taxonID	type	description	language	source
03DFBE05E01BFFF8FF64D265FA21FB66.taxon	description	http: // zoobank. org / E 923 F 896 - 4 AFE- 4006 - 86 B 6 - D 8 FF 1 D 15 EED 6	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E01BFFF8FF64D265FA21FB66.taxon	description	Genotype. Beckhoplia colvillei Dombrow, 2005: 128, original designation by Dombrow (2005). Main characters. Clypeus broader than long, longitudinally depressed laterally, smooth. Margins of clypeus linear and distinctly raised. Pronotum broader than long, punctate with shallow, roundish punctures. Elytra mostly ovate, partly covering to protruding the propygidium. Pygidium vertical, not reaching metathoracic coxae. Prothoracic tibia tridentate. All tibiae with spur. All claws double and vertically split, external claw of metathoracic tarsus not split. Differential diagnosis. Differs from Anisochelus Burmeister, 1844 (Fig. 138). Clypeus of Beckhoplia Dombrow, 2005 (Fig. 1) distinctly longitudinally depressed along lateral margin. Clypeus of Anisochelus (Fig. 2) complanate. Margins of clypeus of Beckhoplia distinctly vertically raised. Margins of clypeus of Anisochelus slightly vertically raised. Main surface of clypeus of Beckhoplia smooth and bare. Main surface of clypeus of Anisochelus dense punctate and loose setose. Parameres of Beckhoplia (Figs. 62 – 97, 100 – 117). Parameres of Anisochelus (Figs. 98 – 99). Redescription Integument: Black; funiculum sometimes pale-brown or seldom brown annulated; elytra mostly transparently palebrown or brown, with more or less blackened margin; very seldom black or yellowish pale-brown. Tegument: Clypeus bare; sparse to loose setose with erect, pale-brown to black setae on the inner side of the vertically reflexed anterior and lateral margin; the latter very seldom bare. Head mostly loose setose with erect, black setae, very seldom whitish or pale-brown setae interspersed. Eye canthus with a loose to dense setose ciliary of black setae. Pronotum without pattern; loose setose with erect, black setae; very seldom whitish or brown setae, sometimes interspersed, very seldom lengthening anteriorly or seldom declining posteriorly; with a loose to dense setose ciliary of black, sometimes whitish setae on lateral margin, very seldom declining posteriorly. Scutellum loose setose with semi-erect to erect, brown to black setae, sometimes shortening apically; with a longitudinal bare strip of various length, sometimes missing. Elytra with or without pattern; mostly sparse to loose setose with semierect or erect, seldom reclinate, brownish or black setae, sometime lengthening anteriorly and / or setae of various length interspersed; seldom whitish setae, scales or squamiform setae interspersed around apical torus; with one or two loose setose longitudinal rows of semi-erect or erect, black setae. If patterned, then additionally with two or three shortened, longitudinal, loose to dense squamose bands of reclinate, white and / or very seldom grey scales. Suture with a loose row of robust, semi-erect to erect, black setae, sometimes lengthening apically. Thorax and pleurites loose, very seldom sparse pilose with long, black, seldom white hairs. Ventrite I – V mostly loose, sometimes sparse setose with reclinate or semi-erect, brownish or black, seldom white setae; very seldom dense squamose with lanceolate, transparently white scales; semi-erect, black setae interspersed. Propygidium loose setose or loose squamose. If setose, then with semi-erect, seldom erect, black, very seldom pale-brown or whitish setae, very seldom squamiform setae; semi-erect, black setae or reclinate, white setae interspersed; sometimes posterior margin with a loose row of reclinate, whitish scales or white setae. If squamose than with reclinate, pale-brown and / or whitish or greyish scales, semi-erect, black setae interspersed. Pygidium mostly sparse setose with erect, black setae, very seldom whitish scales interspersed in upper quarter; tip always with a ciliary of black setae. Prothoracic femur mostly loose pilose, with semi-erect, black, very seldom white or brown hairs; seldom loose setose with semi-erect, black or brown setae; always bare internally. Prothoracic tibia mostly loose setose with reclinate, sometimes and / or semi-erect, pale-brown, whitish or black setae; can be sparser ventrally; sometimes semi-erect or erect white, brown or black setae interspersed; with a sparse setose longitudinal row of erect, black, seldom brown setae dorsally; a loose setose ciliary of black, seldom additionally white setae on internal ridge; sometimes additionally with a sparse, seldom denser setose ciliary of mostly white setae on external ridge; a setaceous ciliary of somewhat brown or black setae on anterior margin; mostly more robust towards internal ridge; a downwards pointing setose ciliary of brownish setae along anterior margin of apical tooth. Mesothoracic femur loose pilose with black, whitish and / or brown hairs; lengthening and / or brownish or white ventrally; seldom erect, black setae interspersed dorsally and externally; always bare internally. Posterior margin with a marginal ciliary of black, very seldom red-brown bristles, sometimes lengthening dorsally, very seldom ventrally; always missing internally. Mesothoracic tibia mostly sparse setose with reclinate or semi-erect, whitish or seldom brownish setae, sometimes sparser internally; semi-erect, robust, black setae interspersed; seldom sparser and normally finer internally. Metathoracic femur loose setose with mostly semi-erect, black, sometimes white setae than black setae interspersed externally; sometimes whitish or brownish ventrally, seldom black hairs interspersed; bare internally. Metathoracic tibia sparse setose with semi-erect, brownish, infrequent black setae; semi-erect, sometimes robust, black setae interspersed; mostly finer internally, sometimes denser. Posterior margin with a normally loose bristled outer marginal ciliary of black bristles, lengthening dorsally and ventrally, frequently more robust ventrally. Tarsomere I – IV of all thoracic legs sparse setose with reclinate, mostly brownish, very seldom black and / or greyish setae or prothoracic and / or mesothoracic tarsomere I – IV bare. Tarsomere I – IV of all thoracic legs with a setaceous, apical, marginal ciliary of black, seldom brownish setae, lengthening towards metathoracic tarsomeres, fine dorsally and more robust ventrally. Pretarsomere of all thoracic legs sparse setose with reclinate or semi-erect, seldom only with a few, brownish or black setae, sometimes lengthening, very seldom also darkening towards metathoracic pretarsomere; mostly with few, semi-erect, black setae at apical margin, shorter and more or less red-brown ventrally. Head: Clypeus broader than long; corner angled; coplanar with frons; distinctly longitudinally depressed laterally; as broad as frons; surface smooth, lustrous; inner surface of reflexed margins punctate with micropunctures. Anterior margin linear; very distinctly vertically reflexed (Fig. 6) and higher than lateral margin; upper edge concave, curly bracket-shaped or broad v-shaped (Figs. 3 – 5) incised (variable character within the species and between the sexes; not used in the species descriptions), following sculpture of clypeus; with a small, triangular-shaped, slightly rounded tooth at each corner. Lateral margin very slightly convex; distinctly reflexed; not merging with eye-canthus; relation to width of head very broad. Suture between head and clypeus indistinct, undulate or linear, complanate seldom crenated or raised, very seldom missing or interrupted medially. Head dense punctate with shallow, very small or small, roundish, sometimes partly merging punctures and sometimes micropunctate within; surface and lateral margin complanate. Eye canthus (Figs. 11 – 14) very long, broad, complanate. Eye above (Figs. 8 – 10) head level. Antenna: Consisting of nine segments (Fig. 15). The long cudgel-shaped scapus and the ± spherical pedicellus followed by a four segmented funiculum: funicle I – III of different shape, funicle IV very short asymmetric frustumshaped and an always three segmented lamellar elliptic to long elliptic clavus (Figs. 16 – 18). Labrum and epipharynx: Labrum strongly sclerotized, vertical, horizontally oriented (Fig. 19); zona lateralis and zona centralis bare; apical margin concave; apical ciliary dense pilose with erect, fine hairs, missing medially; lateral margin distinctly convex; anterior half with a dense setose ciliary of erect, robust setae; clypeo-labral suture linear dorsally, concave ventrally. Epipharynx membranaceous; posterior medial process narrow, complanate; with a dense setose comb, stretching v-shaped from apical corner along border between zona lateralis and zona centralis conceding in front of posterior tormal process, terminating at posterior margin of posterior tormal process; lateral tormal process long, bare. Mandibles: Paired, (Fig. 20) stout, symmetric, sparse setose externally. Lacinia mobilis: narrow, membranaceous, dense pilose towards mola; with a more or less sclerotized tooth. Mola: prominent, sclerotized, dentate; teeth arranged in parallel rows; postmola: membranaceous. Incisivus: stout, sclerotized, somewhat pilose externally. Maxillae: Paired (Fig. 21). Stipes: sparse setose. Maxillar palpus: four-segmented, segment IV bare, more or less thickened, cylindrical, distinctly longer than segment I – III combined; segment III with a ciliary of relatively long setae; segment I – II bare. Galea: sclerotized, with a tuft of hairs above teeth; six-dentate, arranged in two groups: an apical group of three slightly shifted, vertically arranged, elongate, triangular-shaped, pointed teeth; a basal group of three transverse, elongate, pointed, triangular-shaped teeth, grown together in basal three quarters. Lacinia: well developed with a longitudinal, medio-ventral ridge, set with a loose stripe of setae, lengthening from cardo to galea. Cardo: stout, apodeme broadly bi-forked, tip of one joint almost spherical, tip of the second joint ending in three knobs. Labium: Strongly sclerotized, ovate, with a distinct longitudinal medial furrow, longitudinally pulvinate, sparse setose with erect setae. Anterior margin undulate; anterior corner angled with a shortened, loose setose ciliary. Lateral margin distinctly incised in anterior fifth; behind labial pulps distinctly convex, with a loose setose ciliary of longer setae. Posterior margin linear. Labial pulps three-segmented, short and robust; segment III cone-shaped, slightly shorter than segment I – II combined (Fig. 22). Pronotum: Broader than long; widest in anterior two fifth or medially; slightly pulvinate laterally; loose to dense punctate with shallow, roundish punctures, may be denser within medial furrow and towards margins. Anterior margin shallowly concave; anterior corner orthogonal or obtuse, very seldom acute. Lateral margin convex or linearly attenuate anteriorly and subparallel or very seldom slightly convex attenuate posteriorly; surface of lateral margin smooth. With a shortened or complete, shallow, more or less distinct medial furrow. Posterior margin convex; posterior corner obtuse, angled or rounded (Figs. 23 – 24). Scutellum: Buckler-shaped or parabola-shaped (Figs. 25 – 26); longer than broad to broader than long; longitudinally pulvinate, vertically complanate; dense punctate with shallow, roundish punctures, mostly micropunctate within; mostly with a longitudinal, sometimes shortened, more or less broad, median, smooth stripe. Lateral margins slightly convex or linearly attenuate in anterior quarter to two thirds, stronger apically. Tip pointed or rounded (Figs. 27 – 28), not subducting. Elytra: Ovate, seldom oblong (Figs. 29 – 30); longer than broad; widest medially in anterior third or half; protruding or covering, very seldom partly covering the propygidium. Anterior margin slightly concave. Lateral margins slightly convex. Posterior margin rounded and distinctly incised medially. Sculpture more or less distinct; scutellar torus and / or costate sometimes missing; humerus indistinct, complanate; apical torus indistinct; lateral descent distinct; lateral trench sometimes missing; suture raised. Sparse to loose punctate with deep to shallow, roundish sometimes wrinkly punctures, sometimes micropunctures in-between; with two sparse to loose longitudinal rows of big punctures: a juxta-sutural row from anterior margin crossing scutellar torus towards apical torus; and a supra-humeral row from anterior margin between humerus and scutellar torus towards apical torus, sometimes shortened or missing. Thorax: Interspace between mesothoracic coxae narrow. Abdomen: (Fig. 31). Five ventrites visible; not compressed laterally. Ventrite I distinctly to extremely shortened medially; ventrite II – V not shortened medially; ventrite V sometimes broadest and / or slightly constricted ventrally (Figs. 32 – 33). Propygidium with an indistinct or slightly projected stoma at posterior corner. Pygidium vertical, not reaching metathoracic coxae; distinctly broader than long (Fig. 34); slightly to distinctly pulvinate longitudinally and laterally; upper margin slightly to moderately convex, mostly linear medially; lateral margin convex or subparallel in upper fifth or quarter, linearly attenuate apically, or only sometimes linearly attenuate apically (Figs. 36 – 38); tip rounded or linear (Figs. 39 – 40). Prothoracic leg: Femur slender or robust; slightly compressed laterally. Tibia slender or robust; distinctly compressed ventrally; tridentate, teeth robust; with an internal and external ridge (Figs. 45 a, b; 46 a, b); smooth ventrally or with a medio-ventral ridge (Figs. 44, 45 c); loose punctate with shallow, roundish punctures; with a dense longitudinal row of deep, elongate, roundish punctures dorsally (Fig. 41); with spur. Apical tooth elongate, longest, bent forward, outwards, tip slightly pointing downwards. Median tooth elongate, seldom robust, triangular-shaped, slightly shorter than apical tooth, pointing outwards. Basal tooth smallest, sometimes rudimentary. Interspace between teeth almost equal or interspace between anterior and median tooth double in width than between median and basal tooth. Shape of interspace between anterior and median tooth concave or u-shaped and between median and basal tooth concave or linear (Figs. 41 – 43). Articulation point of tarsomere I at base of anterior tooth. Tarsomere I – IV combined extremely short, rarely very short; frustum-shaped; slightly compressed laterally; equal in length or lengthening from tarsomere I – IV (Figs. 47 – 48); almost half in length of tibia. Tarsomere III very rarely with a ventral extension (Fig. 49). Tarsomere IV seldom with a ventral extension (Fig. 49). Pretarsomere slightly to distinctly thickened; smooth or with various structures ventrally (Figs. 50 – 54) followed by a roundish cutting apically; more or less as long as tarsomere I – IV combined. Claw double; both scythe shaped, smooth ventrally, split vertically. Internal claw broadened medially; up to a quarter longer than pretarsomere. External claw nearly three quarters in length of internal claw. Mesothoracic leg: Femur and tibia slender; both distinctly compressed laterally. Tibia cylindrical; slightly convex dorsally; slightly constricted dorsally apically; diagonally truncate; with spur; roughly three quarter in length of femur to slightly longer than femur; loose punctate with shallow and deep, roundish punctures, sparse and shallow internally. Tarsomere I – IV combined extremely short to very short; frustum-shaped; slightly compressed laterally; equal in length; slightly less than 3 / 10 to slightly more than 4 / 5 in length of tibia. Pretarsomere distinctly thickened; smooth or with various structures ventrally, followed by a roundish cutting, sometimes missing (Figs. 55 – 58); roughly one third in length of tibia. more or less as long as tarsomere I – IV combined. Claw double; both scythe shaped, smooth ventrally, split vertically. External claw robust; broadened medially; more or less as long as pretarsomere. Internal claw slender; slightly shorter than external claw. Metathoracic leg: Femur slender; distinctly compressed laterally; without trochanteral spine. Tibia sub-cylindrical; distinctly compressed laterally; almost as long as femur, seldom longer than femur; slightly diagonally truncate; not mucronate; with spur. Loose punctate with shallow, elongate- roundish punctures. Tarsomere I – IV combined distinctly short, very seldom short or very short; frustum shaped; slightly compressed laterally; declining in length from tarsomere I – IV; roughly three quarters in length of tibia. Pretarsomere slender to slightly thickened; slightly compressed laterally; with various structures ventrally (Figs. 59 – 61); almost half in length of tarsomere I – IV combined. Claw double; both smooth ventrally. External claw slender; scythe-shaped; longer than pretarsomere; not split. Internal claw slender; linear in basal two thirds, very slightly bent downwards in apical third; mostly slightly longer than three quarters in length of external claw; split vertically. Sexual dimorphism: Indistinct. Clypeus very slightly depressed laterally; fully punctate; less reflexed (Fig. 7). Eye canthus very seldom full, dividing eye. Clavus of antenna distinctly shorter. Pronotum widest in anterior third to medially (mostly different to the ♂♂ but sometimes equal); posterior corner rounded. Pygidium slightly declined posteriorly. Ventrites distinctly pulvinate (Fig. 35). Tip of pygidium linear. Mesothoracic pretarsomere slender to slightly thickened, Metathoracic tibia distinctly shorter than metathoracic femur; slightly to distinctly extended dorso-apically (variable within species); slightly extended ventro-apically. All thoracic legs including tarsi red-brown and mostly slender.	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E01BFFF8FF64D265FA21FB66.taxon	distribution	Distribution. The genus is endemic to the Northern and Western Cape Provinces of the Republic of South Africa. The centre of its biogeographical distribution falls within the winter-rainfall zone (Fig. 139) at the intersection between the north-western parts of the Fynbos Biome and south-western border of the Succulent Karoo Biome. A northern Succulent Karoo extension is recorded from the central, winter-rainfall, arid Namaqualand region. All species within the genus currently display highly localized distributions confined to specific vegetation type habitats, predominantly those of Renosterveld and succulent Karoo-Shrubland, with a hotspot of diversity centred around the Hantam Bokkeveld Plateau (Fig. 140).	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E011FFE7FF64D126FC62FD48.taxon	description	(Figs. 14, 30, 33, 49, 55, 59, 62 – 63, 100, 119) http: // zoobank. org / 0 A 721 F 95 - 4493 - 47 AD-A 8 A 4 - E 28 D 57 E 6 BE 53	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E011FFE7FF64D126FC62FD48.taxon	materials_examined	Type locality. [RSA, NCP]: nördl. Farm Brandkop, 21 km NO Nieuwoudtville. Type material. [5] Holotype ♂: white label pr « AFRIKA: Südafrika, Kapprovinz | nördl. Farm Brandkop, 21 km NO | Nieuwoudtville, 380 m, 29.08.1989 | 31 ° 14 ’ 30 ” S, 19 ° 12 ’ 08 ” E, leg. R. Fritz » || green label pr « Host plant: | Albuca maxima » || blue label pr « Bioregion: | Trans-Escarpment Succulent Karroo | Vegetation Type: | SKt 2 – Hantam Karroo » || yellow label hw « 54 » pr «. 928 | coll. Dombrow » || red label pr « Holotype | Beckhoplia | nigra sp. n. | det.: H. Dombrow 2019 » (ZSM). – Paratypes 1 ♂, 3 ♀♀: same labels as holotype but || yellow label hw « 54 » pr «. 975 –. 978 | coll. Dombrow » || red label pr « Paratype | Beckhoplia | nigra sp. n. | det.: H. Dombrow 2019 » (ESDW). Condition of holotype. Good. Glued on card. Dissected, parameres and spiculum gastrale glued behind beetle. Left metathoracic leg broken off, glued behind beetle; left metathoracic tarsus missing after tarsomere II.	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E011FFE7FF64D126FC62FD48.taxon	diagnosis	Differential diagnosis. Differs from B. pallidibrunnea: Colour of elytra of B. nigra black. Colour of elytra of B. pallidibrunnea transparently pale-brown. Elytra of B. nigra with squamose pattern of white scales. Elytra of B. pallidibrunnea without scales. Interno-posterior margin of parameres of B. nigra elongate curly bracket-shaped (Figs. 62, 100). Interno-posterior margin of parameres of B. pallidibrunnea parabola-shaped (Figs. 64, 101). Externo-posterior margin of parameres of B. nigra convex (Figs. 62, 100). Externo-posterior margin of parameres of B. pallidibrunnea concave medially (Figs. 64, 101).	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E011FFE7FF64D126FC62FD48.taxon	description	Description Measurements: length 7.5 – 7.6 mm, width 3.7 – 3.9 mm. Integument: Black. Tegument: Clypeus bare; sparse setose with extremely short, erect, pale-brown setae on the inner side of the vertically reflexed anterior and lateral margin. Head loose setose with long, erect, black setae. Eye canthus with a dense setose ciliary of long, black setae. Scapus loose setose with medium-long, black setae, dark-brown ventrally, sparse and shorter on pedicellus, funiculum and clavus sparse setose with very short, semi-erect, black setae. Pronotum loose setose with long, erect, black setae, distinctly long towards anterior margin; with a dense setose ciliary of long to distinctly long, black setae on lateral margin. Scutellum loose setose with short, semi-erect, black setae. Elytra with pattern; sparse setose with medium-long, semi-erect, black setae; lengthening towards anterior margin; with a sparse juxta-sutural row of robust, long, semi-erect, black setae, from anterior margin, crossing scutellar torus towards apical torus; with three shortened, longitudinal, loose squamose bands of reclinate, narrowly-lanceolate, white scales: a supra-lateral one from anterior two fifth terminating beside apical torus; a supra-humeral on from middle terminating in apical third; a juxta-sutural one from apical third termination beside apical torus. Suture with a loose row of short, robust, semi-erect, black setae, lengthening apically. Thorax and pleurites loose pilose with long, black hairs. Ventrite I – V sparse setose with short, reclinate, brown setae; long, semi-erect, black setae interspersed. Propygidium loose setose with medium-long, semi-erect, black setae; posterior margin with a loose row of reclinate, narrowly-lanceolate, whitish scales. Pygidium sparse setose with long, erect, black setae, reclinate, narrowly-lanceolate, whitish scales interspersed in upper quarter; tip with a loose ciliary of medium-long, black setae. Prothoracic femur loose setose with long, semi-erect, black setae; bare internally. Prothoracic tibia sparse setose with short, reclinate and semi-erect, palebrown setae, sparser, erect and longer ventrally; with a sparse setose longitudinal row of short and medium-long, erect, black setae dorsally; a loose setose ciliary of short to medium-long, black setae on internal ridge; a setaceous ciliary of short, brown setae on anterior margin, more robust towards internal ridge; a medium-long, downwards pointing setose ciliary of brown setae along anterior margin of apical tooth. Mesothoracic femur sparse setose with long, semi-erect, black setae, denser, distinctly long and brown ventrally; bare internally. Mesothoracic tibia loose setose with short, semi-erect, pale-brown setae; medium-long, semi-erect, robust, black setae interspersed, sparser and finer internally. Posterior margin with a bristled ciliary of short to medium-long, black bristles; missing internally. Metathoracic femur loose setose with long to distinctly long, semi-erect, black setae; offwhite ventrally; bare internally. Metathoracic tibia sparse setose with medium-long to long, semi-erect, robust black setae; finer internally. Posterior margin with a sparse bristled outer marginal ciliary of short, brown bristles, lengthening dorsally and ventrally. Prothoracic and mesothoracic tarsomere I – IV bare. Metathoracic tarsomere I – IV sparse setose with very short, reclinate, brown setae. Tarsomere I – IV of thoracic legs with a setaceous, apical, marginal ciliary of short, black setae, lengthening and more robust towards metathoracic tarsomeres, fine dorsally and more robust ventrally. Pretarsomere of all thoracic legs sparse setose with short, semi-erect, black setae, lengthening towards metathoracic pretarsomere; few, medium-long, semi-erect, black setae at apical margin, shorter and red-brown ventrally. Head: Suture between head and clypeus undulated, complanate. Head dense punctate with small, shallow, roundish, not merging punctures. Funicle I cylindrical, slightly thickened medially; funicle II – III asymmetric frustum-shaped; clavus elliptic, slightly longer than pedicellus and funiculum combined. Pronotum: Widest medially; loose punctate with very small, shallow, roundish punctures, denser within medial furrow. Anterior corner obtuse. Lateral margin convex. With a distinct, shallow, complete medial furrow. Posterior corner rounded. Scutellum: Parabola-shaped; as broad as long; dense punctate with very small and small, shallow, roundish punctures, micropunctate within. Lateral margin slightly convex. Tip rounded. Elytra: Oblong; longer than broad; widest medially; protruding propygidium. Lateral margin very slightly convex. Sculpture distinct; slightly pulvinate laterally; humerus distinct; with a very broad, roundish torus around scutellum; lateral trench distinct; distinctly depressed along scutellum. Loose punctate with extremely small, shallow, roundish punctures; a sparse juxta-sutural row of small, roundish tubercles from anterior margin, crossing scutellar torus towards apical torus, terminating in apical quarter. Abdomen: Ventrite I distinctly shortened medially; ventrite V broadest, slightly constricted ventrally. Propygidium with a slightly projected stoma at posterior corner. Pygidium distinctly pulvinate longitudinally and laterally; upper margin slightly convex, linear medially; lateral margin linearly attenuate apically; tip rounded. Prothoracic leg: Femur and tibia robust. Tibia smooth ventrally. Median tooth robust. Basal tooth rudimentary. Interspace between anterior and median tooth double in width than between median and basal tooth. Shape of interspace between anterior and median tooth concave and between median and basal tooth linear. Tarsomeres I – IV combined extremely short; lengthening from tarsomere I – IV; slightly less than 1 / 2 in length of tibia. Tarsomeres III – IV with a ventral extension. Pretarsomere distinctly thickened; with a shortened smooth ridge ventrally; slightly longer than tarsomere I – IV combined. Internal claw slightly less than 1 / 10 more in length of pretarsomere. External claw slightly more than 7 / 10 in length of internal claw. Mesothoracic leg: Tibia slightly longer than femur. Tarsomere combined I – IV extremely short; slightly less than 2 / 5 in length of tibia. Pretarsomere with a shortened smooth ridge ventrally followed by a roundish cutting apically; slightly less than 1 / 10 more in length of tarsomere I – IV combined. External claw slightly less than 9 / 10 in length of pretarsomere. Internal claw slightly less than 9 / 10 in length of external claw. Metathoracic leg: Tibia as long as femur. Tarsomere I – IV combined short; 4 / 5 in length of tibia. Pretarsomere slender; with a smooth ridge ventrally; slightly less than 1 / 2 in length of tarsomeres I – IV combined. External claw slightly less than 1 / 10 more in length of pretarsomere. Internal claw slightly less than 9 / 10 in length of external claw. Parameres: (Figs. 62 – 63, 100). ♀♀. Measurements: length 7.7 – 7.9 mm; width 4.0 – 4.2 mm. Integument: Black; colour gradient of elytra from pale-brown medially to black at lateral and posterior margin, margins black; thoracic legs dark-brown, tarsi red-brown. Tegument: Clypeus sparse setose with very short, semi-erect, pale-brown setae. Head loose setose with mediumlong, semi-erect, brown setae. Pronotum without pattern; loose setose with medium-long, semi-erect, brown setae; short, semi-erect, offwhite setae interspersed in medial furrow, denser apically; with a dense setose ciliary of long, brown setae on lateral margin. Setae on scutellum brown. Elytra with pattern; additionally loose setose with short, semi-erect, brown setae; pattern consisting of short, semi-erect, yellowish setae. Hairs on thorax and pleurites yellowish. Ventrite I – V loose setose with short, reclinate, yellowish setae. Propygidium and pygidium loose setose with short to medium-long, semi-erect, brown setae; tip of pygidium with a loose ciliary of medium-long, brown setae. Setae on prothoracic tibia pale-brown. Setae on mesothoracic femur brown and pale-brown ventrally. Interspersed setae of mesothoracic tibia brown, setae of ciliary red-brown. Setae of metathoracic leg brown. Setae of thoracic tarsi red-brown. Head: Clypeus dense punctate with very small to small, shallow, roundish punctures; anterior margin distinctly vertically reflexed. Suture between head and clypeus linear, raised. Punctures on head large. Eye canthus full, dividing eye. Clavus short elliptic. Pronotum: Widest in anterior third; dense punctate with small, shallow, roundish punctures, denser within medial furrow and posteriorly. Lateral margin linearly attenuate in anterior third, slightly convex attenuate posteriorly. Medial furrow indistinct. Scutellum: Size of punctures only very small. Elytra: Ovate; widest in anterior third. Lateral margin slightly convex attenuate apically. Sculpture distinct; humerus indistinct; with a broad, roundish torus around scutellum; slightly depressed along scutellum. Loose punctate with very small, shallow, roundish punctures, micropunctures in-between; a sparse juxta-sutural row of small, shallow, roundish punctures from anterior margin, crossing scutellar torus towards apical torus. Abdomen: Distinctly pulvinate; ventrite II – V broadened medially; ventrite V not constricted ventrally. Pygidium slightly pulvinate longitudinally; slightly declined posteriorly; lateral margin subparallel in upper quarter, linearly attenuate apically; tip linear. Prothoracic leg: Basal tooth of tibia very small. Tarsomere I – IV combined very short; equal in length; slightly less than 7 / 10 in length of tibia. Tarsomere II – IV without ventral extension. Pretarsomere slightly thickened; smooth ventrally, roundish cutting missing; slightly less than 4 / 5 in length of tarsomeres I – IV combined. Internal claw slightly less than 9 / 10 in length of pretarsomere. External claw slightly more than 4 / 5 in length of internal claw. Mesothoracic leg: Tibia as long as femur. Tarsomeres I – IV combined very short; slightly more than 2 / 5 in length of tibia. Pretarsomere slender; with a shortened undulate ridge ventrally, roundish cutting missing; slightly less than 9 / 10 in length of tarsomere I – IV combined. External claw slender; not broadened medially; slightly more than 9 / 10 in length of pretarsomere. Metathoracic leg: Tibia distinctly extended dorso-apically; slightly less than 4 / 5 in length of femur. Tarsomeres I – IV combined slightly more than 4 / 5 in length of tibia. Pretarsomere slightly more than 2 / 5 in length of tarsomeres I – IV combined. External claw slightly more than 1 / 10 more in length of pretarsomere. Internal claw slightly more than 4 / 5 in length of external claw.	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E011FFE7FF64D126FC62FD48.taxon	distribution	Distribution. Northern Cape Province (Fig. 140), only known from the type locality Vegetation Type. Hantam Karroo [SKt 2].	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E011FFE7FF64D126FC62FD48.taxon	biology_ecology	Host Plants. Hyacinthaceae: Albuca canadensis (L.) F. M. Leight.	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E011FFE7FF64D126FC62FD48.taxon	etymology	Etymology. The name derives from the black elytra [Latin: nigr- = black].	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E00EFFE0FF64D70DFAE4FE78.taxon	description	http: // zoobank. org / 725 E 4997 - D 1 E 7 - 4 D 08 - A 3 FF- 95 E 8 A 92913 D 0	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E00EFFE0FF64D70DFAE4FE78.taxon	materials_examined	Type locality. [RSA, NCP]: Farm Glenlyon [now Hantam National Botanical Garden]. Type material. [20] Holotype ♂: yellow label hw « 30 » pr «. 276 | coll. Dombrow » || white label pr « Republic of South Africa | NC, Bokkeveldberge | Farm Glenlyon | 31.23.841 S – 19.08.397 E » bs pr « altitude 732 m | 04.09.2008 | # 522 | leg.: H. & E. Dombrow » || green label pr « Host plant: | Homeria miniata | Arctotis acaulis » || blue label pr « Bioregion: | Shale Renosterveld | Vegetation Type: FRs 2 – | Nieuwoudtville Shale Renosterveld » || yellow label pr « SAM – COL | A 071227 » || red label pr « Holotype | Beckhoplia | pallidibrunnea sp. n. | det.: H. Dombrow 2019 » (SAMC). – Paratypes 1 ♂, 2 ♀♀: same labels as holotype but || yellow label hw « 30 » pr «. 266,. 280,. 287 | coll. Dombrow » || yellow label pr « SAM – COL | A 071227 » || red label pr « Paratype | Beckhoplia | pallidibrunnea sp. n. | det.: H. Dombrow 2019 » (SAMC); 10 ♂♂, 6 ♀♀: same labels as holotype but || yellow label hw « 30 » pr «. 267 –. 268;. 270 –. 271;. 273 –. 275,. 277 –. 279,. 281 –. 286 | coll. Dombrow » || red label pr « Paratype | Beckhoplia | pallidibrunnea sp. n. | det.: H. Dombrow 2019 » (ESDW). Condition of holotype. Very good. Glued on card. Dissected, parameres and spiculum gastrale glued behind beetle.	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E00EFFE0FF64D70DFAE4FE78.taxon	diagnosis	Differential diagnosis. Differs from B. fusca: Scutellum of B. pallidibrunnea as broad as long. Scutellum of B. fusca longer than broad. Pretarsomere of metathoracic tarsus of B. pallidibrunnea slightly thickened. Pretarsomere of metathoracic tarsus of B. fusca slender. Interno-posterior margin of parameres of B. pallidibrunnea parabola-shaped (Figs. 64, 101). Interno-posterior margin of parameres of B. fusca narrowed curly bracket-shaped (Figs. 66, 100). Externo-posterior margin of parameres of B. pallidibrunnea concave medially (Figs. 64, 101). Externo-posterior margin of parameres of B. fusca convex, incised medially (Figs. 66, 100).	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E00EFFE0FF64D70DFAE4FE78.taxon	description	Description Measurements: length 8.5 – 9.0 mm; width 4.3 – 5.0 mm. Integument: Black; funiculum brown annulated; elytra transparently pale-brown, margins sometimes blackened. Tegument: Clypeus bare; sparse setose with very short, erect, black setae on the inner side of the vertically reflexed anterior and lateral margin. Head loose setose with medium-long, erect, black setae. Eye canthus with a loose setose ciliary of long black setae. Scapus loose setose with medium-long, black setae, brown ventrally, sparse and short on pedicellus; funiculum with few very short, semi-erect, brown setae, sparse on clavus. Pronotum loose setose with medium-long and long, erect, black setae; with a loose setose ciliary of long, black setae on lateral margin. Scutellum loose setose with medium-long, erect, black setae. Elytra without pattern. Loose setose with short, semi-erect, brownish setae; medium-long, erect, black setae interspersed, long towards humerus; with a sparse juxtasutural row of robust, long, erect, black setae, shorter in posterior half, from anterior margin, crossing scutellar torus towards apical torus. Suture with a loose row of short to long, robust, erect, black setae. Thorax and pleurites loose pilose with long, black hairs. Ventrite I – V loose setose with short, reclinate, black setae; long, semi-erect, black setae interspersed. Propygidium loose squamose with reclinate, lanceolate, whitish scales; long, semi-erect, black setae interspersed. Pygidium sparse setose with long, erect, black setae; tip of pygidium with a loose ciliary of medium-long, black setae. Prothoracic femur loose pilose with long, black hairs; bare internally. Prothoracic tibia loose setose with short, reclinate, black setae, medium-long and long ventrally; with a sparse setose longitudinal row of short and medium-long, erect, black setae dorsally; a loose setose ciliary of short, black setae on internal ridge; a setaceous ciliary of short, black setae on anterior margin, more robust towards internal ridge; a longer, downwards pointing setose ciliary of brown setae along anterior margin of apical tooth. Mesothoracic femur loose pilose with long, black hairs; bare internally. Mesothoracic tibia loose setose with short, semi-erect, whitish setae; medium-long, semi-erect, black, robust setae interspersed, sparser, finer, medium-long and long internally. Posterior margin with a bristled ciliary of very short and short, black bristles, lengthening dorsally and ventrally, missing internally. Metathoracic femur loose setose with medium-long to distinctly long, erect, black setae; whitish ventrally; bare internally. Metathoracic tibia loose setose with very short, reclinate, brown setae; medium-long and long, robust, semi-erect, black setae interspersed, denser and finer internally. Posterior margin with a loose bristled outer marginal ciliary of short, black bristles, lengthening dorsally and ventrally, more robust ventrally. Prothoracic and mesothoracic tarsomere I – IV bare. Metathoracic tarsomere I – IV sparse setose with very short, reclinate, brown setae. Tarsomere I – IV of all thoracic legs with a setaceous, apical, marginal ciliary of short, black setae, lengthening towards metathoracic tarsomeres, fine dorsally and more robust ventrally. Pretarsomere of all thoracic legs sparse setose with extremely short to very short, semi-erect and reclinate, brown setae; short and semi-erect ventrally; few, medium-long, semi-erect, black setae at apical margin. Head: Suture between head and clypeus undulate, crenated. Head dense punctate with small, shallow, roundish, partly merging punctures, micropunctate within. Funicle I – II cylindrical; funicle III asymmetric frustumshaped; clavus elliptic, slightly longer than pedicellus and funiculum combined. Pronotum: Widest at anterior two fifth; loose punctate with very small and small, shallow, roundish punctures, dense apically, loose within medial furrow. Anterior corner orthogonal. Lateral margin linearly attenuate anteriorly, subparallel posteriorly. With an indistinct, complete, shallow medial furrow. Posterior corner angled. Scutellum: Parabola-shaped; as broad as long; dense punctate with small, shallow, roundish punctures, micropunctate within; with a complete, median longitudinal, narrow, smooth stripe. Lateral margin slightly convex. Tip rounded. Elytra: Ovate; longer than broad; widest medially; protruding propygidium. Lateral margin slightly convex attenuate apically. Sculpture indistinct; slightly pulvinate laterally; humerus distinct; with a roundish torus beside scutellum; lateral trench missing; broadly, slightly, longitudinally depressed along suture. Loose punctate with extremely small, shallow, roundish punctures, micropunctures in-between; a sparse juxta-sutural row of small, roundish punctures from anterior margin, crossing scutellar torus towards apical torus, terminating in apical quarter. Abdomen: Ventrite I extremely shortened medially; ventrite V broadest, not constricted ventrally. Propygidium with a slightly projected stoma at posterior corner. Pygidium distinctly pulvinate longitudinally and laterally; upper margin moderately convex, linear medially; lateral margin slightly convex attenuate apically; tip rounded. Prothoracic leg: Femur and tibia robust. Tibia smooth ventrally. Median tooth elongate. Basal tooth rudimentary. Interspace between anterior and median tooth double in width than between median and basal tooth. Shape of interspace between anterior and median tooth u-shaped and between median and basal tooth concave. Tarsomere I – IV combined extremely short; lengthening from tarsomere I – IV; slightly less than 7 / 10 in length of tibia. Pretarsomere distinctly thickened; with a small, triangular-shaped, median tooth ventrally; slightly longer than tarsomere I – IV combined. Internal claw slightly shorter than pretarsomere. External claw slightly more than 7 / 10 in length of internal claw. Mesothoracic leg: Tibia slightly less than 9 / 10 in length of femur. Tarsomere I – IV combined extremely short; slightly less than 2 / 5 in length of tibia. Pretarsomere with a shortened granulate ridge ventrally, roundish cutting missing; as long as tarsomere I – IV combined. External claw slightly shorter than pretarsomere. Internal claw slightly more than 9 / 10 in length of external claw. Metathoracic leg: Tibia slightly more than 1 / 10 more in length of femur. Tarsomere I – IV combined distinctly short; slightly less than 7 / 10 in length of tibia. Pretarsomere slightly thickened; with an undulate ridge ventrally; slightly more than 1 / 2 in length of tarsomere I – IV combined. External claw slightly less than 1 / 10 more in length of pretarsomere. Internal claw slightly more than 4 / 5 in length of external claw. Parameres: (Figs. 64 – 65, 101). ♀♀. Measurements: length 9.3 – 9.7 mm; width 4.9 – 5.7 mm. Tegument: Clypeus setose with loose, very short, semi-erect brown setae, medium-long on head. Setae on funiculum and clavus pale-brown. Eye canthus with a loose ciliary of medium-long, brown setae. Pronotum without pattern. Loose setose with medium-long and long, erect, brown setae; very short and short, semi-erect, yellowish setae interspersed in medial furrow. With a loose setose ciliary of medium-long and long brown setae on lateral margin. Scutellum loose setose with medium-long, semi-erect, brown setae; short semi-erect, yellowish setae interspersed. Elytra without pattern. Loose setose with short, reclinate, pale-brown setae; long, semi-erect and erect, brown setae interspersed, denser towards humerus. Thorax and pleurites loose pilose with long, brown hairs. Ventrite I – V loose setose with short and medium-long, reclinate, yellowish-white setae; long, semi-erect, brown setae interspersed. Propygidium and pygidium loose setose with short, reclinate, yellowish-white setae; long, semi-erect, brown setae interspersed. Tip of pygidium with a loose setose ciliary of medium-long and long black-brown setae. Prothoracic femur loose pilose with medium-long and long brown hairs; bare internally. Prothoracic tibia loose setose with very short and short semi-erect brown setae; erect and medium-long ventrally. With a loose setose ciliary of medium-long brown setae on internal ridge; anterior margin with a loose setaceous ciliary of short pale-brown setae and a medium-long downwards pointing setose ciliary of pale-brown setae along anterior margin of apical tooth. Mesothoracic femur loose pilose with long black hairs; long and distinctly-long pale-brown ventrally; bare internally. Mesothoracic tibia loose setose with short, semi-erect, yellowish-white setae; sparser internally; medium-long, semi-erect, pale-brown, robust setae interspersed; medium-long and long, finer and darker internally. Fringe of posterior margin as in male but red-brown. Metathoracic femur loose setose with medium-long and long brown setae; paler ventrally; bare internally. Metathoracic tibia loose setose with short, semi-erect, pale-brown setae; medium-long and robust, semi-erect, red-brown setae interspersed; finer, medium-long and long internally. Ciliary of posterior margin as in male but red-brown. Setae of thoracic tarsi red-brown. Head: Clypeus very slightly depressed laterally; dense punctate with large, shallow, roundish punctures. Anterior margin slightly vertically reflexed. Lateral margin slightly reflexed. Head dense punctate with small to large, shallow, roundish punctures, not merging, not micropunctate. Funicle II – III short-asymmetric frustumshaped; clavus short elliptic, as long as pedicellus and funiculum combined. Pronotum: Moderate pulvinate laterally. Dense punctate with small to large, shallow, roundish punctures, denser apically. Lateral margin convex attenuate in posterior three fifth. Posterior corner rounded. Scutellum: With a shortened, median, longitudinal, narrow, smooth stripe from anterior margin terminating in anterior third. Elytra: Widest in anterior third; lateral trench indistinct; scutellar torus missing; not depressed along suture. Abdomen: Distinctly pulvinate; ventrite I distinctly shortened medially; ventrite II – V broadened medially. Pygidium slightly pulvinate longitudinally and laterally; slightly declined posteriorly; lateral margin subparallel in upper quarter, linearly attenuate apically; tip linear. Prothoracic leg: Median tooth broad, triangular-shaped. Basal tooth very small. Shape of interspace between teeth concave. Tarsomere I – IV combined distinctly short; equal in length; slightly more than 3 / 5 in length of tibia. Pretarsomere slightly thickened; with a smooth ridge ventrally; slightly more than 3 / 5 in length of tarsomere I – IV combined. Internal claw slender; not broadened medially; slightly less than 9 / 10 in length of pretarsomere. External claw slightly less than 9 / 10 in length of internal claw. Mesothoracic leg: Tibia 9 / 10 in length of femur. Tarsomere I – IV combined very short; slightly more than 2 / 5 in length of tibia. Pretarsomere with an undulate ridge ventrally, roundish cutting missing; slightly less than 4 / 5 in length of tarsomere I – IV combined. External claw slender; not broadened medially; Internal claw slightly less than 4 / 5 in length of external claw. Metathoracic leg: Femur slightly thickened. Tibia distinctly extended dorso-apically; slightly less than 9 / 10 in length of femur. Tarsomere I – IV combined short; slightly more than 4 / 5 in length of tibia. Pretarsomere slender; slightly more than 2 / 5 in length of tarsomere I – IV combined. External claw slightly shorter than pretarsomere. Internal claw slightly less than 9 / 10 in length of external claw.	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E00EFFE0FF64D70DFAE4FE78.taxon	distribution	Distribution. Northern Cape Province (Fig. 140), only known from the type locality. Vegetation Type. Nieuwoudtville Shale Renosterveld [FRs 2].	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E00EFFE0FF64D70DFAE4FE78.taxon	biology_ecology	Host Plants. Iridaceae: Moraea miniata Andrews; Asteraceae: Arctotis acaulis L.	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E00EFFE0FF64D70DFAE4FE78.taxon	etymology	Etymology. The name derives from the pale-brown elytra [Latin: pallidi- = pale; Latin: brunne- = brown].	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E009FFECFF64D65DFBB9F818.taxon	description	(Figs. 9, 16, 66 – 67, 102) http: // zoobank. org / 9 E 5 F 22 D 0 - AFDD- 4010 - B 842 - BD 1 B 617 E 7533	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E009FFECFF64D65DFBB9F818.taxon	materials_examined	Type locality. [RSA, NCP]: 1 km nördl. Farm Wilemsriver. Type material. [9] Holotype ♂: yellow label hw « 54 », pr «. 920 » | coll. Dombrow || white label pr « AFRIKA: Südafrika, Kapprovinz | 1 km nördl. Farm Wilemsriver | 31 ° 20 ’ 14 ” S, 19 ° 05 ’ 13 ” E, 750 m | 30.08.1989, leg. R. Fritz » || green label pr « Host plant: | Homeria bifida » || blue label pr « Bioregion: | Sandstone Fynbos | Vegetation Type: FFs 1 – | Bokkeveld Sandstone Fynbos » || red label pr « Holotype | Beckhoplia | fusca sp. n. | det.: H. Dombrow 2019 » (ZSM). – Paratypes 8 ♀♀: same labels as holotype but || yellow label hw « 54 », pr «. 982 –. 989 | coll. Dombrow » || red label pr « Paratype | Beckhoplia | fusca sp. n. | det.: H. Dombrow 2019 » (ESDW). Condition of holotype. Excellent. Glued on card. Dissected, parameres and spiculum gastrale glued behind beetle.	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E009FFECFF64D65DFBB9F818.taxon	diagnosis	Differential diagnosis. Differs from B. pallidibrunnea: Scutellum of B. fusca longer than broad. Scutellum of B. pallidibrunnea as broad as long. Pretarsomere of metathoracic tarsus of B. fusca slender. Pretarsomere of metathoracic tarsus of B. pallidibrunnea slightly thickened. Interno-posterior margin of parameres of B. fusca narrowed curly bracket-shaped (Figs. 66, 102). Interno-posterior margin of parameres of B. pallidibrunnea parabola-shaped (Figs. 64, 101). Externo-posterior margin of parameres of B. fusca convex, incised medially (Figs. 66, 102). Externo-posterior margin of parameres of B. pallidibrunnea concave medially (Figs. 64, 101).	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E009FFECFF64D65DFBB9F818.taxon	description	Description Measurements: length 8.2 mm, width 4.4 mm. Integument: Black; elytra transparently brown. Tegument: Clypeus bare; sparse setose with extremely short, erect, brown setae on the inner side of the vertically reflexed anterior and lateral margin. Head loose setose with long, erect, black setae. Eye canthus with a dense setose ciliary of long, black setae. Scapus loose setose with medium-long, black setae, dark-brown ventrally, sparse and short on pedicellus, funiculum and clavus sparse setose with very short and short, semi-erect, brown setae. Pronotum loose setose with medium-long and long, erect, black setae; with a loose setose ciliary of long to distinctly long, black setae on lateral margin. Scutellum loose setose with medium-long, semi-erect, black setae, shortening apically; with a longitudinal, median, bare strip. Elytra without pattern; sparse setose with medium-long, semi-erect, black setae; lengthening towards humerus; with a sparse juxta-sutural row of robust, medium-long, semi-erect, black setae, from anterior margin, crossing scutellar torus towards apical torus; narrowly-lanceolate, semi-erect, offwhite scales interspersed along lateral margin, denser around apical torus. Suture with a loose row of short to long, robust, semi-erect, black setae, lengthening apically. Thorax and pleurites loose pilose with long, black hairs. Ventrite I – V sparse setose with short, reclinate, brown setae; long, semi-erect, black setae interspersed. Propygidium loose setose with long, semi-erect, black setae; posterior margin with a loose row of reclinate, narrowly-lanceolate, whitish scales. Pygidium sparse setose with long, erect, black setae; tip with a loose ciliary of medium-long, black setae. Prothoracic femur loose setose with long, semi-erect, black setae; bare internally. Prothoracic tibia sparse setose with short, reclinate and semi-erect, pale-brown setae, sparser ventrally; with a sparse setose longitudinal row of short and medium-long, erect, black setae dorsally; a loose setose ciliary of short, black setae on internal ridge; a setaceous ciliary of short, dark red-brown setae on anterior margin, more robust towards internal ridge; a medium-long, downwards pointing setose ciliary of brown setae along anterior margin of apical tooth. Mesothoracic femur sparse setose with long, semi-erect, black setae, denser and brown ventrally; bare internally. Mesothoracic tibia loose setose with short, semi-erect, offwhite setae; medium-long, semi-erect, robust, black setae interspersed, sparser and finer internally. Posterior margin with a bristled ciliary of short to medium-long, black bristles; missing internally. Metathoracic femur loose setose with medium-long to distinctly long, semi-erect, black setae; offwhite ventrally; bare internally. Metathoracic tibia sparse setose with medium-long to long, semi-erect, robust black setae; finer internally; very short and short, reclinate, brown setae interspersed. Posterior margin with a loose bristled outer marginal ciliary of short, black bristles, lengthening dorsally and ventrally. Tarsomere I – IV of all thoracic legs sparse setose with very-short, reclinate, brown, setae lengthening towards metathoracic tarsomeres; with a setaceous, apical, marginal ciliary of medium-long, black setae, lengthening and more robust towards metathoracic tarsomeres, fine dorsally and more robust ventrally. Pretarsomere of all thoracic legs sparse setose with very short, reclinate, pale-brown setae; few, medium-long, semi-erect, black setae at apical margin, shorter and red-brown ventrally. Head: Suture between head and clypeus undulated, crenated. Head dense punctate with small, shallow, roundish, not merging punctures, micropunctate within. Funicle I-II cylindrical, slightly thickened medially; funicle III asymmetric frustum-shaped; clavus elliptic, as long as pedicellus and funiculum combined. Pronotum: Widest medially; loose punctate with small, shallow, roundish punctures, dense within medial furrow and posteriorly. Anterior corner orthogonal. Lateral margin linearly attenuate anteriorly, subparallel posteriorly. With an indistinct, complete, shallow, medial furrow. Posterior corner rounded. Scutellum: Parabola-shaped; as broad as long; dense punctate with small, shallow, roundish punctures, micropunctate within; with a complete, median, longitudinal, narrow, smooth stripe. Lateral margin slightly convex. Tip rounded. Elytra: Ovate; as broad as long; widest medially; protruding propygidium. Lateral margin slightly convex attenuate apically. Sculpture indistinct; slightly pulvinate laterally; humerus indistinct; with a roundish torus beside scutellum; lateral trench indistinct; broadly, slightly, longitudinally depressed along suture. Loose punctate with extremely small, shallow, roundish punctures, micropunctures in-between; a sparse juxta-sutural row of large, roundish tubercles from anterior margin, crossing scutellar torus towards apical torus; a sparse supra-humeral row of small, roundish tubercles from anterior margin between humerus and scutellar torus towards apical torus. Abdomen: Ventrite I extremely shortened medially; ventrite V broadest, not constricted ventrally. Propygidium with a slightly projected stoma at posterior corner. Pygidium distinctly pulvinate longitudinally and laterally; upper margin slightly convex, linear medially; lateral margin slightly convex attenuate apically; tip linear. Prothoracic leg: Femur and tibia robust. Tibia smooth ventrally. Median tooth elongate. Basal tooth very small. Interspace between anterior and median tooth double in width than between median and basal tooth. Shape of interspace between anterior and median tooth u-shaped and between median and basal tooth concave. Tarsomere I – IV combined extremely short; lengthening from tarsomere I – IV; slightly more than 2 / 5 in length of tibia. Pretarsomere distinctly thickened; with a small, triangular-shaped, median tooth ventrally; slightly less than 1 / 5 more in length of tarsomere I – IV combined. Internal claw slightly less than 1 / 10 more in length of pretarsomere. External claw slightly less than 4 / 5 in length of internal claw. Mesothoracic leg: Tibia 9 / 10 in length femur. Tarsomere I – IV combined extremely short; slightly more than 3 / 10 in length of tibia. Pretarsomere with a shortened smooth ridge ventrally, followed by a roundish cutting apically; slightly more than 1 / 10 more in length of tarsomere I – IV combined. External claw slightly less than 1 / 10 more in length of pretarsomere. Internal claw slightly less than 9 / 10 in length of external claw. Metathoracic leg: Tibia slightly less than 9 / 10 in length of femur. Tarsomere I – IV combined distinctly short; slightly more than 7 / 10 in length of tibia. Pretarsomere slender; with an undulate ridge ventrally; slightly more than 2 / 5 in length of tarsomere I – IV combined. External claw slightly less than 3 / 10 more in length of pretarsomere. Internal claw slightly more than 4 / 5 in length of external claw. Parameres: (Figs. 66 – 67, 102). ♀♀. Measurements: length 7.9 – 8.9 mm; width 4.4 – 5.2 mm. Integument: Thoracic tarsi dark-brown. Tegument: Clypeus loose setose with short, erect, pale-brown setae, medium-long and brown on head. Eye canthus with a dense setose ciliary of medium-long, brown setae. Setae on scapus brown and pale-brown ventrally; sparse and short on pedicellus. Setae on pronotum including ciliary brown. Setae on scutellum short and brown, with a shortened bare stripe in anterior half. Elytra without pattern; loose setose with short, reclinate, pale-brown setae; long, semi-erect, brown setae interspersed, lengthening towards humerus; with a sparse juxta-sutural row of robust, medium-long, semi-erect, brown setae, from anterior margin, crossing scutellar torus towards apical torus. Suture with a loose row of short to long, robust, semi-erect, brown setae, lengthening apically. Hairs on thorax and pleurites brown. Ventrite I – V loose setose with short, reclinate, yellowish setae; long, semi-erect, brown setae interspersed. Propygidium and pygidium loose setose with short, reclinate, yellowishbrown setae, medium-long to long, semi-erect and erect, pale-brown setae interspersed; tip of pygidium with a loose ciliary of medium-long, brown setae. Setae on prothoracic femur brown. Setae of longitudinal dorsal row of prothoracic tibia brown. Setae on mesothoracic femur brown, pale-brown ventrally. Interspersed setae on mesothoracic tibia brown. Bristles of ciliary red-brown. Metathoracic femur sparse setose with short to medium-long, reclinate, pale-brown setae; long to distinctly long, semi-erect, brown setae interspersed; denser dorsally. Setae of metathoracic tibia brown, interspersed ones offwhite. Bristles of ciliary very short, transparently red-brown and distinctly lengthening dorsally and ventrally. Ciliary of tarsomere I – IV of thoracic legs red-brown. Interspersed setae on pretarsomeres of thoracic legs transparently pale red-brown. Head: Clypeus slightly longitudinally depressed laterally; dense punctate with small, deep, roundish punctures, micropunctate within; anterior margin distinctly reflexed. Head dense punctate with small to large, deep, roundish, partly merging punctures. Eye on head level. Funicle II asymmetric frustum-shaped; funicle III short asymmetric frustum-shaped; clavus short elliptic. Pronotum: Widest in anterior third; moderate pulvinate laterally. Anterior corner obtuse. Lateral margin convex. Scutellum: Dense punctate with very small, deep, roundish punctures; with a shortened, median, longitudinal, narrow, smooth stripe from anterior margin terminating in anterior half. Elytra: Longer than broad; widest in anterior quarter. Sculpture indistinct; scutellar torus missing; complanate along suture. Tubercles of juxta-sutural row of very small; supra-humeral row missing. Abdomen: Distinctly pulvinate; ventrite I distinctly shortened medially; ventrite II – V broadened medially. Propygidium with an indistinct stoma at posterior corner. Pygidium slightly pulvinate longitudinally and laterally; slightly declined posteriorly; lateral margin subparallel in upper quarter, linearly attenuate apically. Prothoracic leg: Median tooth broad triangular-shaped. Shape of interspace between teeth concave. Tarsomere I – IV combined distinctly short; equal in length; slightly more than 3 / 5 in length of tibia. Pretarsomere slender; with smooth ventrally, roundish cutting missing; slightly less than 7 / 10 in length of tarsomere I – IV combined. Internal claw slender; not broadened medially; slightly more than 9 / 10 in length of pretarsomere. External claw slightly less than 3 / 5 in length of internal claw. Mesothoracic leg: Tibia slightly less than 9 / 10 in length femur. Tarsomere I – IV combined very short; slightly more than 2 / 5 in length of tibia. Pretarsomere slender; with a smooth ridge ventrally, roundish cutting missing; slightly less than 4 / 5 in length of tarsomere I – IV combined. External claw slender; not broadened medially; slightly shorter than pretarsomere. Internal claw slightly more than 4 / 5 in length of external claw. Metathoracic leg: Tibia slightly dorso-apically extended; slightly more than 7 / 10 in length of femur. Tarsomere I – IV combined short; slightly less than 4 / 5 in length of tibia. External claw slightly less than 1 / 10 more in length of pretarsomere. Internal claw slightly less than 9 / 10 in length of external claw.	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E009FFECFF64D65DFBB9F818.taxon	distribution	Distribution. Northern Cape Province (Fig. 140), only known from the type locality. Vegetation Type. Bokkeveld Sandstone Fynbos [FFs 1].	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E009FFECFF64D65DFBB9F818.taxon	biology_ecology	Host Plants. Iridaceae: Moraea bifida L. Bolus.	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E009FFECFF64D65DFBB9F818.taxon	etymology	Etymology. The name derives from the brown elytra [Latin: fusc- = brown].	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E006FFEEFF64D48CFB77F818.taxon	description	(Figs. 42, 48, 60, 68 – 69, 103) http: // zoobank. org / 02 F 4 C 75 F-B 9 E 7 - 42 FA- 8 B 31 - 7903 CDD 2680 C	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E006FFEEFF64D48CFB77F818.taxon	materials_examined	Type locality. [RSA, NCP]: 6 km NO Nieuwoudtville. Type material. [1] Holotype ♂: white label pr « AFRIKA: Südafrika, Kapprovinz | 6 km NO Nieuwoudtville | 31 ° 19 ’ 16 ” S, 19 ° 07 ’ 00 ” E | 30.08.1989, leg. R. Fritz » || green label pr « Host plant: | Homeria miniata » || blue label pr « Bioregion: | Sandstone Fynbos | Vegetation Type: FFs 1 – | Bokkeveld Sandstone Fynbos » || yellow label hw « 54 », pr «. 990 » | coll. Dombrow || red label pr « Holotype | Beckhoplia | setosa sp. n. | det.: H. Dombrow 2019 » (ZSM). Condition of holotype. Very good. Glued on card. Dissected, parameres and spiculum gastrale glued behind beetle. Left prothoracic tarsus missing after tarsomere III.	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E006FFEEFF64D48CFB77F818.taxon	diagnosis	Differential diagnosis. Differs from B. castanea: Pronotum of B. setosa without medial furrow. Pronotum of B. castanea with medial furrow. Pretarsomere of metathoracic tarsus of B. setosa with a granulate ridge ventrally. Pretarsomere of metathoracic tarsus of B. castanea with an undulate ridge ventrally. Interno-posterior margin of parameres of B. setosa (Figs. 68, 103) curly bracket-shaped in dorsal view. Interno-posterior margin of parameres of B. castanea rounded, slightly incised medially (Figs. 70, 104). Anterior margin of phallobase of B. setosa (Figs. 68, 103) concave in dorsal view. Anterior margin of phallobase of B. castanea broad v-shaped (Figs. 70, 104).	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E006FFEEFF64D48CFB77F818.taxon	description	Description Measurements: length 7.9 mm; width 4.0 mm. Integument: Black; elytra transparently pale-brown. Tegument: Clypeus bare; sparse setose with very short to short, erect, brown setae on the inner side of the vertically reflexed anterior and lateral margin. Head loose setose with medium-long to long, erect, black setae. Eye canthus with a dense setose ciliary of medium-long, black setae. Scapus loose setose with medium-long, erect, black setae, brown ventrally, sparse and short on pedicellus; funiculum with few very short, erect, brown setae; clavus sparse setose with extremely short and very short, semi-erect, black setae. Pronotum loose setose with medium-long and long, erect, black setae; with a loose setose ciliary of long, black setae on lateral margin. Scutellum loose setose with medium-long, erect, black setae, with a longitudinal, median, bar strip in anterior third. Elytra without pattern; sparse setose with very short, reclinate, brownish setae; short to medium-long, semi-erect, black setae interspersed, lengthening towards anterior margin; with a sparse juxta-sutural row of robust, medium-long, erect, black setae, from anterior margin, crossing scutellar torus towards apical torus. Suture with a loose row of short to long, robust, semi-erect, black setae. Thorax and pleurites loose pilose with long, black hairs. Ventrite I – V sparse setose with short, semi-erect, black setae; medium-long to long, semi-erect, black setae interspersed. Propygidium and pygidium sparse setose with medium-long to long, semi-erect and erect, black setae; tip of pygidium with a sparse ciliary of medium-long, black setae. Prothoracic femur loose pilose with long, black hairs; bare internally. Prothoracic tibia sparse setose with short, semi-erect, pale-brown setae, sparser ventrally; medium-long, semi-erect, brown setae interspersed, long and erect ventrally; with a sparse setose longitudinal row of medium-long, erect, brown setae dorsally; a loose setose ciliary of short, black setae on internal ridge; a setaceous ciliary of short, brown setae on anterior margin, more robust towards internal ridge; a longer, downwards pointing setose ciliary of brown setae along anterior margin of apical tooth. Mesothoracic femur loose pilose with long, black hairs, distinctly long and brown ventrally; bare internally. Mesothoracic tibia sparse setose with short, semi-erect, whitish setae; medium-long, semi-erect, black, robust setae interspersed, sparser, finer, medium-long and long internally. Posterior margin with a bristled ciliary of short, black bristles, lengthening dorsally, missing internally. Metathoracic femur loose setose with medium-long to distinctly long, erect, black setae; pale-brown ventrally; bare internally. Metathoracic tibia loose setose with very short and short, reclinate, brown setae; medium-long and long, robust, semi-erect, black setae interspersed, finer internally. Posterior margin with a loose bristled outer marginal ciliary of short, black bristles, lengthening dorsally and ventrally. Prothoracic and mesothoracic tarsomere I – IV bare. Metathoracic tarsomere I – IV sparse setose with short, reclinate, black setae. Tarsomere I – IV of all thoracic legs with a setaceous, apical, marginal ciliary of short, black setae, lengthening towards metathoracic tarsomeres, fine dorsally and more robust ventrally. Pretarsomere of all thoracic legs sparse setose with very short, reclinate, pale-brown setae; few, medium-long, semi-erect, black setae at apical margin, shorter and red-brown ventrally. Head: Suture between head and clypeus linear, crenated. Head dense punctate with small, shallow, roundish, partly merging punctures, micropunctate within. Funicle I – II cylindrical, slightly thickened medially; funicle III asymmetric frustum-shaped; clavus elliptic, as long as pedicellus and funiculum combined. Pronotum: Widest at anterior two fifth; dense punctate with shallow, very small and small, roundish punctures. Anterior corner orthogonal. Lateral margin linearly attenuate anteriorly, subparallel posteriorly. With an indistinct, shortened, shallow, medial furrow from posterior margin termination behind disk. Posterior corner angled. Scutellum: Parabola-shaped; slightly longer than broad; dense punctate with shallow, small, roundish punctures, micropunctate within; with a shortened, median, longitudinal, narrow, smooth stripe from anterior margin terminating in anterior third. Lateral margin slightly convex. Tip rounded. Elytra: Ovate; longer than broad; widest medially; protruding propygidium. Lateral margin slightly convex attenuate apically. Sculpture indistinct; slightly pulvinate laterally; humerus indistinct; with a broad roundish torus around scutellum; lateral trench missing; complanate beside scutellum and along suture. Loose punctate with very small to extremely small, shallow, roundish punctures, micropunctures in-between; a sparse juxta-sutural row of small, roundish tubercles from anterior margin, crossing scutellar torus towards apical torus; a sparse supra-humeral row of small, shallow, roundish punctures from anterior margin between humerus and scutellar torus towards apical torus. Abdomen: Ventrite I distinctly shortened medially; ventrite V broadest, not constricted ventrally. Propygidium with a slightly projected stoma at posterior corner. Pygidium distinctly pulvinate longitudinally and laterally; upper margin moderately convex; lateral margin subparallel in upper quarter, linearly attenuate apically; tip rounded. Prothoracic leg: Femur and tibia robust. Tibia smooth ventrally. Median tooth robust. Basal tooth rudimentary. Interspace between teeth almost equal. Shape of interspace between anterior and median tooth concave and between median and basal tooth linear. Tarsomere I – IV combined extremely short; lengthening from tarsomere I – IV; slightly more than 2 / 5 in length of tibia. Pretarsomere distinctly thickened; with a small, triangular-shaped, median tooth ventrally; slightly less than 1 / 10 more in length of tarsomere I – IV combined. Internal claw slightly less than 1 / 10 more in length of pretarsomere. External claw slightly less than 4 / 5 in length of internal claw. Mesothoracic leg: Tibia slightly more than 7 / 10 in length of femur. Tarsomere I – IV combined extremely short; slightly more than 3 / 10 in length of tibia. Pretarsomere with a shortened smooth ridge ventrally followed by a roundish cutting apically; slightly less than 1 / 10 more in length of tarsomere I – IV combined. External claw slightly shorter than pretarsomere. Internal claw slightly less than 9 / 10 in length of external claw. Metathoracic leg: Tibia 9 / 10 in length of femur. Tarsomere I – IV distinctly short; slightly more than 7 / 10 in length of tibia. Pretarsomere slightly thickened; with a granulate ridge ventrally; slightly less than 2 / 5 in length of tarsomere I – IV combined. External claw slightly more than 1 / 10 more in length of pretarsomere. Internal claw less more than 4 / 5 in length of external claw. Parameres: (Figs. 68 – 69, 103). ♀: Unknown.	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E006FFEEFF64D48CFB77F818.taxon	distribution	Distribution. Northern Cape Province (Fig. 140), only known from the type locality. Vegetation Type. Bokkeveld Sandstone Fynbos [FFs 1].	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E006FFEEFF64D48CFB77F818.taxon	biology_ecology	Host Plants. Iridaceae: Moraea miniata Andrews.	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E006FFEEFF64D48CFB77F818.taxon	etymology	Etymology. The name derives from the long brown setae on pronotum [Latin: set = a bristle].	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E001FFD5FF64D48CFBA9FF74.taxon	description	(Figs. 18, 23, 47, 70 – 71, 104) http: // zoobank. org / 7 CFAF 1 C 8 - 2140 - 457 D-BC 87 - 6814 D 8 E 18 F 7 B	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E001FFD5FF64D48CFBA9FF74.taxon	materials_examined	Type locality. [RSA, NCP]: Farm Glenlyon [now Hantam National Botanical Garden]. Type material. [9] Holotype ♂: white label pr « RSA, NC, Nieuwoudt- | ville, Glenlyon Farm | 31.24 S – 19.08 E, # 210 | 09.09.1995, leg.: HDO » || red label pr « Paratype | Beckhoplia | colvillei sp. n. | det.: H. Dombrow 2003 » || yellow label pr « SAM – COL | A 071215 » || green label pr « Host plant: | Bulbinella latifolia | ssp. doleritica » || blue label pr « Bioregion: | Shale Renosterveld | Vegetation Type: FRs 2 – | Nieuwoudtville Shale Renosterveld » || yellow label hw « 50 » pr «. 991 | coll. Dombrow » || red label pr « Holotype | Beckhoplia | castanea sp. n. | det.: H. Dombrow 2019 » (ZSM). – Paratypes 3 ♂♂, 5 ♀♀: same labels as holotype but || yellow label hw « 18 » pr «. 840 –. 847 | coll. Dombrow » || red label pr « Paratype | Beckhoplia | castanea sp. n. | det.: H. Dombrow 2019 » (ESDW). Condition of holotype. Excellent. Glued on card. Dissected, parameres and spiculum gastrale glued behind beetle.	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E001FFD5FF64D48CFBA9FF74.taxon	diagnosis	Differential diagnosis. Differs from B. setosa: Pronotum of B. castanea with medial furrow. Pronotum of B. setosa without medial furrow. Pretarsomere of metathoracic tarsus of B. castanea with an undulate ridge ventrally. Pretarsomere of metathoracic tarsus of B. setosa with a granulate ridge ventrally. Interno-posterior margin of parameres of B. castanea rounded, slightly incised medially (Figs. 70, 104). Interno-posterior margin of parameres of B. setosa (Figs. 68, 103) curly bracket-shaped in dorsal view. Anterior margin of phallobase of B. castanea broad v-shaped (Figs. 70, 104). Anterior margin of phallobase of B. setosa (Figs. 68, 103) concave in dorsal view.	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E001FFD5FF64D48CFBA9FF74.taxon	description	Description Measurements: length 8.4 – 9.4 mm; width 4.3 – 4.6 mm. Integument: Black; funiculum pale-brown annulated; elytra transparently pale-brown, outer margins sometimes blackened. Tegument: Clypeus bare; loose setose with very short and short, erect, pale-brown and black setae on the inner side of the reflexed anterior and lateral margin. Head loose setose with medium-long, erect, black setae. Eye canthus with a dense setose ciliary of long, black, setae. Scapus loose setose with long, black setae, brown ventrally, sparser and medium-long on pedicellus; funiculum with few short, semi-erect, brown setae, lengthening and darker towards clavus; clavus sparse setose with short, semi-erect, black and short to very short, semi-erect, brown setae. Pronotum loose setose with medium-long to distinctly long, erect, brown setae; with a dense ciliary of long, black, setae on lateral margin. Scutellum loose setose with short and medium-long, erect, black setae. Elytra without pattern. Loose setose with short, semi-erect, brownish setae; medium-long, semi-erect, black setae interspersed, denser and long in anterior third; loose, long, semi-erect, whitish, squamiform setae around apical torus; with a sparse juxta-sutural row of medium-long, erect, black setae, shorter in posterior half, from anterior margin crossing scutellar torus towards apical torus. Suture with a dense row of robust, short to long semi-erect, black setae. Thorax and pleurites loose pilose with long, black hairs. Ventrite I – V loose setose with short and medium-long, reclinate, brownish setae; medium-long, semi-erect, black setae interspersed. Propygidium loose squamose with reclinate, narrowly-lanceolate, pale-brown and whitish scales; medium-long and long, semi-erect, black setae interspersed. Pygidium sparse setose with long, erect, black setae; tip with a loose ciliary of long, black setae. Prothoracic femur loose pilose with long, black hairs; bare internally. Prothoracic tibia loose setose with short, reclinate, white setae; medium-long, black, semi-erect setae interspersed dorsally, long and black ventrally; with a sparse setose longitudinal row of short and medium-long, erect, black setae dorsally; a loose setose ciliary of short, black setae on internal ridge; a setaceous ciliary of short, brown setae on anterior margin, more robust towards internal ridge; a medium-long downwards pointing setose ciliary of brown setae along anterior margin of apical tooth. Mesothoracic femur loose pilose with long, black hairs; pale-brown ventrally; bare internally. Mesothoracic tibia sparse setose with short, reclinate, white setae; medium-long, semi-erect, robust, black setae interspersed, finer, medium-long and long internally. Posterior margin with a bristled ciliary of very short and short, black bristles; lengthening dorsally and ventrally; missing internally. Metathoracic femur loose setose with long, semi-erect, black setae; long black hairs interspersed near coxa; long pale-brown hairs interspersed ventrally; bare internally. Metathoracic tibia loose setose with short, reclinate, brown setae; medium-long, semi-erect, black setae interspersed, finer and long internally. Posterior margin with a loose bristled outer marginal ciliary of short, black bristles, lengthening dorsally and ventrally, more robust ventrally. Tarsomere I – IV of all thoracic legs sparse setose with very short, reclinate, pale-brown setae; with a setaceous, apical, marginal ciliary of short, dark-brown setae, lengthening towards metathoracic tarsomeres, fine dorsally and more robust ventrally. Pretarsomere of all thoracic legs sparse setose with short, semi-erect, red-brown setae, lengthening towards metathoracic pretarsomere; few, medium-long, semi-erect, black setae at apical margin, shorter and red-brown ventrally. Head: Suture between head and clypeus linear, crenated. Head dense punctate with small, shallow, roundish, partly merging punctures. Funicle I – II cylindrical, slightly thickened medially; funicle III short asymmetric frustum-shaped; clavus long elliptic, slightly longer than pedicellus and funiculum combined. Pronotum: Widest medially; dense punctate with shallow, very small, roundish punctures, denser within medial furrow in posterior fifth. Anterior corner obtuse. Lateral margin convex. With an indistinct, shallow, complete medial furrow. Posterior corner angled. Scutellum: Parabola-shaped; slightly longer than broad; dense punctate with shallow, small, roundish punctures, micropunctate within; with a shortened, median, longitudinal, narrow, smooth stripe from anterior margin terminating in apical fifth. Lateral margin slightly convex. Tip rounded. Elytra: Ovate; longer than broad; widest in anterior third; covering propygidium. Lateral margin slightly convex attenuate apically. Sculpture indistinct; slightly pulvinate laterally; humerus indistinct; with a broad roundish torus around scutellum; lateral trench missing; complanate beside scutellum and along suture. Loose punctate with extremely small, shallow, roundish punctures, micropunctures in-between; a sparse juxta-sutural row of very small, shallow, roundish punctures from anterior margin, crossing scutellar torus towards apical torus; a sparse supra-humeral row of small, shallow, roundish punctures from anterior margin between humerus and scutellar torus towards apical torus. Abdomen: Ventrite I distinctly shortened medially; ventrite V broadest, not constricted ventrally. Propygidium with a slightly projected stoma at posterior corner. Pygidium distinctly pulvinate longitudinally and laterally; upper margin slightly convex, linear medially; lateral margin linear attenuate apically; tip rounded. Prothoracic leg: Femur robust. Tibia slender; smooth ventrally. Median tooth elongate. Basal tooth rudimentary. Interspace between teeth almost equal. Shape of interspace between anterior and median tooth u-shaped and between median and basal tooth concave. Tarsomere I – IV combined extremely short; equal in length; slightly less than 2 / 5 in length of tibia. Pretarsomere slightly thickened; with a small, triangular-shaped, median tooth ventrally; slightly less than 1 / 5 more in length of tarsomere I – IV combined. Internal claw 1 / 10 more in length of pretarsomere. External claw slightly less than 4 / 5 in length of internal claw. Mesothoracic leg: Tibia slightly more than 9 / 10 in length of femur. Tarsomere I – IV combined extremely short; slightly more than 3 / 10 in length of tibia. Pretarsomere with a shortened, smooth ridge ventrally, followed by a roundish cutting apically; slightly less than 1 / 10 more in length of tarsomere I – IV combined. External claw 9 / 10 in length of pretarsomere. Internal claw slightly shorter than external claw. Metathoracic leg: Tibia slightly less than 1 / 10 more in length of femur. Tarsomere I – IV combined distinctly short; slightly more than 3 / 5 in length of tibia. Pretarsomere slightly thickened; with an undulate ridge ventrally; 1 / 2 in length of tarsomere I – IV combined. External claw slightly more than 1 / 10 more in length of pretarsomere. Internal claw slightly less than 4 / 5 in length of external claw. Parameres: (Figs. 70 – 71, 104). ♀♀. Measurements: length 8.2 – 9.3 mm; width 4.5 – 4.9 mm. Integument: Black; flagellum brown; elytra transparently red-brown, outer margins sometimes blackened; thoracic legs dark brown, metathoracic tibia and thoracic tarsi paler. Tegument: Clypeus loose setose with short, erect, pale-brown setae, medium-long and brown on head. Eye canthus with a loose ciliary of medium-long, brown setae. Scapus loose setose with medium-long, brown setae, paler ventrally; funiculum with few extremely short, semi-erect, brown setae, sparse on clavus. Pronotum without pattern. Loose setose with medium-long, erect, brown setae; short, semi-erect, whitish setae interspersed posteriorly; with a loose ciliary of medium-long and long, brown setae on lateral margin. Scutellum loose setose with short and medium-long, semi-erect, whitish and brown setae. Elytra without pattern. Sparse setose with short, semi-erect, pale-brown setae, denser along suture, within lateral trench and around apical torus; long semi-erect brown setae interspersed, erect on humerus and scutellar torus. Suture with a loose row of semi-erect short pale-brown and robust, long erect, brown setae. Thorax and pleurites loose pilose with long, pale-brown hairs. Ventrite I – V loose setose with short and medium-long, reclinate, pale-brown setae; long, erect, brown setae interspersed. Propygidium and pygidium loose setose with short, reclinate, pale-brown setae; medium-long, semi-erect, brown setae interspersed. Tip of pygidium with a loose ciliary of medium-long, brown setae. Prothoracic femur loose pilose with long, brown hairs. Prothoracic tibia loose setose with short, reclinate, pale-brown setae dorsally; erect and medium-long ventrally; with a loose setose ciliary of, short, pale-brown setae on internal ridge; anterior margin with a setaceous ciliary of loose, short, pale-brown setae; a medium-long downwards pointing setose ciliary of pale-brown setae along anterior margin of apical tooth. Mesothoracic femur loose pilose with long, brown hairs; paler ventrally. Mesothoracic tibia sparse setose with short, reclinate, pale-brown setae; long, semi-erect, robust, brown setae interspersed; finer, paler, medium-long and long internally. Posterior margin with a short and medium-long bristled marginal ciliary of redbrown bristles; missing internally. Metathoracic femur loose setose with medium-long and long, whitish and brown setae. Metathoracic tibia sparse setose with very short and short, semi-erect, white setae; medium-long, semi-erect, pale-brown setae interspersed, long internally. Ciliary of posterior margin as in male but pale-brown. Setae of thoracic tarsi pale red-brown. Head: Clypeus very slightly depressed laterally; dense punctate with large, shallow, roundish punctures. Anterior margin of clypeus distinctly reflexed vertically. Lateral margin very slightly reflexed. Suture between head and clypeus undulate. Head dense punctate with very small to small, deep, roundish, not merging punctures, micropunctate within. Eye on head level. Clavus short elliptic, as long as pedicellus and funiculum combined. Pronotum: Widest in anterior third; moderate pulvinate laterally; size of punctures very small and small; lateral margin linearly attenuate in anterior third, slightly convex attenuate posteriorly. Posterior corner rounded. Scutellum: Broader than long; dense punctate with very small, deep, roundish punctures; with a median, smooth spot at anterior margin. Elytra: Widest in anterior quarter. Lateral margin slightly convex attenuate apically. Sculpture indistinct; with a broad roundish torus beside scutellum; a shortened, broad juxta-sutural, costa from anterior margin beside scutellum terminating behind disk; lateral trench indistinct. Loose punctate with very small to small, shallow, roundish punctures, micropunctures in-between; a sparse juxta-sutural row of small, shallow, roundish punctures from anterior margin, crossing scutellar torus towards apical torus; a sparse supra-humeral row of small, shallow, roundish punctures from anterior margin between humerus and scutellar torus towards apical torus. Abdomen: Distinctly pulvinate; ventrite I extremely shortened medially; ventrite II – IV not shortened medially; ventrite V broadest, broadened medially, not constricted ventrally. Pygidium slightly pulvinate longitudinally and laterally; slightly declined posteriorly; lateral margin subparallel in upper quarter, linear attenuate apically; tip linear. Prothoracic leg: Median tooth broad, triangular-shaped. Basal tooth very small. Shape of interspace between teeth concave. Tarsomere I – IV combined distinctly short; slightly more than 7 / 10 in length of tibia. Pretarsomere slender; with an undulate ridge ventrally; slightly more than 3 / 5 more in length of tarsomere I – IV combined. Internal claw slender; not broadened medially; slightly less than 9 / 10 in length of pretarsomere. External claw slightly less than 4 / 5 in length of internal claw. Mesothoracic leg: Tibia slightly less than 9 / 10 in length of femur. Tarsomere I – IV combined distinctly short; slightly less than 1 / 2 in length of tibia. Pretarsomere slightly thickened; with a smooth ridge ventrally, roundish cutting missing; slightly less than 4 / 5 in length of tarsomere I – IV combined. External claw slender; not broadened medially; slightly more than 4 / 5 in length of pretarsomere. Internal claw slightly less than 9 / 10 in length of external claw. Metathoracic leg: Tibia slightly extended dorso-apically; slightly more than 7 / 10 in length of femur. Tarsomere I – IV combined slightly more than 4 / 5 in length of tibia. Pretarsomere slender; slightly less than 1 / 2 in length of tarsomere I – IV combined. External claw slightly shorter than pretarsomere.	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E001FFD5FF64D48CFBA9FF74.taxon	distribution	Distribution. Northern Cape Province (Fig. 140), only known from the type locality. Vegetation Type. Nieuwoudtville Shale Renosterveld [FRs 2].	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E001FFD5FF64D48CFBA9FF74.taxon	biology_ecology	Host Plants. Asphodelaceae: Bulbinella latifolia subsp. doleritica (P. L. Perry) P. L. Perry.	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E001FFD5FF64D48CFBA9FF74.taxon	etymology	Etymology. The name derives from the brown elytra [Latin: castane- = brown].	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E03CFFD6FF64D551FCFAFF74.taxon	description	(Figs. 1, 3 – 5, 8, 15, 19 – 22, 34 – 35, 41, 44 – 45, 51, 72 – 73, 105) http: // zoobank. org / 1 B 2 F 14 BA- 5603 - 4601 - AB 77 - 79 A 8273 CC 0 E 9	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E03CFFD6FF64D551FCFAFF74.taxon	materials_examined	Type locality. [RSA, NC]: Farm Glenlyon [now Hantam National Botanical Garden]. Type material. [45] Holotype ♂: white label: pr « RSA, NCP, Nieuwoudtv., | Farm Glenlyon, 31.24 S – | 19.08 E, 22 / 08 / 1998, | leg.: J. Colville » || yellow label hw « 10168 » | pr « coll. Dombrow » || green label pr « Host plant: | Gazania rigida » || blue label pr « Bioregion: | Shale Renosterveld | Vegetation Type: FRs 2 – | Nieuwoudtville Shale Renosterveld » || yellow label pr « SAM – COL | A 071226 » || red type label pr « Holotype | Beckhoplia | colvillei sp. n. | det.: Dombrow 2003 » (SAMC). – Paratypes 1 ♂, 2 ♀♀: same labels as holotype but || yellow label hw « 18 », pr «. 811,. 817,. 848 | coll. Dombrow » || yellow label pr « SAM – COL | A 071226 » || red label pr « Paratype | Beckhoplia | colvillei sp. n. | det.: Dombrow 2003 » (SAMC); 36 ♂♂, 5 ♀♀: same labels as holotype but || yellow label hw « 18 », pr «. 774,. 776 –. 810,. 812 –. 816 | coll. Dombrow » || red label pr « Paratype | Beckhoplia | colvillei sp. n. | det.: Dombrow 2003 » (ESDW). Condition of holotype. Good. Pinned. Dissected, parameres and spiculum gastrale glued on separate card below beetle. Right prothoracic tarsus and right metathoracic tarsus missing after tarsomere I.	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E03CFFD6FF64D551FCFAFF74.taxon	discussion	Remark. The following specimens from the original type series are transferred to several other species, because of the distinct morphological differences see descriptions of the target species: Specimens from the locality “ Nieuwoudtville Wildflower Reserve ” are transferred to B. nigrosetosa sp. n.; specimens with the locality “ Farm Glenlyon … # 210 ” are transferred to B. castanea; specimens from the locality “ Hantam, Nieuwoudtville ” to B. suturalis sp. n.; and specimens from the locality “ Biedouw Valley ” to B. dolichiocnemis sp. n.	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E03CFFD6FF64D551FCFAFF74.taxon	diagnosis	Differential diagnosis. Differs from B. pulchra: Lateral margin of pronotum of B. colvillei widest in anterior half. Lateral margin of pronotum of B. pulchra widest at posterior corner. Externo-posterior margin of parameres of B. colvillei linear with a median convex extension in dorsal view (Figs. 72, 105). Externo-posterior margin of parameres of B. pulchra undulate in dorsal view (Figs. 74, 106). Interno-posterior margin of parameres of B. colvillei very narrow curly bracket-shaped in dorsal view (Figs. 72, 105). Interno-posterior margin of parameres of B. pulchra broad curly bracket-shaped in dorsal view (Figs. 74, 106).	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E03CFFD6FF64D551FCFAFF74.taxon	description	Redescription Measurements: length 6.7 – 8.4 mm; width 3.5 – 4.5 mm. Integument: Black; funiculum pale-brown annulated; elytra transparently pale-brown, margins sometimes blackened. Tegument: Clypeus bare; loose setose with short, erect, brown setae on the inner side of the reflexed anterior and lateral margin. Head loose setose with long, erect, black setae. Eye canthus with a dense setose ciliary of long, black setae. Scapus loose setose with medium-long, black setae, pale-brown ventrally, sparse and short on pedicellus; funiculum with few short, semi-erect, black setae; sparse and brown on clavus. Pronotum loose setose with medium-long and long, erect, brown setae; with a loose setose ciliary of long and distinctly-long, black setae on lateral margin. Scutellum loose setose with short and medium-long, erect, black setae. Elytra without pattern. Loose setose with short, semi-erect, brown setae; medium-long, semi-erect, black setae interspersed, long on humerus and along anterior margin; few short, semi-erect, whitish, squamiform setae interspersed around apical torus; with two sparse rows of medium-long, erect, black setae, slightly shorter in posterior half: a juxta-sutural row from anterior margin, crossing scutellar torus to apical torus; a juxta-humeral row from beside humerus to apical torus. Suture with a loose row of robust, medium-long to distinctly-long, semi-erect, black setae. Thorax and pleurites loose pilose with long, black hairs. Ventrite I – V loose setose with short, reclinate, brownish setae; medium-long and long, semi-erect, black setae interspersed. Propygidium loose squamose with reclinate, narrowly-lanceolate, pale-brown and whitish scales; medium-long and long, semi-erect, black setae interspersed. Pygidium sparse setose with long and distinctlylong, erect, black setae; tip with a loose ciliary of medium-long and long, black setae. Prothoracic femur loose pilose with long, black hairs; bare internally. Prothoracic tibia loose setose with short, reclinate, white setae; short, semi-erect, black setae interspersed dorsally; medium-long, erect, black setae interspersed ventrally; with a sparse setose longitudinal row of short and medium-long, erect, black setae dorsally; a loose setose ciliary of short, black setae on internal ridge; a setaceous ciliary of short, black setae on anterior margin, more robust towards internal ridge; a medium-long, downwards pointing setose ciliary of pale-brown setae along anterior margin of apical tooth. Mesothoracic femur loose pilose with long, black hairs; bare internally. Mesothoracic tibia loose setose with short, reclinate, white setae, sparser internally; medium-long, semi-erect, robust, black setae interspersed, long internally. Posterior margin with a bristled ciliary of very short and short, black bristles; missing internally. Metathoracic femur loose setose with long, erect, black setae, long black hairs interspersed; bare internally. Metathoracic tibia loose setose with short, reclinate, brown setae; medium-long and long, semi-erect, black setae interspersed, denser and long internally. Posterior margin with a loose bristled outer marginal ciliary of short and medium-long, brown bristles, lengthening dorsally and ventrally, more robust ventrally. Prothoracic and mesothoracic tarsomere I – IV bare. Metathoracic tarsomere I – IV sparse setose with short, reclinate, brown setae. Tarsomere I – IV of all thoracic legs with a setaceous, apical, marginal ciliary of short, black setae, lengthening towards metathoracic tarsomeres, fine dorsally and more robust ventrally. Pretarsomere of all thoracic legs sparse setose with short, semi-erect, pale-brown setae; few, medium-long, semi-erect, red-brown setae at apical margin, shorter and paler ventrally. Head: Suture between head and clypeus undulate, crenated. Head dense punctate with small, shallow, roundish, not merging punctures, micropunctate within. Funicle I – II cylindrical, slightly thickened medially; funicle III short asymmetric frustum-shaped; clavus elliptic, slightly longer than pedicellus and funiculum combined. Pronotum: Widest medially; dense punctate with small, shallow, roundish punctures, loose on disk. Anterior corner orthogonal. Lateral margin linearly attenuate anteriorly, subparallel posteriorly. With an indistinct, complete, shallow medial furrow. Posterior corner angled. Scutellum: Buckler-shaped; longer than broad; dense punctate with small to large, shallow, roundish punctures, micropunctate within; with a shortened, median, longitudinal, very narrow, smooth stripe from anterior margin terminating in anterior third. Lateral margin linearly attenuate in anterior two thirds, stronger apically. Tip pointed. Elytra: Ovate; longer than broad; widest medially; protruding propygidium. Lateral margin slightly convex attenuate apically. Sculpture indistinct; slightly pulvinate laterally; humerus indistinct; with a broad roundish torus around scutellum; lateral trench missing; complanate beside scutellum and along suture. Loose punctate with extremely small to very small, shallow, roundish punctures, micropunctures in-between; a sparse juxta-sutural row of small, shallow, roundish punctures from anterior margin, crossing scutellar torus towards apical torus; a sparse supra-humeral row of small, shallow, roundish punctures from anterior margin between humerus and scutellar torus towards apical torus. Abdomen: Ventrite I extremely shortened medially; ventrite V broadest, not constricted ventrally. Propygidium with a slightly projected stoma at posterior corner. Pygidium slightly pulvinate longitudinally and laterally; upper margin moderately convex; lateral margin linearly attenuate apically; tip rounded. Prothoracic leg: Femur robust. Tibia slender; with a longitudinal, medio-ventral ridge. Median tooth elongate. Basal tooth very small. Interspace between teeth almost equal. Shape of interspace between teeth concave. Tarsomere I – IV combined extremely short; lengthening from tarsomere I – IV; slightly more than 2 / 5 in length of tibia. Pretarsomere slightly thickened; with a shortened smooth ridge ventrally; as long as tarsomere I – IV combined. Internal claw slightly less than 1 / 10 more in length of pretarsomere. External claw slightly less than 7 / 10 in length of internal claw. Mesothoracic leg: Tibia slightly more than 4 / 5 in length of femur. Tarsomere I – IV combined extremely short; slightly less than 2 / 5 in length of tibia. Pretarsomere with a shortened smooth ridge ventrally, followed by a roundish cutting apically; 9 / 10 in length of tarsomere I – IV combined. External claw slightly less than 9 / 10 in length of pretarsomere. Internal claw slightly less than 4 / 5 in length of external claw. Metathoracic leg: Tibia slightly more than 9 / 10 in length of femur. Tarsomere I – IV combined distinctly short; slightly more than 7 / 10 in length of tibia. Pretarsomere slightly thickened; with a smooth ridge ventrally; slightly less than 1 / 2 in length of tarsomere I – IV combined. External claw slightly less than 1 / 5 more in length of pretarsomere. Internal claw slightly more than 7 / 10 in length of external claw. Parameres: (Figs. 72 – 73, 105). ♀♀. Measurements: length 8.5 – 9.3 mm; width 4.2 – 5.1 mm. Integument: Black; flagellum brown; thoracic legs dark brown; metathoracic tibia and all thoracic tarsi paler. Tegument: Clypeus loose setose with short, erect, brown setae, medium-long on head. Eye canthus with a loose setose ciliary of medium-long, brown setae. Setae on funiculum and clavus extremely short. Pronotum without pattern. Loose setose with short, reclinate and long, semi-erect, brown setae; with a loose setose ciliary of long, brown setae on lateral margin. Scutellum loose setose with short and medium-long, semi-erect, whitish and brown setae. Elytra loose setose with very short and short, semi-erect, pale-brown setae; medium-long, semi-erect, brown setae interspersed; few long, semi-erect, pale-brown, squamiform setae within lateral trench and around apical torus. Suture with a loose row of short and medium-long, semi-erect, robust, brown setae. Thorax and pleurites loose pilose with long, pale-brown hairs. Ventrite I – V loose setose with short, reclinate, pale-brown setae; medium-long and long, erect, brown setae interspersed. Propygidium and pygidium loose setose with short, reclinate, pale-brown setae; medium-long, semi-erect, brown setae interspersed; tip of pygidium with a loose ciliary of short and mediumlong, brown setae. Prothoracic femur loose pilose with short and medium-long, brown hairs. Prothoracic tibia loose setose with short, reclinate, pale-brown setae dorsally, erect, medium-long and long ventrally; with a setaceous ciliary of short, pale-brown setae on anterior margin. Mesothoracic femur loose pilose with long, brown hairs; paler ventrally. Mesothoracic tibia loose setose with short, reclinate, pale-brown setae; sparser internally; medium-long, semi-erect, robust, red-brown setae interspersed, longer internally. Posterior margin with a bristled ciliary of very short and short, red-brown bristles; lengthening dorsally and ventrally; missing internally. Metathoracic femur loose setose with medium-long and long, whitish and brown setae. Metathoracic tibia sparse setose with short, semi-erect, white setae; medium-long and long, robust, semi-erect, brown setae interspersed, loose and finer internally. Posterior margin with a loose bristled outer marginal ciliary of very short, red-brown bristles, lengthening dorsally and ventrally, more robust ventrally. Ciliary of tarsomere I – IV of thoracic legs red-brown. Pretarsomere of thoracic legs sparse setose with extremely short, semi-erect, pale-brown setae. Head: Clypeus very slightly depressed laterally; dense punctate with large, shallow roundish punctures. Anterior margin of clypeus slightly vertically reflexed. Lateral margin very slightly reflexed. Head dense punctate with small to large, shallow, roundish, partly merging punctures. Eye below head level. Clavus as long as pedicellus and funiculum combined. Pronotum: Moderate pulvinate laterally. Anterior corner orthogonal. Lateral margin convex. Posterior corner rounded. Scutellum: Parabola-shaped; as broad as long; dense punctate with small, shallow, roundish punctures. Lateral margin convex. Tip rounded. Elytra: Widest in anterior quarter; covering propygidium. Sculpture indistinct; scutellar torus missing; lateral trench indistinct. Dense punctate with very small, shallow, roundish punctures, micropunctures in-between. Abdomen: Distinctly pulvinate; ventrite I distinctly shortened medially; ventrite II – V broadened medially; ventrite V not constricted ventrally. Pygidium slightly declined posteriorly; tip linear. Prothoracic leg: Median tooth broad triangular-shaped. Tarsomere I – IV combined very short; equal in length; 7 / 10 in length of tibia. Pretarsomere slender; smooth ventrally, roundish cutting missing; slightly less than 3 / 5 in length of tarsomere I – IV combined. Internal claw slender; not broadened; as long as pretarsomere. External claw slightly less than 4 / 5 in length of internal claw. Mesothoracic leg: Tibia 4 / 5 in length of femur. Tarsomere I – IV combined distinctly short; slightly less than 1 / 2 in length of tibia. Pretarsomere slightly thickened; smooth ventrally, roundish cutting missing; slightly more than 7 / 10 in length of tarsomere I – IV combined. External claw slender; not broadened medially; as long as pretarsomere. Internal claw slightly less than 9 / 10 in length of external claw. Metathoracic leg: Femur slightly thickened. Tibia slightly extended dorso-apically; slightly more than 7 / 10 in length of femur. Tarsomere I – IV combined short; slightly less than 7 / 10 in length of tibia. Pretarsomere slender; with a granulate ridge ventrally. Internal claw slightly less than 4 / 5 in length of external claw.	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E03CFFD6FF64D551FCFAFF74.taxon	distribution	Distribution. Northern Cape Province (Fig. 140), only known from the type locality. Vegetation Type. Nieuwoudtville Shale Renosterveld (FRs 2).	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E03CFFD6FF64D551FCFAFF74.taxon	biology_ecology	Host Plants. Asteraceae: Gazania rigida (Burm. f.) Roessler.	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E03FFFD1FF64D551FC32F818.taxon	description	(Figs. 53, 74 – 75, 106) http: // zoobank. org / AB 9 FCC 8 F-A 5 DA- 4004 - 8143 - DE 08345 C 578 B	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E03FFFD1FF64D551FC32F818.taxon	materials_examined	Type locality. [RSA, NCP]: 2 km SW Farm Papkuilsfontein. Type material. [1] Holotype ♂: white label pr « AFRIKA: Südafrika, Kapprovinz | 2 km SW Farm Papkuilsfontein | 31 ° 34 ’ 21 ” S, 19 ° 10 ’ 52 ” E, 680 m | 30.08.1989, leg. Fritz » || green label pr « Host plant: | Babiana vanziliae » || blue label pr « Bioregion: | Trans-Escarpment Succulent Karroo | Vegetation Type: SKt 2 – | Hantam Karroo » || yellow label hw « 54 », pr «. 921 | coll. Dombrow » || red label pr « Holotype | Beckhoplia | pulchra sp. n. | det.: H. Dombrow 2019 » (ZSM). Condition of holotype. Excellent. Glued on card. Dissected, parameres and spiculum gastrale glued behind beetle.	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E03FFFD1FF64D551FC32F818.taxon	diagnosis	Differential diagnosis. Differs from B. colvillei: Lateral margin of pronotum of B. pulchra widest at posterior corner. Lateral margin of pronotum of B. colvillei widest in anterior half. Externo-posterior margin of parameres of B. pulchra undulate in dorsal view (Figs. 74, 106). Externo-posterior margin of parameres of B. colvillei linear with a median convex extension in dorsal view (Figs. 72, 105). Interno-posterior margin of parameres of B. pulchra broad curly bracket-shaped in dorsal view (Figs. 74, 106). Interno-posterior margin of parameres of B. colvillei very narrow curly bracket-shaped in dorsal view (Figs. 72, 105).	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E03FFFD1FF64D551FC32F818.taxon	description	Description Measurements: length 7.8 mm; width 4.0 mm. Integument: Black; elytra transparently brown. Tegument: Clypeus bare; sparse setose with very short, erect, brown setae on the inner side of the vertically reflexed anterior and lateral margin. Head loose setose with medium-long to long, erect, black setae. Eye canthus with a loose setose ciliary of medium-long, black setae. Scapus loose setose with medium-long, erect, black setae, brown ventrally, sparse and short on pedicellus; funiculum with few very short, erect, black setae; clavus sparse setose with extremely short, semi-erect, brown setae. Pronotum loose setose with medium-long and long, erect, black setae; with a loose setose ciliary of long to distinctly long, black setae on lateral margin. Scutellum loose setose with short to medium-long, semi-erect, brown setae, with a longitudinal, median, bar strip in anterior half. Elytra without pattern; sparse setose with very short, reclinate, brownish setae; short to medium-long, semi-erect, black setae interspersed, lengthening towards anterior margin; reclinate, narrowly-lanceolate, white scales interspersed medially behind disk and around apical torus; with a sparse juxta-sutural row of robust, long, erect, black setae, from anterior margin, crossing scutellar torus towards apical torus. Suture with a loose row of short to long, robust, semi-erect, black setae. Thorax and pleurites loose pilose with long, black hairs. Ventrite I – V sparse setose with short, semi-erect, white setae; medium-long to long, semi-erect, black setae interspersed. Propygidium sparse setose with medium-long, semi-erect, black setae; short, reclinate, white setae interspersed at posterior margin. Pygidium loose setose with medium-long and long, erect, black setae; tip with a sparse ciliary of medium-long, black setae. Prothoracic femur loose pilose with long, black hairs; bare internally. Prothoracic tibia sparse setose with short, semi-erect, offwhite setae, sparser ventrally; medium-long, semi-erect, brown setae interspersed, longer and erect ventrally; with a sparse setose longitudinal row of medium-long, erect, black setae dorsally; a loose setose ciliary of short, black setae on internal ridge; a setaceous ciliary of short, brown setae on anterior margin; a longer, downwards pointing setose ciliary of brown setae along anterior margin of apical tooth. Mesothoracic femur loose pilose with long, black hairs, distinctly long and brown ventrally; bare internally. Mesothoracic tibia sparse setose with short, reclinate, whitish setae; medium-long, semi-erect, black, robust setae interspersed, sparser, finer, medium-long internally. Posterior margin with a bristled ciliary of short, black bristles, lengthening dorsally, missing internally. Metathoracic femur loose setose with long to distinctly long, erect, black setae; pale-brown ventrally; bare internally. Metathoracic tibia sparse setose with very short and short, reclinate, brown setae; medium-long and long, robust, semi-erect, black setae interspersed, finer and longer internally. Posterior margin with a loose bristled outer marginal ciliary of short and medium-long, brown bristles, lengthening dorsally and ventrally. Tarsomere I – IV of all thoracic legs sparse setose with extremely short, reclinate, black setae; with a setaceous, apical, marginal ciliary of short, black setae, lengthening towards metathoracic tarsomeres, fine dorsally and more robust ventrally. Pretarsomere of all thoracic legs sparse setose with very short, reclinate, pale-brown setae; few, medium-long to long, semi-erect, black setae at apical margin, shorter and red-brown ventrally. Head: Suture between head and clypeus linear, complanate. Head dense punctate with small, shallow, roundish, not merging punctures. Funicle I – II cylindrical, slightly thickened medially; funicle III asymmetric frustum-shaped; clavus elliptic, slightly longer than pedicellus and funiculum combined. Pronotum: Widest at anterior two fifth; loose punctate with shallow, very small, roundish punctures, dense in posterior third of medial furrow. Anterior corner obtuse. Lateral margin linearly attenuate anteriorly, subparallel posteriorly. With an indistinct, complete, shallow medial furrow. Posterior corner rounded. Scutellum: Parabola-shaped; longer than broad; dense punctate with shallow, small to large, roundish punctures, micropunctate within; with a shortened, median, longitudinal, narrow, smooth stripe from anterior margin terminating medially. Lateral margin slightly convex. Tip pointed. Elytra: Ovate; broader than long; widest in anterior third; protruding propygidium. Lateral margin slightly convex attenuate apically. Sculpture indistinct; slightly pulvinate laterally; humerus indistinct; with a very broad roundish torus around scutellum; lateral trench missing; complanate beside scutellum and along suture. Loose punctate with very small, shallow, roundish punctures, micropunctures in-between; a sparse juxta-sutural row of small, shallow, roundish punctures from anterior margin, crossing scutellar torus towards apical torus; a sparse supra-humeral row of small, shallow, roundish punctures from anterior margin between humerus and scutellar torus towards apical torus. Abdomen: Ventrite I extremely shortened medially; ventrite V broadest, not constricted ventrally. Propygidium with an indistinct stoma at posterior corner. Pygidium slightly pulvinate longitudinally and laterally; upper margin slightly convex; lateral margin subparallel in upper quarter, linearly attenuate apically; tip rounded. Prothoracic leg: Femur and tibia slender. Tibia smooth ventrally. Median tooth elongate. Basal tooth very small. Interspace between teeth almost equal. Shape of interspace between teeth concave. Tarsomere I – IV combined extremely short; lengthening from tarsomere I – IV; 2 / 5 in length of tibia. Pretarsomere slightly thickened; with two small, triangular-shaped, median teeth consecutive ventrally; slightly more than 2 / 5 in length of tarsomere I – IV combined. Internal claw as long as pretarsomere. External claw 7 / 10 in length of internal claw. Mesothoracic leg: Tibia 7 / 10 in length of femur. Tarsomere I – IV combined very short; slightly more than 2 / 5 in length of tibia. Pretarsomere with a shortened granulate ridge ventrally, followed by a roundish cutting apically; 9 / 10 in length of tarsomere I – IV combined. External claw slightly less than 9 / 10 in length of pretarsomere. Internal claw slightly more than 9 / 10 in length of external claw. Metathoracic leg: Tibia slightly more than 9 / 10 in length of femur. Tarsomere I – IV combined distinctly short; slightly more than 7 / 10 in length of tibia. Pretarsomere slender; with an undulate ridge ventrally; slightly less than 1 / 2 in length of tarsomere I – IV combined. External claw slightly more than 1 / 10 more in length of pretarsomere. Internal claw slightly more than 4 / 5 in length of external claw. Parameres: (Figs. 74 – 75, 106). ♀: Unknown.	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E03FFFD1FF64D551FC32F818.taxon	distribution	Distribution. Northern Cape Province (Fig. 140), only known from the type locality. Vegetation Type. Hantam Karroo (SKt 2).	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E03FFFD1FF64D551FC32F818.taxon	biology_ecology	Host Plants. Iridaceae: Babiana vanziliae L. Bolus.	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E03FFFD1FF64D551FC32F818.taxon	etymology	Etymology. The name derives from the [Latin: pulchr- = beautiful].	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E039FFD2FF64D48CFF4CF93F.taxon	description	(Figs. 46, 50, 76 – 77, 107) http: // zoobank. org / 3 C 324320 - AB 78 - 4 D 58 - 872 C- 649569 AFBFFD	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E039FFD2FF64D48CFF4CF93F.taxon	materials_examined	Type locality. [RSA, NC]: 5 km E Nieuwoudtville. Type material. [11] Holotype ♂: yellow label hw « 50 » pr « 448 | coll. Dombrow » || white label pr « AFRICA: Republic of South Africa | Northern Cape, Nieuwoudtville | 5 km E, 31 ° 17 ’ 39 ’’ S – 19 ° 08 ’ 08 ’’ E | 27.8.2012, lg. J. Colville / A. Melin » || green label pr « Host plant: | Didelta spinosa » || blue label pr « Bioregion: | Sandstone Fynbos | Vegetation Type: FFs 1 – | Bokkeveld Sandstone Fynbos » || yellow label pr « SAM – COL | A 071225 » || red label pr « Holotype | Beckhoplia | nigrofasciata sp. n. | det.: H. Dombrow 2019 » (SAMC). - Paratypes 2 ♂♂, 7 ♀♀: same labels as holotype, but || yellow label hw « 50 » pr «. 449 –. 457 | coll. Dombrow » || red label pr « Paratype | Beckhoplia | nigrofasciata sp. n. | det.: H. Dombrow 2019 » (ESDW); 1 ♀: same labels as holotype but || yellow label hw « 50 » pr «. 458 | coll. Dombrow » || yellow label pr « SAM – COL | A 071225 » || red label pr « Paratype | Beckhoplia | nigrofasciata sp. n. | det.: H. Dombrow 2019 » (SAMC). Condition of holotype. Very good. Glued on card. Dissected, parameres and spiculum gastrale glued behind beetle.	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E039FFD2FF64D48CFF4CF93F.taxon	diagnosis	Differential diagnosis. Differs from B. caliginosa: Posterior corner of pronotum of B. nigrofasciata angled. Posterior corner of pronotum of B. caliginosa rounded. Elytra of B. nigrofasciata longitudinally uni-costae. Elytra of B. caliginosa longitudinally bi-costae. Externo-posterior margin of parameres of B. nigrofasciata convex in dorsal view (Figs. 76, 107). Externo-posterior margin of parameres of B. caliginosa undulate in dorsal view (Figs. 78, 108).	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E039FFD2FF64D48CFF4CF93F.taxon	description	Description Measurements: length 7.8 – 8.1 mm; width 3.9 – 4.0 mm. Integument: Black; elytra transparently pale-brown, outer margins blackened. Tegument: Clypeus bare; loose setose with short, erect, pale-brown to black setae on the inner side of the reflexed anterior and lateral margins. Head loose setose with medium-long and long, erect, black setae. Eye canthus with a dense setose ciliary of long, black setae. Scapus loose setose with medium-long, black setae, brown ventrally, sparser and shorter on pedicellus; funiculum with few very short, semi-erect, black setae, sparse and brown on clavus. Pronotum loose setose with long, erect, black setae; with a loose setose ciliary of long, black, setae on lateral margin. Scutellum loose setose with short and medium-long, semi-erect, brown setae. Elytra with pattern. Sparse setose with short, reclinate and semi-erect, black setae; long, erect, black setae interspersed, denser towards humerus and lateral margin; with a loose juxta-sutural row of long, erect, black setae from anterior margin crossing scutellar torus towards apical torus; with two sparse to loose shortened longitudinal squamose stripes of reclinate, narrowly-lanceolate, white scales; a sutural stripe in posterior quarter; a very short juxta-sutural stripe behind middle; a stripe around apical torus. Suture with a loose row of robust, short to long erect, black setae. Thorax and pleurites loose pilose with long, black hairs. Ventrite I – V loose setose with short, reclinate, white and black setae; medium-long and long semi-erect, black setae interspersed. Propygidium sparse squamose with reclinate, narrowlylanceolate, whitish scales; long, semi-erect, black setae interspersed. Pygidium sparse setose with long, erect, black setae; short, semi-erect, black setae interspersed; tip with a sparse ciliary of long, black, setae. Prothoracic femur loose pilose with long, black hairs; bare internally. Prothoracic tibia loose setose with short, reclinate, black setae, medium-long ventrally; short, semi-erect, white setae interspersed dorsally; with a sparse setose longitudinal row of short and medium-long, erect, black setae dorsally; a loose setose ciliary of short, black setae on internal ridge; a setaceous ciliary of short, brown setae on anterior margin, more robust towards internal ridge; a medium-long downwards pointing setose ciliary of brown setae along anterior margin of apical tooth. Mesothoracic femur loose pilose with long, black hairs; bare internally. Mesothoracic tibia loose setose with short semi-erect, whitish setae, sparser internally; medium-long, semi-erect, robust, black setae interspersed, long and finer internally. Posterior margin with a bristled ciliary of very short and short, red-brown bristles; lengthening dorsally and ventrally; missing internally. Metathoracic femur loose setose with medium-long and long, erect, black setae; bare internally. Metathoracic tibia sparse setose with very short and short reclinate, brown setae; long, robust, semi-erect, black setae interspersed, finer internally. Posterior margin with a loose bristled outer marginal ciliary of short, black bristles, lengthening dorsally and ventrally, more robust ventrally. Tarsomere I – IV of all thoracic legs sparse setose with short, reclinate, brown setae; with a setaceous, apical, marginal ciliary of medium-long, black setae, lengthening towards metathoracic tarsomeres, fine dorsally and more robust ventrally. Pretarsomere of all thoracic legs sparse setose with short, semi-erect, brown setae, lengthening towards metathoracic pretarsomere; few, long, semi-erect, black setae at apical margin, shorter and red-brown ventrally. Head: Suture between head and clypeus missing. Head dense punctate with very small, shallow, roundish, not merging punctures. Funicle I – II cylindrical; funicle III asymmetric frustum-shaped; clavus elliptic, slightly longer than pedicellus and funiculum combined. Pronotum: Widest medially; loose punctate with very small, shallow, roundish punctures. Anterior corner obtuse. Lateral margin linearly attenuate anteriorly and posteriorly. With an indistinct, shallow, complete medial furrow. Posterior corner angled. Scutellum: Parabola-shaped; as broad as long; dense punctate with small, shallow, roundish punctures, micropunctate within; with a shortened, median, longitudinal, narrow, smooth stripe from anterior margin terminating in anterior third. Lateral margin slightly convex. Tip rounded. Elytra: Ovate; longer than broad; widest medially; protruding propygidium. Lateral margin slightly convex attenuate apically. Sculpture indistinct; slightly pulvinate laterally; humerus indistinct; with a broad roundish torus around scutellum; lateral trench missing; complanate beside scutellum and along suture. Loose punctate with extremely small, shallow, roundish punctures, micropunctures in-between; a sparse juxta-sutural row of very small, shallow, roundish punctures from anterior margin, crossing scutellar torus towards apical torus; a sparse supra-humeral row of very small, shallow, roundish punctures from anterior margin between humerus and scutellar torus towards apical torus. Abdomen: Ventrite I extremely shortened medially; ventrite V not constricted ventrally. Propygidium with a slightly projected stoma at posterior corner. Pygidium distinctly pulvinate longitudinally and laterally; upper margin slightly convex; lateral margin subparallel in upper fifth, linearly attenuate apically; tip rounded. Prothoracic leg: Femur and tibia slender. Tibia smooth ventrally. Median tooth elongate. Basal tooth very small. Interspace between teeth almost equal. Shape of interspace between teeth concave. Tarsomere I – IV combined very short; equal in length; slightly more than 1 / 2 in length of tibia. Tarsomere IV with a single ventral extension. Pretarsomere slightly thickened; with a small, triangular-shaped, median teeth ventrally, followed by a roundish cutting apically; slightly more than 9 / 10 in length of tarsomere I – IV combined. Internal claw slightly less than 1 / 10 more in length of pretarsomere. External claw slightly less than 9 / 10 in length of internal claw. Mesothoracic leg: Tibia 3 / 5 in length of femur. Tarsomere I – IV combined extremely short; slightly more than 2 / 5 in length of tibia. Pretarsomere with a shortened smooth ridge ventrally followed by a roundish cutting apically; slightly more than 4 / 5 in length of tarsomere I – IV combined. External claw as long as pretarsomere. Internal claw slightly more than 9 / 10 in length of external claw. Metathoracic leg: Tibia 9 / 10 in length of femur. Tarsomere I – IV combined distinctly short; slightly more than 7 / 10 in length of tibia. Pretarsomere slightly thickened; with an undulate ridge ventrally; slightly more than 2 / 5 in length of tarsomere I – IV combined. External claw slightly more than 1 / 5 more in length of pretarsomere. Internal claw slightly more than 4 / 5 in length of external claw. Parameres: (Figs. 76 – 77, 107). ♀♀. Measurements: length 7.9 – 8.7 mm; width 4.0 – 4.9 mm. Integument: Black; flagellum brown; elytra transparently pale-brown, lateral margins blackened; thoracic legs dark brown, metathoracic tibia and thoracic tarsi paler. Tegument: Clypeus sparse setose with short, semi-erect, brown setae, long, denser and erect on head. Eye canthus with a loose ciliary of long, black setae. Scapus with loose setose with long, brown setae, paler ventrally, very short and pale-brown on pedicellus; funiculum with few extremely short, semi-erect, pale-brown setae, sparse and very short on clavus. Pronotum without pattern. Loose setose with long, erect, brown setae, short towards posterior margin; loose, short, semi-erect, yellowish-white setae in posterior half of medial furrow; with a loose setose ciliary of long brown setae on lateral margin. Scutellum loose setose with short, semi-erect, pale-brown and whitish setae. Elytra with pattern. Sparse setose with short, reclinate and semi-erect, brown setae; long, erect, black setae interspersed; with three longitudinal, sparse to loose setose bands of short, reclinate, yellowish-white setae: a sutural band from anterior margin to posterior margin; a juxta-sutural band from middle, terminating in apical quarter; a lateral band from beside humerus around apical torus, sometimes shortened anteriorly. Thorax and pleurites loose pilose with long, pale-brown hairs. Ventrite I – V loose setose with short, reclinate, yellowish setae; medium-long, semi-erect, brown setae interspersed. Propygidium and pygidium loose setose with short, reclinate, yellowish setae; medium-long, semi-erect, pale-brown setae interspersed; tip of pygidium with a loose setose ciliary of medium-long, brown setae. Prothoracic femur loose pilose with long, pale-brown hairs. Prothoracic tibia loose setose with short, reclinate, brown setae; medium-long ventrally; a loose setose ciliary of short, pale-brown setae on internal ridge; anterior margin with a setaceous ciliary of loose short pale-brown setae; a medium-long downwards pointing ciliary of pale-brown setae along anterior margin of apical tooth. Mesothoracic femur loose pilose with long, brown hairs; yellowish ventrally; bare internally. Mesothoracic tibia sparse setose with short, reclinate, yellowish setae; sparser internally; medium-long, semi-erect, robust, brown setae interspersed, long and finer internally. Metathoracic femur loose setose with long, semi-erect, brown and yellowish setae; bare internally. Metathoracic tibia sparse setose with short, reclinate, pale-brown setae; long and robust, semi-erect, dark brown and red brown setae interspersed, denser and finer internally. Ciliary of posterior margin as in male but pale-brown. Setae of thoracic tarsi red-brown. Head: Clypeus very slightly depressed laterally; dense punctate with large, shallow, roundish punctures. Anterior margin of clypeus slightly reflexed vertically. Lateral margin very slightly reflexed. Suture between head and clypeus undulate, crenated. Head dense punctate with small, shallow, roundish. Eye below head level. Funicle III short asymmetric frustum-shaped; clavus short elliptic, as long as pedicellus and funiculum combined. Pronotum: Moderate pulvinate laterally; dense punctate with small, shallow, roundish punctures, denser in posterior third of medial furrow. Lateral margin convex. Posterior corner rounded. Scutellum: Dense punctate with very small and small, deep, roundish punctures; without smooth stripe. Elytra: Lateral trench indistinct. Size of punctures of juxta-sutural row small. Abdomen: Distinctly pulvinate; ventrite II – IV not shortened medially; ventrite V broadest, broadened medially. Propygidium with an indistinct stoma at posterior corner. Pygidium slightly pulvinate longitudinally and laterally; slightly declined posteriorly; lateral margin subparallel in upper fifth, linearly attenuate apically; tip linear. Prothoracic leg: Femur robust. Median tooth of tibia broad triangular-shaped. Interspace between anterior and median tooth double in width than between median and basal tooth. Tarsomere I – IV combined 7 / 10 in length of tibia. Tarsomere IV without ventral extension. Pretarsomere slender; smooth ventrally, roundish cutting missing; slightly more than 3 / 5 in length of tarsomere I – IV combined. Internal claw slender; not broadened medially; slightly less than 9 / 10 in length of pretarsomere. External claw slightly more than 4 / 5 in length of internal claw. Mesothoracic leg: Tibia 4 / 5 in length of femur. Tarsomere I – IV combined very short; slightly less than 1 / 2 in length of tibia. Pretarsomere slender; roundish cutting missing; slightly more than 7 / 10 in length of 1 tarsomere I – IV combined. External claw slender; not broadened medially; slightly less than 9 / 10 in length of pretarsomere. Internal claw slightly less than 9 / 10 in length of external claw. Metathoracic leg: Femur slightly thickened. Tibia slightly extended dorso-apically; slightly more than 4 / 5 in length of femur. Tarsomere I – IV combined short; slightly more than 4 / 5 in length of tibia. Pretarsomere slender. External claw slightly less than 1 / 10 more in length of pretarsomere. Internal claw slightly less than 9 / 10 in length of external claw.	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E039FFD2FF64D48CFF4CF93F.taxon	distribution	Distribution. Northern Cape Province (Fig .. 140), only known from the type locality. Vegetation Type. Bokkeveld Sandstone Fynbos (FFs 1).	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E039FFD2FF64D48CFF4CF93F.taxon	biology_ecology	Host Plants. Asteraceae: Didelta spinosa (L. f.) W. T. Aiton.	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E039FFD2FF64D48CFF4CF93F.taxon	etymology	Etymology. The name derives from the black band on the outer margins [Latin: nigr- = black; Latin: fasciat- = banded]	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E03BFFDBFF64D318FB3BFE04.taxon	description	(Figs. 17, 37, 78 – 79, 100, 108, 118) http: // zoobank. org / D 024 B 073 - 9088 - 4 B 4 D- 823 B- 5 DAC 1 ADDCDAA	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E03BFFDBFF64D318FB3BFE04.taxon	materials_examined	Type locality. [RSA, NCP]: Botterkloof Pass. Type material. [42] Holotype ♂: yellow label hw « 54 » pr «. 992 | coll. Dombrow » || white label pr « RSA, WC, Botterkloof Pass | 31 ° 49,978 ’ S – 19 ° 16,093 ’ E | 524 m, 29.08.2012 | leg: J. F. Colville & A. Melin » || green label pr « Host plant: | Didelta spinosa » || blue label pr « Bioregion: | Rainshadow Valley Karroo | Vegetation Type: SKv 3 – | Agter-Sederberg Shrubland » || yellow label pr « SAM – COL | A 071224 » || red label pr « Holotype | Beckhoplia | caliginosa sp. n. | det.: H. Dombrow 2019 » (SAMC). – Paratypes 7 ♂♂, 22 ♀♀: same labels as holotype but || yellow label hw « 50 » pr «. 379 –. 407 | coll. Dombrow » || red label pr « Paratype | Beckhoplia | caliginosa sp. n. | det.: H. Dombrow 2019 » (ESDW); 4 ♀♀: same labels as holotype but || yellow label hw « 50 » pr «. 408 –. 411 | coll. Dombrow » || yellow label pr « SAM – COL | A 071224 » || red label pr « Paratype | Beckhoplia | caliginosa sp. n. | det.: H. Dombrow 2019 » (SAMC); 6 ♀♀: white label pr « RSA, NCP, Botterkloof | Pass, 31.48 S – 19.16 E | 25 / 08 / 1999 | leg.: J. Colville » || green label pr « Host plant: | Didelta spinosa » || blue label pr « Bioregion: | Rainshadow Valley Karroo | Vegetation Type: SKv 3 – | Agter-Sederberg Shrubland » || yellow label hw « 18 » pr «. 471 –. 476 | coll. Dombrow » || red label pr « Paratype | Beckhoplia | occidentalis sp. n. | det.: Dombrow 2003 » || red label pr « Paratype | Beckhoplia | caliginosa sp. n. | det.: H. Dombrow 2019 » (ESDW); 2 ♀♀: same labels but || yellow label hw « 54 » pr «. 993 –. 994 | coll. Dombrow » || yellow label pr « SAM – COL | A 071224 » || red label pr « Paratype | Beckhoplia | occidentalis sp. n. | det.: H. Dombrow 2003 » || red label pr « Paratype | Beckhoplia | caliginosa sp. n. | det.: H. Dombrow 2019 » (SAMC). Condition of holotype. Very good. Glued on card. Dissected, parameres and spiculum gastrale glued behind beetle.	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E03BFFDBFF64D318FB3BFE04.taxon	diagnosis	Differential diagnosis. Differs from B. nigrofasciata: Posterior corner of pronotum of B. caliginosa rounded. Posterior corner of pronotum of B. nigrofasciata angled. Elytra of B. caliginosa longitudinally bi-costae. Elytra of B. nigrofasciata longitudinally uni-costae. Externo-posterior margin of parameres of B. caliginosa undulate in dorsal view (Figs. 78, 108). Externo-posterior margin of parameres of B. nigrofasciata convex in dorsal view (Figs. 76, 107).	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E03BFFDBFF64D318FB3BFE04.taxon	description	Description Measurements: length 6.9 – 7.8 mm; width 3.5 – 4.0 mm. Integument: Black; scutellar torus on elytra sometimes brown. Tegument: Clypeus bare; sparse setose with short, erect, black setae on the inner side of the reflexed anterior and lateral margins. Head loose setose with long and distinctly long, erect, black setae. Eye canthus with a dense setose ciliary of long, black setae. Scapus loose setose with long, black setae, brown ventrally, sparser and shorter on pedicellus; funiculum with few very short, semi-erect, black setae, sparse on clavus. Pronotum loose setose with long and distinctly long, erect, black setae; with a setose ciliary of distinctly long black setae, dense in anterior half, loose in posterior half. Scutellum loose setose with short and medium-long, erect, black setae. Elytra with pattern. Loose setose with short, semi-erect, black setae; long to distinctly long, erect, black setae interspersed, denser anteriorly; with two loose longitudinal rows of long, erect, black setae: a juxta-sutural row from anterior margin crossing scutellar torus towards apical torus, shorter in posterior third and a supra-humeral row from anterior margin between humerus and scutellar torus towards apical torus terminating towards middle; with three shortened, longitudinal, loose squamose stripes of reclinate, narrowly-lanceolate, grey and white scales: a sutural stripe in posterior third; a juxta-sutural stripe from behind middle, terminating towards apical quarter; a lateral stripe from anterior third, within lateral trench, less dense around apical torus. Suture with a loose setose row of medium-long to distinctly long, robust, erect, black setae. Thorax and pleurites loose pilose with long, black hairs. Ventrite I – V loose setose with short, reclinate and long, semi-erect, black setae. Propygidium loose setose with short, reclinate, black setae; long, semi-erect, black setae interspersed. Pygidium sparse setose with medium-long and long, erect, black setae; tip with a loose setose ciliary of medium-long and long, black, setae. Prothoracic femur loose pilose with long, black hairs; bare internally. Prothoracic tibia loose setose with short, semi-erect, brown setae; erect medium-long and long ventrally; with a sparse setose longitudinal row of short and medium-long, erect, black setae dorsally; a loose setose ciliary of short, black setae on internal ridge; a setaceous ciliary of short, brown setae on anterior margin; a medium-long downwards pointing setose ciliary of brown setae along anterior margin of apical tooth. Mesothoracic femur loose pilose with long and distinctly long, black hairs; bare internally. Mesothoracic tibia sparse setose with short, reclinate, brown setae; mediumlong, semi-erect, robust, black setae interspersed, finer, medium-long and long internally. Posterior margin with a bristled ciliary of short and medium-long, black bristles; missing internally. Metathoracic femur loose setose with long, erect, black setae; whitish ventrally; bare internally. Metathoracic tibia sparse setose with short, semierect, black setae; medium-long and long, erect, black setae interspersed; sparser and finer internally. Posterior margin with a loose bristled outer marginal ciliary of short, black bristles, distinctly lengthening and more robust dorsally and ventrally. Tarsomere I – IV of all thoracic legs with sparse setose with short, reclinate, pale-brown setae, lengthening and darker towards metathoracic tarsomeres; a setaceous, apical, marginal ciliary of short, black setae, lengthening towards metathoracic tarsomeres, fine dorsally and more robust ventrally. Pretarsomere of all thoracic legs sparse setose with short, semi-erect, black setae, lengthening towards metathoracic pretarsomere; few, long, semi-erect, black setae at apical margin, shorter ventrally. Head: Suture between head and clypeus undulate, interrupted medially, crenated. Head dense punctate with small, shallow, roundish, not merging punctures, micropunctate within. Funicle I – II cylindrical, slightly thickened medially; funicle III short asymmetric frustum-shaped; clavus elliptic, slightly longer than pedicellus and funiculum combined. Pronotum: Widest at anterior two fifth; loose punctate with small, shallow, roundish punctures, dense within anterior and posterior fifth of medial furrow. Anterior corner obtuse. Lateral margin convex. With an indistinct, complete, shallow medial furrow. Posterior corner rounded. Scutellum: Buckler-shaped; as broad as long; dense punctate with small, shallow, roundish punctures. Lateral margin linearly attenuate in anterior quarter, stronger apically. Tip pointed. Elytra: Ovate; longer than broad; widest in anterior third; protruding propygidium. Lateral margin slightly convex attenuate apically. Sculpture distinct; slightly pulvinate laterally; humerus distinct; with a broad roundish torus around scutellum; lateral trench distinct; with a distinct juxta-sutural costa from anterior margin beside scutellum towards apical torus terminating in posterior quarter; an indistinct supra-humeral costa from anterior margin beside humerus towards apical torus terminating on disk; slightly narrowly depressed along scutellum; complanate along suture. Loose punctate with very small, shallow, roundish punctures, micropunctures in-between; a sparse juxta-sutural row of very small, shallow, roundish punctures from anterior margin, crossing scutellar torus towards apical torus; a sparse supra-humeral row of very small, shallow, roundish punctures from anterior margin between humerus and scutellar torus towards apical torus. Abdomen: Ventrite I extremely shortened medially; ventrite V slightly constricted ventrally. Propygidium with a slightly projected stoma at posterior corner. Pygidium distinctly pulvinate longitudinally and laterally; upper margin slightly convex; lateral margin linearly attenuate apically; tip rounded. Prothoracic leg: Femur and tibia slender. Tibia smooth ventrally. Median tooth elongate. Basal tooth rudimentary. Interspace between teeth almost equal. Shape of interspace between anterior and median tooth u-shaped and between median and basal tooth slightly concave. Tarsomere I – IV combined very short; equal in length; slightly less than 7 / 10 in length of tibia. Tarsomere III – IV with a single ventral extension. Pretarsomere distinctly thickened; smooth ventrally; slightly more than 7 / 10 in length of tarsomere I – IV combined. Internal claw slightly longer than pretarsomere. External claw slightly less than 7 / 10 in length of internal claw. Mesothoracic leg: Tibia slightly less than 9 / 10 in length of femur. Tarsomere I – IV combined extremely short; slightly less than 2 / 5 in length of tibia. Pretarsomere with a shortened smooth ridge ventrally followed by a roundish cutting apically; slightly less than 1 / 10 more in length of tarsomere I – IV combined. External claw slightly more than 4 / 5 in length of pretarsomere. Internal claw slightly shorter than external claw. Metathoracic leg: Tibia as long as femur. Tarsomere I – IV combined distinctly short; slightly more than 4 / 5 in length of tibia. Pretarsomere slightly thickened; with a shortened, undulate ridge ventrally; slightly less than 1 / 2 in length of tarsomere I – IV combined. External claw slightly longer than pretarsomere. Internal claw less more than 9 / 10 in length of external claw. Parameres: (Figs. 78 – 79, 108). ♀♀. Measurements: length 6.8 - 8.1 mm; width 3.9 - 4.4 mm. Integument: Black; elytra transparently pale to dark brown, margins and costae blackened; thoracic tarsi pale red-brown to red-brown. Tegument: Clypeus sparse setose with short, erect, pale-brown setae. Head loose setose with short and medium-long, erect, brownish setae. Setae on funiculum and clavus brown. Eye canthus with a loose setose ciliary of long, brown setae. Scapus loose setose with medium-long, brown setae, paler ventrally. Pronotum without pattern. Loose setose with medium-long and long, erect, brown setae; short, semi-erect, pale-brown setae interspersed in posterior half of medial furrow; with a setose ciliary of long and distinctly long brown setae, dense in anterior half, loose in posterior half on lateral margin. Scutellum loose setose with short and medium-long, erect, brown setae. Elytra with pattern. Loose setose with short, semi-erect, brown setae; long and distinctly long, erect, brown setae interspersed, denser anteriorly; with three shortened, longitudinal, loose setose stripes of short, reclinate, yellowishorange setae: a sutural stripe in posterior third, a juxta-sutural stripe from behind middle, terminating towards apical third; an indistinct lateral stripe from anterior third, within lateral trench, sometimes around apical torus. Suture with a loose setose row of short to long, robust, semi-erect, brown setae. Thorax and pleurites loose pilose with long, brown hairs. Ventrite I – V loose setose with short to medium-long, reclinate and long semi-erect, pale-brown setae; long, semi-erect, pale-brown and dark-brown setae interspersed. Propygidium and pygidium loose setose with short, reclinate, brown setae; medium-long and long, semi-erect, pale-brown setae interspersed; tip of pygidium with a loose setose ciliary of medium-long and long, brown setae. Prothoracic femur loose pilose with long, brown hairs. Prothoracic tibia loose setose with short, semi-erect, brown setae; medium-long and pale-brown ventrally; a loose setose ciliary of short, pale-brown setae on internal ridge; anterior margin with a setaceous ciliary of loose short pale-brown setae; a medium-long downwards pointing setose ciliary of pale-brown setae along anterior margin of apical tooth. Mesothoracic femur loose pilose with long, brown hairs; distinctly long and pale-brown ventrally. Mesothoracic tibia sparse setose with short, reclinate, brown setae; medium-long, semi-erect, robust, brown setae interspersed, long and finer internally. Posterior margin as in male but red-brown. Metathoracic femur loose setose with long, semi-erect, brown setae; distinctly long and pale-brown ventrally. Metathoracic tibia sparse setose with short, semi-erect, brown setae; medium-long and long, semi-erect, brown setae interspersed; sparse setose with, long, erect, brown setae internally. Posterior margin as in male but setae red-brown. Setae of thoracic tarsi pale red-brown. Head: Clypeus very slightly depressed laterally; loose to dense punctate with very small, shallow, roundish punctures. Anterior margin slightly vertically reflexed. Lateral margin very slightly reflexed. Head dense punctate with very small and small, shallow, roundish, partly merging punctures. Eye on head level. Funicle III very short asymmetric frustum-shaped; clavus short elliptic. Pronotum: Moderate pulvinate laterally; dense punctate with very small and small, shallow, roundish punctures, denser within medial furrow and posteriorly. Scutellum: Parabola-shaped; broader than long; dense punctate with very small, deep, roundish punctures, micropunctate within; with a shortened, median, longitudinal, narrow, smooth stripe from anterior margin terminating medially. Lateral margin convex. Elytra: Sculpture indistinct; humerus indistinct; lateral trench indistinct; costae indistinct. Dense punctate with very small, shallow, roundish punctures, micropunctures in-between; loose and small on costae. Abdomen: Distinctly pulvinate; ventrite II – V broadened medially; ventrite V not constricted ventrally. Pygidium slightly pulvinate longitudinally and laterally; slightly declined posteriorly; upper margin slightly convex, linear medially; tip linear. Prothoracic leg: Femur robust. Median tooth of tibia broad triangular-shaped. Basal tooth very small. Shape between teeth concave. Tarsomere I – IV combined distinctly short; equal in length; slightly more than 9 / 10 in length of tibia. Tarsomere III – IV without ventral extension. Pretarsomere slightly thickened; without roundish cutting apically; slightly more than 3 / 5 in length of tarsomere I – IV combined. Internal claw slender; not broadened medially; slightly less than 9 / 10 in length of pretarsomere. External claw slightly more than 7 / 10 in length of internal claw. Mesothoracic leg: Tibia slightly more than 7 / 10 in length of femur. Tarsomere I – IV combined very short; slightly more than 1 / 2 in length of tibia. Pretarsomere slightly thickened; with a smooth ridge ventrally, roundish cutting missing; slightly less than 7 / 10 in length of tarsomere I – IV combined. External claw slender; not broadened medially; slightly more than 7 / 10 in length of pretarsomere. Internal claw slightly more than 9 / 10 in length of external claw. Metathoracic leg: Tibia slightly extended dorso-apically; slightly less than 7 / 10 in length of femur. Tarsomere I – IV combined short; slightly longer than tibia. Pretarsomere with a granulate ridge ventrally; slightly less than 2 / 5 in length of tarsomere I – IV combined. External claw as long as pretarsomere. Internal claw slightly more than 9 / 10 in length of external claw.	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E03BFFDBFF64D318FB3BFE04.taxon	distribution	Distribution. Western Cape Province (Fig. 140), only known from the type locality. Vegetation Type. Agter-Sederberg Shrubland (SKv 3).	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E03BFFDBFF64D318FB3BFE04.taxon	biology_ecology	Host Plants. Asteraceae: Didelta spinosa (L. f.) W. T. Aiton.	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E03BFFDBFF64D318FB3BFE04.taxon	etymology	Etymology. The name derives from the black colour of the beetle [Latin: caligin- = dark].	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E032FFC4FF64D5C1FB39FD7C.taxon	description	(Figs. 25, 80 – 81, 109) http: // zoobank. org / B 1 E 5 A 044 - A 23 B- 41 FA-BED 0 - 9 D 951 DDD 50 DC	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E032FFC4FF64D5C1FB39FD7C.taxon	materials_examined	Type locality. [RSA, NC]: Farm Nuwedam. Type material. [12] Holotype ♂: white label pr « SOUTH AFRICA Western | Cape, Vanrhynsdorp, Farm | Nuwedam, 31.48 S – 18.41 E | 18.08.2002, leg:. J. Colville » || yellow label hw « 18 » pr «. 482 | coll. Dombrow » || green label pr « Host plant: | Didelta spinosa | Berkheya fruticosa | Arctotheca calendula » || blue label pr « Bioregion: | Sandstone Fynbos | Vegetation Type: FFs 1 – | Bokkeveld Sandstone Fynbos » || yellow label pr « SAM – COL | A 071223 » || red label pr « Holotype | Beckhoplia | pumilla sp. n. | det.: H. Dombrow 2019 » (SAMC). – Paratypes 1 ♂: same labels as holotype but || yellow label hw « 18 » pr «. 484 | coll. Dombrow » || red label pr « Paratype | Beckhoplia | pumilla sp. n. | det.: H. Dombrow 2019 » (ESDW); 4 ♀♀: same labels as holotype but || yellow label hw « 54 » pr «. 789 –. 792 | coll. Dombrow » || red label pr « Paratype | Beckhoplia | pumilla sp. n. | det.: H. Dombrow 2019 » (ESDW); 1 ♀: same labels but || yellow label hw « 54 » pr «. 793 | coll. Dombrow » || yellow label pr « SAMCOL | A 071223 » || red label pr « Paratype | Beckhoplia | pumilla sp. n. | det.: H. Dombrow 2019 » (SAMC); 4 ♀♀: same labels as holotype, but | 01.09.2000, leg. J. Colville » || yellow label hw « 18 » pr «. 477 –. 478,. 480 –. 481 | coll. Dombrow » || red label pr « Paratype | Beckhoplia | pumilla sp. n. | det.: H. Dombrow 2019 » (ESDW); 1 ♀: same labels but || yellow label hw « 18 » pr «. 479 | coll. Dombrow » || yellow label pr « SAM – COL | A 071223 » || red label pr « Paratype | Beckhoplia | pumilla sp. n. | det.: H. Dombrow 2019 » (SAMC). Condition of holotype. Good. Glued on a card. Dissected, parameres and spiculum gastrale glued behind beetle. Left prothoracic tarsus missing.	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E032FFC4FF64D5C1FB39FD7C.taxon	diagnosis	Differential diagnosis. Differs from B. gifbergensis: Pronotum of B. pumilla widest in anterior two fifth. Pronotum of B. gifbergensis widest medially. Elytra of B. pumilla yellowish pale-brown. Elytra of B. gifbergensis transparently pale-brown. Elytra of B. pumilla protruding propygidium. Elytra of B. gifbergensis covering propygidium. Externo-posterior margin of parameres of B. pumilla undulate in dorsal view (Figs. 80, 109). Externo-posterior margin of parameres of B. gifbergensis slightly convex in dorsal view (Figs. 82, 110). Anterior margin of phallobase of B. pumilla deep v-shaped incised in dorsal view (Figs. 80, 109). Anterior margin of phallobase of B. gifbergensis concave in dorsal view (Figs. 82, 110).	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E032FFC4FF64D5C1FB39FD7C.taxon	description	Description Measurements: length 6.5 – 7.1 mm; width 3.4 – 3.5 mm. Integument: Black; elytra yellowish pale-brown, lateral and posterior margin blackened. Tegument: Clypeus bare; sparse setose with very short and short, erect, black setae on the inner side of the reflexed anterior and lateral margins. Head loose setose with long, erect, black and whitish setae. Eye canthus with a loose setose ciliary of very long, black setae. Scapus loose setose with, long, black setae, pale-brown ventrally, sparser and shorter on pedicellus; funiculum with few extremely short and short, erect, pale-brown setae, sparse and semi-erect on clavus. Pronotum loose setose with medium-long and long, erect, black and whitish setae; with a loose setose ciliary of long black setae. Scutellum loose setose with short and medium-long, semi-erect, brown setae. Elytra with pattern. Sparse setose with short, reclinate, black setae, denser laterally; long, erect, black setae interspersed; with a very sparse juxta-sutural row of long, erect, black setae from anterior margin crossing scutellar torus towards apical torus; two shortened, longitudinal, loose squamose stripes of reclinate narrowly-lanceolate, white scales: a sutural stripe in posterior third; a lateral stripe in posterior quarter and around apical torus. Suture with a loose row of long, robust, erect, black setae. Thorax and pleurites loose pilose with long, whitish hairs; black on prothorax. Ventrite I – V sparse setose with medium-long and long, reclinate, black setae; reclinate narrowly-lanceolate white scales interspersed at upper margin of ventrite V. Propygidium loose squamose with reclinate, narrowly-lanceolate, white scales; long, semi-erect, black setae interspersed. Pygidium sparse setose with medium-long and long, semi-erect, black setae; tip with a loose setose ciliary of long, black setae. Prothoracic femur sparse pilose with medium-long and long, black hairs; bare internally. Prothoracic tibia loose setose with short, reclinate, black and whitish setae, medium-long and long ventrally; with a sparse setose longitudinal row of short and medium-long, erect, black setae dorsally; a loose setose ciliary of short, black setae on internal ridge; a setaceous ciliary of short, black setae on anterior margin, more robust towards internal ridge; a medium-long downwards pointing setose ciliary of pale-brown setae along anterior margin of apical tooth. Mesothoracic femur sparse pilose with long, black hairs; distinctly long and whitish ventrally; bare internally. Mesothoracic tibia sparse setose with short, reclinate, white and black setae; medium-long, semi-erect, robust, black setae interspersed, long and finer internally. Posterior margin with a bristled ciliary of short, black bristles; missing internally. Metathoracic femur loose setose with long, semi-erect, black setae; distinctly long and whitish ventrally; bare internally. Metathoracic tibia sparse setose with short, reclinate, black setae; medium-long, semi-erect, black setae interspersed, finer internally. Posterior margin with a loose bristled outer marginal ciliary of short, black bristles, lengthening dorsally and ventrally. Prothoracic and mesothoracic tarsomere I – IV bare. Metathoracic tarsomere I – IV sparse setose with short, reclinate, pale-brown setae. Tarsomere I – IV of all thoracic legs with a setaceous, apical, marginal ciliary of short, dark-brown setae, lengthening towards metathoracic tarsomeres, fine dorsally and more robust ventrally. Pretarsomere of all thoracic legs with few, medium-long, semi-erect, black setae at apical margin, shorter ventrally. Pretarsomere of metathoracic leg additionally sparse setose with very short, reclinate, pale-brown setae. Head: Suture between head and clypeus linear, crenated. Head dense punctate with very small, shallow, roundish, not merging punctures, micropunctate within. Funicle I – II cylindrical, slightly thickened medially; funicle III short asymmetric frustum-shaped; clavus long elliptic, slightly longer than pedicellus and funiculum combined. Pronotum: Widest at anterior two fifth; loose punctate with small, shallow, roundish punctures denser within medial furrow. Anterior corner obtuse. Lateral margin linearly attenuate anteriorly and posteriorly. With an indistinct complete, shallow, medial furrow. Posterior corner angled. Scutellum: Buckler-shaped; slightly broader than long; dense punctate with small, shallow, roundish punctures, micropunctate within; with a shortened, median, longitudinal, broad, smooth stripe from anterior margin terminating in apical fifth. Lateral margin linearly attenuate in anterior third, stronger apically. Tip pointed. Elytra: Ovate; longer than broad; widest in anterior third; protruding propygidium. Lateral margin slightly convex attenuate apically. Sculpture indistinct; very slightly pulvinate laterally; humerus indistinct; with a very broad roundish torus around scutellum; lateral trench indistinct; complanate along scutellum and suture. Sparse punctate with extremely small, shallow, roundish punctures, micropunctures in-between; a sparse juxta-sutural row of very small, shallow and deep, roundish punctures from anterior margin, crossing scutellar torus towards apical torus; a sparse supra-humeral row of very small, shallow and deep, roundish punctures from anterior margin between humerus and scutellar torus towards apical torus. Abdomen: Ventrite I distinctly shortened medially; ventrite V broadest, slightly constricted ventrally. Propygidium with a slightly projected stoma at posterior corner. Pygidium distinctly pulvinate longitudinally and laterally; upper margin very slightly convex, linear medially; lateral margin linearly attenuate apically; tip rounded. Prothoracic leg: Femur and tibia robust. Tibia smooth ventrally. Median tooth elongate. Basal tooth rudimentary. Interspace between anterior and median tooth double in width than between median and basal tooth. Shape of interspace between anterior and median tooth u-shaped and between median and basal tooth slightly concave. Tarsomere I – IV combined extremely short; equal in length; slightly more than 2 / 5 in length of tibia. Tarsomere IV with a single ventral extension. Pretarsomere distinctly thickened; with a shortened smooth ventrally; slightly less than 1 / 10 more in length of tarsomere I – IV combined. Internal claw slightly more than 1 / 10 more in length of pretarsomere. External claw slightly less than 3 / 5 in length of internal claw. Mesothoracic leg: Tibia slightly less than 4 / 5 in length of femur. Tarsomere I – IV combined extremely short; slightly more than 3 / 10 in length of tibia. Pretarsomere with a shortened smooth ridge ventrally, roundish cutting missing; as long as tarsomere I – IV combined. External claw as long as pretarsomere. Internal claw slightly less than 9 / 10 in length of external claw. Metathoracic leg: Tibia slightly less than 9 / 10 in length of femur. Tarsomere I – IV combined very short; slightly less than 4 / 5 in length of tibia. Pretarsomere slightly thickened; with an undulate ridge ventrally; slightly less than 1 / 2 in length of tarsomere I – IV combined. External claw slightly less than 1 / 10 more in length of pretarsomere. Internal claw 4 / 5 in length of external claw. Parameres: (Figs. 80 – 81, 109). ♀♀. Measurements: length 6.8 – 8.2 mm; width 3.8 – 4.4 mm. Integument: Black; lateral and posterior margin of elytra darkened; metathoracic tibia and thoracic tarsi redbrown. Tegument: Clypeus sparse setose with short, erect, brown setae, loose and medium-long on head. Eye canthus with a loose setose ciliary of medium-long, black setae. Setae on funiculum and clavus very short. Pronotum with pattern. Loose setose with long, erect, black setae, short in posterior half; with three sparse longitudinal setose bands of short semi-erect white setae in posterior half: a medial band within medial furrow; a juxta-median band on each side. Elytra with pattern. Sparse setose with very short, semi-erect, brown setae, denser laterally; medium-long, erect, brown setae interspersed, denser towards humerus; two shortened, sparse, longitudinal, squamose stripes of reclinate narrowly-lanceolate, white scales: a sutural stripe in posterior third; a lateral stripe in posterior half of lateral trench and around apical torus. Thorax and pleurites loose pilose with long, yellowish-white hairs; brown on prothorax. Ventrite I – V loose setose with short and medium-long, reclinate, whitish setae; long, semi-erect, black setae interspersed. Propygidium and pygidium loose setose with short and medium-long, reclinate, yellowish-white setae. Tip of pygidium with a loose setose ciliary of long, brown, setae. Prothoracic femur sparse pilose with long, brown hairs. Prothoracic tibia loose setose with short and medium-long, reclinate and semi-erect, brown setae; medium-long and long ventrally; a loose setose ciliary of short, pale-brown setae on internal ridge; anterior margin with a setaceous ciliary of loose, short, red-brown setae; a medium-long downwards pointing setose ciliary of red-brown setae along anterior margin of apical tooth. Mesothoracic femur sparse pilose with long, brown hairs. Mesothoracic tibia sparse setose with short, reclinate, whitish setae. Posterior margin as in male but red-brown. Metathoracic femur loose setose with medium-long and long, semi-erect, whitish and brown setae; only whitish and long ventrally. Metathoracic tibia sparse setose with short, reclinate, pale-brown setae; medium-long semi-erect, red-brown setae interspersed; sparse setose with medium-long, semi-erect, brown setae internally, lengthening towards tarsus. Posterior margin with a loose bristled outer marginal ciliary of very short, red-brown bristles, lengthening and more robust dorsally and ventrally. Setae of thoracic tarsi pale red-brown. Head: Clypeus very slightly depressed laterally; dense punctate with small, shallow, roundish punctures. Anterior margin of clypeus distinctly vertically reflexed. Lateral margin very slightly reflexed. Suture between head and clypeus undulate. Head dense punctate with small and large, shallow, roundish, not micropunctate within. Eye on head level. Funicle I – II cylindrical; clavus short elliptic. Pronotum: Widest medially; moderate pulvinate laterally; loose punctate with very small and small, shallow, roundish punctures, micropunctate within, dense within medial furrow. Anterior corner orthogonal. Lateral convex. With an indistinct shortened, shallow, medial furrow from posterior margin terminating in anterior third. Posterior corner rounded. Scutellum: Without smooth stripe. Tip rounded. Elytra: Sculpture indistinct; slightly pulvinate laterally; with a broad roundish torus around scutellum. Size of punctures of juxta-sutural and supra-humeral row small. Abdomen: Distinctly pulvinate; ventrite II – V broadened medially; ventrite V not constricted ventrally. Propygidium with an indistinct stoma at posterior corner. Pygidium slightly pulvinate longitudinally and laterally; slightly declined posteriorly; tip linear. Prothoracic leg: Median tooth of tibia broad triangular-shaped. Basal tooth very small. Shape of interspace between teeth concave. Tarsomere I – IV combined distinctly short; tarsomere I petiolate chalice-shaped; tarsomere I longest, tarsomere II – IV equal in length; slightly less than 4 / 5 in length of tibia. Tarsomere IV without ventral extension. Pretarsomere slightly thickened; smooth ventrally, roundish cutting missing; 3 / 5 in length of I – IV tarsomere combined. Internal claw slender; not broadened medially; slightly less than 4 / 5 in length of pretarsomere. External claw slightly less than 9 / 10 in length of internal claw. Mesothoracic leg: Tarsomere I – IV combined distinctly short; slightly less than 3 / 5 in length of tibia. Pretarsomere slightly thickened; with an undulate ridge ventrally; slightly less than 7 / 10 in length of tarsomere I – IV combined. External claw slender; not broadened medially; slightly more than 4 / 5 in length of pretarsomere. Internal claw slightly more than 4 / 5 in length of external claw. Metathoracic leg: Femur slightly thickened. Tibia distinctly broadened dorso-apically; 7 / 10 in length of femur. Tarsomere I – IV combined distinctly short; slightly less than 9 / 10 in length of tibia. Pretarsomere slightly more than 2 / 5 in length of tarsomere I – IV combined. External claw slightly longer than pretarsomere. Internal claw slightly less than 4 / 5 in length of external claw.	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E032FFC4FF64D5C1FB39FD7C.taxon	distribution	Distribution. Western Cape Province (Fig. 140), only known from the type locality. Vegetation Type. Bokkeveld Sandstone Fynbos (FFs 1).	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E032FFC4FF64D5C1FB39FD7C.taxon	biology_ecology	Host Plants. Asteraceae: Didelta spinosa (L. f.) W. T. Aiton, Berkheya fruticosa (L.) Ehrh., Arctotheca calendula (L.) Levyns.	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E032FFC4FF64D5C1FB39FD7C.taxon	etymology	Etymology. The name derives from the small size of the species [Latin: pumili- = dwarf].	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E02DFFC1FF64D759FC54FCB1.taxon	description	(Figs. 43, 82 – 83, 110) http: // zoobank. org / 3 A 279973 - E 69 B- 4 F 02 - B 0 F 9 - C 287 FEB 5 CD 93	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E02DFFC1FF64D759FC54FCB1.taxon	materials_examined	Type locality. [RSA, WCP]: Vanrhynsdorp Rd. to Gifberg. Type material. [7] Holotype ♂: white label pr « AFRICA: Republic of South Africa | Western Cape, Vanrhynsdorp | Road to Gifberg | 31 ° 38 ꞌ 53 ꞌꞌS – 18 ° 44 ꞌ 10 ꞌꞌS | Host plant: Berkheya spinosa » | 29.08.2011, leg.: J. F. Colville » || blue label pr « Bioregion: | Knersvlakte | Vegetation Type: SKk 5 – | Vanrhynsdorp Gannabosveld » || yellow label pr « SAM – COL | A 071222 » || red label pr « Holotype | Beckhoplia gifbergensis sp. n. | det.: H. Dombrow 2019 » (SAMC). – Paratypes 1 ♂, 5 ♀♀: same labels as holotype but || yellow label hw « 37 » pr «. 563 –. 569 | coll. Dombrow » || red label pr « Paratype | Beckhoplia gifbergensis sp. n. | det.: H. Dombrow 2019 » (ESDW). Condition of holotype. Very good. Pinned. Dissected, parameres glued on separate card below beetle. Right mesothoracic leg removed for genetic analysis.	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E02DFFC1FF64D759FC54FCB1.taxon	diagnosis	Differential diagnosis. Differs from B. pumilla: Pronotum of B. gifbergensis widest medially. Pronotum of B. pumilla widest in anterior two fifth. Elytra of B. gifbergensis transparently pale-brown. Elytra of B. pumilla yellowish pale-brown. Elytra of B. gifbergensis covering propygidium. Elytra of B. pumilla protruding propygidium. Externo-posterior margin of parameres of B. gifbergensis slightly convex in dorsal view (Figs. 82, 110). Externoposterior margin of parameres of B. pumilla undulate in dorsal view (Figs. 80, 109). Anterior margin of phallobase of B. gifbergensis concave in dorsal view (Figs. 82, 110). Anterior margin of phallobase of B. pumilla deep v-shaped incised in dorsal view (Figs. 80, 109).	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E02DFFC1FF64D759FC54FCB1.taxon	description	Description Measurements: length 6.6 – 7.1 mm; width 3.4 – 3.6 mm. Integument: Black, slightly lustrous; elytra transparently pale-brown, lateral and posterior margin broadly blackened. Tegument: Clypeus bare; sparse setose with short, erect, black setae on the inner side of the reflexed anterior and lateral margins. Head sparse setose with erect, short, pale-brown and medium-long black setae. Eye canthus with a loose setose ciliary of medium-long, black setae. Scapus loose setose with medium-long, black setae, palebrown ventrally, sparser and short on pedicellus; funicle I – II bare, funicle III – IV and clavus sparse setose with short, semi-erect, pale-brown and dark-brown setae. Pronotum loose setose with erect, short, whitish and long brown setae; with a loose setose ciliary of long, black setae on lateral margin. Scutellum loose setose with short and mediumlong, semi-erect, brown setae. Elytra with pattern. Sparse setose with short, reclinate, black setae, denser laterally; medium-long, erect, brown setae interspersed in anterior third; a sparse juxta-sutural row of short, semi-erect, black setae from anterior margin crossing scutellar torus towards apical torus; with three shortened, loose, longitudinal, squamose stripes of reclinate, narrowly-lanceolate, white scales: a sutural stripe in apical quarter; a juxta-sutural stripe from behind middle, terminating towards apical third; a sparse sometimes interrupted supra-lateral stripe in apical half and around apical torus. Suture a loose row of medium-long and long, robust, erect, black setae. Thorax and pleurites sparse pilose with long, white hairs. Ventrite I – V sparse setose with very short, reclinate, whitish setae; medium-long, semi-erect, black setae interspersed; upper corner loose squamose with reclinate, narrowly-lanceolate, white scales. Propygidium loose squamose with reclinate, narrowly-lanceolate, white scales. Pygidium sparse setose with long, erect, black setae; tip with a loose setose ciliary of medium-long, black setae. Prothoracic femur loose setose with short brown setae; medium-long brown hairs interspersed ventrally; bare internally. Prothoracic tibia loose setose with short, reclinate, whitish setae; erect medium-long and long ventrally; with a sparse setose longitudinal row of medium-long, erect, brown setae dorsally; a loose setose ciliary of short, black and white setae on internal ridge; a setaceous ciliary of short, transparently brown setae on anterior margin, more robust towards internal ridge; a medium-long, downwards pointing setose ciliary of pale-brown setae along anterior margin of apical tooth. Mesothoracic femur loose pilose with long, whitish and brown hairs; bare internally. Mesothoracic tibia sparse setose with short, reclinate, white setae; medium-long, semi-erect, robust, black setae interspersed ventrally and externally. Posterior margin with a bristled ciliary of very short and short, black bristles; lengthening dorsally; missing internally. Metathoracic femur loose setose with long, semi-erect white setae; medium-long erect, black setae interspersed externally; bare internally. Metathoracic tibia sparse setose with short, semi-erect, black setae; medium-long, semi-erect, robust, black setae interspersed; sparse setose with long, erect, black setae internally. Posterior margin with a loose bristled outer marginal ciliary of short, brown bristles, lengthening dorsally and ventrally, more robust ventrally. Prothoracic and mesothoracic tarsomere I – IV bare. Metathoracic tarsomere I – IV sparse setose with very short, reclinate, black and greyish setae. Tarsomere I – IV of all thoracic legs with a setaceous, apical, marginal ciliary of short, black setae, lengthening towards metathoracic tarsomeres, fine dorsally and more robust ventrally. Pretarsomere of thoracic legs sparse setose with extremely short, reclinate, black setae, semi-erect and short ventrally, lengthening towards metathoracic pretarsomere. Head: Suture between head and clypeus linear, raised. Head dense punctate with small, shallow, roundish, not merging punctures, micropunctate within. Funicle I – II cylindrical, slightly thickened medially; funicle III short asymmetric frustum-shaped; clavus long elliptic, slightly longer than pedicellus and funiculum combined. Pronotum: Widest medially; loose punctate with small and extremely small, shallow, roundish punctures, dense within medial furrow and towards margins. Anterior corner orthogonal. Lateral margin linearly attenuate anteriorly and posteriorly. With an indistinct, shortened, shallow medial furrow from posterior margin terminating in anterior fifth, broadening at disk. Posterior corner angled. Scutellum: Buckler-shaped; as broad as long; dense punctate with very small and small, shallow, roundish punctures, micropunctate within; with a complete, median, longitudinal, narrow, smooth stripe. Lateral margin linearly attenuate in anterior third, stronger apically. Tip pointed. Elytra: Ovate; longer than broad; widest in anterior third; covering propygidium. Lateral margin slightly convex attenuate apically. Sculpture indistinct; very slightly pulvinate laterally; humerus indistinct; with a broad roundish torus around scutellum; lateral trench missing; complanate along scutellum and suture. Sparse punctate with very small, shallow, roundish punctures, micropunctures in-between; a loose juxta-sutural row of small, deep, roundish punctures from anterior margin, crossing scutellar torus to apical torus; a sparse supra-humeral row of small, deep, roundish punctures from anterior margin between humerus and scutellar torus to apical torus. Abdomen: Ventrite I distinctly shortened medially; ventrite V slightly constricted ventrally. Propygidium with a slightly projected stoma at posterior corner. Pygidium distinctly pulvinate longitudinally and laterally; upper margin moderately convex; lateral margin subparallel in upper fifth, linearly attenuate apically; tip rounded. Prothoracic leg: Femur and tibia slender. Tibia smooth ventrally. Median tooth elongate Basal tooth very small. Interspace between teeth almost equal. Shape of interspace between anterior and median tooth u-shaped and between median and basal tooth linear. Tarsomere I – IV combined extremely short; equal in length; slightly less than 1 / 2 in length of tibia. Pretarsomere distinctly thickened; smooth ventrally; slightly less than 9 / 10 in length of tarsomere I – IV combined. Internal claw slightly more than 9 / 10 in length of pretarsomere. External claw 9 / 10 in length of internal claw. Mesothoracic leg: Tibia slightly more than 4 / 5 in length of femur. Tarsomere I – IV combined extremely short; slightly less than 2 / 5 in length of tibia. Pretarsomere with a shortened smooth ridge ventrally followed by a roundish cutting; slightly shorter than tarsomere I – IV combined. External claw as long as pretarsomere. Internal claw 9 / 10 in length of external claw. Metathoracic leg: Tibia 9 / 10 in length of femur. Tarsomere I – IV combined short; slightly more than 9 / 10 in length of tibia. Pretarsomere slightly thickened; with an undulate ridge ventrally; slightly more than 2 / 5 in length of tarsomere I – IV combined. External claw slightly less than 1 / 5 more in length of pretarsomere. Internal claw slightly less than 4 / 5 in length of external claw. Parameres: (Figs. 82 – 83, 110). ♀♀. Measurements: length 7.2 – 7.4 mm; width 3.9 – 4.1 mm. Integument: Black; elytra pale-brown, lateral margin broadly blackened; thoracic tarsi brown. Tegument: Clypeus sparse setose with short, erect, red-brown setae, loose and medium-long on head. Funiculum with few very short, semi-erect and erect, pale-brown setae, sparse and semi-erect on clavus. Pronotum with pattern. Loose setose with short and long, erect, brown setae; sparse, short, reclinate, white setose interspersed, denser in posterior half of medial furrow and at posterior margin near corner; with a loose setose ciliary of medium-long and long, brown setae on lateral margin. Scutellum loose setose with short and medium-long, semi-erect, brown and whitish setae. Elytra with pattern. Sparse setose with short, reclinate, brown setae; denser laterally; long, erect, brown setae interspersed at humerus; with two shortened, longitudinal loose setose stripes of short, reclinate, white squamiform setae; a sutural stripe in apical quarter; a sparse supra-lateral stripe in apical half and around apical torus. Suture with a loose row of short, robust, erect, black setae. Thorax and pleurites sparse pilose with long, yellowish-white hairs. Ventrite I – V sparse setose with short, reclinate, white setae; medium-long, semi-erect, white setae interspersed. Propygidium loose setose with short, reclinate, brown setae and white squamiform setae; medium-long, semi-erect, brown and white setae interspersed. Pygidium sparse setose with short, reclinate, brown and whitish setae; long, erect, brown setae interspersed; tip with a loose ciliary of medium-long and long, brown setae. Prothoracic tibia loose setose with short, reclinate, yellowish setae; erect, medium-long and pale-brown ventrally; a loose setose ciliary of short, brown setae on internal ridge; a loose setose ciliary of short and medium-long, black and yellowish setae on external ridge; anterior margin with a loose setaceous ciliary of short, pale-brown setae on anterior margin; a medium-long downwards pointing setose ciliary of pale-brown setae along anterior margin of apical tooth. Mesothoracic tibia sparse setose with short and medium-long, reclinate, whitish setae; medium-long, semi-erect, robust, brown setae interspersed, long and finer internally; Fringe of posterior margin as in male but redbrown. Metathoracic femur loose setose with long, semi-erect, whitish and brown setae. Metathoracic tibia sparse setose with short, semi-erect, brown setae; long, semi-erect, robust, brown setae interspersed, longer and finer internally. Posterior margin with a loose bristled outer marginal ciliary of very short, red-brown bristles, lengthening dorsally and ventrally, more robust ventrally. Setae of thoracic tarsi red-brown. Head: Clypeus very slightly depressed laterally; loose punctate with very small, shallow, roundish punctures. Anterior margin of clypeus slightly vertically reflexed. Lateral margin slightly reflexed. Suture between head and clypeus undulated, crenated. Eye on head level. Funicle I – II cylindrical; funicle III asymmetric frustum-shaped; clavus elliptic, as long as pedicellus and funiculum combined. Pronotum: Moderate pulvinate laterally; dense punctate with small, shallow, roundish punctures, denser in posterior third of medial furrow. Lateral margin convex. With an indistinct, shortened, shallow medial furrow from posterior margin terminating on disk. Posterior corner rounded. Scutellum: Dense punctate with small, shallow, roundish punctures, not micropunctate within. Elytra: Lateral trench indistinct. Loose punctate with extremely small and very small, shallow, roundish punctures, micropunctures in-between; a loose juxta-sutural row of small, deep, roundish punctures from anterior margin, crossing scutellar torus to apical torus; a sparse supra-humeral row of small, deep, roundish punctures from anterior margin between humerus and scutellar torus to apical torus. Abdomen: Distinctly pulvinate; ventrite II – V broadened medially; ventrite V not constricted ventrally. Propygidium with a slightly projected stoma at posterior corner. Pygidium slightly pulvinate longitudinally and laterally; slightly declined posteriorly; upper margin moderately convex, linear medially; tip linear. Prothoracic leg: Femur robust. Median tooth of tibia broad triangular-shaped. Shape of interspace between teeth concave. Tarsomere I – IV combined distinctly short; tarsomere I petiolate chalice-shaped; tarsomere I longest, tarsomere II – IV equal in length; slightly less than 7 / 10 in length of tibia. Pretarsomere slightly thickened; without roundish cutting apically; slightly less than 3 / 5 in length of tarsomere I – IV combined. Internal claw slender; not broadened medially; slightly more than 7 / 10 in length of pretarsomere. External claw slightly more than 3 / 5 in length of internal claw. Mesothoracic leg: Tibia slightly less than 4 / 5 in length of femur. Tarsomere I – IV combined very short; slightly less than 1 / 2 in length of tibia. Pretarsomere slender; with a full undulate ridge ventrally, roundish cutting missing; slightly more than 7 / 10 in length of tarsomere I – IV combined. External claw slender; not broadened medially; slightly shorter than pretarsomere. Internal claw slightly more than 7 / 10 in length of external claw. Metathoracic leg: Femur slightly thickened. Tibia distinctly extended dorso-apically; slightly less than 4 / 5 in length of femur. Tarsomere I – IV combined slightly more than 4 / 5 in length of tibia; petiolate chalice-shaped. Pretarsomere slender. External claw slightly less than 1 / 10 more in length of pretarsomere.	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E02DFFC1FF64D759FC54FCB1.taxon	distribution	Distribution. Western Cape Province (Fig. 140), only known from the type locality. Vegetation Type. Vanrhynsdorp Gannabosveld (SKk 5).	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E02DFFC1FF64D759FC54FCB1.taxon	biology_ecology	Host Plants. Asteraceae: Didelta spinosa (L. f.) W. T. Aiton.	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E02DFFC1FF64D759FC54FCB1.taxon	etymology	Etymology. The specific name derives from the resemblance to B. julianae.	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E028FFC3FF64D797FD46FB9D.taxon	description	(Figs. 27, 84 – 85, 111) http: // zoobank. org / 61 F 58 CC 0 - 24 C 7 - 4 CAC- 909 A- 42 B 901 BBCD 2 C	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E028FFC3FF64D797FD46FB9D.taxon	materials_examined	Type locality. [RSA, NCP]: Kamieskroon 11 km S. Type material. [3] Holotype ♂: white label pr « SOUTH AFRICA: Northern | Cape, Namaqualand | Kamieskroon 11 km S at | 30.16.26 S- 17.53.48 E » bs pr « Time: 14: 30, 04. Sept. 2001 | alt. 722 m, ASTERACEAE: | Didelta carnosa | leg. Holger Dombrow || yellow label hw « 54 » pr «. 995 | coll. Dombrow » || blue label pr « Bioregion: | Namaqualand Hardeveld | Vegetation Type: SKn 3 – | Namaqualand Blomveld » || yellow label pr « SAM – COL | A 071216 » || red label pr « Holotype | Beckhoplia | julianae sp. n. | det.: Dombrow 2003 » (SAMC). – Paratypes: 2 ♂♂: same labels as holotype but || yellow label hw « 18 » pr «. 468 –. 469 | coll. Dombrow » || red label pr « Paratype | Beckhoplia | julianae sp. n. | det.: Dombrow 2003 » (ESDW). Condition of holotype. Very good. Glued on card. Dissected, parameres glued behind beetle. Taxonomic note. In consequence of the fact that the only # f of the type series is not from the type locality. It will not be evaluated as a # f of B. julianae. Because a new species B. elkeae close allied to B. julianae occur at Groenvlei Pass which is very near to Kilian’s Pass. Compared to the # ♀ f of B. elkeae no differences could be found. This led to the result that the specimen is a # f of B. elkeae and will be listed there as additional material.	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E028FFC3FF64D797FD46FB9D.taxon	diagnosis	Differential diagnosis. Differs from B. elkeae: Lateral margin of pronotum of B. julianae slightly concave attenuate anteriorly, linearly attenuate posteriorly. Lateral margin of pronotum of B. elkeae linearly attenuate anteriorly, slightly convex attenuate posteriorly. Interno-posterior margin of parameres of B. julianae broad parabola-shaped in dorsal view (Figs. 84, 111). Interno-posterior margin of parameres of B. elkeae broad curly bracket-shaped in dorsal view (Figs. 86, 112). Externo-posterior margin of parameres of B. julianae convex in dorsal view (Figs. 84, 111). Externo-posterior margin of parameres of B. elkeae distinctly undulate in dorsal view (Figs. 86, 112).	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E028FFC3FF64D797FD46FB9D.taxon	description	Redescription Measurements: length 7.6 – 8.2 mm; width 3.8 – 4.4 mm. Integument: Black; elytra transparently pale-brown, lateral and posterior margin broadly blackened. Tegument: Clypeus bare; sparse setose with short, erect, black setae on the inner side of the reflexed anterior and lateral margins. Head loose setose with medium-long and long, erect, black setae. Eye canthus with a loose setose ciliary of medium-long, black, setae. Scapus loose setose with medium-long, black setae, pale-brown ventrally, sparse and short on pedicellus; funiculum with few very short, erect, brown setae, sparse on clavus. Pronotum loose setose with medium-long and long, erect, whitish setae; long, erect, dark brown setae interspersed in anterior half; with a loose setose ciliary of long, black and whitish setae on lateral margin. Scutellum loose setose with short and medium-long, semi-erect, whitish setae. Elytra with pattern. Sparse setose with short, reclinate, black setae, denser in darkened areas; long, erect, black setae interspersed in anterior third; with two loose, longitudinal rows of distinctly long, erect, black setae: a juxta-sutural row from anterior margin crossing scutellar torus towards apical torus, shorter in apical quarter; a supra-humeral row from anterior margin between humerus and scutellar torus towards apical torus, shorter in apical half; with three dense squamose shortened, longitudinal stripes of reclinate, lanceolate, white scales: a sutural stripe in apical third; a juxta-sutural stripe from behind middle, terminating towards apical third; a lateral stripe beside apical torus. Suture with a loose row of long and distinctly long, robust, erect, black setae. Thorax and pleurites loose pilose with long, white hairs. Ventrite I – V dense squamose with reclinate, lanceolate, transparently white scales; medium-long, semi-erect, black setae interspersed. Propygidium dense squamose with reclinate, oblanceolate, transparently white scales; medium-long, semi-erect, black setae interspersed. Pygidium sparse setose with medium-long, erect, black setae; tip with a loose setose ciliary of long, black setae. Prothoracic femur loose pilose with long, white hairs; bare internally. Prothoracic tibia loose setose with short, reclinate, white setae; medium-long, semi-erect, black setae interspersed, long ventrally; with a sparse setose longitudinal row of medium-long, erect, black setae dorsally; a loose setose ciliary of short, black and white setae on internal ridge; a sparse setose ciliary of short, transparently white setae on external ridge; a setaceous ciliary of short, black setae on anterior margin, more robust towards internal ridge; a medium-long, downwards pointing setose ciliary of pale-brown setae along anterior margin of apical tooth. Mesothoracic femur loose pilose with long, whitish hairs; distinctly long ventrally; bare internally. Mesothoracic tibia sparse setose with short, reclinate white setae; mediumlong, semi-erect, robust, black setae interspersed, long and fine internally. Posterior margin with a bristled ciliary of very short and short, black bristles; lengthening dorsally; missing internally. Metathoracic femur loose setose with medium-long, semi-erect, white setae, long dorsally; medium-long, erect, robust, black setae interspersed externally; bare internally. Metathoracic tibia sparse setose with short, semi-erect, black setae; medium-long, semi-erect, black setae interspersed; sparse setose with long, erect, black setae internally. Posterior margin with a loose bristled outer marginal ciliary of short, black bristles, lengthening dorsally and ventrally. Tarsomere I – IV of all thoracic legs sparse setose with very short, reclinate, pale-brown setae; with a setaceous, apical, marginal ciliary of short, black setae, lengthening towards metathoracic tarsomeres, fine dorsally and more robust ventrally. Pretarsomere of all thoracic legs sparse setose with very short, reclinate, pale-brown setae, lengthening and darkening towards metathoracic pretarsomere; few, medium-long, semi-erect, black setae at apical margin, shorter and dark red-brown ventrally. Head: Suture between head and clypeus linear, crenated. Head dense punctate with very small, shallow, roundish, not merging punctures. Funicle I – II cylindrical, slightly thickened medially; funicle III very short asymmetric frustum-shaped; clavus elliptic, as long as pedicellus and funiculum combined. Pronotum: Widest in anterior two fifth; loose punctate with very small, shallow, roundish punctures. Anterior corner acute. Lateral margin slightly concave attenuate anteriorly, linearly attenuate posteriorly. With an indistinct, shortened, shallow medial furrow from posterior margin terminating in anterior fifth. Posterior corner rounded. Scutellum: Parabola-shaped; as broad as long; dense punctate with very small, shallow, roundish punctures, micropunctate within; with a complete, median, longitudinal, broad, smooth stripe. Lateral margin slightly convex. Tip pointed. Elytra: Ovate; longer than broad; widest in anterior third; covering propygidium. Lateral margin slightly convex attenuate apically. Sculpture distinct; dorsally not pulvinate laterally; humerus distinct; with a very broad roundish torus around scutellum; lateral trench distinct; complanate along scutellum; slightly, broadly depressed along suture. Loose punctate with extremely small, shallow, roundish punctures, micropunctures in-between; a loose juxta-sutural row of small, deep, roundish punctures from anterior margin, crossing scutellar torus to apical torus; a sparse supra-humeral row of very small, shallow and deep, roundish punctures from anterior margin between humerus and scutellar torus to apical torus, terminating medially. Abdomen: Ventrite I distinctly shortened medially; ventrite V slightly constricted ventrally. Propygidium with an indistinct stoma at posterior corner. Pygidium slightly pulvinate longitudinally and laterally; upper margin moderately convex, linear medially; lateral margin subparallel in upper fifth, linearly attenuate apically; tip rounded. Prothoracic leg: Femur and tibia slender. Tibia smooth ventrally. Median tooth robust Basal tooth rudimentary. Interspace between teeth almost equal. Shape of interspace between anterior and median tooth u-shaped and between median and basal tooth slightly concave. Tarsomere I – IV combined extremely short; equal in length; 2 / 5 in length of tibia. Tarsomere IV with a single ventral extension. Pretarsomere distinctly thickened; smooth ventrally; slightly longer than length of tarsomere I – IV combined. Internal claw slightly less than 1 / 5 more in length of pretarsomere. External claw slightly less than 4 / 5 in length of internal claw. Mesothoracic leg: Tibia slightly shorter than femur. Tarsomere I – IV combined extremely short; slightly less than 3 / 10 in length of tibia. Pretarsomere with a shortened undulate ridge ventrally, followed by a roundish cutting apically; slightly less than 1 / 5 more in length of tarsomere I – IV combined. External claw as long as pretarsomere. Internal claw slightly less than 9 / 10 in length of external claw. Metathoracic leg: Tibia slightly more than 1 / 10 more in length of femur. Tarsomere I – IV combined distinctly short; slightly more than 3 / 5 in length of tibia. Pretarsomere slightly thickened; with an undulate ridge ventrally; slightly less than 1 / 2 in length of tarsomere I – IV combined. External claw slightly more than 1 / 5 more in length of pretarsomere. Internal claw slightly less than 9 / 10 in length of external claw. Parameres: (Figs. 84 – 85, 111). ♀: Unknown.	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E028FFC3FF64D797FD46FB9D.taxon	distribution	Distribution. Northern Cape Province (Fig. 140), only known from the type locality. Vegetation Type. Namaqualand Blomveld (SKn 3).	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E028FFC3FF64D797FD46FB9D.taxon	biology_ecology	Host Plants. Asteraceae: Didelta carnosa (L. f.) Aiton.	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E02AFFCCFF64D079FBAFFA80.taxon	description	(Figs. 24, 54, 86 – 87, 112) http: // zoobank. org / 57 D 792 FF- 0 ECF- 4470 - A 44 C-A 14640 DD 9 F 96	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E02AFFCCFF64D079FBAFFA80.taxon	materials_examined	Type locality. [RSA, NCP]: Grootvlei Pass. Type material. [153] Holotype ♂: white label pr « RSA, NC, Grootvlei Pass W | side, 595 m, 30 ° 13,007 ’ S – | 17 ° 46,156 ’ E, 30.08.2012, | leg.: H & E. Dombrow # 615 » bs pr « Host plants: | Tripteris clandestina | Tripteris oppositifolium | Didelta spinosum # 615 » || yellow label hw « 43 » pr «. 876 | coll. Dombrow » || blue label pr « Bioregion: | Namaqualand Hardeveld | Vegetation type: SKn 1 - | Namaqualand Klipkoppe Shrubland » » || yellow label pr « SAM – COL | A 071221 » || red label pr « Holotype | Beckhoplia | elkeae sp. n. | det.: H. Dombrow 2019 » (SAMC). – Paratypes: 5 ♂♂, 10 ♀♀: same labels as holotype but || yellow label hw « 43 » pr «. 960 –. 974 | coll. Dombrow » || yellow label pr « SAM – COL | A 071221 » || red label pr « Paratype | Beckhoplia | elkeae sp. n. | det.: H. Dombrow 2019 » (SAMC); 83 ♂♂, 54 ♀♀: same labels as holotype but || yellow label hw « 43 » pr «. 877 –. 959;. 975 –. 999 » & hw « 44 », pr «. 000 –. 001;. 003 –. 028 | coll. Dombrow » || red label pr « Paratype | Beckhoplia | elkeae sp. n. | det.: H. Dombrow 2019 » (ESDW). Additional material examined. 1 ♀: white label pr « RSA, CP, Namaquald. | # 0003 Kilianꞌs Pass | 30.10.33 S – 17.40.60 E | 500 m, leg. P. Desmet » || pr « # 0003, 14.08.1994 | Stoeberia frutescens | Ruschia sp., Arctotis sp. » || yellow label hw « 18 » pr «. 470 | coll. Dombrow » || red label pr « Paratype | Beckhoplia | julianae sp. n. | det.: Dombrow 2003 » (ESDW). Condition of holotype. Very good. Glued on card. Dissected, parameres and spiculum gastrale glued behind beetle.	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E02AFFCCFF64D079FBAFFA80.taxon	diagnosis	Differential diagnosis. Differs from B. julianae: Lateral margin of pronotum of B. elkeae linearly attenuate anteriorly, slightly convex attenuate posteriorly. Lateral margin of pronotum of B. julianae slightly concave attenuate anteriorly, linearly attenuate posteriorly. Interno-posterior margin of parameres of B. elkeae broad curly bracketshaped in dorsal view (Figs. 86, 112). Interno-posterior margin of parameres of B. julianae broad parabola-shaped in dorsal view (Figs. 84, 111). Externo-posterior margin of parameres of B. elkeae distinctly undulate in dorsal view (Figs. 86, 112). Externo-posterior margin of parameres of B. julianae convex in dorsal view (Figs. 84, 111).	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E02AFFCCFF64D079FBAFFA80.taxon	description	Description Measurements: length 7.9 – 8.4 mm; width 4.1 – 4.4 mm. Integument: Black; elytra transparently pale-brown, lateral and posterior margin broadly blackened, suture sometimes blackened. Tegument: Clypeus bare; sparse setose with short, erect, pale-brown setae on the inner side of the reflexed anterior and lateral margins. Head loose setose with medium-long and long, erect, black setae. Eye canthus with a loose setose ciliary of medium-long, black, setae. Scapus loose setose with medium-long, black setae, pale-brown ventrally, sparse and short on pedicellus; funiculum with few very short, erect, brown setae, sparse on clavus. Pronotum loose setose with medium-long and long, erect, whitish setae; long, erect, dark-brown setae interspersed; with a loose setose ciliary of long, black setae on lateral margin. Scutellum loose setose with short and mediumlong, semi-erect, whitish setae, shortening apically. Elytra with pattern. Sparse setose with short, reclinate, black setae, denser in darkened areas; long, erect, black setae interspersed in anterior third; with two loose, longitudinal rows of distinctly long, erect, black setae: a juxta-sutural row from anterior margin crossing scutellar torus towards apical torus, shortening apically; a supra-humeral row from anterior margin between humerus and scutellar torus towards apical torus, shortening apically; with three dense squamose shortened, longitudinal stripes of reclinate, oblanceolate, white scales: a sutural stripe in apical third; a juxta-sutural stripe from behind middle, terminating towards apical third; a lateral stripe beside apical torus. Suture with a loose row of long and distinctly long, robust, erect, black setae. Thorax and pleurites loose pilose with long, white hairs. Ventrite I – V dense squamose with reclinate, oblanceolate, transparently white scales; medium-long, semi-erect, black setae interspersed. Propygidium dense squamose with reclinate, oblanceolate, transparently white scales; medium-long, semi-erect, black setae interspersed. Pygidium sparse setose with medium-long, erect, black setae; tip with a loose setose ciliary of long, black setae. Prothoracic femur loose pilose with long, white hairs; bare internally. Prothoracic tibia loose setose with short, reclinate, white setae; medium-long, semi-erect, black setae interspersed, long ventrally; with a sparse setose longitudinal row of medium-long, erect, robust, black setae dorsally; a loose setose ciliary of short, black setae on internal ridge; a sparse setose ciliary of short, transparently white setae on external ridge; a setaceous ciliary of short, black setae on anterior margin, more robust towards internal ridge; a medium-long, downwards pointing setose ciliary of pale-brown setae along anterior margin of apical tooth. Mesothoracic femur loose pilose with long, whitish hairs; distinctly long ventrally; bare internally. Mesothoracic tibia sparse setose with short, semi-erect, white setae; medium-long, semi-erect, robust, black setae interspersed, long and fine internally. Posterior margin with a bristled ciliary of very short and short, black bristles; lengthening dorsally and ventrally; missing internally. Metathoracic femur loose setose with medium-long to long, semi-erect, white setae; distinctly long ventrally; medium-long, erect, robust, black setae interspersed externally; bare internally. Metathoracic tibia sparse setose with short, semi-erect, black setae; medium-long, semi-erect, black setae interspersed; sparse setose with long, erect, black setae internally. Posterior margin with a loose bristled outer marginal ciliary of short, black bristles, lengthening dorsally and ventrally. Tarsomere I – IV of all thoracic legs sparse setose with very short, reclinate, pale-brown setae; with a setaceous, apical, marginal ciliary of short, black setae, lengthening towards metathoracic tarsomeres, fine dorsally and more robust ventrally. Pretarsomere of all thoracic legs sparse setose with very short, reclinate, pale-brown setae, lengthening and darkening towards metathoracic pretarsomere; few, medium-long, semi-erect, black setae at apical margin, shorter ventrally. Head: Suture between head and clypeus linear, crenated. Head dense punctate with very small, shallow, roundish, not merging punctures. Funicle I – II cylindrical; funicle III asymmetric frustum-shaped; clavus elliptic, slightly longer than pedicellus and funiculum combined. Pronotum: Widest in anterior two fifth; sparse to loose punctate with extremely small to very small, shallow, roundish punctures, dense within medial furrow at posterior margin. Anterior corner acute. Lateral margin linearly attenuate anteriorly, slightly convex attenuate posteriorly. With an indistinct, complete, shallow medial furrow from posterior to anterior margin. Posterior corner rounded. Scutellum: Parabola-shaped; as broad as long; dense punctate with very small, shallow, roundish punctures, micropunctate within; with a shortened, median, longitudinal, broad, smooth stripe, from anterior margin terminating medially. Lateral margin slightly convex. Tip pointed. Elytra: Ovate; longer than broad; widest in anterior third; protruding propygidium. Lateral margin slightly convex attenuate apically. Sculpture distinct; dorsally not pulvinate laterally; humerus distinct; with a very broad roundish torus around scutellum, extending apically, terminating on disk; lateral trench distinct; complanate along scutellum; slightly, broadly depressed along suture. Loose punctate with extremely small, shallow, roundish punctures, micropunctures in-between; a loose juxta-sutural row of small, deep, roundish punctures from anterior margin, crossing scutellar torus to apical torus; a sparse supra-humeral row of very small, shallow and deep, roundish punctures from anterior margin between humerus and scutellar torus to apical torus, terminating medially. Abdomen: Ventrite I distinctly shortened medially; ventrite I – IV broadest, slightly constricted ventrally. Propygidium with an indistinct stoma at posterior corner. Pygidium distinctly pulvinate longitudinally and laterally; upper margin moderately convex; lateral margin linearly attenuate apically; tip rounded. Prothoracic leg: Femur and tibia slender. Tibia smooth ventrally. Median tooth robust. Basal tooth rudimentary. Interspace between teeth almost equal. Shape of interspace between teeth concave. Tarsomere I – IV combined extremely short; equal in length; slightly more than 2 / 5 in length of tibia. Tarsomere IV without ventral extension. Pretarsomere distinctly thickened; with a shortened granulate ridge ventrally; slightly more than 9 / 10 in length of tarsomere I – IV combined. Internal claw slightly more than 1 / 10 more in length of pretarsomere. External claw slightly less than 4 / 5 in length of internal claw. Mesothoracic leg: Tibia slightly longer than femur. Tarsomere I – IV combined extremely short; slightly less than 3 / 10 in length of tibia. Pretarsomere with a shortened smooth ridge ventrally followed by a roundish cutting apically; as long as tarsomere I – IV combined. External claw as long as pretarsomere. Internal claw slightly more than 4 / 5 in length of external claw. Metathoracic leg: Tibia as long as femur. Tarsomere I – IV combined distinctly short; slightly less than 7 / 10 in length of tibia. Pretarsomere slightly thickened; with an undulate ridge ventrally; slightly more than 2 / 5 in length of tarsomere I – IV combined. External claw slightly less than 1 / 5 more in length of pretarsomere. Internal claw slightly more than 4 / 5 in length of external claw. Parameres: (Figs. 86 – 87, 112). ♀♀. Measurements: length 7.6 – 8.7 mm; width 4.2 – 4.9 mm. Integument: Black; elytra transparently pale red-brown, darkening, sometimes blackened towards lateral margin, suture sometimes red-brown; mesothoracic femur and tibia, prothoracic tarsus as well as metathoracic femur dark red-brown; metathoracic tibia, mesothoracic and metathoracic tarsus red-brown. Tegument: Clypeus sparse setose with short, erect, pale-brown setae. Setae on head, eye canthus and dorsal part of scapus brown. Pronotum with pattern. Loose setose with medium-long erect, brown setae, shortening towards posterior margin; with a loose, squamose, longitudinal band of reclinate, lanceolate, whitish scales in posterior half of medial furrow; with a loose squamose spot of reclinate, narrowly-elliptic, yellowish scales at posterior margin near corner; setae of ciliary brown. Scutellum loose setose with short, semi-erect, pale-brown setae, semierect, lanceolate, yellowish scales interspersed in apical half. Setae of elytra brown, interspersed setae extended to apical margin; scales of stripes lanceolate and offwhite, juxta-sutural stripe shorter and with an additional shortened, median, supra-lateral stripe within lateral trench. Hairs on thorax and pleurites offwhite. Scales on ventrite I – V narrowly-lanceolate and offwhite; setae yellowish. Scales on propygidium lanceolate and yellowish as well as the setae. Pygidium dense squamose with reclinate, ovate, yellowish scales, short to medium-long, erect, yellowish setae interspersed; tip with a loose setose ciliary of medium-long, pale-brown setae. Hairs of prothoracic femur yellowish. Prothoracic tibia sparse setose with short, semi-erect, pale-brown to brown setae; medium-long to long and erect ventrally; setae of longitudinal, dorsal row brown; setae of ciliary on internal ridge brown; setae of ciliary on external ridge reclinate and transparently pale-brown; setae of ciliary on anterior margin red-brown and not robust. Hairs of mesothoracic femur offwhite. Setae of mesothoracic tibia offwhite. Ciliary on posterior margin red-brown. Metathoracic femur only loose pilose with medium-long to long, semi-erect, offwhite hairs. Setae of metathoracic tibia pale-brown; interspersed setae red-brown; medium-long to long and brown internally. Bristles of ciliary on posterior margin less robust and transparently pale red-brown. Setae of ciliary of tarsomere I – IV of thoracic legs red-brown; pale red-brown in between on mesothoracic and metathoracic tarsomeres. Setae on pretarsomere of thoracic legs red-brown. Head: Clypeus dense punctate with deep, very large, roundish punctures. Anterior margin distinctly vertically reflexed. Suture between head and clypeus undulate. Head dense punctate with large, deep, roundish punctures. Funicle I – II cylindrical, slightly thickened medially; funicle III short asymmetric frustum-shaped; clavus short elliptic. Pronotum: Widest in anterior third; loose punctate with very small, shallow, roundish punctures, denser within medial furrow. Anterior corner obtuse. Lateral margin linearly attenuate in anterior third, slightly convex attenuate posteriorly. Scutellum: Broader than long; without smooth stripe. Tip rounded. Elytra: Almost square; as long as broad. Sculpture indistinct; lateral trench distinct. Abdomen: Distinctly pulvinate; ventrite II – V slightly broadened medially; ventrite V broadest, not constricted ventrally. Propygidium with a slightly projected stoma at posterior corner. Pygidium slightly declined posteriorly; slightly pulvinate longitudinally and laterally; lateral margin subparallel in upper fifth, linearly attenuate apically; tip linear. Prothoracic leg: Tarsomere I – IV combined short; tarsomere I longest, tarsomere II – IV equal in length; slightly than 9 / 10 in length of tibia. Pretarsomere slightly thickened; with a shortened smooth ridge ventrally, roundish cutting missing; slightly less than 1 / 2 in length of tarsomere I – IV combined. Internal claw slightly less than 9 / 10 in length of pretarsomere. External claw slightly less than 9 / 10 in length of internal claw. Mesothoracic leg: Tibia slightly less than 4 / 5 in length of femur. Tarsomere I – IV combined short; slightly less than 1 / 2 in length of tibia. Pretarsomere slightly thickened; slightly more than 7 / 10 in length of tarsomere I – IV combined. Metathoracic leg: Femur slightly thickened. Tibia slightly less than 9 / 10 in length of femur. Tarsomere I – IV combined short; slightly less than 9 / 10 in length of tibia. Pretarsomere slender; with a smooth ridge ventrally. External claw as long as pretarsomere. Internal claw slightly more than 9 / 10 in length of external claw.	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E02AFFCCFF64D079FBAFFA80.taxon	distribution	Distribution. Northern Cape Province (Fig. 140). Vegetation Type. Namaqualand Klipkoppe Shrubland (SKn 1).	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E02AFFCCFF64D079FBAFFA80.taxon	biology_ecology	Host Plants. Asteraceae: Tripteris clandestina (Harv.) Norl., Tripteris oppositifolia (Aiton) B. Nord.; Didelta spinosa (L. f.) Aiton. – Aizoaceae: Stoeberia frutescens (L. Bolus) Van Jaarsv.	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E02AFFCCFF64D079FBAFFA80.taxon	etymology	Etymology. The species is dedicated to Elke Dombrow, wife of the first author.	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E025FFC9FF64D145FC41FB48.taxon	description	(Figs. 29, 88 – 89, 113) http: // zoobank. org / 5 E 04 AD 0 A- 2014 - 42 C 8 - 8242 - 2179490 AA 908	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E025FFC9FF64D145FC41FB48.taxon	materials_examined	Type locality. [RSA, NCP]: Nieuwoudtville. Type material. [49] Holotype ♂: white label pr « SOUTH AFRICA, Northern | Cape, Hantam | Nieuwoudtville at | 31.22.33 S – 19.07.43 E » // pr « Time 12: 30, 15. Sept. 2001 | alt. 733 m, IRIDACEAE: | Homeria miniata | leg.: Holger Dombrow # 327 » || yellow label hw « 18 » pr «. 485 | coll. Dombrow » || red label pr « Paratype | Beckhoplia | colvillei sp. n. | det.: H. Dombrow 2003 » || blue label pr « Bioregion: | Shale Renosterveld | Vegetation Type: FRs 2 – | Nieuwoudtville Shale Renosterveld » || yellow label pr « SAM – COL | A 071220 » || red label pr « Holotype | Beckhoplia | suturalis sp. n. | det.: H. Dombrow 2019 » (SAMC). – Paratypes 29 ♂♂, 13 ♀♀: same labels as holotype but || yellow label hw « 18 » pr «. 486,. 488 –. 514,. 517 –. 529,. 534 –. 553 | coll. Dombrow » || red label pr « Paratype | Beckhoplia | suturalis sp. n. | det.: H. Dombrow 2019 » (ESDW); 2 ♂♂, 4 ♀♀: same labels as holotype but || yellow label hw « 18 » pr «. 515 –. 516,. 530 –. 533 | coll. Dombrow » || yellow label pr « SAM – COL | A 071220 » || red label pr « Paratype | Beckhoplia | suturalis sp. n. | det.: H. Dombrow 2019 » (SAMC). Condition of holotype. Excellent. Glued on a card. Dissected, parameres glued behind beetle.	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E025FFC9FF64D145FC41FB48.taxon	diagnosis	Differential diagnosis. Differs from B. nigrosetosa: Elytra of B. suturalis widest in anterior third. Elytra of B. nigrosetosa widest medially. Lateral margin of elytra of B. suturalis broadly blackened. Lateral margin of elytra of B. nigrosetosa narrowly darkened. Interno-posterior margin of parameres of B. suturalis rounded in dorsal view (Figs. 88, 113). Interno-posterior margin of parameres of B. nigrosetosa parabola in dorsal view (Figs. 90, 114). Externoposterior margin of parameres of B. suturalis convex in dorsal view (Figs. 88, 113). Externo-posterior margin of parameres of B. nigrosetosa linear with a convex median extension in dorsal view (Figs. 90, 114). Anterior margin of phallobase of B. suturalis curly bracket-shaped in dorsal view (Figs. 88, 113). Anterior margin of phallobase of B. nigrosetosa broad parabola-shaped in dorsal view (Figs. 90, 114).	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E025FFC9FF64D145FC41FB48.taxon	description	Description Measurements: length 7.8 – 9.6 mm; width 4.0 – 4.7 mm. Integument: Black; funiculum brown annulated; elytra transparently brown, lateral and posterior margin blackened. Tegument: Clypeus bare; loose setose with short, erect, black setae on the inner side of the reflexed anterior and lateral margins. Head loose setose with long, erect, black setae. Eye canthus with a dense setose ciliary of long, black setae. Scapus loose setose with long, black setae, whitish ventrally, sparse and short on pedicellus; funicle I bare; funicle II – IV with few extremely short, semi-erect, pale-brown setae; clavus sparse setose with extremely short, reclinate pale-brown setae, very short, semi-erect, black setae interspersed. Pronotum loose setose with long, erect, black setae, shorter towards posterior margin; with a loose setose ciliary of long, black setae. Scutellum loose setose with short and medium-long, semi-erect, brown setae. Elytra without pattern. Sparse setose with short, reclinate and semi-erect, black setae; few short, reclinate, white setae interspersed below apical torus; with a loose juxtasutural row of long, erect, black setae from anterior margin crossing scutellar torus towards apical torus. Suture with a loose row of medium-long and long, robust, erect, black setae. Thorax and pleurites loose pilose with long, black hairs. Ventrite I – V loose setose with short, reclinate, black setae; medium-long and long, semi-erect, black setae and short reclinate white setae interspersed. Propygidium and pygidium loose setose with long, reclinate, black setae; short reclinate white setae interspersed on propygidium; tip of pygidium with a sparse setose ciliary of mediumlong and long, black setae. Prothoracic femur loose pilose with long black hairs; bare internally. Prothoracic tibia loose setose with short, reclinate, greyish setae; medium-long, semi-erect, black setae interspersed; erect and long ventrally; with a sparse setose longitudinal row of medium-long, erect, black setae dorsally; a loose setose ciliary of short, black setae on internal ridge; a setaceous ciliary of short, brown setae on anterior margin, more robust towards internal ridge; a medium-long downwards pointing setose ciliary of pale-brown setae along anterior margin of apical tooth. Mesothoracic femur loose pilose with long black hairs; bare internally. Mesothoracic tibia loose setose with short, reclinate, whitish setae, sparser internally; medium-long, semi-erect, robust, black setae interspersed, long and finer internally. Posterior margin with a bristled ciliary of short, black bristles; missing internally. Metathoracic femur loose setose with long, semi-erect, black setae; distinctly long and whitish ventrally; bare internally. Metathoracic tibia sparse setose with short, reclinate, brown setae; medium-long, semi-erect, robust, black setae interspersed, long, denser and finer internally. Posterior margin with a loose bristled outer marginal ciliary of short, brown bristles, lengthening dorsally and ventrally, more robust ventrally. Prothoracic tarsomere I – IV bare. Mesothoracic and metathoracic tarsomere I – IV sparse setose with short, reclinate, brown setae. Tarsomere I – IV of all thoracic legs with a setaceous, apical, marginal ciliary of short, black setae lengthening towards metathoracic tarsomeres, fine dorsally and more robust ventrally. Pretarsomere of all thoracic legs sparse setose with short, semi-erect, brown setae, lengthening towards metathoracic pretarsomere; few, long, semi-erect, black setae at apical margin, shorter and red-brown ventrally. Head: Suture between head and clypeus linear, raised. Head dense punctate with small, shallow, roundish, not merging punctures. Funicle I – II cylindrical, slightly thickened medially; funicle III asymmetric frustum-shaped; clavus elliptic, slightly longer than pedicellus and funiculum combined. Pronotum: Widest medially; loose punctate with small, shallow, roundish punctures, dense posteriorly and within medial furrow. Anterior corner orthogonal. Lateral margin linearly attenuate anteriorly and subparallel posteriorly. With an indistinct, shortened, shallow medial furrow from posterior margin terminating on disk. Posterior corner rounded. Scutellum: Parabola-shaped; as broad as long; dense punctate with small to large, shallow, roundish punctures, micropunctate within; with a shortened, median, longitudinal, narrow, smooth stripe from anterior margin terminating in apical third. Lateral margin slightly convex. Tip pointed. Elytra: Ovate; longer than broad; widest in anterior third; protruding propygidium. Lateral margin slightly convex attenuate apically. Sculpture indistinct; almost complanate laterally; humerus indistinct; with a broad roundish torus beside scutellum, expanding towards disk; lateral trench indistinct; complanate along scutellum and suture. Loose punctate with extremely small to very small, shallow, roundish punctures, micropunctures in-between; a loose juxta-sutural row of small, deep, roundish punctures from anterior margin, crossing scutellar torus to apical torus. Abdomen: Ventrite I distinctly shortened medially; ventrite V not constricted ventrally. Propygidium with a slightly projected stoma at posterior corner. Pygidium distinctly pulvinate longitudinally and laterally; upper margin slightly convex, linear medially; lateral margin slightly convex; tip rounded. Prothoracic leg: Femur and tibia slender. Tibia smooth ventrally. Median tooth elongate. Basal tooth very small. Interspace between teeth almost equal. Shape of interspace between teeth concave. Tarsomere I – IV combined extremely short; equal in length; slightly less than 1 / 2 in length of tibia. Pretarsomere distinctly thickened; smooth ventrally; slightly less than 9 / 10 length of tarsomere I – IV combined. Internal claw as long as pretarsomere. External claw slightly more than 4 / 5 in length of internal claw. Mesothoracic leg: Tibia slightly shorter than femur. Tarsomere I – IV combined extremely short; slightly less than 2 / 5 in length of tibia. Pretarsomere with a shortened smooth ridge ventrally followed by a roundish cutting apically; slightly shorter than tarsomere I – IV combined. External claw slightly more than 4 / 5 in length of pretarsomere. Internal claw slightly more than 9 / 10 in length of external claw. Metathoracic leg: Tibia slightly less than 3 / 10 more in length of femur. Tarsomere I – IV combined distinctly short; slightly less than 7 / 10 in length of tibia. Pretarsomere slightly thickened; with an undulate ridge ventrally; slightly less than 1 / 2 in length of tarsomere I – IV combined. External claw slightly less than 1 / 5 more in length of pretarsomere. Internal claw slightly more than 4 / 5 in length of external claw. Parameres: (Figs. 88 – 89, 113). ♀♀. Measurements: length 8.6 – 9.5 mm; width 4.8 – 5.2 mm. Integument: Lateral margins of elytra darkened; thoracic legs dark red-brown; metathoracic tibia and thoracic tarsi pale red-brown. Tegument: Clypeus loose setose with short, erect, pale-brown setae; medium-long and brown on head. Eye canthus with a loose setose ciliary of medium-long, brown setae. Scapus loose setose with medium-long, brown setae, paler ventrally, shorter and pale-brown on pedicellus; funicle I bare, funicle II – IV with extremely short, semierect, pale-brown setae, reclinate on clavus. Pronotum without pattern. Loose setose with short and long, erect brown setae; with a loose setose ciliary of long, brown setae. Scutellum loose setose with short, semi-erect, whitish setae. Elytra without pattern. Loose setose with short, reclinate, yellowish-brown setae; long, semi-erect, brown setae interspersed, denser towards lateral margin. Suture with a loose row of short and long, robust, semi-erect, black setae. Thorax and pleurites loose pilose with long, yellowish hairs. Ventrite I – V loose setose with short, reclinate, yellowish setae; long, semi-erect, yellowish and brown setae interspersed. Propygidium and pygidium loose setose with short and medium-long, reclinate, yellowish setae; medium-long, semi-erect, brown setae interspersed; tip of pygidium with a sparse setose ciliary long, brown setae. Prothoracic femur loose pilose with long, brown hairs. Prothoracic tibia loose setose with short, reclinate, brown setae; erect and long ventrally; a loose setose ciliary of short and medium-long, black and yellowish setae on internal ridge; with a setaceous ciliary of loose short pale-brown setae on anterior margin; a medium-long downwards pointing setose ciliary of pale-brown setae along anterior margin of apical tooth. Mesothoracic femur loose pilose with long, yellowish-brown hairs; distinctly long ventrally; long semi-erect brown setae interspersed externally. Mesothoracic tibia sparse setose with short, reclinate, whitish setae; medium-long, semi-erect, robust, brown setae interspersed, long and finer internally; ciliary of posterior margin as in male but red-brown. Metathoracic femur loose setose with long, semi-erect, yellowish-brown setae; paler ventrally. Metathoracic tibia sparse setose with short, reclinate, whitish setae; medium-long and robust, semi-erect, red-brown setae interspersed, loose, long and finer internally. Posterior margin with a bristled outer marginal ciliary of short, loose, transparently red-brown bristles, lengthening dorsally and ventrally and more robust ventrally. Setae of thoracic tarsi pale red-brown. Head: Clypeus very slightly depressed laterally; dense punctate with large and very large, deep, roundish punctures. Anterior margin of clypeus slightly vertically reflexed. Lateral margin slightly reflexed. Suture between head and clypeus undulate, crenated. Head dense punctate with small and large, deep, roundish punctures. Eye on head level. Funicle II – II cylindrical; clavus as long as pedicellus and funiculum combined. Pronotum: Widest in anterior third; moderate pulvinate; loose punctate with very small, shallow, roundish punctures, dense in posterior half of medial furrow. Anterior corner obtuse. Lateral margin convex. With an indistinct, complete, shallow, narrow medial furrow. Elytra: Covering propygidium. Abdomen: Distinctly pulvinate; ventrite II – V broadened medially. Pygidium slightly pulvinate longitudinally and laterally; slightly declined posteriorly; lateral margin subparallel in upper fifth, linearly attenuate apically; tip linear. Prothoracic leg: Femur robust. Median tooth of tibia broad triangular-shaped. Tarsomere I – IV combined very short; slightly more than 7 / 10 in length of tibia. Pretarsomere slightly thickened; with a shortened smooth ridge ventrally, roundish cutting missing; slightly less than 7 / 10 length of tarsomere I – IV combined. Internal claw slender; not broadened medially; slightly more than 9 / 10 length of pretarsomere. External claw 4 / 5 in length of internal claw. Mesothoracic leg: Tibia 9 / 10 in length of femur. Tarsomere I – IV combined distinctly short; slightly more than 1 / 2 in length of tibia. Pretarsomere slightly thickened; roundish cutting missing; slightly more than 7 / 10 in length of than tarsomere I – IV combined. External claw slender; not broadened medially; as long as pretarsomere. Internal claw 4 / 5 in length of external claw. Metathoracic leg: Femur slightly thickened. Tibia slightly extended dorso-apically; slightly less than 9 / 10 in length of femur. Tarsomere I – IV combined slightly more than 4 / 5 in length of tibia. Pretarsomere slender; with a smooth ridge ventrally; slightly more than 1 / 2 in length of tarsomere I – IV combined. External claw slightly more than 1 / 10 more in length of pretarsomere. Internal claw slightly less than 4 / 5 in length of external claw.	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E025FFC9FF64D145FC41FB48.taxon	distribution	Distribution. Northern Cape Province (Fig. 140), only known from the type locality. Vegetation Type. Nieuwoudtville Shale Renosterveld (FRs 2).	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E025FFC9FF64D145FC41FB48.taxon	biology_ecology	Host Plants. Iridaceae: Moraea miniata (Andrews).	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E025FFC9FF64D145FC41FB48.taxon	etymology	Etymology. The name derives from the elevated suture [Latin: sutur = seam].	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E020FFCAFF64D10DFEB1FB2C.taxon	description	(Figs. 7, 13, 26, 28, 56, 90 – 91, 114) http: // zoobank. org / 2 C 34 F 785 - 38 C 8 - 4265 - B 59 D-DE 94 C 9023 A 05	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E020FFCAFF64D10DFEB1FB2C.taxon	materials_examined	Type locality. [RSA, NCP]: Nieuwoudtville Wildflower Reserve. Type material. [269] Holotype ♂: white label pr « RSA, NCP, Nieuwoudt- | ville, Wildflower Res. | 31.24 S – 19.08 E | 22 / 08 / 98, J. Colville » || yellow label hw « 18 » pr «. 534 | coll. Dombrow » || red label pr « Paratype | Beckhoplia | colvillei sp. n. | det.: Dombrow 2003 » || green label pr « Host plant: | Gazania rigida | Senecio erosus » || blue label pr « Bioregion: | Shale Renosterveld | Vegetation Type: FRs 2 – | Nieuwoudtville Shale Renosterveld » || yellow label pr « SAM – COL | A 071219 » || red label pr « Holotype | Beckhoplia | nigrosetosa sp. n. | det.: H. Dombrow 2019 » (SAMC). – Paratypes 179 ♂♂, 40 ♀♀: same labels as holotype but || yellow label hw « 18 » pr «. 535 –. 713,. 734 –. 773 | coll. Dombrow » || red label pr « Paratype | Beckhoplia | colvillei sp. n. | det.: Dombrow 2003 » || red label pr « Paratype | Beckhoplia | nigrosetosa sp. n. | det.: H. Dombrow 2019 » (ESDW); 39 ♂♂ 10 ♀♀: same labels as holotype but || yellow label hw « 55 » pr «. 708 –. 756 | coll. Dombrow » || yellow label pr « SAM – COL | A 071219 » || red label pr « Paratype | Beckhoplia | nigrosetosa sp. n. | det.: H. Dombrow 2019 » (SAMC). Condition of holotype. Very good. Pinned. Dissected, parameres glued on separate card below beetle.	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E020FFCAFF64D10DFEB1FB2C.taxon	diagnosis	Differential diagnosis. Differs from B. suturalis: Elytra of B. nigrosetosa widest medially. Elytra of B. suturalis widest in anterior third. Lateral margin of elytra of B. nigrosetosa narrowly darkened. Lateral margin of elytra of B. suturalis broadly blackened. Interno-posterior margin of parameres of B. nigrosetosa parabola in dorsal view (Figs. 90, 114). Interno-posterior margin of parameres of B. suturalis rounded in dorsal view (Figs. 88, 113). Externo-posterior margin of parameres of B. nigrosetosa linear with a convex median extension in dorsal view (Figs. 90, 114). Externo-posterior margin of parameres of B. suturalis convex in dorsal view (Figs. 88, 113). Anterior margin of phallobase of B. nigrosetosa broad parabola-shaped in dorsal view (Figs. 90, 114). Anterior margin of phallobase of B. suturalis curly bracket-shaped in dorsal view (Figs. 88, 113).	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E020FFCAFF64D10DFEB1FB2C.taxon	description	Description: Measurements: length 7.6 – 8.4 mm; width 4.5 – 4.9 mm. Integument: Black; funiculum pale-brown annulated; elytra transparently pale-brown, lateral and posterior margins blackened. Tegument: Clypeus bare; loose setose with short, erect, black setae on the inner side of the reflexed anterior and lateral margins. Head loose setose with long, erect, black setae. Eye canthus with a loose setose ciliary of long, black, setae. Scapus loose setose with long, black setae, pale-brown ventrally, sparse and medium-long on pedicellus; funiculum with few very-short, erect, brown setae, shorter and semi-erect on clavus. Pronotum loose setose with long, erect, brown setae, shorter towards posterior margin; with a loose setose ciliary of long, brown setae on lateral margin. Scutellum loose setose with short and medium-long, semi-erect, brown setae. Elytra without pattern. Sparse setose with short, semi-erect, pale-brown and black setae, medium-long at humerus; few short, reclinate, whitish setae interspersed below apical torus; with two loose, longitudinal rows of medium-long, semi-erect, black setae: a juxta-sutural row from anterior margin crossing scutellar torus towards apical torus; a juxta-humeral row from anterior margin between humerus and scutellar torus towards apical torus terminating towards middle. Suture with a loose row of short and long, robust, semi-erect, black setae. Thorax and pleurites loose pilose with long, black hairs. Ventrite I – V loose setose with short, reclinate, brownish setae; long, semi-erect, black setae interspersed. Propygidium and pygidium loose setose with short, reclinate, pale-brown and whitish setae; long, semi-erect, black setae interspersed on propygidium, erect on pygidium; tip of pygidium with a loose setose ciliary of long, black setae. Prothoracic femur loose pilose with long, brown hairs; bare internally. Prothoracic tibia loose setose with short, reclinate, pale-brown setae dorsally; erect, medium-long and black ventrally; medium-long, black, semi-erect setae interspersed dorsally; with a sparse setose longitudinal row of medium-long erect, black setae dorsally; a loose setose ciliary of short, black setae on internal ridge; whitish and very short at external ridge; a setaceous ciliary of short, brown setae on anterior margin; a medium-long downwards pointing setose ciliary of pale-brown setae along anterior margin of apical tooth. Mesothoracic femur loose pilose with long, brown hairs; distinctly long, brown ventrally; long, erect, black setae interspersed dorsally and externally; bare internally. Mesothoracic tibia sparse setose with short, reclinate, white and pale-brown setae, sparser internally; medium-long, semi-erect, robust, black setae interspersed, long and finer internally. Posterior margin with a bristled ciliary of medium-long, black bristles; missing internally. Metathoracic femur loose setose with medium-long and long, semi-erect, black setae; long, palebrown setae interspersed ventrally; bare internally. Metathoracic tibia loose setose with short, reclinate, pale-brown setae; medium-long, semi-erect, black setae interspersed; denser, finer and long internally. Posterior margin with a loose bristled outer marginal ciliary of short, black bristles, lengthening dorsally and ventrally. Tarsomere I – IV of all thoracic legs sparse setose with very short, reclinate, brown setae; with a setaceous, apical, marginal ciliary of short, dark red-brown setae lengthening towards metathoracic tarsomeres, fine dorsally and more robust ventrally. Pretarsomere of all thoracic legs with few, medium-long, semi-erect, red-brown setae at apical margin, shorter ventrally. Mesothoracic and metathoracic pretarsomere additionally sparse setose with short, reclinate, pale-brown setae, lengthening towards metathoracic pretarsomere. Head: Suture between head and clypeus linear, crenated. Head dense punctate with small, shallow, roundish, partly merging punctures, micropunctate within. Funicle I – II cylindrical, slightly thickened medially; funicle III short asymmetric frustum-shaped; clavus elliptic, slightly longer than pedicellus and funiculum combined. Pronotum: Widest medially; loose punctate with small, shallow, roundish punctures, dense within medial furrow and posteriorly. Anterior corner orthogonal. Lateral margin linearly attenuate anteriorly and posteriorly. With an indistinct, shallow, shortened medial furrow from posterior margin terminating on disk. Posterior corner rounded. Scutellum: Parabola-shaped; as broad as long; dense punctate with small to large, shallow, roundish punctures, micropunctate within; with a shortened, median, longitudinal, narrow, smooth stripe from anterior margin terminating in apical third. Lateral margin slightly convex. Tip pointed. Elytra: Ovate; longer than broad; widest in anterior third; protruding propygidium. Lateral margin slightly convex attenuate apically. Sculpture indistinct; very slightly pulvinate laterally; humerus indistinct; with a broad roundish torus beside scutellum, expanding towards disk; lateral trench missing; complanate along scutellum and suture. Loose punctate with extremely small to very small, shallow, roundish punctures, micropunctures in-between; with two loose, longitudinal rows of small, deep, roundish punctures: a juxta-sutural row from anterior margin crossing scutellar torus to apical torus; a supra-humeral row from anterior margin between humerus and scutellar torus towards apical torus terminating medially. Abdomen: Ventrite I extremely shortened medially; ventrite V not constricted ventrally. Propygidium with a slightly projected stoma at posterior corner. Pygidium slightly pulvinate longitudinally and laterally; upper margin slightly convex, linear medially; lateral margin linearly attenuate apically; tip linear. Prothoracic leg: Femur and tibia slender. Tibia with a medio-ventral ridge. Median tooth elongate. Basal tooth rudimentary. Interspace between teeth almost equal. Shape of interspace between teeth concave. Tarsomere I – IV combined extremely short; equal in length; slightly less than 1 / 2 in length of tibia. Pretarsomere distinctly thickened; with a shortened granulate ridge ventrally; slightly more than 1 / 10 more in length of tarsomere I – IV combined. Internal claw slightly longer than pretarsomere. External claw slightly more than 7 / 10 in length of internal claw. Mesothoracic leg: Tibia slightly less than 9 / 10 in length of femur. Tarsomere I – IV combined extremely short; slightly less than 2 / 5 in length of tibia. Pretarsomere with a shortened granulate ridge ventrally, roundish cutting missing; slightly longer than tarsomere I – IV combined. External claw slightly less than 1 / 5 more in length of pretarsomere. Internal claw slightly less than 7 / 10 in length of external claw. Metathoracic leg: Tibia slightly shorter than femur. Tarsomere I – IV combined distinctly short; slightly less than 4 / 5 in length of tibia. Pretarsomere slightly thickened; with an undulate ridge ventrally; slightly less than 1 / 2 in length of tarsomere I – IV combined. External claw slightly more than 1 / 10 more in length of pretarsomere. Internal claw slightly more than 4 / 5 in length of external claw. Parameres: (Figs. 90 – 91, 114). ♀♀. Measurements: length 8.4 – 9.4 mm; width 4.5 – 5.3 mm. Integument: Black; elytra pale-brown, lateral margins darkened; thoracic legs dark red-brown; metathoracic tibia and thoracic tarsi red-brown. Tegument: Clypeus loose setose with short, erect, black setae, medium-long and brown on head. Eye canthus with a loose setose ciliary of medium-long, brown setae. Setae on scapus and pedicellus red-brown, pale-brown on funiculum and clavus. Pronotum loose setose with short and long, semi-erect, brown setae; short, reclinate, whitish setae sometimes interspersed on disk, in posterior half of medial furrow and at posterior margin; with a loose setose ciliary of long, brown setae on lateral margin. Scutellum loose setose with short, semi-erect, brown setae; short, reclinate, white setae interspersed. Elytra loose setose with short, reclinate, pale-brown setae; long, semi-erect and erect, black setae interspersed, distinctly long anteriorly; sometimes few, short, reclinate, whitish setae below apical torus. Thorax and pleurites loose pilose with long, pale-brown hairs. Ventrite loose setose with short, reclinate, yellowish-white setae; medium-long and long, semi-erect, brown setae interspersed. Propygidium and pygidium loose setose with short, reclinate, yellowish-white setae; medium-long, semi-erect, pale-brown setae interspersed. Tip of pygidium with a loose setose ciliary of long, brown, setae. Prothoracic femur loose pilose with long, brown hairs, paler towards tibia. Prothoracic tibia loose setose with short, semi-erect, pale-brown setae, erect and medium-long ventrally; a loose setose ciliary of short, pale-brown setae on internal ridge and external ridge; a setaceous ciliary of short, pale-brown setae on anterior margin. Mesothoracic tibia sparse setose with short, reclinate white setae; robust, medium-long, semi-erect, pale-brown setae interspersed. Ciliary of posterior margin as in male but red-brown. Metathoracic femur loose setose with long, semi-erect, pale-brown setae. Metathoracic tibia loose setose with medium-long and long, semi-erect, red-brown setae; more robust dorsally; denser, fine and long internally. Fringe of posterior margin as in male but red-brown. Setae of thoracic tarsi pale red-brown. Head: Clypeus very slightly depressed laterally; dense punctate with large, deep, roundish punctures, micropunctate within. Anterior margin of clypeus slightly vertically reflexed. Lateral margin very slightly reflexed. Suture between head and clypeus undulate. Head dense punctate with very large, deep, roundish, not merging punctures. Eye below head level. Funicle I cylindrical; clavus short elliptic. Pronotum: Widest in anterior third; moderate pulvinate laterally; punctures additionally with a micropunctuation within. Lateral margin linearly attenuate in anterior third, slightly convex attenuate posteriorly. With an indistinct, shallow, complete medial furrow. Scutellum: Dense punctate with small, shallow, roundish punctures; with a shortened, median, longitudinal, narrow, smooth stripe from anterior margin terminating medially. Tip rounded. Elytra: Covering propygidium. Scutellar torus missing; lateral trench indistinct. Abdomen: Distinctly pulvinate; ventrite I distinctly shortened medially; ventrite II – V broadened medially; ventrite V broadest. Pygidium slightly pulvinate longitudinally and laterally; slightly declined posteriorly; lateral margin subparallel in upper fifth, linearly attenuate apically. Prothoracic leg: Femur and tibia robust. Tibia smooth ventrally. Median tooth broad triangular-shaped. Basal tooth very small. Tarsomere I – IV combined distinctly short; tarsomere I petiolate chalice-shaped, longest, tarsomere II – IV equal in length; slightly less than 7 / 10 in length of tibia. Pretarsomere slightly thickened; smooth ventrally, roundish cutting missing; slightly more than 3 / 5 in length of tarsomere I – IV combined. Internal claw slender; not broadened medially; as long as pretarsomere. External claw slightly less than 4 / 5 in length of internal claw. Mesothoracic leg: Tibia slightly less than 4 / 5 in length of femur. Tarsomere I – IV combined very short; slightly less than 1 / 2 in length of tibia. Pretarsomere slender; with a shortened smooth ridge ventrally; slightly less than 7 / 10 in length of tarsomere I – IV combined. External claw slender; not broadened medially; slightly less than 1 / 10 more in length of pretarsomere. Internal claw slightly less than 4 / 5 in length of external claw. Metathoracic leg: Femur slightly thickened. Tibia distinctly extended dorso-apically; slightly less than 4 / 5 in length of femur. Tarsomere I – IV combined short. Pretarsomere slender; slightly more than 1 / 2 in length of tarsomere I – IV combined. External claw slightly shorter than pretarsomere. Internal claw 4 / 5 in length of external claw.	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E020FFCAFF64D10DFEB1FB2C.taxon	distribution	Distribution. Northern Cape Province (Fig. 140), only known from the type locality. Vegetation Type. Nieuwoudtville Shale Renosterveld [FRs 2].	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E020FFCAFF64D10DFEB1FB2C.taxon	biology_ecology	Host Plants. Asteraceae: Gazania rigida (Burm. f.) Roessler; Senecio erosus L. f.	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E020FFCAFF64D10DFEB1FB2C.taxon	etymology	Etymology. The name derives from the black setae, especially on the pronotum [Latin: nigr- = black; Latin: set = a bristles].	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E023FFB7FF64D0E9FD3BFC28.taxon	description	(Figs. 31, 39, 58, 92 - 93, 115) http: // zoobank. org / 6 B 0920 E 2 - 5 C 48 - 4648 - A 210 - C 41 A 7378 EBD 2	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E023FFB7FF64D0E9FD3BFC28.taxon	materials_examined	Type locality. [RSA, NCP]: Nieuwoudtville 5 km E. Type material. [1] Holotype ♂: white label pr « SOUTH AFRICA | Northern Cape, Hantam | Nieuwoudtville 5 km S at | 31.22.37 S – 19.09.13 E » bs pr « Time 14: 30, 17. Sept. 2001 | alt. 752 m, ASTERACEAE | Arctotis acaulis | leg. Holger Dombrow # 339 » || blue label pr « Bioregion: | Shale Renosterveld | Vegetation Type: FRs 2 – | Nieuwoudtville Shale Renosterveld » || yellow label hw « 18 » pr «. 721 | coll. Dombrow » || red label pr « Holotype | Beckhoplia | occidentalis sp. n. | det.: H. Dombrow 2003 » (ZSM). Additional material examined. 7 ♂♂, 6 ♀♀: white label pr « Republic of South Africa, | Northern Cape, Nieuwoudtville, | 31.24 S – 19.08 E, 04. – 07.09.2000, | leg. Ingrid Nänni » || yellow label hw « 55 » pr «. 721 | coll. Dombrow » (ESDW). Taxonomic note. The following specimens from the original type series are transferred to several other species, because of the distinct morphological differences see descriptions of the target species: The 6 ♀♀ from the locality Botterkloof Pass are transferred to B. caliginosa sp. n.; the 3 ♂♂ and 10 ♀♀ from the locality Farm Nuwedam are transferred to B. pumilla sp. n. The locality has to be adjusted because the coordinate is located 5 km E of Nieuwoudtville. Condition of holotype. Very good: Glued on a card. Dissected, parameres glued behind beetle.	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E023FFB7FF64D0E9FD3BFC28.taxon	diagnosis	Differential diagnosis. Differs from B. dolichiocnemis: Elytra of B. occidentalis without a longitudinal depression along suture. Elytra of B. dolichiocnemis with a longitudinal depression along suture. Elytra of B. occidentalis partly covering propygidium. Elytra of B. dolichiocnemis protruding the propygidium. Interno-posterior margin of parameres of B. occidentalis curly bracket-shaped in dorsal view (Figs. 92, 115). Interno-posterior margin of parameres of B. dolichiocnemis parabola-shaped in dorsal view (Figs. 96, 117). Externo-posterior margin of parameres of B. occidentalis curly bracket-shaped, convex medially in dorsal view (Figs. 92, 115). Externo-posterior margin of parameres of B. dolichiocnemis undulate in dorsal view (Figs. 96, 117).	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E023FFB7FF64D0E9FD3BFC28.taxon	description	Redescription Measurements: length 7.6 – 8.8 mm; width 4.0 – 4.5 mm. Integument: Black; funiculum brown annulated; elytra transparently brown, margins blackened. Tegument: Clypeus bare; inner side of the reflexed anterior and lateral margins loose setose with short, erect, black setae. Head loose setose with long, erect, black setae. Eye canthus with a loose setose ciliary of long, black setae. Scapus loose setose with long, black setae, whitish ventrally, sparse and medium-long on pedicellus; funicle I – II bare, funicle II – IV with few very short and short, erect, brown setae; clavus sparse setose with short, reclinate, black and very short, semi-erect, pale-brown setae. Pronotum loose setose with long, erect, black setae, shorter towards posterior margin; with a loose setose ciliary of distinctly long, black setae on lateral margin. Scutellum loose setose with short and medium-long, semi-erect, brown setae. Elytra without pattern. Sparse setose with short, reclinate and semi-erect, black setae; with two loose, longitudinal rows of medium-long, semi-erect, black setae: a juxta-sutural row from anterior margin crossing scutellar torus towards apical torus; a juxta-humeral row from anterior margin between humerus and scutellar torus towards apical torus. Suture with a loose row of short and long, robust, erect, black setae. Thorax and pleurites loose pilose with long, black hairs. Ventrite I – V loose setose with short, reclinate, black setae; medium-long and long, semi-erect, black setae interspersed. Propygidium loose setose with long, reclinate, black setae, erect on pygidium. Tip of pygidium with a sparse setose ciliary of medium-long and long, black setae. Prothoracic femur loose pilose with long, black hairs; bare internally. Prothoracic tibia loose setose with short, reclinate and semi-erect, black setae; erect and medium-long ventrally; short, semi-erect, white setae interspersed dorsally; with a sparse setose longitudinal row of medium-long, erect, black setae dorsally; with a loose setose ciliary of medium-long, black setae on internal ridge; a setaceous ciliary of short, black setae on anterior margin, more robust towards internal ridge; a medium-long downwards pointing setose ciliary of pale-brown setae along anterior margin of apical tooth. Mesothoracic femur loose pilose with long, black hairs; distinctly long ventrally; bare internally. Mesothoracic tibia loose setose with short and medium-long, reclinate, whitish setae, sparser internally; medium-long, semi-erect, robust, black setae interspersed, long and finer internally. Posterior margin with a bristled ciliary of short, black bristles; lengthening dorsally; missing internally. Metathoracic femur loose setose with short and long, semi-erect, black, setae; bare internally. Metathoracic tibia sparse setose with very short and short, reclinate, brown setae; medium-long, semi-erect, robust, black setae interspersed; denser, long and finer internally. Posterior margin with a loose bristled outer marginal ciliary of short, black bristles, lengthening dorsally and ventrally, more robust ventrally. Prothoracic tarsomere I – IV bare. Mesothoracic and metathoracic tarsomere I – IV sparse setose with very short, reclinate, pale-brown setae. Tarsomere I – IV of all thoracic legs with a setaceous, apical, marginal ciliary of short, dark red-brown setae, lengthening towards metathoracic tarsomeres, fine dorsally and more robust ventrally. Pretarsomere of all thoracic legs sparse setose with very short, reclinate, pale-brown setae; few, long, semi-erect, black setae at apical margin, shorter and dark red-brown ventrally. Head: Suture between head and clypeus linear, crenated. Head dense punctate with small, shallow, roundish, not merging punctures. Funicle I – II cylindrical; funicle III asymmetric frustum-shaped; clavus elliptic, slightly longer than pedicellus and funiculum combined. Pronotum: Widest medially; dense punctate with shallow, very small and small, roundish punctures, denser within medial furrow. Anterior corner orthogonal. Lateral margin linearly attenuate anteriorly in anterior quarter and slightly convex attenuate posteriorly. With an indistinct, shortened, shallow medial furrow from posterior margin terminating on disk. Posterior corner rounded. Scutellum: Parabola-shaped; slightly longer than broad; dense punctate with shallow, small to large, roundish punctures, micropunctate within. Lateral margin slightly convex. Tip rounded. Elytra: Ovate; longer than broad; widest in anterior third; covering propygidium. Lateral margin slightly convex attenuate apically. Sculpture indistinct; slightly pulvinate laterally; humerus indistinct; with a broad roundish torus around scutellum; lateral trench missing; complanate along scutellum and suture. Loose punctate with extremely small, shallow, roundish punctures, micropunctures in-between; with two sparse, longitudinal rows of small, deep, roundish punctures: a juxta-sutural row from anterior margin crossing scutellar torus to apical torus; a supra-humeral row from anterior margin between humerus and scutellar torus to apical torus. Abdomen: Ventrite I distinctly shortened medially; ventrite V slightly constricted ventrally. Propygidium with an indistinct stoma at posterior corner. Pygidium slightly pulvinate longitudinally and laterally; upper margin moderately convex; lateral margin subparallel in upper fifth, linearly attenuate apically; tip linear. Prothoracic leg: Femur and tibia slender. Tibia smooth ventrally. Median tooth elongate. Basal tooth rudimentary. Interspace between apical and median tooth double in width than between median and basal tooth. Shape of interspace between teeth concave. Tarsomere I – IV combined extremely short; equal in length; slightly more than 2 / 5 in length of tibia. Tarsomere IV with a single ventral extension. Pretarsomere distinctly thickened; smooth ventrally; slightly less than 1 / 10 more in length of tarsomere I – IV combined. Internal claw slightly more than 1 / 10 more in length of pretarsomere. External claw slightly more than 7 / 10 in length of internal claw. Mesothoracic leg: Tibia slightly less than 2 / 5 in length of femur. Tarsomere I – IV combined very short; slightly less than 2 / 5 in length of tibia. Pretarsomere with a shortened smooth ridge ventrally, roundish cutting missing; as long as tarsomere I – IV combined. External claw slightly shorter than pretarsomere. Internal claw slightly less than 4 / 5 in length of external claw. Metathoracic leg: Tibia slightly longer than femur. Tarsomere I – IV combined distinctly short; slightly more than 7 / 10 in length of tibia. Pretarsomere slightly thickened; with an undulate ridge ventrally; slightly more than 1 / 2 in length of tarsomere I – IV combined. External claw slightly more than 1 / 10 more in length of pretarsomere. Internal claw 4 / 5 in length of external claw. Parameres: (Figs. 92 – 93, 115). ♀♀. Measurements: length 7.4 – 9.2 mm; width 4.4 – 5.4 mm. Integument: Only lateral margin of elytra blackened; mesothoracic, metathoracic tibia and all tarsi pale redbrown. Tegument: Clypeus sparse setose with short, erect, pale-brown setae. Colour of setae on head and eye canthus pale-brown. Funicle with few very short semi-erect, whitish setae; denser on clavus. Colour of setae on pronotum brown. Scutellum loose setose with very short, semi-erect, brown setae; extremely short to very short, semi-erect, whitish setae interspersed medially and towards tip. Colour of setae on elytra brown. Colour of hairs on thorax and pleurites pale-brown. Colour of setae on ventrites yellowish; interspersed setae long and pale-brown. Propygidium and pygidium loose setose with short, semi-erect, yellowish setae; long, semi-erect, pale-brown setae interspersed. Tip of pygidium with a loose setose ciliary of medium-long, brown setae. Colour of hairs on prothoracic femur palebrown; medium-long, erect, brown setae interspersed ventrally; bare internally. Colour of hairs on mesothoracic femur pale-brown to brown. Interspersed setae on mesothoracic tibia red-brown; additionally with few semi-erect, medium-long, transparently red-brown bristles dorsally. Posterior margin with a bristled ciliary of medium-long, transparently red-brown bristles; not lengthening dorsally; missing internally. Metathoracic femur sparse setose with medium-long, erect, whitish, setae; long, erect, pale-brown setae interspersed; bare internally. Colour of setae on metathoracic tibia whitish; interspersed setae transparently red-brown. Metathoracic tibia sparse setose with very short and short, reclinate, brown setae; medium-long, semi-erect, robust, black setae interspersed; denser, long and finer internally. Colour of bristles on ciliary transparently red-brown. Tarsomere I – IV of thoracic legs with few very short, reclinate, whitish setae; colour of bristles on ciliary transparently red-brown. Colour of setae on pretarsomere of all thoracic legs transparently pale red-brown. Head: Clypeus very slightly depressed laterally; dense punctate with large, deep, roundish punctures. Anterior margin of clypeus slightly vertically reflexed. Lateral margin slightly reflexed. Suture between head and clypeus undulate. Head dense punctate with very large, deep, roundish punctures. Eye below head level. Funicle I – II cylindrical, slightly thickened medially; funicle III short asymmetric frustum-shaped. Pronotum: Lateral margin linearly attenuate in anterior two fifth, convex posteriorly. Medial furrow missing. Scutellum: As broad as long. Elytra: Lateral trench indistinct. Abdomen: Distinctly pulvinate; ventrite II – V broadened medially; ventrite I – IV broadest. Propygidium with a slightly projected stoma at posterior corner. Pygidium slightly declined posteriorly. Prothoracic leg: Femur and tibia robust. Median tooth broad triangular-shaped. Basal tooth very small. Tarsomere I – IV combined distinctly short; slightly less than 3 / 5 in length of tibia; tarsomere IV without ventral extension. Pretarsomere slightly thickened; with a shortened smooth ridge ventrally, roundish cutting missing; slightly less than 7 / 10 in length of tarsomere I – IV combined. Internal claw slender; not broadened medially; as long as pretarsomere. External claw slightly less than 4 / 5 in length of internal claw. External claw slightly more than 4 / 5 in length of internal claw. Mesothoracic leg: Tibia slightly more than 9 / 10 in length of femur. Tarsomere I – IV combined distinctly short. Pretarsomere slightly thickened; slightly more than 7 / 10 in length of tarsomere I – IV combined. External claw slender; not broadened medially; as long as pretarsomere. Internal claw slightly more than 4 / 5 in length of external claw. Metathoracic leg: Femur slightly thickened. Tibia cornet-shaped; 7 / 10 in length of femur. Tarsomere I – IV combined very short; slightly less than 4 / 5 in length of tibia. Pretarsomere slender; with a shortened smooth ridge ventrally; slightly less than 4 / 5 in length of tarsomere I – IV combined. External claw 2 / 5 more in length of pretarsomere. Internal claw slightly more 3 / 5 in length of external claw.	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E023FFB7FF64D0E9FD3BFC28.taxon	distribution	Distribution. Northern Cape Province (Fig. 140), only known from the type locality. Vegetation Type. Nieuwoudtville Shale Renosterveld [FRs 2].	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E023FFB7FF64D0E9FD3BFC28.taxon	biology_ecology	Host Plants. Asteraceae: Arctotis acaulis L.	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E05EFFB1FF64D7EDFAA7FBF1.taxon	description	(Figs. 40, 94 – 95, 116 http: // zoobank. org / 4 CC 61 FC 4 - 17 AB- 4996 - 98 E 5 - 0 BE 7 A 186 C 663	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E05EFFB1FF64D7EDFAA7FBF1.taxon	materials_examined	Type locality. [RSA, NCP]: R 357, Nieuwoudtville to Loeriesfontein. Type material. [1] Holotype ♂: white label pr « RSA, NCP, R 357 to Loeriesfont. | 5 km E Nieuwoudtville | 31 ° 17 ’ 39 ’’ S – 19 ° 08 ’ 08 ’’ E | 27.8.2012, lg. Colville / Melin » || green label pr « Host plant: | Didelta spinosa » || blue label pr « Bioregion: | Sandstone Fynbos | Vegetation Type: FFs 1 – | Bokkeveld Sandstone Fynbos » || yellow label hw « 50 » pr « 447 | coll. Dombrow » || yellow label pr « SAM – COL | A 071218 » || red label pr « Holotype | Beckhoplia | bicolor sp. n. | det.: H. Dombrow 2019 » (SAMC). Condition of holotype. Good. Glued on a card. Dissected, parameres and spiculum gastrale glued behind beetle. Left mesothoracic tibia with tarsus missing.	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E05EFFB1FF64D7EDFAA7FBF1.taxon	discussion	Remark. The locality description requires an adjustment, as the GPS coordinate on the locality label actually indicates a locality ~ 10 km NE Nieuwoudtville on the R 357 to Loeriesfontein.	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E05EFFB1FF64D7EDFAA7FBF1.taxon	diagnosis	Differential diagnosis. Differs from B. dolichiocnemis: Elytra of B. bicolor partly covering propygidium. Elytra of B. dolichiocnemis protruding propygidium. Elytra of B. bicolor transparently pale-brown with broadly blackened lateral margins. Elytra of B. dolichiocnemis transparently pale red brown with narrowly blackened lateral margins. Interno-posterior margin of parameres of B. bicolor u-shaped in dorsal view (Figs. 94, 116). Interno-posterior margin of parameres of B. dolichiocnemis parabola-shaped in dorsal view (Figs. 96, 117). Externo-posterior margin of parameres of B. bicolor concave with a broad convex median extension in dorsal view (Figs. 94, 116). Externo-posterior margin of parameres of B. dolichiocnemis undulate in dorsal view (Figs. 96, 117).	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E05EFFB1FF64D7EDFAA7FBF1.taxon	description	Description Measurements: length 8.3 mm; width 3.8 mm. Integument: Black; elytra transparently yellowish pale-brown, lateral and posterior margin broadly blackened. Tegument: Clypeus and lateral margin bare; inner side of reflexed anterior margin sparse setose with short, erect, black setae. Head loose setose with medium-long, erect, black setae. Eye canthus with a loose setose ciliary of long, black setae. Scapus loose setose with long, black setae, pale-brown ventrally, sparse and medium-long on pedicellus; funiculum with few very short, erect, brown and black setae, shorter and semi-erect on clavus. Pronotum loose setose with long, erect, black setae, shorter towards posterior margin; with loose setose ciliary of distinctly long, black setae on lateral margin. Scutellum loose setose with short and medium-long, semi-erect, brown setae. Elytra without pattern. Loose setose with short, reclinate, brown setae; medium-long, erect, black setae interspersed; few short, reclinate, whitish setae below apical torus; with two loose, longitudinal rows of long, erect, black setae: a juxta-sutural row from anterior margin crossing scutellar torus towards apical torus, shorter in posterior third; a juxta-humeral row from anterior margin between humerus and scutellar torus towards apical torus. Suture with a loose row of short and long, robust, semi-erect, black setae. Thorax and pleurites loose pilose with long, black hairs. Ventrite I – V loose setose with short, reclinate, white and medium-long, black setae; long, semi-erect, black setae interspersed. Propygidium loose squamose with reclinate, narrowly-lanceolate, greyish scales; long, semi-erect, black setae interspersed. Pygidium sparse setose with long, erect, black setae; tip with a loose setose ciliary of long, black setae. Prothoracic femur loose pilose with long, black hairs; bare internally. Prothoracic tibia loose setose with short, reclinate, greyish setae; medium-long, semi-erect, black setae interspersed; long ventrally; with a sparse setose longitudinal row of medium-long, erect, black setae dorsally; with a loose ciliary of mediumlong, black and whitish setae on internal ridge; a sparse ciliary of short pale-brown setae on external ridge; a setaceous ciliary of short, black setae on anterior margin, more robust towards internal ridge; a medium-long downwards pointing setose ciliary of pale-brown setae along anterior margin of apical tooth. Mesothoracic femur loose pilose with long, black hairs; distinctly long ventrally; bare internally. Mesothoracic tibia sparse setose with short, reclinate, white setae; medium-long, semi-erect, robust, black setae interspersed, long and finer internally. Posterior margin with a bristled ciliary of short, black bristles; lengthening dorsally; missing internally. Metathoracic femur loose setose with long, semi-erect, black setae; bare internally. Metathoracic tibia sparse setose with short, reclinate, brown and black setae; medium-long, semi-erect, robust, black setae interspersed; long and fine internally. Posterior margin with a loose bristled outer marginal ciliary of short, black bristles, lengthening dorsally and ventrally, more robust ventrally. Prothoracic tarsomeres I – IV bare. Mesothoracic and metathoracic tarsomeres I – IV sparse setose with very short, reclinate, pale-brown setae. Tarsomere I – IV of all thoracic legs with a setaceous, apical, marginal ciliary of short, black setae, lengthening towards metathoracic tarsomeres, fine dorsally and more robust ventrally. Pretarsomere of all thoracic legs with few, medium-long, semi-erect, black setae at apical margin, shorter and dark-brown ventrally. Mesothoracic and metathoracic pretarsomere additionally sparse setose with short, reclinate, brown setae. Head: Suture between head and clypeus undulate, crenated. Head dense punctate with small, shallow, roundish, partly merging punctures, micropunctate within. Funicle I cylindrical, slightly thickened medially; funicle II cylindrical; funicle III short asymmetric frustum-shaped; clavus long elliptic, slightly shorter than scapus, pedicellus and funiculum combined. Pronotum: Widest in anterior two fifth; loose punctate with shallow, very small and small, roundish punctures, dense laterally and posteriorly and within posterior half of medial furrow. Anterior corner obtuse. Lateral margin convex. With an indistinct, shortened, shallow medial furrow from posterior margin terminating on disk. Posterior corner rounded. Scutellum: Buckler-shaped; as broad as long; dense punctate with shallow, very small and small, roundish punctures, partly merging, micropunctate within; with a shortened, median, longitudinal, narrow, smooth stripe from anterior margin terminating in anterior fifth. Lateral margin linearly attenuated in anterior quarter, stronger apically. Tip rounded. Elytra: Ovate; longer than broad; widest in anterior third; partly covering propygidium. Lateral margin slightly convex attenuate apically. Sculpture indistinct; almost complanate laterally; humerus indistinct; with a broad roundish torus beside scutellum; lateral trench missing; complanate along scutellum and suture. Loose punctate with extremely small to very small, shallow, roundish punctures, micropunctures in-between; with two loose, longitudinal rows of small, deep, roundish punctures: a juxta-sutural row from anterior margin crossing scutellar torus to apical torus; a supra-humeral row from anterior margin between humerus and scutellar torus to apical torus. Abdomen: Ventrite I extremely shortened medially; ventrite V broadest, not constricted ventrally. Propygidium with a slightly projected stoma at posterior corner. Pygidium slightly pulvinate longitudinally and laterally; upper margin slightly convex, linear medially; lateral margin linearly attenuate apically; tip rounded. Prothoracic leg: Femur and tibia slender. Tibia smooth ventrally. Median tooth robust. Basal tooth rudimentary. Interspace between teeth almost equal. Shape of interspace between teeth concave. Tarsomere I – IV combined extremely short; equal in length; slightly more than 1 / 2 in length of tibia. Tarsomere IV with a single ventral extension. Pretarsomere distinctly thickened; smooth ventrally; slightly more than 9 / 10 in length of tarsomere I – IV combined. Internal claw as long as pretarsomere. External claw slightly less than 9 / 10 in length of internal claw. Mesothoracic leg: Tibia slightly more than 4 / 5 in length of femur. Tarsomere I – IV combined extremely short; slightly less than 2 / 5 in length of tibia. Pretarsomere with a shortened smooth ridge ventrally, followed by a roundish cutting apically; slightly less than 1 / 10 more in length of tarsomere I – IV combined. External claw slightly less than 9 / 10 in length of pretarsomere. Internal claw shorter than external claw. Metathoracic leg: Tibia slightly longer than femur. Tarsomere I – IV combined distinctly short; slightly more than 7 / 10 in length of tibia. Pretarsomere slightly thickened; with an undulate ridge ventrally; slightly less than 1 / 2 in length of tarsomere I – IV combined. External claw slightly less than 1 / 10 more in length of pretarsomere. Internal claw slightly more than 4 / 5 in length of external claw. Parameres: (Figs. 94 – 95, 116). ♀: Unknown.	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E05EFFB1FF64D7EDFAA7FBF1.taxon	distribution	Distribution. Northern Cape Province (Fig. 140), only known from the type locality. Vegetation Type. Bokkeveld Sandstone Fynbos [FFs 1].	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E05EFFB1FF64D7EDFAA7FBF1.taxon	biology_ecology	Host Plants. Asteraceae: Didelta spinosa (L. f.) W. T. Aiton.	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E05EFFB1FF64D7EDFAA7FBF1.taxon	etymology	Etymology. The name derives from the bicoloured elytra [Latin: bi = two; Latin: color = colour].	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E058FFB3FF64D0D2FDD1F8FC.taxon	description	(Figs. 6, 10, 36, 96 – 97, 117) http: // zoobank. org / 497 F 3 A 0 F- 0 D 97 - 4 B 46 - A 0 FD- 49 C 67 A 87159 A	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E058FFB3FF64D0D2FDD1F8FC.taxon	materials_examined	Type locality. [RSA, WCP]: Biedouw Valley. Type material. [23] Holotype ♂: white label pr « RSA, WCP, Biedouw | Valley 32.05 S – 19.05 E | 08 / 09 / 98 | J. Colville » || yellow label hw « 18 » pr «. 827 | coll. Dombrow » || red label pr « Paratype | Beckhoplia | colvillei sp. n. | det.: Dombrow 2003 » || green label pr « Host plant: | Dimorphotheca sinuata | Ursinia anthemoides | Arctotheca calendula » || blue label pr « Bioregion: | Sandstone Fynbos | Vegetation Type: FFs 4 – | Cederberg Sandstone Fynbos » || yellow label pr « SAM – COL | A 071217 » || red label pr « Holotype | Beckhoplia | dolichiocnemis sp. n. | det.: H. Dombrow 2019 » (SAMC). – Paratypes 20 ♂ ♂: same labels as holotype but || yellow label hw « 18 » pr «. 818 –. 826,. 828 –. 836 | coll. Dombrow » || red label pr « Paratype | Beckhoplia | colvillei sp. n. | det.: Dombrow 2003 » || red label pr « Paratype | Beckhoplia | dolichiocnemis sp. n. | det.: H. Dombrow 2019 » (ESDW); 2 ♂♂: same labels as holotype but || yellow label hw « 18 » pr «. 837 –. 838 | coll. Dombrow » || red label pr « Paratype | Beckhoplia | colvillei sp. n. | det.: Dombrow 2003 » || yellow label pr « SAM – COL | A 071217 » || red label pr « Paratype | Beckhoplia | dolichiocnemis sp. n. | det.: H. Dombrow 2019 » (SAMC). Condition of holotype. Very good. Pinned. Dissected, parameres and spiculum gastrale glued on separate card below beetle.	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E058FFB3FF64D0D2FDD1F8FC.taxon	discussion	Remark. After consulting the collector, the coordinate of the locality has to be adjusted to 32 ° 08 ’ S, 19 ° 11 ’ E.	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E058FFB3FF64D0D2FDD1F8FC.taxon	diagnosis	Differential diagnosis. Differs from B. occidentalis: Elytra of B. dolichiocnemis with a longitudinal depression along suture. Elytra of B. occidentalis without a longitudinal depression along suture. Elytra of B. dolichiocnemis protruding the propygidium. Elytra of B. occidentalis partly covering propygidium. Interno-posterior margin of parameres of B. dolichiocnemis parabola-shaped in dorsal view (Figs. 96, 117). Interno-posterior margin of parameres of B. occidentalis curly bracket-shaped in dorsal view (Figs. 92, 115). Externo-posterior margin of parameres of B. dolichiocnemis undulate in dorsal view (Figs. 96, 117). Externoposterior margin of parameres of B. occidentalis curly bracket-shaped, convex medially in dorsal view (Figs. 92, 115). Differs from B. bicolor: Elytra of B. dolichiocnemis protruding propygidium. Elytra of B. bicolor partly covering propygidium. Elytra of B. dolichiocnemis transparently pale red brown with narrowly blackened lateral margins. Elytra of B. bicolor transparently pale-brown with broadly blackened lateral margins. Interno-posterior margin of parameres of B. dolichiocnemis parabola-shaped in dorsal view (Figs. 96, 117). Interno-posterior margin of parameres of B. bicolor u-shaped in dorsal view (Figs. 94, 116). Externo-posterior margin of parameres of B. dolichiocnemis undulate in dorsal view (Figs. 96, 117). Externo-posterior margin of parameres of B. bicolor concave with a broad convex median extension in dorsal view (Figs. 94, 116).	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E058FFB3FF64D0D2FDD1F8FC.taxon	description	Description Measurements: length 4.6 – 5.7 mm; width 2.1 – 2.8 mm. Integument: Black; elytra transparently pale-brown, lateral and posterior margin blackened. Tegument: Clypeus bare; inner side of the reflexed anterior and lateral margins loose setose with short, erect, black setae. Head loose setose with long, erect, black setae. Eye canthus with a loose setose ciliary of long, black setae. Scapus loose setose with long, black setae, pale-brown ventrally, sparse and medium-long on pedicellus; funiculum with few very short and short, erect, brow setae, sparse and semi-erect on clavus. Pronotum loose setose with long, erect, black setae, slightly shorter towards posterior margin; with a loose setose ciliary of distinctly long, black setae on lateral margin. Scutellum loose setose with short and mediumlong, semi-erect, brown setae. Elytra without pattern. Loose setose with short, reclinate, brown setae; long, erect, black setae interspersed, denser around humerus; with two loose setose longitudinal rows of mediumlong, semi-erect, black, setae: a juxta-sutural row from anterior margin crossing scutellar torus towards apical torus; a juxta-humeral row from anterior margin between humerus and scutellar torus towards apical torus terminating towards middle. Suture with a loose row of short and long, robust, semi-erect, black setae. Thorax and pleurites loose pilose with long, black hairs. Ventrite I – V loose setose with short, reclinate, brownish setae; medium-long, semi-erect, black setae interspersed. Propygidium loose setose with reclinate, short, whitish, squamiform setae; long, semi-erect, black setae interspersed. Pygidium sparse setose with long, erect, black setae; tip with a loose setose ciliary of long, black setae. Prothoracic femur loose pilose with long, black hairs; bare internally. Prothoracic tibia loose setose with short, reclinate, whitish and brownish setae dorsally; sparse setose with erect, medium-long whitish and brownish and long black setae ventrally; medium-long, black, semi-erect setae interspersed dorsally; with a sparse setose longitudinal row of mediumlong, erect, black setae dorsally; with a loose ciliary of short, black and whitish setae on internal ridge; whitish and short at external ridge; a setaceous ciliary of short, brown setae on anterior margin; a medium-long downwards pointing setose ciliary of pale-brown setae along anterior margin of apical tooth. Mesothoracic femur loose pilose with long, brown hairs; distinctly long brown ventrally; long, erect, black setae interspersed dorsally and externally; bare internally. Mesothoracic tibia sparse setose with short, reclinate, white setae, sparser internally; medium-long, semi-erect, robust, black setae interspersed, long and finer internally. Posterior margin with a bristled ciliary of medium-long, black bristles; missing internally. Metathoracic femur loose setose with medium-long and long, semi-erect, black setae; distinctly long, pale-brown setae interspersed ventrally; bare internally. Metathoracic tibia loose setose with short, reclinate, pale-brown setae; medium-long, semi-erect, black setae interspersed; denser, finer and long internally. Posterior margin with a loose bristled outer marginal ciliary of short, black bristles, lengthening dorsally and ventrally and more robust ventrally. Prothoracic and mesothoracic tarsomere I – IV bare. Metathoracic tarsomere I – IV sparse setose with very short, reclinate, pale-brown setae. Tarsomere I – IV of all thoracic legs with a setaceous, apical, marginal ciliary of short, black setae lengthening towards metathoracic tarsomeres, fine dorsally and more robust ventrally. Pretarsomere of all thoracic legs sparse setose with very short, semi-erect, brown setae; few, long, semi-erect, black setae at apical margin, shorter and dark red-brown ventrally. Head: Suture between head and clypeus linear, crenated. Head dense punctate with very small, shallow, roundish, partly merging punctures, micropunctate within. Funicle I – II cylindrical; funicle III short asymmetric frustum-shaped; clavus long elliptic, slightly longer than pedicellus and funiculum combined. Pronotum: Widest in anterior two fifth; dense punctate with shallow, small, roundish punctures, denser laterally and within posterior third of medial furrow. Anterior corner obtuse. Lateral margin linearly attenuate anteriorly, subparallel posteriorly. With an indistinct, shortened medial furrow from posterior margin terminating on disk. Posterior corner rounded. Scutellum: Parabola-shaped; as broad as long; dense punctate with shallow, very small to large, roundish punctures, micropunctate within; with a shortened, median, longitudinal, narrow, smooth stripe from anterior margin terminating in anterior fifth. Lateral margin slightly convex. Tip pointed. Elytra: Ovate; longer than broad; widest in anterior third; protruding propygidium. Lateral margin slightly convex attenuate apically. Sculpture indistinct; slightly pulvinate laterally; humerus indistinct; with a broad roundish torus beside scutellum, expanding towards disk; lateral trench missing; complanate along scutellum and suture. Loose punctate with extremely small to very small, shallow, roundish punctures, micropunctures in-between; with two loose, longitudinal rows of small, deep, roundish punctures: a juxta-sutural row from anterior margin crossing scutellar torus to apical torus; a supra-humeral row from anterior margin between humerus and scutellar torus to apical torus. Abdomen: Ventrite extremely shortened medially; ventrite V broadest, slightly constricted ventrally. Propygidium with a slightly projected stoma at posterior corner. Pygidium slightly pulvinate longitudinally and laterally; upper margin moderately convex, linear medially; lateral margin subparallel in upper fifth, linearly attenuate apically; tip rounded. Prothoracic leg: Femur and tibia robust. Tibia smooth ventrally. Median tooth elongate. Basal tooth rudimentary. Interspace between teeth almost equal. Shape of interspace between teeth concave. Tarsomere I – IV combined extremely short; equal in length; slightly more than 2 / 5 in length of tibia. Pretarsomere slightly thickened; smooth ventrally; slightly more than 1 / 10 more in length of tarsomere I – IV combined. Internal claw slightly longer than pretarsomere. External claw slightly less than 4 / 5 in length of internal claw. Mesothoracic leg: Tibia slightly less than 9 / 10 in length of femur. Tarsomere I – IV combined extremely short; slightly less than 3 / 10 in length of tibia. Pretarsomere with a shortened smooth ridge ventrally followed by a roundish cutting; slightly less than 1 / 5 more in length of tarsomere I – IV combined. External claw slightly shorter than pretarsomere. Internal claw slightly less than 9 / 10 in length of external claw. Metathoracic leg: Tibia slightly more than 9 / 10 in length of femur. Tarsomere I – IV combined distinctly short; slightly less than 4 / 5 in length of tibia. Pretarsomere slender; with a smooth ridge ventrally; slightly less than 1 / 2 in length of tarsomere I – IV combined. External claw slightly less than 1 / 5 more in length of pretarsomere. Internal claw slightly more than 4 / 5 in length of external claw. Parameres: (Figs. 96 – 97, 117). ♀: Unknown.	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E058FFB3FF64D0D2FDD1F8FC.taxon	distribution	Distribution. Western Cape Province (Fig. 140), only known from the type locality. Vegetation Type. Cederberg Sandstone Fynbos (FFs 4).	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E058FFB3FF64D0D2FDD1F8FC.taxon	biology_ecology	Host Plants. Asteraceae: Dimorphotheca sinuata DC.; Ursinia anthemoides (L.) Poir.; Arctotheca calendula (L.) Levyns.	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
03DFBE05E058FFB3FF64D0D2FDD1F8FC.taxon	etymology	Etymology. The specific name derives from the long, elongate hind legs [Greek: δολῐχóς (dolikhós-) = long; Greek: κνήμη (kn ḗmç) = the shin bone]	en	Dombrow, Holger E., Colville, Jonathan F. (2020): Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Zootaxa 4823: 1-79, DOI: 10.11646/zootaxa.4823.1.1
