growth from 1.5 width of selenizone below selenizone, to mid base (~ 3 selenizone widths below selenizone). Base convex, evenly curving into umbilicus, barely constricted below selenizone; sculpture becoming weaker with growth and from selenizone towards base; reticulate sculpture with raised points at intersection below selenizone, subsequently approximately 13 low spiral steps. Aperture rounded, adumbilical portion of peristome flared. Selenizone slightly above periphery; keels of moderate strength, broken in early portion of whorl, appearing to have horizontal orientation, on last 0.5 – 0.66 whorls orientation of keels parallel to coiling axis; slit open, margins parallel. Animal unknown. Differential diagnosis. Anatoma aedonia Watson, 1886, from the Caribbean has a similar overall shape and similar protoconch. However, TI is of 1 whorl and has a distinct spiral cord in the position of the selenizone, the sculpture on TII remains the same throughout growth, with distinct axial cords, and very fine spiral lines, and the axial sculpture on the base extends to the umbilicus. Differential diagnosis based on examination of types in BMNH and NMW by SEM.	en	Geiger, Daniel L., Mclean, James H. (2010): New species and records of Scissurellidae and Anatomidae from the Americas (Mollusca: Gastropoda: Vetigastropoda). Zootaxa 2356: 1-35, DOI: 10.5281/zenodo.275645
03DCBC11C24AFFAA0ED21AF0FD2D6586.taxon	distribution	Distribution. Known only from type locality.	en	Geiger, Daniel L., Mclean, James H. (2010): New species and records of Scissurellidae and Anatomidae from the Americas (Mollusca: Gastropoda: Vetigastropoda). Zootaxa 2356: 1-35, DOI: 10.5281/zenodo.275645
03DCBC11C24CFFAA0ED21C4FFBE1648E.taxon	materials_examined	Type species. Scissurella amoena Thiele, 1912 (OD).	en	Geiger, Daniel L., Mclean, James H. (2010): New species and records of Scissurellidae and Anatomidae from the Americas (Mollusca: Gastropoda: Vetigastropoda). Zootaxa 2356: 1-35, DOI: 10.5281/zenodo.275645
03DCBC11C24CFFAA0ED21C4FFBE1648E.taxon	diagnosis	Diagnosis. Anatomidae with open slit, protoconch with reticulate sculpture.	en	Geiger, Daniel L., Mclean, James H. (2010): New species and records of Scissurellidae and Anatomidae from the Americas (Mollusca: Gastropoda: Vetigastropoda). Zootaxa 2356: 1-35, DOI: 10.5281/zenodo.275645
03DCBC11C24CFFA40ED21CBDFBF967C6.taxon	materials_examined	Type material. Holotype (USNM 181820. Lost: McLean 1967: 405, Geiger pers. obs. 2. 2004), 5.5 × 6 mm (H × W). Neotype here designated (SBMNH 3215: see remarks). Type locality. California at U. S. Fish Commission Station 4353 [32.700 ˚ N, 117.233 ˚ W], also off San Diego (OD), 639 fms [= 1169 m], 15 mls from Pt. Loma, 47 ˚ W (USNM label). Neotype locality: 183 m, off La Jolla, California, USA [32.843 ˚ N, 117.392 ˚ W].	en	Geiger, Daniel L., Mclean, James H. (2010): New species and records of Scissurellidae and Anatomidae from the Americas (Mollusca: Gastropoda: Vetigastropoda). Zootaxa 2356: 1-35, DOI: 10.5281/zenodo.275645
03DCBC11C24CFFA40ED21CBDFBF967C6.taxon	etymology	Etymology. Named in honor of Prof. F. W. Kelsey, of San Diego (OD).	en	Geiger, Daniel L., Mclean, James H. (2010): New species and records of Scissurellidae and Anatomidae from the Americas (Mollusca: Gastropoda: Vetigastropoda). Zootaxa 2356: 1-35, DOI: 10.5281/zenodo.275645
03DCBC11C24CFFA40ED21CBDFBF967C6.taxon	description	Description. Shell large (to 6 mm: fide Dall, 1905), trochiform, biconical, white. Protoconch of 0.75 whorls, with reticulate sculpture, apertural varix barely connected to embryonic cap, apertural margin straight. TI of 0.75 whorls, with approximately 38 axial cordlets, one spiral cordlet in position of selenizone, interstices with flocculent sculpture. TII of 3.125 whorls in 3.2 mm specimen, space between suture and selenizone as wide as selenizone showing approximately the uppermost three spirals. Shoulder somewhat convex, suture lightly impressed, axials approximately of same density and strength as on TI, approximately 22 equispaced fine spiral lines running over cordlets. Base with slight constriction below selenizone, axials of same strength and density as on shoulder, crossed with approximately 32 equispaced spiral lines, interstices with fine irregular growth lines. Umbilicus very narrow, evenly curving with base, funiculus present. Selenizone slightly above periphery, keels of moderate strength and moderate elevation, growth marks rather weak; slit open with parallel margins. Aperture rounded, adumbilical portion flared, roof overhanging. Operculum corneous, thin, multispiral with central nucleus. Radula. Rachidian tooth trapezoid, central denticle largest, approximately 6 denticles on each side. Lateral teeth 1 – 3 similar, apical denticle largest; 3 – 4 denticles on outer cutting edge; 1 – 2 denticles on along inner edge, half size of outer denticles. Lateral tooth 4 reduced, hookshaped, apical denticle largest, 3 – 4 minute denticles on inner edge. Lateral tooth 5 enlarged by elongation, apical denticle largest, approximately 12 denticles along inner edge, approximately 4 denticles along distal half of cusp’s outer edge. Inner marginal teeth with triangular cusp on long shafts, apical denticle largest, approximately 6 denticles on each side. Outer marginal teeth thin, broadened, with many small bristles along edge of cusp. Radular interlock of central field moderate. Body. Tentacles in anterior-posterior order: cephalic tentacles papillate; ocular stalk without pigmented eye; two distinct, non-papillate tentacles in neck region in anterior-posterior alignment with cephalic tentacle and first epipodial tentacle; bifid papillate epipodial tentacles with single posterior blunt, stubby non-papillate structure about half as long as second epipodial tentacle, most likely elongated epipodial sense organ (ESO); two single papillate epipodial tentacles. Operculum attached to discrete fine flap of metapodium. Mantle with papillate tentacle at posterior junction of slit near anal papilla. Two monopectinate gills, approximately 25 – 30 gill leaflets each, distinct bottle-shaped bursicles in distal half of each filament, opening of bursicles in middle of filament. Differential Diagnosis. Anatoma keenae (McLean, 1966) [= A. epicharis (McLean, 1966)] from the Panamic province has a protoconch with flocculent sculpture (reticulate in T. kelseyi), on TI lacks a spiral cord in the position of the selenizone (present in T. kelseyi), and forms small but distinct points at the intersection of spiral and axial cords on TII (no points in T. kelseyi). Thieleella baxteri from the northeastern Pacific has a TI of approximately 0.5 whorls (0.75 in T. kelseyi) and axial sculpture consisting of raised lamellae (simple cords in T. kelseyi). Anatoma lyra (Berry, 1947), from the northeastern Pacific has a protoconch with flocculent sculpture (reticulate in T. kelseyi), overall stronger and less dense sculpture, and a distinct funiculus in the umbilicus (absent in T. kelseyi).	en	Geiger, Daniel L., Mclean, James H. (2010): New species and records of Scissurellidae and Anatomidae from the Americas (Mollusca: Gastropoda: Vetigastropoda). Zootaxa 2356: 1-35, DOI: 10.5281/zenodo.275645
03DCBC11C24CFFA40ED21CBDFBF967C6.taxon	distribution	Distribution. Queen Charlotte Islands, British Columbia, Canada, to South Coronado Island, northern Baja California, Mexico (Dall 1921). Specimen records. USA: California. 1503 m, US Fish Comm. St. 2923, off San Diego (USNM 209412, 4). 1033 – 1244 m, 16 miles SW off Point Loma, 32.700 ˚ N, 117.246 ˚ W (USNM 211166, 4). San Diego Trough: 1171 – 1241 m, 32.417 ˚ N, 117.467 ˚ W (SIO M 1974, 1: complete). 1166 – 1222 m, 32.450 ˚ N, 117.467 ˚ W (SIO M 1991, 1, 5: complete). 1207 – 1234 m, SIO 69 - 487, 32.450 ˚ N, 117.525 ˚ W (SIO M 2070, 3: complete). 1183 – 1216 m, R- 28, 32.583 ˚ N, 117.558 ˚ W (SIO 2171, 1: complete). 1241 m, R- 42, 32.440 ˚ N, 117.483 ˚ W (SIO M 1778, 1: complete). 1199 – 1241 m, R- 17, 32.447 ˚ N, 117.467 ˚ W (SIO M 1834, 2: complete). San Clemente Island: 390 m, off Tanner Bank, 32.683 ˚ N, 119.234 ˚ W (LACM 76 - 313, 6). 1047 m, 32.585 ˚ N, 118.965 ˚ W (LACM 76 - 378, 6). 933 m, 30.8 mi, 65 ˚ T of China Point, 32.583 ˚ N, 118.982 ˚ W (LACM 76 - 377, 2). 973 m, California, USA, 32.569 ˚ N, 118.984 ˚ W (LACM 76 - 381, 5). 292 m, 32.566 ˚ N, 119.131 ˚ W (LACM 76 - 387, 4). 201 m, 32.772 ˚ N, 118.379 ˚ W (LACM AHF 1082, 1). Between Tanner Bank and Cortez Bank, 32.450 ˚ N, 119.250 ˚ W (SBMNH 33350, 1). 1200 m, Off N. Coronado Island, 32.750 ˚ N, 117.500 ˚ W (USNM 208901, 3; USNM 209054, 3). 183 m, off La Jolla, 32.8430 ˚ N, 117.392 ˚ W (SBMNH 3215, 1). Santa Rosa Island: 70 m, 33.531 ˚ N, 119.982 ˚ W (LACM 77 - 303, 1). 115 m, 33.865 ˚ N, 119.917 ˚ W (LACM 75 - 300, 6). 61 m, 33.865 ˚ N, 120.030 ˚ W (LACM 75 - 306, 1). 105 m, 33.934 ˚ N, 119.901 ˚ W (LACM 75 - 480, 1). 442 m, 33.884 ˚ N, 120.316 ˚ W (LACM 76 - 213, 8). 421 m, 33.850 ˚ N, 120.183 ˚ W (LACM 23129, 2: complete). 492 m, Off Point Sal, California, USA, 34.970 ˚ N, 121.298 ˚ W (SBMNH 350392, 1; SBMNH 350392, 2). USA: Oregon. 1580 m, 45.828 ˚ N, 125.233 ˚ W (LACM 67 - 167, 1). Canada: British Columbia. 2176 m, Estevan Point Light, Hesquiat Peninsula, Noetka Sound, Vancouver Island, 48.635 ˚ N, 126.967 ˚ W (LACM 71 - 374.7, 3: complete). 1602 m, off Queen Charlotte Sound, 51.500 ˚ N, 129.000 ˚ W (USNM 206690, 1). 55 m, Naden Harbor, N side of Graham Island, Queen Charlotte Island, 54.000 ˚ N, 132.667 ˚ W (LACM 66 - 66, 0). 260 m, Dixon Entrance, N of Graham Island, 54.483 ˚ N, 133.200 ˚ W (LACM 67 - 28, 2). USA: Alaska. 88 m, NW of Gibson Island, off NE side Attu Island, Near Islands, 52.958 ˚ N, 173.237 ˚ W (LACM 86 - 312, 5). 179 m, S of Coronation Island, W of N end Dall Island, 55.143 ˚ N, 134.210 ˚ W (LACM 87 - 377, 2). Baranof Island: 278 m, W of gulf of Esquibel, S of Byron Bay, 55.537 ˚ N, 134.891 ˚ W (LACM 87 - 369, 27). 205 m, W of N end Baker Island, S of Cape Ommany, 55.394 ˚ N, 134.675 ˚ W (LACM 87 - 370, 20). 443 m, W of Nation Point, Coronation Island, SW of Cape Ommaney, 55.925 ˚ N, 135.438 ˚ W (LACM 87 - 365, 8). 252 m, SW of Cape Ommaney, S end, 56.000 ˚ N, 135.000 ˚ W (LACM 87 - 357, 2). 364 m, W of Pt. Howard, Kulu Island, S of Sitka, 56.037 ˚ N, 135.307 ˚ W (LACM 87 - 364, 13). 362 m, W of Pt. Crowley, Kulu Island, S. of Necker Bay, 56.090 ˚ N, 135.150 ˚ W (LACM 87 - 363, 9). 240 m, S of Kruzof Island, W of Necker Bay, 56.598 ˚ N, 135.782 ˚ W (LACM 87 - 358, 100). W of Whale Bay, SSW of Biorka Island, 56.514 ˚ N, 135.661 ˚ W (LACM 87 - 395, 29). 390 m, SW of mouth of Sitka Sound, SSW of Cape Edgecumbe, S end of Kruzof Island, 56.778 ˚ N, 135.970 ˚ W (LACM 87 - 356, 25). 274 m, S of Cape Edward, W of Gilmer Bay, Kruzof Island, 57.219 ˚ N, 136.234 ˚ W (LACM 87 - 354, 17). 408 m, S of Icy Point, W of Point Theodore, Yakobi Island, 57.853 ˚ N, 137.111 ˚ W (LACM 87 - 346, 1). 214 m, SSW of Cape Fairweather, Chichagof Island, W of Cape Bingham, 58.073 ˚ N, 138.476 ˚ W (LACM 87 - 330, 3). 384 m, W of Yakobi Island, S of Yakutat Bay, 58.000 ˚ N, 140.000 ˚ W (LACM 87 - 335, 1). 178 m, SE of Icy Bay, SW of Yakutat Bay, 59.453 ˚ N, 141.150 ˚ W (LACM 87 - 336, 1). 353 m, S of Icy Bay, E of Dry Bay, 59.118 ˚ N, 141.495 ˚ E (LACM 87 - 333, 10).	en	Geiger, Daniel L., Mclean, James H. (2010): New species and records of Scissurellidae and Anatomidae from the Americas (Mollusca: Gastropoda: Vetigastropoda). Zootaxa 2356: 1-35, DOI: 10.5281/zenodo.275645
03DCBC11C24CFFA40ED21CBDFBF967C6.taxon	discussion	Remarks. Although well figured by McLean (1996), the identification of this familiar species must be changed, as it is clearly not conspecific with A. crispata (Fleming, 1828) with type locality in Europe. Given the controversial nature of the name, it is prudent for taxon stabilization to designate a neotype. A specimen collected near to the type locality is here so designated. The specimen agrees in all aspects with the original description, though the latter lacks much specific detail. Specimens in the original type repository (USNM) originally identified as kelseyi were examined, and the best specimen was imaged by SEM (USNM 209054). Unfortunately, its protoconch is eroded, which makes generic assignment impossible (see Geiger 2003 for discussion of genera in Anatomidae). A specimen with preserved protoconch sculpture, agreeing in all aspects of teleoconch morphology with the USNM kelseyi specimens and obtained from near the original type locality, was chosen instead. The generic placement of kelseyi follows from the reticulate sculpture of the protoconch.	en	Geiger, Daniel L., Mclean, James H. (2010): New species and records of Scissurellidae and Anatomidae from the Americas (Mollusca: Gastropoda: Vetigastropoda). Zootaxa 2356: 1-35, DOI: 10.5281/zenodo.275645
03DCBC11C24CFFA40ED21CBDFBF967C6.taxon	description	The two smooth tentacles located between the cephalic tentacle and the first epipodial tentacles are most likely homologous to what Quinn (1983), Sasaki (1998) and Kano (2009) called the accessory cephalic tentacle. A single suboptic tentacle can be found in some species, but it is more closely associated with the eye, and usually significantly smaller. As these accessory cephalic tentacles are paired, and as fertilization is most likely external and no ciliary tracks are associated with them, it is unlikely that they have any penial function. The existence of an epipodial sense organ (ESO) in Anatomidae has been uncertain. Here, the nonpapillate stubby projection associated with the first two epipodial tentacles is interpreted as the ESO. Its nonpapillate, blunt nature agrees with typical ESO, which usually, however, are located ventrally to epipodial tentacles and are only about has high as wide (here 3 – 4 times as high as wide). Sasaki et al. (in press) interpreted the homologous structure in Anatoma n. sp. also as an ESO. The presumably chemosensory bursicles (see Geiger et al. 2008 for review) were found by Sasaki (1998) in Anatomidae and their presence in the family is here confirmed. The bursicles are distinct, with their typical bottle shape and the opening to the chamber could be seen in several cases.	en	Geiger, Daniel L., Mclean, James H. (2010): New species and records of Scissurellidae and Anatomidae from the Americas (Mollusca: Gastropoda: Vetigastropoda). Zootaxa 2356: 1-35, DOI: 10.5281/zenodo.275645
03DCBC11C242FFA10ED21E10FCAD66A8.taxon	materials_examined	Type material. Holotype (UF 383672, shell, mounted soft parts with radula removed, mounted radula). Four paratypes (UF 424420, fluid preserved shells with preserved bodies), from type locality. Type locality. 991 – 1015 m, off Northern Peru [approximately 175 km WNW of Trujillo], Peru-Chile Trench, Peru, 7.983 ˚ S, 80.617 ˚ W.	en	Geiger, Daniel L., Mclean, James H. (2010): New species and records of Scissurellidae and Anatomidae from the Americas (Mollusca: Gastropoda: Vetigastropoda). Zootaxa 2356: 1-35, DOI: 10.5281/zenodo.275645
03DCBC11C242FFA10ED21E10FCAD66A8.taxon	etymology	Etymology. Named after its provenance from off Peru.	en	Geiger, Daniel L., Mclean, James H. (2010): New species and records of Scissurellidae and Anatomidae from the Americas (Mollusca: Gastropoda: Vetigastropoda). Zootaxa 2356: 1-35, DOI: 10.5281/zenodo.275645
03DCBC11C242FFA10ED21E10FCAD66A8.taxon	description	Description. Shell medium size for genus (up to 3.0 mm), trochoid globular. Protoconch of 0.75 whorls, reticulate sculpture, (see arrows on Fig. 20) condition of apertural varix and shape of apertural margin unknown. TI of 1 whorl, approximately 42 fine axial cords, spiral cord in position of selenizone. TII of 2.5 whorls, suture impressed, below periphery of previous whorl. Shoulder convex, approximately 43 axial cords on first whorl, spiral lines starting at onset of selenizone, approximately 10 after 1 whorl, 12 at apertural margin of holotype, slightly irregularly spaced between suture and selenizone. Base with slight constriction below selenizone, 1.5 times as many axials cords as on shoulder, approximately 28 spiral lines stronger than on shoulder, forming points at intersection with axials. Umbilicus narrow, weak funiculus. Aperture D-shaped with columellar portion almost straight. Selenizone slightly above periphery; keels moderately strong, moderately elevated; lunules irregular, distinct; slit with parallel margins. Operculum covering aperture, corneous, thin, round, multispiral, nucleus central. Radula: rachidian tooth triangular, cusp with central denticle larges, 4 – 5 denticles on each side, decreasing in size laterally. Lateral teeth 1 – 3 similar, decreasing in size two fold, 4,4,3 denticles on outer edge of cusp, respectively. Lateral tooth 4 reduced, hook-shaped. Lateral tooth 5 enlarged by broadening, cusp with penultimate denticle largest, one smaller one on outer edge, five on inner edge, decreasing in size towards base. Marginal teeth hook-shaped with many small denticles on each side. Radular interlock of central field moderate. Body pale cream in color. Head with non-papillate cephalic tentacles (ct), no eyestalk or eye, suboptic tentacle (ct) present, accessory cephalic tentacle (act). Four epipodial tentacles (et), biserially papillate on anterior and posterior edge, apical papillary tuft. Possible epipodial sense organ (ESO) on ventral base of second epipodial tentacle [or base of broken secondary ramus of epipodial tentacle?]. Opercular attachment (oa) with scale-like muscle fibres. Two mantle tentacles at posterior convergence of mantle slit. Differential diagnosis. This is the first record of any anatomid from the Peruvian coastline. It differs from T. baxteri from the Pacific Northwest by the turreted shape (lenticular in T. baxteri), TI of 1 whorl (0.66 in T. baxteri), and the less distinct axial cords (raised lamellae in T. baxteri). The comparison is based on examination of the holotype of T. baxteri by SEM, plus conspecific material as identified by comparison to holotype. Thieleella kelseyi from the Pacific North-West shares the overall trochoid-turreted shape, and the overall sculptural pattern. The difference in TI whorls (0.66 in T. peruviana, 0.75 in T. kelseyi) is not diagnostic as intraspecific variability is known to be ± 0.125 whorls (Geiger 2003). The overall shell shape is less turreted in T. kelseyi, with the suture being only one width of the selenizone below the selenizone, whereas in T. peruviana that space is up to three times as wide. Although the position of the suture descends with growth, the distinction holds particularly for larger specimens (3 mm). The axials are about twice as dense in T. kelseyi, particularly towards the apertural margin of large specimens (3 mm). Furthermore, the radula shows some striking dissimilarities, particularly in the shape of lateral tooth 5. While it is broad with five denticles on inner margin in T. peruviana, it is more elongated with approximately 12 denticles on the inner margin in T. kelseyi. Although the individual distinctions may each appear slight, in conjunction they allow for a clear separation of the two taxa. Thieleella bathypacifica n. sp. from deep water west of Central America, has on TI a strong spiral cord in the position of the selenizone where the shell profile shows a distinct angle and has fewer (15 vs. 42) and stronger axial cords on TI, while T. peruviana has only a fine spiral line, does not show an angle in the shell’s profile, and has more (42 vs. 15) and finer axial cords on TI.	en	Geiger, Daniel L., Mclean, James H. (2010): New species and records of Scissurellidae and Anatomidae from the Americas (Mollusca: Gastropoda: Vetigastropoda). Zootaxa 2356: 1-35, DOI: 10.5281/zenodo.275645
03DCBC11C247FF9F0ED21EA5FDA564A6.taxon	materials_examined	Type material. Holotype (FMNH 307884), shell, operculum and radula mounted on SEM stubs. RV Atlantis, submersible grab, leg. J. Voight. Type locality. 2572 m, ALVIN Dive 3938, Genesis, East Pacific Rise, 12.811 ˚ N, 103.940 ˚ W [approximately midway between Acapulco, Mexico, and Clipperton Island]. Sample with anemone and serpulid rocks.	en	Geiger, Daniel L., Mclean, James H. (2010): New species and records of Scissurellidae and Anatomidae from the Americas (Mollusca: Gastropoda: Vetigastropoda). Zootaxa 2356: 1-35, DOI: 10.5281/zenodo.275645
03DCBC11C247FF9F0ED21EA5FDA564A6.taxon	etymology	Etymology. Bathy- referring to the deep-sea (bathyal) habitat of the species; - pacifica referring the Pacific Ocean; adjective.	en	Geiger, Daniel L., Mclean, James H. (2010): New species and records of Scissurellidae and Anatomidae from the Americas (Mollusca: Gastropoda: Vetigastropoda). Zootaxa 2356: 1-35, DOI: 10.5281/zenodo.275645
03DCBC11C247FF9F0ED21EA5FDA564A6.taxon	description	Description. Shell trochiform globular to 1.85 mm. Protoconch of 0.75 whorls, reticulate sculpture, no apertural varix, apertural margin straight. TI of 0.875 whorls, 15 indistinct axial cords, prominent spiral cord forming ridge in position of selenizone, at intersection forming thickenings. TII of at least 1.5 whorls. Shoulder convex, 26 distinct raised axial cords, first spiral line after 0.33 TII whorls, 7 spiral lines after 1.5 whorls on outer two thirds of shoulder, spiral lines running over axial cords. Base convex, barely constricted below selenizone, evenly curving into umbilicus, same density of axial cords as on shoulder, crossed by spiral lines, 23 after 1.25 TII whorls, spiral lines running over axial cords. Selenizone at periphery, keels of moderate strength, high (higher than width of selenizone), regular growth lunules; slit open, with parallel margins. Operculum round, thin, multispiral, central nucleus, covering aperture. Radula (Fig. 20). Rachidian trapezoid, central denticle often largest, approximately five denticles on each side (Fig. 20 A – C: R). Lateral teeth 1 – 3 similar, 6,4,4 cusps respectively (Fig. 20 B – C: L 1 – 3). Lateral tooth 4 reduced, hook-shaped, apical denticle largest, two small denticles on inner cutting edge (Fig. 20 C: L 4). Lateral tooth 5 enlarged by broadening, inner edge with 4 – 5 denticles, outer edge with three denticles (Fig. 20 B – C: L 5). Inner marginal teeth triangular with approximately 3 – 4 denticles on each side (Fig. 20 D – E); with posterior projection on upper shaft (except marginal tooth 1: Fig. 20 C: M 1); outer marginal teeth spoon or paddle shaped with many fine denticles along apical edge (Fig. 20 E – F). Differential diagnosis. Thieleella baxteri from the northeastern Pacific has elevated lamellar axial sculpture. Anatoma janetae Geiger, 2006, from deep water of the eastern Pacific has a protoconch with flocculent sculpture in spiral orientation, the spiral cord on TI does not form a marked angulation, and shows a marked change of sculpture from predominant axial elements to exclusively spiral elements after 1.25 TII whorls. Thieleella peruviana from deep water off Peru, on TI has only a week spiral cord not forming an angulation on the shell’s profile, and has more (42 vs. 15) and weaker axial cords on TI.	en	Geiger, Daniel L., Mclean, James H. (2010): New species and records of Scissurellidae and Anatomidae from the Americas (Mollusca: Gastropoda: Vetigastropoda). Zootaxa 2356: 1-35, DOI: 10.5281/zenodo.275645
03DCBC11C247FF9F0ED21EA5FDA564A6.taxon	discussion	Remarks. The only specimen at hand is most likely not fully mature, because the final quarter whorl does not markedly descend along the coiling axis. The specimen is sufficiently distinct to warrant description as new, and material from that depth is unlikely to be collected in the foreseeable future. The specimen was preserved with the body in the shell, hence, was collected live in its natural deep-sea habitat. The radula of the species shows a thus far unknown detail, namely the posterior projection on the apical portion of the shaft of the marginal teeth from marginal tooth 2 onwards. It is comparable to the food groove known from Tegula funebralis (A. Adams, 1855) and Liotiidae (see Morris & Hickman 1981, Hickman & McLean 1990). The occurrence of such a food groove in thus far a single species of Anatomidae demonstrates further the by now fairly well-documented variability of the radula within Anatomidae (Geiger 2006 a, Geiger & Sasaki 2008, Sasaki et al. in press).	en	Geiger, Daniel L., Mclean, James H. (2010): New species and records of Scissurellidae and Anatomidae from the Americas (Mollusca: Gastropoda: Vetigastropoda). Zootaxa 2356: 1-35, DOI: 10.5281/zenodo.275645
