taxonID	type	description	language	source
03DF87B01F5D8E0786C5FA7D2434FB71.taxon	materials_examined	Holotype: E MALAYSIA: 3, two labels: " SABAH \ Crocker Ra. \ 1550 - 1650 m \ 16. V. 1987 \ Burckhardt - Löbl " [white, printed]; " CEPHENNULA \ densepunctata m. \ det. P. Jałoszyński, 2011 \ HOLOTYPUS " [red, printed] (MHNG).	en	Jałoszyński, Paweł (2011): Ten new species of Cephennula Jałoszyński (Coleoptera, Staphylinidae, Scydmaeninae) from Malaysia. Zootaxa 3113: 36-52, DOI: 10.5281/zenodo.279337
03DF87B01F5D8E0786C5FA7D2434FB71.taxon	diagnosis	Diagnosis. Body reaching 1 mm in length; antennal club strongly flattened and compact; dorsum of head densely covered with large and deep punctures; punctures on frons and vertex nearly equally distributed; median part of pronotum with dense punctures; sublateral pronotal carinae as long as pronotum; subhumeral carinae slightly shorter than 1 / 3 x EL, humeral carinae distinctly shorter; aedeagus in ventral view drop-shaped, with triangular and pointed apex of median lobe; endophallus with two pairs of elongate sclerites — median pair strongly curved laterally and lateral pair nearly straight.	en	Jałoszyński, Paweł (2011): Ten new species of Cephennula Jałoszyński (Coleoptera, Staphylinidae, Scydmaeninae) from Malaysia. Zootaxa 3113: 36-52, DOI: 10.5281/zenodo.279337
03DF87B01F5D8E0786C5FA7D2434FB71.taxon	description	Description. Body of male (Fig. 3) strongly convex, elongate, with shallow but distinct constriction between pronotum and elytra, moderately glossy, dark brown, covered with yellowish vestiture, legs and antennae slightly lighter. BL 1.05 mm. Head broadest at large, coarsely faceted and strongly convex eyes, HL 0.13 mm, HW 0.24 mm; vertex and frontoclypeal region convex; supraantennal tubercles barely marked. Punctures on head dorsum large, deep and sharply defined, equally distributed and separated by spaces about as wide as half puncture diameter; setae short and sparse, suberect. Antennae with strongly enlarged, strongly flattened and compactly assembled antennomeres X – XI and moderately enlarged antennomere IX, so that club appears two-segmented, AnL 0.35 mm; antennomere I about 1.6 x as long as broad; II 2.3 x as long as broad; III 1.2 x as long as broad; IV – VII subequal in length and width, each 1.6 x as long as broad; VIII slightly broader but distinctly shorter than VII, 1.2 x as long as broad; IX clearly broader and longer than VIII, as long as broad; X much longer and broader than IX, slightly transverse; XI slightly narrower but distinctly longer than X, 1.6 x as long as broad. Pronotum approximately semielliptical, broadest slightly anterior to middle; PL 0.33 mm, PW 0.45 mm, anterior margin weakly rounded; lateral margins distinctly microserrate, strongly rounded in anterior half, very weakly concave in posterior 1 / 4 and distinctly convergent towards sharp and acute hind angles; posterior margin feebly bisinuate; sublateral carinae distinct and as long as pronotum. Transverse ante-basal groove shallow but sharply marked and slightly arcuate; sublateral ante-basal foveae small and shallow; lateral foveae deep and large. Punctures on median part of pronotal disc larger than those on head, deep and sharply marked, separated by spaces as wide as half puncture diameter, punctures between sublateral pronotal carinae and lateral margins of pronotum slightly larger and denser than those in middle; setae sparse, moderately long, suberect. Elytra strongly convex, oval, broadest between middle and anterior 1 / 3; EL 0.60 mm, EW 0.50 mm, EI 1.20; base of each elytron with rudimentary fovea located much closer to scutellum than to base of humeral carina; subhumeral carinae distinct, slightly shorter than 1 / 3 of EL; humeral carinae slightly less distinct and slightly shorter. Punctures on elytra very fine and shallow but with sharp margins, separated by spaces 1.5 – 2 x as wide as puncture diameter; setae similar as those on pronotum. Legs moderately long and slender; all tibiae nearly straight. Aedeagus (Figs. 13, 14) in ventral view drop-shaped with narrow, subtriangular and pointed apex; AeL 0.24 mm. Endophallus darkly sclerotized, with two pairs of elongate sclerites — shorter and strongly curved median pair and longer lateral, nearly straight pair directed towards apex of aedeagus; parameres slender, not exceeding apex of median lobe, each bearing single apical seta. Female. Unknown.	en	Jałoszyński, Paweł (2011): Ten new species of Cephennula Jałoszyński (Coleoptera, Staphylinidae, Scydmaeninae) from Malaysia. Zootaxa 3113: 36-52, DOI: 10.5281/zenodo.279337
03DF87B01F5D8E0786C5FA7D2434FB71.taxon	distribution	Distribution. East Malaysia (Borneo: Sabah).	en	Jałoszyński, Paweł (2011): Ten new species of Cephennula Jałoszyński (Coleoptera, Staphylinidae, Scydmaeninae) from Malaysia. Zootaxa 3113: 36-52, DOI: 10.5281/zenodo.279337
03DF87B01F5D8E0786C5FA7D2434FB71.taxon	etymology	Etymology. Locotypical, after the state Sarawak of Malaysia.	en	Jałoszyński, Paweł (2011): Ten new species of Cephennula Jałoszyński (Coleoptera, Staphylinidae, Scydmaeninae) from Malaysia. Zootaxa 3113: 36-52, DOI: 10.5281/zenodo.279337
03DF87B01F5C8E0186C5FB232784FC75.taxon	materials_examined	Holotype: E MALAYSIA: 3, two labels: " SABAH \ Mt. Kinabalu \ 1450 - 1550 m \ 23. V. 1987 \ Burckhardt - Löbl " [white, printed]; " CEPHENNULA \ densepunctata m. \ det. P. Jałoszyński, 2011 \ HOLOTYPUS " [red, printed] (MHNG). Paratypes (3 33, 13 ƤƤ): 1 3, 3 ƤƤ, same data as holotype; 1 3, 4 ƤƤ, same data except 1500 m and 21. V. 1987; 1 3, 1 Ƥ, same data except 1500 m and 30. IV. 1987; 3 ƤƤ, same data except 1430 m and 22. V. 1987; 1 Ƥ, " SABAH: E Mt. Kinabalu \ 1150 m, rte Ranau-Kota \ Kinabalu, 24. V. 1987 \ Burckhardt - Löbl " [white, printed]; 1 Ƥ, " BORNEO SABAH Mt. \ Kinabalu Nat. Pk. \ HQ Liwagu River \ 1490 m 18. V. 1987 \ A. Smetana " (MHNG, PCPJ).	en	Jałoszyński, Paweł (2011): Ten new species of Cephennula Jałoszyński (Coleoptera, Staphylinidae, Scydmaeninae) from Malaysia. Zootaxa 3113: 36-52, DOI: 10.5281/zenodo.279337
03DF87B01F5C8E0186C5FB232784FC75.taxon	diagnosis	Diagnosis. Body reaching or nearly reaching 1 mm in length; antennal club strongly flattened and compact; punctures on head unevenly distributed, those on sides of vertex, on frons and clypeus separated by spaces 0.5 – 2 x as wide as puncture diameter, vertex with impunctate median area; median part of pronotum with dense punctures; sublateral pronotal carinae as long as pronotum; subhumeral carinae about as long as 1 / 3 x EL and distinctly longer than humeral carinae; aedeagus in ventral view drop-shaped, with triangular and pointed apex of median lobe; endophallus with four pairs of elongate sclerites, of which three pairs are curved laterally and one nearly straight.	en	Jałoszyński, Paweł (2011): Ten new species of Cephennula Jałoszyński (Coleoptera, Staphylinidae, Scydmaeninae) from Malaysia. Zootaxa 3113: 36-52, DOI: 10.5281/zenodo.279337
03DF87B01F5C8E0186C5FB232784FC75.taxon	description	Description. Body of male (Fig. 4) externally nearly identical as C. densepunctata, only differences are described below. BL 0.98 – 1.00 mm (mean 0.98 mm). Head unevenly covered with moderately large and deep, sharply marked punctures, on sides of vertex, on frons and clypeus separated by spaces 0.5 – 2 x as wide as puncture diameter, vertex in middle impunctate; HL 0.11 – 0.13 mm (mean 0.12 mm), HW 0.21 – 0.24 mm (mean 0.22 mm). AnL 0.35 – 0.38 mm (mean 0.36 mm). Median part of pronotum covered with punctures similar to those on head and becoming gradually larger and denser towards sides, those on median part of disc separated by spaces approximately equal to puncture diameter or slightly wider, punctures between sublateral carinae and lateral margins of pronotum distinctly larger and denser than those in middle, nearly adjacent to each other; PL 0.30 – 0.31 mm (mean 0.30 mm), PW 0.43 – 0.45 mm (mean 0.43 mm). Elytra covered with fine and sparse punctures, EL 0.55 – 0.58 mm (mean 0.56 mm), EW 0.48 – 0.50 mm (mean 0.49 mm), EI 1.13 – 1.18 (mean 1.15); subhumeral carinae about as long as 1 / 3 x EL; humeral carinae less distinct and much shorter. Aedeagus (Figs. 15, 16) in ventral view drop-shaped with narrow, subtriangular and pointed apex; AeL 0.23 mm. Endophallus darkly sclerotized, with four pairs of elongate sclerites — three pairs strongly curved laterally and one median pair nearly straight and directed towards apex of aedeagus; parameres slender, not exceeding apex of median lobe, each bearing two apical setae. Female. Externally indistinguishable from males. BL 1.00 – 1.01 mm (mean 1.01 mm); HL 0.11 – 0.13 mm (mean 0.12 mm), HW 0.23 – 0.24 mm (mean 0.24 mm), AnL 0.34 – 0.38 mm (mean 0.35 mm); PL 0.31 – 0.33 mm (mean 0.32 mm), PW 0.45 – 0.48 mm (mean 0.46 mm); EL 0.56 – 0.58 mm (mean 0.57 mm), EW 0.50 mm, EI 1.13 – 1.15 (mean 1.15).	en	Jałoszyński, Paweł (2011): Ten new species of Cephennula Jałoszyński (Coleoptera, Staphylinidae, Scydmaeninae) from Malaysia. Zootaxa 3113: 36-52, DOI: 10.5281/zenodo.279337
03DF87B01F5C8E0186C5FB232784FC75.taxon	distribution	Distribution. East Malaysia (Borneo: Sabah).	en	Jałoszyński, Paweł (2011): Ten new species of Cephennula Jałoszyński (Coleoptera, Staphylinidae, Scydmaeninae) from Malaysia. Zootaxa 3113: 36-52, DOI: 10.5281/zenodo.279337
03DF87B01F5C8E0186C5FB232784FC75.taxon	etymology	Etymology. The name is derived from Latin soror gemina (a twin sister) and refers to the high similarity of this species to C. densepunctata.	en	Jałoszyński, Paweł (2011): Ten new species of Cephennula Jałoszyński (Coleoptera, Staphylinidae, Scydmaeninae) from Malaysia. Zootaxa 3113: 36-52, DOI: 10.5281/zenodo.279337
03DF87B01F5C8E0186C5FB232784FC75.taxon	discussion	Remarks. This species is externally nearly identical to C. densepunctata. In addition to the clearly different aedeagus C. gemina can be distinguished also by distinctly smaller and sparser punctures on the head and pronotum.	en	Jałoszyński, Paweł (2011): Ten new species of Cephennula Jałoszyński (Coleoptera, Staphylinidae, Scydmaeninae) from Malaysia. Zootaxa 3113: 36-52, DOI: 10.5281/zenodo.279337
03DF87B01F5A8E0286C5FC20245FFCE6.taxon	materials_examined	Holotype: E MALAYSIA: 3, two labels: " SARAWAK \ Semengoh For. \ 11 miSW Kuching \ R. Taylor, 5.68 " [white, printed]; " CEPHENNULA \ minutissima m. \ det. P. Jałoszyński, 2011 \ HOLOTYPUS " [red, printed] (MHNG).	en	Jałoszyński, Paweł (2011): Ten new species of Cephennula Jałoszyński (Coleoptera, Staphylinidae, Scydmaeninae) from Malaysia. Zootaxa 3113: 36-52, DOI: 10.5281/zenodo.279337
03DF87B01F5A8E0286C5FC20245FFCE6.taxon	diagnosis	Diagnosis. Body length below 0.7 mm; antennal club slightly flattened and compact; dorsum of head nearly evenly and densely covered with large and deep punctures; punctures on median part of pronotum moderately large and dense; sublateral pronotal carinae fine and obliterated in anterior 1 / 4 of pronotum; subhumeral carinae nearly as long as half EL and about twice as long as humeral carinae; aedeagus in ventral view drop-shaped, with triangular and blunt apex; endophallus with one pair of elongate sclerites directed towards apex of aedeagus.	en	Jałoszyński, Paweł (2011): Ten new species of Cephennula Jałoszyński (Coleoptera, Staphylinidae, Scydmaeninae) from Malaysia. Zootaxa 3113: 36-52, DOI: 10.5281/zenodo.279337
03DF87B01F5A8E0286C5FC20245FFCE6.taxon	description	Description. Body of male (Fig. 5) strongly convex, elongate, with very shallow constriction between pronotum and elytra, moderately glossy, yellowish-brown, covered with vestiture only slightly lighter than cuticle, legs and antennae slightly lighter. BL 0.68 mm. Head broadest at large, coarsely faceted and strongly convex eyes, HL 0.08 mm, HW 0.18 mm; vertex and frontoclypeal region convex; supraantennal tubercles barely marked. Punctures on dorsum of head large, deep and dense, nearly equally distributed (minimally sparser in median part of frons); setae very short, sparse and nearly recumbent. Antennae with strongly enlarged, slightly flattened and compactly assembled antennomeres X – XI and only slightly enlarged antennomere IX, so that club appears two-segmented, AnL 0.25 mm; antennomere I about 1.3 x as long as broad; II 2.2 x as long as broad; III slightly transverse; IV – VI subequal in length and width, each about 1.2 x as long as broad; VII slightly broader than VI, about as long as broad; VIII slightly shorter and narrower than VII, slightly transverse; IX distinctly broader and longer than VIII, distinctly transverse; X much longer and broader than IX, strongly transverse; XI about as broad as X, 1.4 x as long as broad. Pronotum approximately semielliptical, broadest between middle and anterior 1 / 3; PL 0.20 mm, PW 0.28 mm, anterior margin strongly rounded; lateral margins distinctly microserrate, strongly rounded in anterior 1 / 3, straight in posterior half and slightly convergent towards sharp and acute hind angles; posterior margin very weakly bisinuate; sublateral carinae very fine and obliterated near anterior 1 / 4 of pronotum. Transverse ante-basal groove very shallow and indistinct, slightly arcuate; sublateral and lateral ante-basal foveae equally small and shallow. Punctures on median part of pronotal disc slightly smaller than those on head, deep and dense, punctures between sublateral pronotal carinae and lateral margins of pronotum distinctly larger and denser, nearly adjacent to each other but not coarse; setae sparse and short, nearly recumbent except for suberect setae along lateral margins. Elytra strongly convex, oval, broadest between middle and anterior 1 / 3; EL 0.40 mm, EW 0.33 mm, EI 1.23; base of each elytron with very small but distinct basal fovea located much closer to scutellum than to base of humeral carina; subhumeral carinae distinct, nearly as long as 1 / 2 x EL; humeral carinae similarly distinct but about half shorter. Punctures on elytra distinctly smaller than those on head and pronotum, very shallow and with diffused margins, dense; setae very short, sparse, nearly recumbent. Legs moderately long and slender; pro- and mesotibiae nearly straight, metatibiae slightly curved. Aedeagus (Figs. 17, 18) in ventral view drop-shaped with moderately broad, subtriangular and rounded apex; AeL 0.18 mm. Endophallus darkly sclerotized, with one pair of weakly curved elongate sclerites directed towards apex of aedeagus; parameres slender, not exceeding apex of median lobe, each bearing single apical seta. Female. Unknown.	en	Jałoszyński, Paweł (2011): Ten new species of Cephennula Jałoszyński (Coleoptera, Staphylinidae, Scydmaeninae) from Malaysia. Zootaxa 3113: 36-52, DOI: 10.5281/zenodo.279337
03DF87B01F5A8E0286C5FC20245FFCE6.taxon	distribution	Distribution. East Malaysia (Borneo: Sarawak).	en	Jałoszyński, Paweł (2011): Ten new species of Cephennula Jałoszyński (Coleoptera, Staphylinidae, Scydmaeninae) from Malaysia. Zootaxa 3113: 36-52, DOI: 10.5281/zenodo.279337
03DF87B01F5A8E0286C5FC20245FFCE6.taxon	etymology	Etymology. The specific epithet “ minutissima ”, Latin " the smallest ", refers to the extremely small body of this species.	en	Jałoszyński, Paweł (2011): Ten new species of Cephennula Jałoszyński (Coleoptera, Staphylinidae, Scydmaeninae) from Malaysia. Zootaxa 3113: 36-52, DOI: 10.5281/zenodo.279337
03DF87B01F5A8E0286C5FC20245FFCE6.taxon	discussion	Remarks. This is the smallest species of Cephennula known so far, and one of two with strikingly short, nearly recumbent vestiture, especially on the elytra. The other Cephennula with such unusually short setae is C. penrissenana, which clearly differs in genital characters.	en	Jałoszyński, Paweł (2011): Ten new species of Cephennula Jałoszyński (Coleoptera, Staphylinidae, Scydmaeninae) from Malaysia. Zootaxa 3113: 36-52, DOI: 10.5281/zenodo.279337
03DF87B01F598E0386C5FC5B2413FD1D.taxon	materials_examined	Holotype: E MALAYSIA: 3, two labels: " E MALAYSIA: Sarawak \ Gn. Penrissen, 1000 m \ 23. V. 1994, edge prim. \ montane for., # 9 a \ Löbl & Burckhardt " [white, printed]; " CEPHENNULA \ penrissenana m. \ det. P. Jałoszyński, 2011 \ HOLOTYPUS " [red, printed] (MHNG).	en	Jałoszyński, Paweł (2011): Ten new species of Cephennula Jałoszyński (Coleoptera, Staphylinidae, Scydmaeninae) from Malaysia. Zootaxa 3113: 36-52, DOI: 10.5281/zenodo.279337
03DF87B01F598E0386C5FC5B2413FD1D.taxon	diagnosis	Diagnosis. Body between 0.7 and 0.8 mm in length; antennal club strongly flattened and compact; head covered with moderately large and unevenly distributed punctures, those on vertex and sides of frons dense, punctures on median part of frons distinctly sparser; median part of pronotum with small and dense punctures; sublateral pronotal carinae obliterated in anterior 1 / 5 of pronotum; subhumeral carinae distinct, as long as 1 / 3 x EL and much longer than rudimentary humeral carinae; aedeagus in ventral view drop-shaped, with broad, nearly subrectangular apex and nearly straight apical margin; endophallus with three pairs of strongly curved elongate sclerites.	en	Jałoszyński, Paweł (2011): Ten new species of Cephennula Jałoszyński (Coleoptera, Staphylinidae, Scydmaeninae) from Malaysia. Zootaxa 3113: 36-52, DOI: 10.5281/zenodo.279337
03DF87B01F598E0386C5FC5B2413FD1D.taxon	description	Description. Body of male (Fig. 6) strongly convex, elongate, with shallow but distinct constriction between pronotum and elytra, moderately glossy, light brown, covered with yellowish vestiture, legs and antennae slightly lighter. BL 0.74 mm. Head broadest at large, coarsely faceted and strongly convex eyes, HL 0.08 mm, HW 0.18 mm; vertex and frontoclypeal region convex; supraantennal tubercles barely marked. Punctures on dorsum of head unevenly distributed, moderately large and deep, on vertex and sides of frons dense, median part of frons with sparser punctures, separated by spaces 1.5 – 2 x wider than puncture diameter, punctures on clypeus separated by spaces as wide as puncture diameter; setae very short and sparse, nearly recumbent. Antennae with strongly enlarged, strongly flattened and compactly assembled antennomeres X – XI and moderately enlarged antennomere IX, so that club appears two-segmented, AnL 0.25 mm; antennomere I about 1.3 x as long as broad; II 2 x as long as broad; III as long as broad; IV – VII subequal in length and width, each about 1.2 x as long as broad; VII as broad as VII but distinctly shorter, minimally broader than long; IX only slightly longer and much broader than VIII, strongly transverse; X much longer and broader than IX, distinctly transverse; XI slightly narrower than X, 1.5 x as long as broad. Pronotum approximately semielliptical, broadest between middle and anterior 1 / 3; PL 0.24 mm, PW 0.33 mm, anterior margin weakly rounded; lateral margins distinctly microserrate, strongly rounded in anterior 1 / 3, nearly straight in posterior 1 / 3 and distinctly convergent towards sharp and acute hind angles; posterior margin very weakly bisinuate; sublateral carinae fine but distinct, obliterated in anterior 1 / 5 of PL. Transverse ante-basal groove very shallow and indistinct, slightly biarcuate; sublateral ante-basal foveae small and shallow; lateral ante-basal foveae slightly larger but also shallow. Punctures on median part of pronotal disc slightly smaller than those on head but deep and dense, punctures between sublateral pronotal carinae and lateral margins of pronotum distinctly larger and denser than those in middle, nearly adjacent to each other but not coarse; setae sparse, very short, nearly recumbent. Elytra strongly convex, oval, broadest between middle and anterior 1 / 3; EL 0.43 mm, EW 0.34 mm, EI 1.26; base of each elytron with rudimentary basal fovea located much closer to scutellum than to base of humeral carina; subhumeral carinae distinct, about as long as 1 / 3 x EL; humeral carinae rudimentary, shorter than 1 / 7 x EL. Punctures on elytra much smaller than those on head and pronotum, fine and shallow but distinct and dense; setae similar as those on pronotum. Legs moderately long and slender; pro- and mesotibiae nearly straight, metatibiae slightly curved. Aedeagus (Figs. 19, 20) in ventral view drop-shaped with relatively broad, subrectangular apex and nearly straight apical margin; AeL 0.18 mm. Endophallus darkly sclerotized, with three pairs of elongate sclerites — very short median pair curved ventrally, very long sublateral pair strongly curved laterally and very short lateral pair strongly curved laterally; parameres slender, nearly reaching apex of median lobe, each bearing single apical seta. Female. Unknown.	en	Jałoszyński, Paweł (2011): Ten new species of Cephennula Jałoszyński (Coleoptera, Staphylinidae, Scydmaeninae) from Malaysia. Zootaxa 3113: 36-52, DOI: 10.5281/zenodo.279337
03DF87B01F598E0386C5FC5B2413FD1D.taxon	distribution	Distribution. East Malaysia (Borneo: Sarawak).	en	Jałoszyński, Paweł (2011): Ten new species of Cephennula Jałoszyński (Coleoptera, Staphylinidae, Scydmaeninae) from Malaysia. Zootaxa 3113: 36-52, DOI: 10.5281/zenodo.279337
03DF87B01F598E0386C5FC5B2413FD1D.taxon	etymology	Etymology. Locotypical, after Mount Penrissen in Borneo.	en	Jałoszyński, Paweł (2011): Ten new species of Cephennula Jałoszyński (Coleoptera, Staphylinidae, Scydmaeninae) from Malaysia. Zootaxa 3113: 36-52, DOI: 10.5281/zenodo.279337
03DF87B01F588E0C86C5FCC225ABFDC6.taxon	materials_examined	Holotype: E MALAYSIA: 3, two labels: " SABAH: Poring Hot \ Springs, 550 - 600 m \ 9. V. 1987 \ Burckhardt - Löbl " [white, printed]; " CEPHENNULA \ poringana m. \ det. P. Jałoszyński, 2011 \ HOLOTYPUS " [red, printed] (MHNG).	en	Jałoszyński, Paweł (2011): Ten new species of Cephennula Jałoszyński (Coleoptera, Staphylinidae, Scydmaeninae) from Malaysia. Zootaxa 3113: 36-52, DOI: 10.5281/zenodo.279337
03DF87B01F588E0C86C5FCC225ABFDC6.taxon	diagnosis	Diagnosis. Body between 0.8 and 0.9 mm in length; antennal club strongly flattened and compact; vertex with median impunctate area; punctures on median part of frons sparser than those on sides; median part of pronotum with moderately large punctures separated by spaces equal to and wider than puncture diameter; sublateral pronotal carinae as long as pronotum; subhumeral carinae nearly as long as 1 / 2 x EL and nearly twice as long as humeral carinae; aedeagus in ventral view drop-shaped, with moderately narrow, triangular and blunt apex; endophallus with two pairs of elongate sclerites — short and nearly straight median pair and long lateral pair strongly curved laterally.	en	Jałoszyński, Paweł (2011): Ten new species of Cephennula Jałoszyński (Coleoptera, Staphylinidae, Scydmaeninae) from Malaysia. Zootaxa 3113: 36-52, DOI: 10.5281/zenodo.279337
03DF87B01F588E0C86C5FCC225ABFDC6.taxon	description	Description. Body of male (Fig. 7) strongly convex, elongate, with shallow but distinct constriction between pronotum and elytra, moderately glossy, light brown, covered with yellowish vestiture, legs and antennae slightly lighter. BL 0.85 mm. Head broadest at large, coarsely faceted and strongly convex eyes, HL 0.13 mm, HW 0.24 mm; vertex and frontoclypeal area convex; supraantennal tubercles barely marked. Punctures on dorsum of head unevenly distributed, large and deep, on sides of vertex and frons dense, median part of frons and clypeus with sparser punctures, separated by spaces 1.5 – 2 x wider than puncture diameter, median part of vertex impunctate; setae short and sparse, suberect. Antennae with strongly enlarged, flattened and compactly assembled antennomeres X – XI and moderately enlarged antennomere IX, so that club appears two-segmented, AnL 0.45 mm; antennomere I about 1.3 x as long as broad; II 2.2 x as long as broad; III as long as broad; IV 1.2 x as long as broad; V – VII subequal in length and width, each 1.3 x as long as broad; VIII slightly broader but shorter than VII, 1.1 x as long as broad; IX distinctly broader and slightly longer than VIII, about as long as broad; X much longer and broader than IX, distinctly transverse; XI slightly narrower than X, 1.5 x as long as broad. Pronotum approximately semielliptical, broadest near middle; PL 0.25 mm, PW 0.40 mm, anterior margin weakly rounded; lateral margins distinctly microserrate, strongly rounded in anterior half, nearly straight in posterior 1 / 4 and slightly convergent towards sharp and acute hind angles; posterior margin very weakly bisinuate; sublateral carinae as long as pronotum. Transverse ante-basal groove shallow but sharply marked and slightly arcuate; sublateral ante-basal foveae small and shallow; lateral ante-basal foveae slightly deeper and larger. Punctures on median part of pronotal disc slightly smaller than those on head, deep and sparse, punctures between sublateral pronotal carinae and lateral margins of pronotum distinctly larger and denser than those in middle, nearly adjacent to each other and coarse; setae sparse, moderately long, suberect. Elytra strongly convex, oval, broadest between middle and anterior 1 / 3; EL 0.50 mm, EW 0.43 mm, EI 1.18; base of each elytron with rudimentary basal fovea located slightly closer to scutellum than to base of humeral carina; subhumeral carinae distinct, nearly as long as 1 / 2 x EL and nearly twice as long as slightly less distinct humeral carinae. Punctures on elytra much smaller than those on head and pronotum, fine and shallow but distinct, separated by spaces 1 – 2 x as wide as puncture diameter; setae similar as those on pronotum. Legs moderately long and slender; pro- and mesotibiae nearly straight, metatibiae slightly curved. Aedeagus (Figs. 21, 22) in ventral view drop-shaped with moderately narrow, subtriangular and blunt apex; AeL 0.18 mm. Endophallus darkly sclerotized, with two pairs of elongate sclerites — very short and nearly straight median pair and very long lateral pair strongly curved laterally; parameres slender, nearly reaching apex of median lobe, each bearing single apical seta. Female. Unknown.	en	Jałoszyński, Paweł (2011): Ten new species of Cephennula Jałoszyński (Coleoptera, Staphylinidae, Scydmaeninae) from Malaysia. Zootaxa 3113: 36-52, DOI: 10.5281/zenodo.279337
03DF87B01F588E0C86C5FCC225ABFDC6.taxon	distribution	Distribution. East Malaysia (Borneo: Sabah).	en	Jałoszyński, Paweł (2011): Ten new species of Cephennula Jałoszyński (Coleoptera, Staphylinidae, Scydmaeninae) from Malaysia. Zootaxa 3113: 36-52, DOI: 10.5281/zenodo.279337
03DF87B01F588E0C86C5FCC225ABFDC6.taxon	etymology	Etymology. Locotypical, after the Poring resort.	en	Jałoszyński, Paweł (2011): Ten new species of Cephennula Jałoszyński (Coleoptera, Staphylinidae, Scydmaeninae) from Malaysia. Zootaxa 3113: 36-52, DOI: 10.5281/zenodo.279337
03DF87B01F578E0886C5FBE926AAFBE5.taxon	materials_examined	Holotype: W MALAYSIA: 3, two labels: " W. Malaysia: Pahang \ Genting Highlands # 28 \ Awana, 950 m, 4. IV. 93 \ Löbl & Calame " [white, printed]; " CEPHENNULA \ titiwangsana m. \ det. P. Jałoszyński, 2011 \ HOLOTYPUS " [red, printed] (MHNG). Paratype: 3, same data as holotype (MHNG).	en	Jałoszyński, Paweł (2011): Ten new species of Cephennula Jałoszyński (Coleoptera, Staphylinidae, Scydmaeninae) from Malaysia. Zootaxa 3113: 36-52, DOI: 10.5281/zenodo.279337
03DF87B01F578E0886C5FBE926AAFBE5.taxon	diagnosis	Diagnosis. Body nearly reaching 1 mm in length; antennal club nearly cylindrical and loosely assembled; head unevenly covered with moderately large and deep punctures, those on vertex and sides of frons separated by spaces 0.5 – 2 x as wide as puncture diameter; frons with median impunctate area; median part of pronotum with punctures about as fine and sparse as those on elytra; sublateral pronotal carinae weakly marked and developed only in posterior half; subhumeral carinae slightly shorter than 1 / 4 x EL and slightly shorter than humeral carinae; aedeagus in ventral view bottle-shaped with apical margin rounded; endophallus with median and apical group of lightly pigmented elongate sclerites.	en	Jałoszyński, Paweł (2011): Ten new species of Cephennula Jałoszyński (Coleoptera, Staphylinidae, Scydmaeninae) from Malaysia. Zootaxa 3113: 36-52, DOI: 10.5281/zenodo.279337
03DF87B01F578E0886C5FBE926AAFBE5.taxon	description	Description. Body of male (Fig. 8) strongly convex, elongate, with shallow but distinct constriction between pronotum and elytra, moderately glossy, moderately dark brown, covered with yellowish vestiture, legs and antennae slightly lighter. BL 0.91 – 0.94 mm. Head broadest at large, coarsely faceted and strongly convex eyes, HL 0.13 mm, HW 0.24 mm; vertex and frontoclypeal area convex; supraantennal tubercles barely marked. Punctures on dorsum of head unevenly distributed, those on vertex and sides of frons and clypeus moderately large, deep and separated by spaces 0.5 – 2 x as wide as puncture diameter, median area on frons and clypeus nearly impunctate; setae short and sparse, suberect. Antennae with distinct, loosely assembled and only minimally flattened 3 - segmented club, AnL 0.43 mm; antennomere I about 1.5 x as long as broad; II twice as long as broad; III as long as broad; IV 1.1 x as long as broad; V – VI each 1.2 x as long as broad; VII distinctly longer and broader than VI, 1.5 x as long as broad; VIII slightly shorter and narrower than VII, 1.2 x as long as broad; IX clearly longer and broader than VIII, 1.1 x as long as broad; X much broader and longer than IX, slightly transverse; XI nearly as broad as X and 1.6 x as long as broad. FIGURES 13 – 24. Aedeagus in ventral (13, 15, 17, 19, 21, 23) and lateral (14, 16, 18, 20, 22, 24) views. 13, 14 ― Cephennula densepunctata sp. n.; 15, 16 ― Cephennula gemina sp. n.; 17, 18 ― Cephennula minutissima sp. n.; 19, 20 ― Cephennula penrissenana sp. n.; 21, 22 ― Cephennula poringana sp. n.; 23, 24 ― Cephennula titiwangsana sp. n. Scale bars: 0.05 mm. Pronotum nearly rectangular, broadest near anterior 1 / 3; PL 0.26 mm, PW 0.39 – 0.40 mm, anterior margin weakly rounded; lateral margins distinctly microserrate, strongly rounded only in anterior 1 / 3, then very weakly concave or nearly straight and slightly convergent towards nearly right and acute hind angles; posterior margin very weakly bisinuate; sublateral carinae very fine, developed only in posterior half. Transverse ante-basal groove moderately deep, sharply marked and slightly biarcuate; sublateral ante-basal foveae small but deep and nearly fused with similarly deep but slightly larger lateral foveae. Punctures on entire pronotum, including area between sublateral carinae and lateral margins, much finer than those on sides of frons, relatively sharply marked and sparse; setae sparse, moderately long, suberect. Elytra strongly convex, oval, broadest near anterior 1 / 3; EL 0.53 – 0.55 mm, EW 0.48 mm, EI 1.11 – 1.16; base of each elytron with rudimentary basal fovea located slightly closer to scutellum than to base of humeral carina; subhumeral carinae distinct, slightly shorter than 1 / 4 x EL; humeral carina distinct and slightly longer than subhumeral carina but also shorter than 1 / 4 x EL. Punctures on elytra about as fine as those on median part of pronotum, small and shallow but distinct and with slightly raised margins, separated by spaces 1.5 – 2 x as wide as puncture diameter; setae similar as those on pronotum. Legs moderately long and slender; pro- and mesotibiae nearly straight, metatibiae slightly recurved. Aedeagus (Figs. 23, 24) in ventral view bottle-shaped with broad, subtrapezoidal and rounded apex; AeL 0.16 mm. Endophallus moderately darkly sclerotized, with one pair of broad and dark and three pairs of slender and light, indistinct elongate sclerites directed towards apex of aedeagus, additionally apical part of endophallus with two bunches of lightly pigmented elongate sclerites; parameres relatively broad, not exceeding apex of median lobe, each bearing one apical and one subapical seta. Female. Unknown.	en	Jałoszyński, Paweł (2011): Ten new species of Cephennula Jałoszyński (Coleoptera, Staphylinidae, Scydmaeninae) from Malaysia. Zootaxa 3113: 36-52, DOI: 10.5281/zenodo.279337
03DF87B01F578E0886C5FBE926AAFBE5.taxon	distribution	Distribution. West Malaysia (Pahang).	en	Jałoszyński, Paweł (2011): Ten new species of Cephennula Jałoszyński (Coleoptera, Staphylinidae, Scydmaeninae) from Malaysia. Zootaxa 3113: 36-52, DOI: 10.5281/zenodo.279337
03DF87B01F578E0886C5FBE926AAFBE5.taxon	etymology	Etymology. Locotypical, after the Titiwangsa Mountains, where the Genting Highlands are located.	en	Jałoszyński, Paweł (2011): Ten new species of Cephennula Jałoszyński (Coleoptera, Staphylinidae, Scydmaeninae) from Malaysia. Zootaxa 3113: 36-52, DOI: 10.5281/zenodo.279337
03DF87B01F578E0886C5FBE926AAFBE5.taxon	discussion	Remarks. The aedeagus of this species shows characters intermediary between those of the minuta and multicarinata species groups. The general shape of the aedeagus is more similar to that found in the multicarinata group, while the presence of the median complex of elongate sclerites is characteristic of the minuta group. In C. titiwangsana the sclerites are lightly pigmented, mostly indistinct, except for one median pair. However, these two sclerites are different in shape from structures typical for the minuta group and form the median complex together with slender, needle-like and lightly pigmented structures. Additionally, the endophallus has the apical complex composed of two densely assembled bunches of elongate sclerites. These structures are not known in any species of the minuta group. Cephennula titiwangsana is also unusual by having nearly cylindrical, loosely assembled antennal club, while in majority of species the club is strongly flattened and more (usually strongly) compact. Only C. porcata from Thailand has a similar antennal club; this species also has the endophallus with apical sclerites, but without any median complex.	en	Jałoszyński, Paweł (2011): Ten new species of Cephennula Jałoszyński (Coleoptera, Staphylinidae, Scydmaeninae) from Malaysia. Zootaxa 3113: 36-52, DOI: 10.5281/zenodo.279337
03DF87B01F538E0986C5FB5026F5FAA6.taxon	materials_examined	Holotype: W MALAYSIA: 3, two labels: " W. Malaysia: Pahang \ Cameron Highlands \ 1500 m, trails 4, 13: # 15 \ Löbl & Calame, 23.3.93 " [white, printed]; " CEPHENNULA \ medioglabra m. \ det. P. Jałoszyński, 2011 \ HOLO- TYPUS " [red, printed] (MHNG). Paratypes (533, 2 ƤƤ): 13, " W. Malaysia: Pahang # 16 a \ Cameron Highls 1600 m \ Bukit Mentiga, trail 14 \ Löbl & Calame, 23.3.93 " [white, printed]; 333, 2 ƤƤ, " W. Malaysia: Pahang # 18 b \ Cameron Highls 1550 m \ Gunung Jasar, trail 11 \ Löbl & Calame, 24.3.93 "; 13, " W. Malaysia: Pahang \ Cameron Highls 1720 m \ Gunung Jasar # 19 b \ Löbl & Calame, 25.3.93 " (MHNG, PCPJ).	en	Jałoszyński, Paweł (2011): Ten new species of Cephennula Jałoszyński (Coleoptera, Staphylinidae, Scydmaeninae) from Malaysia. Zootaxa 3113: 36-52, DOI: 10.5281/zenodo.279337
03DF87B01F538E0986C5FB5026F5FAA6.taxon	diagnosis	Diagnosis. Body length ca. 0.8 mm, not reaching 0.9 mm; antennal club strongly flattened and loosely assembled; head unevenly covered with large and deep punctures, dense on sides of vertex and frons; median part of vertex and frons impunctate; median part of pronotum with large, deep and dense punctures; sublateral pronotal carinae very fine and obliterated in anterior 1 / 3; subhumeral carinae about as long as 1 / 5 x EL, slightly shorter than humeral carinae; males with dense subapical setal field on each elytron; aedeagus in ventral view bottle-shaped with nearly straight apical margin, endophallus very lightly pigmented except of distinct pair of apical lobes slightly curved mesally.	en	Jałoszyński, Paweł (2011): Ten new species of Cephennula Jałoszyński (Coleoptera, Staphylinidae, Scydmaeninae) from Malaysia. Zootaxa 3113: 36-52, DOI: 10.5281/zenodo.279337
03DF87B01F538E0986C5FB5026F5FAA6.taxon	description	Description. Body of male (Fig. 9) moderately convex, elongate, with barely marked constriction between pronotum and elytra, moderately glossy, light brown to yellowish-brown, covered with yellowish vestiture, legs and antennae slightly lighter. BL 0.79 – 0.83 mm (mean 0.81 mm). Head broadest at large, coarsely faceted and strongly convex eyes, HL 0.09 – 0.10 mm (mean 0.09 mm), HW 0.19 – 0.20 mm (mean 0.19 mm); vertex and frontoclypeal area convex; supraantennal tubercles barely marked. Punctures on dorsum of head very large, deep and sharply defined, those on sides of vertex and frons dense, narrow median area on vertex and frons impunctate; setae short and sparse, suberect. Antennae with strongly enlarged, strongly flattened, loosely assembled and distinctly 3 - segmented club, AnL 0.33 mm; antennomere I about 1.4 x as long as broad; II 2.2 x as long as broad; III slightly broader than long; IV – VII subequal in length and width, each about 1.4 x as long as broad; VIII as broad as VII but distinctly shorter, 2.2 x as long as broad; IX much broader and longer than VIII, slightly transverse; X much broader and longer than IX, strongly transverse; XI nearly as broad as X, 1.6 x as long as broad. Pronotum approximately trapezoidal, broadest near anterior 1 / 3; PL 0.23 – 0.25 mm (mean 0.24 mm), PW 0.33 – 0.34 mm (mean 0.33 mm), anterior margin broadly rounded; lateral margins distinctly microserrate, strongly rounded in anterior 1 / 3, distinctly concave in posterior 1 / 3 and nearly parallel near nearly straight but acute hind angles; posterior margin very weakly bisinuate; sublateral carinae very fine and obliterated in anterior 1 / 3. Transverse ante-basal groove shallow but sharply marked and nearly straight; sublateral and lateral ante-basal foveae equally small and relatively deep. Punctures on median part of pronotal disc similar in diameter and depth to those on sides of frons, dense; punctures between sublateral carinae and lateral margins of pronotum distinctly larger and denser than those in middle, nearly adjacent to each other, but not coarse; setae sparse, moderately long, suberect. Elytra moderately convex, oval, broadest near anterior 1 / 3; EL 0.46 – 0.48 mm (mean 0.47 mm), EW 0.38 – 0.39 mm (mean 0.38 mm), EI 1.23 – 1.27 (mean 1.24); base of each elytron with very small but distinct basal fovea located nearly in middle between scutellum and base of humeral carina; subhumeral carinae very fine, about as long as 1 / 5 x EL; humeral carinae slightly more distinct and slightly longer. Punctures on elytra smaller and shallower than those on head and pronotum, but relatively distinct and dense, especially on circumsutural area in anterior half; setae similar as those on pronotum, except for small subapical field on each elytron composed of dense and nearly recumbent setae directed towards suture. Legs moderately long and slender; pro- and mesotibiae nearly straight, metatibiae slightly recurved. Aedeagus (Figs. 25, 26) in ventral view bottle-shaped with broad, subtrapezoidal apex and nearly straight apical margin; AeL 0.18 mm. Endophallus lightly pigmented, with two broad and rounded elongate apical lobes curved mesally; parameres slender and very short, not exceeding apex of median lobe, each bearing single apical seta. Female. Externally differs from male in lacking subapical setal field on each elytron and slightly more slender body. BL 0.85 mm; HL 0.10 mm, HW 0.18 mm, AnL 0.33 mm; PL 0.25 mm, PW 0.34 mm; EL 0.50 mm, EW 0.39 mm, EI 1.29.	en	Jałoszyński, Paweł (2011): Ten new species of Cephennula Jałoszyński (Coleoptera, Staphylinidae, Scydmaeninae) from Malaysia. Zootaxa 3113: 36-52, DOI: 10.5281/zenodo.279337
03DF87B01F538E0986C5FB5026F5FAA6.taxon	distribution	Distribution. West Malaysia (Pahang).	en	Jałoszyński, Paweł (2011): Ten new species of Cephennula Jałoszyński (Coleoptera, Staphylinidae, Scydmaeninae) from Malaysia. Zootaxa 3113: 36-52, DOI: 10.5281/zenodo.279337
03DF87B01F538E0986C5FB5026F5FAA6.taxon	etymology	Etymology. The specific epithet “ medioglabra ” refers to the impunctate (Latin glaber, " smooth ") median part of the vertex and frons.	en	Jałoszyński, Paweł (2011): Ten new species of Cephennula Jałoszyński (Coleoptera, Staphylinidae, Scydmaeninae) from Malaysia. Zootaxa 3113: 36-52, DOI: 10.5281/zenodo.279337
03DF87B01F538E0986C5FB5026F5FAA6.taxon	discussion	Remarks. This is the only known species of Cephennula (and the Cephennomicrus group of genera) with modified elytra in males, bearing a dense subapical setal field. Similar modifications in Cephenniini were known previously only in some species of Cephennodes Reitter, 1884. Cephennula medioglabra has the aedeagus most similar to that of C. scaphisoma, also occurring in Pahang (Jałoszyński 2008), but details of the endophallus clearly differ in the latter species.	en	Jałoszyński, Paweł (2011): Ten new species of Cephennula Jałoszyński (Coleoptera, Staphylinidae, Scydmaeninae) from Malaysia. Zootaxa 3113: 36-52, DOI: 10.5281/zenodo.279337
03DF87B01F528E0A86C5FA1B242AFB71.taxon	materials_examined	Holotype: E MALAYSIA: 3, two labels: " Borneo Sabah \ Sepilok Sibuga Forest Res. \ nr. Sandakan 9. vi. 1968 \ 68.378 \ R. W. Tazlor " [white, partly printed, “ Sepilok ” crossed out, “ Sibuga ” and numbers handwritten in black ink]; " CEPHENNULA \ sibugana m. \ det. P. Jałoszyński, 2011 \ HOLOTYPUS " [red, printed] (MHNG).	en	Jałoszyński, Paweł (2011): Ten new species of Cephennula Jałoszyński (Coleoptera, Staphylinidae, Scydmaeninae) from Malaysia. Zootaxa 3113: 36-52, DOI: 10.5281/zenodo.279337
03DF87B01F528E0A86C5FA1B242AFB71.taxon	diagnosis	Diagnosis. Body length between 0.7 and 0.8 mm; antennal club strongly flattened and compact; head nearly evenly covered with large, deep and dense punctures; punctures on median part of pronotum moderately large, dense; sublateral pronotal carinae as long as pronotum; ante-basal pronotal groove with distinct median fovea; subhumeral carinae nearly as long as 1 / 2 x EL, humeral carinae less distinct and about half shorter; aedeagus in ventral view bottle-shaped with subtrapezoidal and rounded apex; endophallus composed of lightly pigmented elongate tubular structure.	en	Jałoszyński, Paweł (2011): Ten new species of Cephennula Jałoszyński (Coleoptera, Staphylinidae, Scydmaeninae) from Malaysia. Zootaxa 3113: 36-52, DOI: 10.5281/zenodo.279337
03DF87B01F528E0A86C5FA1B242AFB71.taxon	description	Description. Body of male (Fig. 10) strongly convex, elongate, with shallow but distinct constriction between pronotum and elytra, moderately glossy, dark brown, covered with yellowish vestiture, legs and antennae slightly lighter. BL 0.75 mm. Head broadest at large, coarsely faceted and strongly convex eyes, HL 0.10 mm, HW 0.19 mm; vertex and frontoclypeal area convex; supraantennal tubercles barely marked. Punctures on head dorsum nearly equally distributed, large, deep and dense; setae very short and sparse, nearly recumbent. Antennae with strongly enlarged, strongly flattened and compactly assembled antennomeres X – XI and only slightly enlarged antennomere IX, so that club appears two-segmented, AnL 0.25 mm; antennomere I about 1.3 x as long as broad; II 2.2 x as long as broad; III and IV each about 1.1 x as long as broad; V 1.2 x as long as broad; VI 1.1 x as long as broad; VII 1.2 x as long as broad; VIII slightly broader but shorter than VII, as long as broad; IX distinctly broader and slightly longer than VII, distinctly transverse; X much broader and longer than IX, strongly transverse; XI about as broad as X but only 1.1 x as long as broad. Pronotum approximately semielliptical, broadest between middle and anterior 1 / 3; PL 0.25 mm, PW 0.31 mm, anterior margin weakly rounded; lateral margins distinctly microserrate, broadly rounded in anterior 2 / 3, straight and parallel in posterior 1 / 4; hind angles sharp and acute; posterior margin very weakly bisinuate; sublateral carinae distinct and as long as pronotum .. Transverse ante-basal groove shallow and indistinct, slightly arcuate and with distinct median foveae; sublateral ante-basal foveae small and indistinct; lateral ante-basal foveae larger and deeper. Punctures on median part of pronotal disc slightly smaller and shallower than those on head but distinct and dense, areas between sublateral carinae and lateral margins of pronotum covered with much larger and denser, coarse punctures; setae sparse, moderately long, only slightly suberect. Elytra strongly convex, oval, broadest between middle and anterior 1 / 3; EL 0.40 mm, EW 0.35 mm, EI 1.14; base of each elytron with rudimentary basal fovea located much closer to scutellum than to base of humeral carina; subhumeral carinae very distinct and nearly as long as 1 / 2 x EL, nearly twice as long as less distinct humeral carinae. Punctures on elytra very fine and very shallow, barely discernible under 100 x magnification; setae similar as those on pronotum. Legs moderately long and slender; pro- and mesotibiae nearly straight, metatibiae distinctly curved. Aedeagus (Figs. 27, 28) in ventral view bottle-shaped with moderately narrow, subtrapezoidal apex and strongly rounded apical margin; AeL 0.13 mm. Endophallus lightly pigmented, with median tubular structure; parameres slender, not exceeding apex of median lobe, each bearing two apical and one subapical seta. Female. Unknown.	en	Jałoszyński, Paweł (2011): Ten new species of Cephennula Jałoszyński (Coleoptera, Staphylinidae, Scydmaeninae) from Malaysia. Zootaxa 3113: 36-52, DOI: 10.5281/zenodo.279337
03DF87B01F528E0A86C5FA1B242AFB71.taxon	distribution	Distribution. East Malaysia (Borneo: Sabah).	en	Jałoszyński, Paweł (2011): Ten new species of Cephennula Jałoszyński (Coleoptera, Staphylinidae, Scydmaeninae) from Malaysia. Zootaxa 3113: 36-52, DOI: 10.5281/zenodo.279337
03DF87B01F528E0A86C5FA1B242AFB71.taxon	etymology	Etymology. Locotypical, after the Sibuga Forest in Borneo.	en	Jałoszyński, Paweł (2011): Ten new species of Cephennula Jałoszyński (Coleoptera, Staphylinidae, Scydmaeninae) from Malaysia. Zootaxa 3113: 36-52, DOI: 10.5281/zenodo.279337
03DF87B01F518E0B86C5FB232318FB15.taxon	materials_examined	Holotype: E MALAYSIA: 3, two labels: " Malaisie. Selangor \ Gombak. 29. VIII. 72 \ 13 miles Kuala Lumpur \ T. Jaccoud " [white, handwritten in black ink]; " CEPHENNULA \ gombakiana m. \ det. P. Jałoszyński, 2011 \ HOLO- TYPUS " [red, printed] (MHNG).	en	Jałoszyński, Paweł (2011): Ten new species of Cephennula Jałoszyński (Coleoptera, Staphylinidae, Scydmaeninae) from Malaysia. Zootaxa 3113: 36-52, DOI: 10.5281/zenodo.279337
03DF87B01F518E0B86C5FB232318FB15.taxon	diagnosis	Diagnosis. Body between 0.7 and 0.8 mm in length; antennal club strongly flattened and compact; head unevenly covered with large and deep punctures, vertex with very small median impunctate area, punctures on sides of vertex and frons dense, those on median part of frons and on clypeus sparser; median part of pronotum with small, shallow and dense punctures; sublateral pronotal carinae as long as pronotum; ante-basal transverse groove with median fovea; subhumeral carinae slightly longer than 1 / 3 x EL and much longer than humeral carinae; aedeagus in ventral view approximately bottle-shaped with slightly broadened apex and broadly arcuate apical margin; endophallus lightly pigmented, composed of median bell-shaped structure.	en	Jałoszyński, Paweł (2011): Ten new species of Cephennula Jałoszyński (Coleoptera, Staphylinidae, Scydmaeninae) from Malaysia. Zootaxa 3113: 36-52, DOI: 10.5281/zenodo.279337
03DF87B01F518E0B86C5FB232318FB15.taxon	description	Description. Body of male (Fig. 11) strongly convex, elongate, with barely marked constriction between pronotum and elytra, moderately glossy, light brown, covered with yellowish vestiture, legs and antennae slightly lighter. BL 0.76 mm. Head broadest at large, coarsely faceted and strongly convex eyes, HL 0.09 mm, HW 0.20 mm; vertex and frontoclypeal area convex; supraantennal tubercles barely marked. Punctures on dorsum of head unevenly distributed, large and deep, on sides of vertex and frons dense, median part of frons and clypeus with sparser punctures, small median area on vertex impunctate; setae short and sparse, suberect. Antennae with strongly enlarged, strongly flattened and compactly assembled antennomeres X – XI and moderately enlarged antennomere IX, so that club appears two-segmented, AnL 0.28 mm; antennomere I about 1.2 x as long as broad; II 2 x as long as broad; III about as long as broad; IV – VI subequal in length and width, each about 1.2 x as long as broad; VII slightly longer than VI, 1.3 x as long as broad; VIII distinctly shorter and hardly noticeably broader than VII, about as long as broad; IX distinctly broader and slightly longer than VIII, slightly transverse; X much longer and broader than IX, strongly transverse; XI slightly narrower than X, 1.4 x as long as broad. Pronotum approximately semielliptical, broadest slightly anterior to middle; PL 0.23 mm, PW 0.35 mm, anterior margin weakly rounded; lateral margins distinctly microserrate, strongly rounded in anterior half, nearly straight in posterior half and slightly convergent towards sharp and acute hind angles; posterior margin very weakly bisinuate; sublateral carinae as long as pronotum. Transverse ante-basal groove shallow but sharply marked and slightly arcuate, with small but distinct median fovea; sublateral ante-basal foveae small and shallow; lateral antebasal foveae distinctly deeper and larger. Punctures on median part of pronotum slightly smaller and shallower than those on head but distinct and dense, punctures between sublateral pronotal carinae and lateral margins of pronotum much larger and denser than those in middle, nearly adjacent to each other and coarse; setae sparse, moderately long, suberect. Elytra strongly convex, oval, broadest between middle and anterior 1 / 3; EL 0.45 mm, EW 0.38 mm, EI 1.20; base of each elytron with rudimentary basal fovea located slightly closer to scutellum than to base of humeral carina; subhumeral carinae distinct, slightly longer than 1 / 3 x EL; humeral carinae indistinct and very short, shorter than 1 / 5 x EL. Punctures on elytra much smaller than those on head and pronotum, very fine and shallow but relatively distinct, sparse; setae similar as those on pronotum but slightly less suberect. Legs moderately long and slender; protibiae nearly straight, meso- and metatibiae slightly curved. Aedeagus (Figs. 29, 30) in ventral view approximately bottle-shaped with broad, subrectangular and slightly broadened apex and broadly, regularly arcuate apical margin; AeL 0.10 mm. Endophallus lightly pigmented, with median bell-shaped structure; parameres slender, very short, each bearing single apical seta. Female. Unknown.	en	Jałoszyński, Paweł (2011): Ten new species of Cephennula Jałoszyński (Coleoptera, Staphylinidae, Scydmaeninae) from Malaysia. Zootaxa 3113: 36-52, DOI: 10.5281/zenodo.279337
03DF87B01F518E0B86C5FB232318FB15.taxon	distribution	Distribution. West Malaysia (Selangor).	en	Jałoszyński, Paweł (2011): Ten new species of Cephennula Jałoszyński (Coleoptera, Staphylinidae, Scydmaeninae) from Malaysia. Zootaxa 3113: 36-52, DOI: 10.5281/zenodo.279337
03DF87B01F518E0B86C5FB232318FB15.taxon	etymology	Etymology. Locotypical, after the district Gombak of the Selangor State of Malaysia.	en	Jałoszyński, Paweł (2011): Ten new species of Cephennula Jałoszyński (Coleoptera, Staphylinidae, Scydmaeninae) from Malaysia. Zootaxa 3113: 36-52, DOI: 10.5281/zenodo.279337
03DF87B01F518E0B86C5FB232318FB15.taxon	discussion	Remarks. The aedeagus of this species is very similar to that of C. secunda, also known to occur in Selangor (Jałoszyński 2008). Cephennula gombakiana differs in having an additional, median ante-basal fovea on the pronotum (in middle of the ante-basal groove) and the length of the humeral elytral carinae, which in this species are shorter than 1 / 5 x EL, while in C. secunda the humeral carinae are as long as 1 / 3 x EL.	en	Jałoszyński, Paweł (2011): Ten new species of Cephennula Jałoszyński (Coleoptera, Staphylinidae, Scydmaeninae) from Malaysia. Zootaxa 3113: 36-52, DOI: 10.5281/zenodo.279337
03DF87B01F508E1486C5FAC027A1FBAD.taxon	materials_examined	Holotype: W MALAYSIA: 3, two labels: " W. Malaysia: Pahang \ Batu caves N Kuala \ Lumpur, 31.3.1993 \ Löbl & Calame, nr 24 " [white, printed]; " CEPHENNULA \ micropunctata m. \ det. P. Jałoszyński, 2011 \ HOLOTY- PUS " [red, printed] (MHNG).	en	Jałoszyński, Paweł (2011): Ten new species of Cephennula Jałoszyński (Coleoptera, Staphylinidae, Scydmaeninae) from Malaysia. Zootaxa 3113: 36-52, DOI: 10.5281/zenodo.279337
03DF87B01F508E1486C5FAC027A1FBAD.taxon	diagnosis	Diagnosis. Body between 0.7 and 0.8 mm in length; antennal club strongly flattened and compact; head covered with small and unevenly distributed punctures, those on sides of frons and vertex separated by spaces 0.5 – 2 x as wide as puncture diameter, punctures on median part of frons sparser; median part of pronotum with small punctures separated by spaces 1 – 3 x as wide as puncture diameter; sublateral pronotal carinae as long as pronotum; antebasal pronotal groove with small median fovea; subhumeral carinae slightly longer than 1 / 3 x EL and much longer than humeral carinae; aedeagus in ventral view approximately bottle-shaped, very slender, with strongly broadened apex and rounded apical margin with distinct median point; endophallus indistinct, bell-shaped.	en	Jałoszyński, Paweł (2011): Ten new species of Cephennula Jałoszyński (Coleoptera, Staphylinidae, Scydmaeninae) from Malaysia. Zootaxa 3113: 36-52, DOI: 10.5281/zenodo.279337
03DF87B01F508E1486C5FAC027A1FBAD.taxon	description	Description. Body of male (Fig. 12) strongly convex, elongate, with shallow but distinct constriction between pronotum and elytra, moderately glossy, light brown, covered with yellowish vestiture, legs and antennae slightly lighter. BL 0.76 – 0.78 mm. Head broadest at large, coarsely faceted and strongly convex eyes, HL 0.08 mm, HW 0.20 mm; vertex and frontoclypeal area convex; supraantennal tubercles barely marked. Punctures on dorsum of head unevenly distributed, small but deep and distinct, on sides of vertex and frons dense, on median part of frons and clypeus distinctly sparser; setae short and sparse, suberect. Antennae with strongly enlarged, strongly flattened and compactly assembled antennomeres X – XI and moderately enlarged antennomere IX, so that club appears two-segmented, AnL 0.28 mm; antennomere I about 1.2 x as long as broad; II 2.2 x as long as broad; III 1.1 x as long as broad; IV – VIII subequal in length and width, each about 1.2 x as long as broad; IX slightly longer and distinctly broader than VIII, about as long as broad; X much longer and broader than IX, distinctly transverse; XI slightly narrower than X, 1.5 x as long as broad. Pronotum approximately semielliptical, broadest between middle and anterior 1 / 3; PL 0.25 mm, PW 0.34 mm, anterior margin weakly rounded; lateral margins distinctly microserrate, strongly rounded in anterior 1 / 3 and feebly rounded in posterior half, slightly convergent towards sharp and acute hind angles; posterior margin very weakly bisinuate; sublateral carinae as long as pronotum. Transverse ante-basal groove shallow but sharply marked and slightly arcuate, with shallow and small median fovea; sublateral ante-basal foveae small and shallow; lateral antebasal foveae slightly deeper and larger. Punctures on median part of pronotum slightly smaller than those on head, shallow but distinct, separated by spaces 1 – 3 x as wide as puncture diameter, punctures between sublateral pronotal carinae and lateral margin of pronotum distinctly larger and denser than those in middle, nearly adjacent to each other but not coarse; setae sparse, moderately long, suberect. Elytra strongly convex, oval, broadest in anterior 1 / 3; EL 0.44 – 0.45 mm, EW 0.35 – 0.38 mm, EI 1.20 – 1.25; base of each elytron with rudimentary basal fovea located slightly closer to scutellum than to base of humeral carina; subhumeral carinae distinct, slightly longer than 1 / 3 x EL; humeral carinae less distinct, only as long as about 1 / 6 x EL. Punctures on elytra much smaller than those on head and pronotum, fine and shallow but relatively distinct, sparse; setae similar as those on pronotum. Legs moderately long and slender; pro- and mesotibiae nearly straight, metatibiae slightly curved. Aedeagus (Figs. 31, 32) in ventral view approximately bottle-shaped with broad, subtrapezoidal apex strongly expanded laterally and with rounded apical margin with clearly marked median point; AeL 0.18 mm. Endophallus lightly pigmented, with median bell-shaped structure; parameres slender and very short, each bearing two apical setae. Female. Unknown.	en	Jałoszyński, Paweł (2011): Ten new species of Cephennula Jałoszyński (Coleoptera, Staphylinidae, Scydmaeninae) from Malaysia. Zootaxa 3113: 36-52, DOI: 10.5281/zenodo.279337
03DF87B01F508E1486C5FAC027A1FBAD.taxon	distribution	Distribution. West Malaysia (Pahang).	en	Jałoszyński, Paweł (2011): Ten new species of Cephennula Jałoszyński (Coleoptera, Staphylinidae, Scydmaeninae) from Malaysia. Zootaxa 3113: 36-52, DOI: 10.5281/zenodo.279337
03DF87B01F508E1486C5FAC027A1FBAD.taxon	etymology	Etymology. The specific epithet “ micropunctata ” refers to small punctures on the head and pronotum of this species.	en	Jałoszyński, Paweł (2011): Ten new species of Cephennula Jałoszyński (Coleoptera, Staphylinidae, Scydmaeninae) from Malaysia. Zootaxa 3113: 36-52, DOI: 10.5281/zenodo.279337
