taxonID	type	description	language	source
03DC87CAFF8DFFD2FF2F13F8FF12FC85.taxon	description	Description. Adult, winged, slender and elongated. Female, defined on the basis of the large last sternite as wide as last tergite and apically sinuous and concave in the middle. Body length: 3.7 mm; elytra: 3.1 mm; antennae: approximately 2.8 mm. Entirely dark brown including legs and antennae. Head almost completely exposed, rounded, with thin and shallow punctation. Eyes rounded, convex, inserted laterally to the head, inter-ocular dorsal distance about 2.4 times greater than eye diameter. Mandibles not visible. Maxillary palps 4 - segmented, unequal in length, with the last palpomere securiform. Labial palps 3 - segmented, last segment securiform. Antennae filiform, 11 - segmented and with all segments pubescent, relative short, surpassing half of elytra; antennomere I (scape) robust, elongated, club – shaped; antennomere II robust, short, 1.7 times shorter than scape; antennomeres III – V elongated, robust (particularly the third), slightly shorter than first article; antennomeres VI – IX filiform, subequal, thinner and around 0.1 – 0.2 times shorter than previous three articles; antennomere X filiform, approximately 0.3 times shorter than previous one; antennomere XI filiform as long as antennomeres VI – IX. Pronotum larger basally, trapezoidal, larger than head, posterior corners rounded, posterior margin strongly bordered, anterior margin and sides slightly bordered, sides concave at half length, surface irregular and equipped with short, thick pubescence. Scutellum subquadrate with apex truncate and straight. Elytra slender, parallel-sided, very elongated and completely covering and slightly surpassing the last abdominal segments, wider than pronotum, apex rounded, surface rugose and equipped with many short setae. Posterior wings present, almost completely covered by elytra. Metasternum dark brown, rounded posteriorly, equipped with setae; sternites short, transverse, wrinkled; last sternite wide, rounded and sinuous at apex with central concavity. Legs short and pubescent; coxae massive; trochanters rounded; femora slightly enlarged; tibiae thin, cylindrical, longer than femora. All tarsi 5 - segmented; first segment elongated; second tarsomere elongated and longer than first tarsomere; third about 2.3 times shorter than second, straight at apex; fourth tarsomere bilobed at sides; fifth flat and elongated; claws apparently bifid at apex without denticles at base.	en	Fanti, Fabrizio, Pankowski, Mary K. (2018): Three new species of soldier beetles from Baltic amber (Coleoptera, Cantharidae). Zootaxa 4455 (3): 513-524, DOI: 10.11646/zootaxa.4455.3.7
03DC87CAFF8DFFD2FF2F13F8FF12FC85.taxon	etymology	Etymology. Species named after Mary L. Pankowski, paternal grandmother of the second author, who provides immense love to her family, supports her community in numerous ways and inspires all with her indomitable spirit of optimism.	en	Fanti, Fabrizio, Pankowski, Mary K. (2018): Three new species of soldier beetles from Baltic amber (Coleoptera, Cantharidae). Zootaxa 4455 (3): 513-524, DOI: 10.11646/zootaxa.4455.3.7
03DC87CAFF8DFFD2FF2F13F8FF12FC85.taxon	materials_examined	Holotype. Female, adult specimen included in Baltic amber, accession No. USNM PAL 712534 in the USNM. Type locality. Yantarny settlement (formerly Palmnicken), Sambian (Samland) Peninsula, Kaliningrad Region, Russia. Type strata. Baltic amber, Upper Eocene, Prussian Formation (Priabonian). Estimated age: 37.8 – 33.9 Mya. Syninclusions. Wood remains.	en	Fanti, Fabrizio, Pankowski, Mary K. (2018): Three new species of soldier beetles from Baltic amber (Coleoptera, Cantharidae). Zootaxa 4455 (3): 513-524, DOI: 10.11646/zootaxa.4455.3.7
03DC87CAFF8DFFD2FF2F13F8FF12FC85.taxon	diagnosis	Differential diagnosis. The new species belongs to the genus Rhagonycha Eschscholtz, 1830 based on its bifid claws and for its anteriorly narrowed pronotum. Two species of the genus Rhagonycha have previously been described from inclusions in Baltic amber: R. kryshtofovichi (Yablokov-Khnzorian, 1960) and R. sucinobaltica (Poinar & Fanti, 2016). Rhagonycha kryshtofovichi initially attributed to the genus Malchinus Kiesenwetter, 1863 was then transferred to Rhagonycha by Kazantsev (2013) based on the anteriorly narrowed pronotum, the fourth palpomere securiform and unmodified terminalia. In the genus Rhagonycha males and females possess the same pronotal shape. Thus, because the pronotal shape of R. maryae sp. nov. is different from R. kryshtofovichi and R. sucinobaltica, it can be distinguished as a new species. (It’s worth noting here that the specimen of R. maryae sp. nov. is a female because its last sternite is as wide as its last tergite, and it has an apical margin sinuous and concave in the middle. In the males of Rhagonycha, the last tergite is triangular and narrower than the last tergite.) R. kryshtofovichi is easily distinguishable from R. maryae sp. nov. based on its larger size (with a body length of 7 mm compared to 3.7 mm for R. maryae), different pronotum and different length of the antennomeres (Yablokov- Khnzorian 1960). While the other known species R. sucinobaltica Poinar & Fanti, 2016 has similar dimensions (a body length of 3.5 mm), it possesses a different pronotum that is more curved at the sides and less bordered at the posterior margin (Poinar & Fanti 2016). Rhagonycha maryae sp. nov. also differs from other inclusions in Baltic amber known at a generic level as well as specimens known from mold impression and compression fossils in various rocky sediments (Fanti 2017 a).	en	Fanti, Fabrizio, Pankowski, Mary K. (2018): Three new species of soldier beetles from Baltic amber (Coleoptera, Cantharidae). Zootaxa 4455 (3): 513-524, DOI: 10.11646/zootaxa.4455.3.7
03DC87CAFF8DFFD2FF2F13F8FF12FC85.taxon	discussion	Remarks. The amber piece measures 19 x 19 x 4 mm and weighs 0.8 grams. It has a round shape and is rather flat.	en	Fanti, Fabrizio, Pankowski, Mary K. (2018): Three new species of soldier beetles from Baltic amber (Coleoptera, Cantharidae). Zootaxa 4455 (3): 513-524, DOI: 10.11646/zootaxa.4455.3.7
03DC87CAFF8FFFD7FF2F13BCFD93FD80.taxon	description	Description. Adult, winged, elongated, rather robust. Male, defined on the basis of the last sternite very small, narrow, rounded at apex, and separated and away from the last tergite. Body length: 4.5 mm; elytra: 3.3 mm; antennae: approximately 2.5 mm. Head black, elytra blackish to dark brown, pronotum with antennae and legs reddish to dark brown. Head very elongated and narrow, prognathous, completely exposed, wrinkled with shallow punctation and short setae. Eyes subelliptical, convex, inserted dorsally and near the sides of the head. Mandibles robust, elongated, slightly falciform, without teeth or denticles. Maxillary palps 4 - segmented and unequal in length; first palpomere massive; second palpomere flat, robust, rounded and enlarged in the middle; third small, thin and short; last palpomere securiform, elongated, rounded externally, apically thin but with a bulbous tip. Labial palps 3 - segmented, very elongated; first segment massive; second segment thin; last segment elongated, very slightly securiform with the apex robustly pointed. Antennae 12 - segmented, inserted dorsally and between the eyes, very short, reaching and slightly surpassing the humeral zone, all segments pubescent along all margins; antennomere I (scape) filiform and robust, elongated, not club-shaped, slightly enlarged apically; antennomere II robust, short, 2.0 times shorter than scape, concave in the middle on the external side; antennomere III robust, slightly shorter than second article and not concave in the middle; antennomeres IV – V subequal, very slightly shorter than antennomere III, enlarged apically (dentate) on the external side; antennomere VI elongated, robust, approximately 1.5 times longer than antennomeres IV – V and not dentate on the external side; antennomere VII robust and the shortest, concave in the middle on the internal side; antennomere VIII filiform but slightly dentate at the external apex; antennomeres IX – XII filiform, unequal in length with the antennomere XII rounded at apex and the longest. Pronotum subquadrate, slightly longer than wide, very slightly wider than head, corners rounded, margins bordered, in the middle of the pronotum there is a large and very raised (evident) area slightly concave in the center, sides and margins flat after the raised area with shallow punctation and short pubescence. Scutellum not well visible. Elytra very elongated, slender, parallel-sided, completely covering the last abdominal segments, wider than pronotum, apex rounded, surface equipped with long and erected setae. Posterior wings completely covered by elytra. Metasternum blackish, punctate, elongated, posteriorly slightly rounded; sternites and tergites wide, punctate; last tergite short, wide and strongly bent at sides; last sternite small, narrow, rounded at apex. Anterior and median legs short, posterior legs longer; coxae massive, wide at base, elongated; trochanters elongated, posteriorly triangular; profemora strongly enlarged and slightly curved, meso- and metafemora thin; pro- and mesotibiae shorter than pro- and mesofemora, cylindrical, thin, with two spurs (one very short) and setae; metatibiae longer than metafemora, cylindrical, thin, with spurs and setae. Tarsi 5 - segmented and with setae; pro – and mesotarsi with the first segment elongated, 1.5 times longer than second; metatarsi with the first tarsomere approximately 1.9 times longer than second; third tarsomere shorter than second and slightly enlarged apically; fourth tarsomere bilobed at sides with the lobes very long, curved and rounded at apex; fifth tarsomere elongated, thin; claws simple, long and without lobes or denticles.	en	Fanti, Fabrizio, Pankowski, Mary K. (2018): Three new species of soldier beetles from Baltic amber (Coleoptera, Cantharidae). Zootaxa 4455 (3): 513-524, DOI: 10.11646/zootaxa.4455.3.7
03DC87CAFF8FFFD7FF2F13BCFD93FD80.taxon	etymology	Etymology. Species named after Frank and Joanne Samsock, maternal grandparents of the second author, who worked hard, dedicated their lives to their family and were excellent role models for their children and grandchildren.	en	Fanti, Fabrizio, Pankowski, Mary K. (2018): Three new species of soldier beetles from Baltic amber (Coleoptera, Cantharidae). Zootaxa 4455 (3): 513-524, DOI: 10.11646/zootaxa.4455.3.7
03DC87CAFF8FFFD7FF2F13BCFD93FD80.taxon	materials_examined	Holotype. Male, adult specimen included in Baltic amber, accession No. USNM PAL 712533 in the USNM. Type locality. Yantarny settlement (formerly Palmnicken), Sambian (Samland) Peninsula, Kaliningrad Region, Russia. Type strata. Baltic amber, Upper Eocene, Prussian Formation (Priabonian). Estimated age: 37.8 – 33.9 Mya. Syninclusions. Many wood remains and masses of plant fragments (detritus).	en	Fanti, Fabrizio, Pankowski, Mary K. (2018): Three new species of soldier beetles from Baltic amber (Coleoptera, Cantharidae). Zootaxa 4455 (3): 513-524, DOI: 10.11646/zootaxa.4455.3.7
03DC87CAFF8FFFD7FF2F13BCFD93FD80.taxon	diagnosis	Differential diagnosis. Antennae with 12 antennomeres and the claws without lobes of the new species suggest that it belongs to the genus Sucinorhagonycha. (For more details, see the key below.) The genus is easily distinguishable from the fossils of the family Cantharidae Imhoff, 1856 with its 12 - segmented antennae present only in Cacomorphocerus Schaufuss, 1892 (for the differences, see Discussion section below) and the extant Australian genus Heteromastix Boheman, 1858 (Subfamily Dysmorphocerinae Brancucci, 1980), which also sometimes has 12 antennal articles (Brancucci 1980; Fanti 2017 c), with the antennomeres X – XI strongly modified (Brancucci 1980; Kuśka 1996). Only one other species of Sucinorhagonycha has been described, S. kulickae Kuśka, 1996, which is known on the basis of a male and a female found in different Baltic amber pieces (Kuśka 1996; Kubisz 2000; Fanti 2017 a). Sucinorhagonycha samsockorum sp. nov. differs from both sexes of S. kulickae because its antennal articles IV – V are slightly dentate (compared to filiform in S. kulickae), and because its pronotum is not transverse and has two evident central thickenings (while in S. kulickae the pronotum has a longitudinal central linear depression and protuberances on the sides). In addition, the male of S. kulickae has shorter antennomeres than S. samsockorum sp. nov.	en	Fanti, Fabrizio, Pankowski, Mary K. (2018): Three new species of soldier beetles from Baltic amber (Coleoptera, Cantharidae). Zootaxa 4455 (3): 513-524, DOI: 10.11646/zootaxa.4455.3.7
03DC87CAFF8FFFD7FF2F13BCFD93FD80.taxon	discussion	Remarks. The yellow amber piece has a spherical shape measuring 15 x 14 x 11 mm and weighs 1.3 grams. The inclusion is complete and well visible.	en	Fanti, Fabrizio, Pankowski, Mary K. (2018): Three new species of soldier beetles from Baltic amber (Coleoptera, Cantharidae). Zootaxa 4455 (3): 513-524, DOI: 10.11646/zootaxa.4455.3.7
03DC87CAFF8AFFD9FF2F13C8FC84FC85.taxon	description	Description. Adult, winged. Male, based on the last abdominal segments modified and the long antennae. Body length: 2.8 mm; elytra: 1.2 mm; antennae: 2.2 mm. Entirely dark brown including legs and antennae, without yellow spots on elytra. Head completely exposed, strongly rounded, larger than pronotum, equipped with shallow punctation. Eyes rounded, very large and prominent, inserted laterally to the head. Mandibles elongated, falciform, large at base and thin apically, concave on the internal side. Maxillary palps 4 - segmented, unequal in length, with the terminal palpomere globular and distally pointed. Labial palps 3 - segmented with the last segment globular and pointed. Antennae filiform, 11 - segmented, long, surpassing the elytra, not reaching the last abdominal segments; antennomere I (scape) elongated, not club-shaped; antennomeres II – III filiform and approximately 1.4 times shorter than scape; antennomeres IV – X elongated, filiform, subequal in length, slightly longer than antennomeres II – III; antennomere XI thinner and longer than previous one, apically rounded; antennomeres with small setae. Pronotum large, transverse, surface undulating and with shallow punctation, posterior margin straight, anterior margin strongly bordered, sides sinuous, corners rounded. Scutellum triangular-shaped. Elytra wider than pronotum, short, slender, parallel-sided, reaching until the fifth abdominal segment, wide at base and narrower posteriorly to the humeral zone, apex rounded, surface pubescent. Posterior wings wide and long, dark, surpassing the elytra and the abdominal segments. Anterior and median legs very short, posterior legs long; coxae robust; trochanters elongated; femora enlarged; pro- and mesotibiae as long as pro- and mesofemora, metatibiae longer than metafemora, all the tibiae cylindrical and very thin. Tarsi 5 - segmented; tarsomere I long and more of two times longer than second; tarsomere III slightly shorter than second; tarsomere IV bilobed; tarsomere V slender; claws simple. Penultimate tergite (tergite 9) elongated and slightly bent at sides; last tergite (tergite 10) short and wide with the sides bent, apically the margin is straight; last sternite (sternite 9) long and flat, large at base and curved after few millimeters, from which start two elongated and separated lobes with rounded apex; two wide and irregular with rounded apex urophysis, each inserted by the penultimate sternite (sternite 8) and located on both sides of the sternite 9. Sternal surface of abdomen punctate and with pubescence.	en	Fanti, Fabrizio, Pankowski, Mary K. (2018): Three new species of soldier beetles from Baltic amber (Coleoptera, Cantharidae). Zootaxa 4455 (3): 513-524, DOI: 10.11646/zootaxa.4455.3.7
03DC87CAFF8AFFD9FF2F13C8FC84FC85.taxon	etymology	Etymology. Species named after Joseph M. Pankowski, paternal grandfather of the second author, who always puts his family first, is generous beyond words and who is a friend to all.	en	Fanti, Fabrizio, Pankowski, Mary K. (2018): Three new species of soldier beetles from Baltic amber (Coleoptera, Cantharidae). Zootaxa 4455 (3): 513-524, DOI: 10.11646/zootaxa.4455.3.7
03DC87CAFF8AFFD9FF2F13C8FC84FC85.taxon	materials_examined	Holotype. Male, adult specimen included in Baltic amber, accession No. USNM PAL 712536 in the USNM. Type locality. Yantarny settlement (formerly Palmnicken), Sambian (Samland) Peninsula, Kaliningrad Region, Russia. Type strata. Baltic amber, Upper Eocene, Prussian Formation (Priabonian). Estimated age: 37.8 – 33.9 Mya. Syninclusions. Wood remains and air bubbles.	en	Fanti, Fabrizio, Pankowski, Mary K. (2018): Three new species of soldier beetles from Baltic amber (Coleoptera, Cantharidae). Zootaxa 4455 (3): 513-524, DOI: 10.11646/zootaxa.4455.3.7
03DC87CAFF8AFFD9FF2F13C8FC84FC85.taxon	diagnosis	Differential diagnosis. Characters such as the globular and pointed last palpomere, the short elytra and the strongly modified last abdominal segments (both tergites and sternites) suggest that the new species belongs to the genus Malthodes. No other fossil of Malthodes shows the same features of the terminalia of the new species (Fanti & Vitali 2017; Fanti & Michalski 2018), and no extant species of Central-North Europe and Alps is similar. Only the group of M. alpicola Kiesenwetter, 1852 (M. alpicola, M. guttifer Kiesenwetter, 1852, M. spretus Kiesenwetter, 1852, M. bertolinii Fiori, 1905) might be seen as similar, although not from a strictly anatomical point of view, because of the last sternite (sternite 9) that is shaped as a wide, apically forked membrane (Liberti 2011, 2015, 2016) and possessing a similar last tergite (tergite 10). However, these species are much bigger in size, and tendentially have a yellow spot (more or less evident) on each elytral apex. The group of M. minimus (Linnaeus, 1758) is only vaguely similar, with the last tergite lobe-shaped and little developed, and the last sternite without median membrane. The important diagnostic characters of Malthodes are based on the male’s terminalia and habitus, as the morphology of the fossils is nearly identical to the living species. It is, however, interesting to note that, out of all the fossil species known, only M. perkovskyi Kazantsev from Rovno amber (Kazantsev 2010) has the elytra with two yellow spots apically (Förster 1891; Kuśka & Kupryjanowicz 2005; Kuśka & Kania 2010; Kazantsev & Perkovsky 2014; Fanti 2017 b; Fanti & Vitali 2017; Fanti & Michalski 2018). These are indeed present, even if sometimes difficult to see, in many living species (Fender 1951; Wittmer 1970; Liberti 2011, 2015, 2016, 2017).	en	Fanti, Fabrizio, Pankowski, Mary K. (2018): Three new species of soldier beetles from Baltic amber (Coleoptera, Cantharidae). Zootaxa 4455 (3): 513-524, DOI: 10.11646/zootaxa.4455.3.7
03DC87CAFF8AFFD9FF2F13C8FC84FC85.taxon	discussion	Remarks. The amber piece measures 28 x 22 x 10 mm, is rectangularly shaped and weighs 2.8 grams. The amber is opaque yellow and there are scratches on the surface.	en	Fanti, Fabrizio, Pankowski, Mary K. (2018): Three new species of soldier beetles from Baltic amber (Coleoptera, Cantharidae). Zootaxa 4455 (3): 513-524, DOI: 10.11646/zootaxa.4455.3.7
