identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03DE082705458A03FF12FB1CFE2CFAB2.text	03DE082705458A03FF12FB1CFE2CFAB2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cephalomalthinus Pic 1921	<div><p>Genus Cephalomalthinus Pic, 1921</p><p>Type species. Cephalomalthinus ocularis Pic, 1921 .</p><p>Diagnosis (see Li et al. 2016).</p></div>	https://treatment.plazi.org/id/03DE082705458A03FF12FB1CFE2CFAB2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hsiao, Yun;Tsai, Cheng-Lung	Hsiao, Yun, Tsai, Cheng-Lung (2019): Cephalomalthinus simplicicornis (Wittmer, 1993) rev. stat. et comb. n.: a resurrected soldier beetle (Coleoptera, Cantharidae) from Taiwan based on morphological and molecular data. Zootaxa 4619 (2): 297-310, DOI: 10.11646/zootaxa.4619.2.6
03DE082705458A04FF12FA06FD6CFAFE.text	03DE082705458A04FF12FA06FD6CFAFE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cephalomalthinus formosanus (Pic 1910)	<div><p>Cephalomalthinus formosanus (Pic, 1910)</p><p>(Figs 2–4, 11–12, 15–16, 21–23, 29–30)</p><p>Rhagonycha formosana Pic, 1910: 271 .</p><p>Podabrus curvatipes Pic, 1922: 4 . [Synonymized by Wittmer, 1986: 131.]</p><p>Rhagonycha obscurioripes Pic, 1926: 235 . [Synonymized by Wittmer, 1982: 129.]</p><p>Micropodabrus formosanus: Wittmer, 1982: 129 .</p><p>Fissocantharis formosana: Yang et al. 2012: 47 .</p><p>Type material examined. Type ♂ of Rhagonycha formosana Pic, 1910 (MNHN): [handwritten] Fainan [= Tainan] / Formosa IV // type // [printed] TYPE // [h] Rhagonycha / formosana / Pic // 66 F // Micropodabrus / formosanus / (Pic) / [p] det. W. Wittmer; Type ♂ of Podabrus curvatipes Pic, 1922 (MNHN): [h] Fainan [= Tainan] / Formosa IV // type // [printed] TYPE // [h] Podabrus / curvatipes / Pic // Micropodabrus / formosanus / (Pic) / [p] det. W. Wittmer; Holotype ♀ of Rhagonycha obscurioripes Pic, 1926 (MNHN): [p] Formosa // [h] R. obscurioripes / Pic // Rhagonycha / obscurioripes / n. sp. // [p] HOLOTYPUS // [h] Micropodabrus / formosanus / (Pic) / [p] det. W. Wittmer.</p><p>Other material examined. TAIWAN: 2 ♂, 3 ♀ (NMNS), Kukuan, Taichung, 11–12.iv.1986, C.-S. Lin leg. // NMNS ENT 64-246; NMNS ENT 64-247; NMNS ENT 64-255; NMNS ENT 64-256; NMNS ENT 64-257; 2 ♂, 1 ♀ (NMNS), Chingshan, Tainan, 18.iv.1992, Sweeping Net, C.-Y. Li leg. // NMNS ENT 1446-3328; NMNS ENT 1446-3415; NMNS ENT 1446-3456; 1 ♂ (NMNS), Hoawanta, Jenai, Nantou, 30.iv.1992, Sweeping Net, W.-T.Yang leg. // 1448-131 (NMNS); 1 ♂ (NMNS), Shihtyutou, Nantou, 2.viii.1994, C.-C. Lo leg. // NMNS ENT 4463-2097; 1 ♂ (NMNS), Tona, Maolin, Kaohsiung, 29–31.iii.2005, UV Light trap, W.-T. Yang leg. // NMNS ENT 4880-175; 1 ♂ (NMNS), Lienhuachih, Nantou, 11.iv.–2.v.2005, Malaise trap (KCN), C.-S. Lin &amp; W.-T. Yang leg. // NMNS ENT 6817-769; 2 ♂ (NMNS), Panchiu Station, Peinan, Taitung, 4.i.–9.ii.2006, Malaise trap (KCN), C.-S. Lin &amp; W.-T. Yang leg. // NMNS ENT 5884-486; NMNS ENT 5884-487; 2 ♂, 2 ♀ (NMNS), Panchiu Station, Peinan, Taitung, 8.ii.–14.iii.2007, Malaise trap (KCN), C.-S. Lin &amp; W.-T. Yang leg. // NMNS ENT 7130-459; NMNS ENT 7130-460; NMNS ENT 7130-461; NMNS ENT 7130-462; 1 ♂, 1 ♀ (YHC), Wushe, Nantou, 31.iii.2012, C.-H. Wu leg. ; 2 ♀ (YHC), Nanshanshi, Nantou, 1.iv.2012, C.-H. Wu leg. ; 1 ♂ (YHC), Fuli ( Nihoushan), Hualien, 1.i.2014, W.-C. Huang leg. ; 1 ♀ (YHC), Jenai, Nantou, 1.iii.2014, C.-H. Wu leg. ; 1 ♂ (YHC), Majia, Pingtung, 12.iii.2014, Y.-T. Chung leg. ; 4 ♂, 1 ♀ (3 ♂ in NCHU; 1 ♂, 1 ♀ in YHC), Kantoushan, Tainan, 29.iii.2014, W.-C. Liao leg. ; 4 ♂, 6 ♀ (2 ♂, 1 ♀ in NCHU; 2 ♂, 5 ♀ in YHC), Huisun Forest Field, Nantou, 4.iv.2014, Y. Hsiao leg. ; 1 ♂ (YHC), Chung Cheng Camp Zone, Taichung, 11.iv.2014, W.-R. Liang leg. ; 1 ♂, 2 ♀ (YHC), Shanping Park, Kaohsiung, 12.iv.2014, W.-C. Liao leg. ; 1 ♂, 2 ♀ (YHC), Meling, Nanshi, Nantou, 19.iv.2014, W.-C. Liao leg. ; 11 ♂, 16 ♀ (4 ♂, 10 ♀ in NCHU; 7 ♂, 6 ♀ in YHC), Chunyang, Nantou, 4.v.2014, Y. Hsiao leg. ; 1 ♀ (YHC), Shupishan, Chiayi, 26.iv.2015, collector unknown ; 1 ♂, 3 ♀ (NCHU), Malunshan, Taichung, 25.iv.2016, Y. Li leg.</p><p>Supplementary description. Male genitalia: inner sac enlarged, with basal part as wide as median lobe, slightly shorter than tegmen, elongated ventro-posteriorly, obtuse apically; ventral surface with rounded, swollen process basally and paired clusters of acuminate setae in apical one-third; dorsal surface with rounded, swollen process basally and horn-like process apically, elongated dorso-anteriorly (Figs 21–23).</p><p>Diagnosis. This species is closely similar to C. pingtungensis (Wittmer, 1982) and C. simplicicornis (Wittmer, 1993) rev. stat. et comb. n. in the color pattern and filiform antennae. It can be distinguished from C. pingtungensis by the simple and unmodified male antennae (antennomeres III–VII with small rounded excavation medially in C. pingtungensis) and conjoint dorsal plate of aedeagal parameres 0.5 times longer than ventral process of each paramere (conjoint dorsal plate extremely short in C. pingtungensis). From C. simplicicornis, it differs in the large, prominent eyes (Figs. 11–12; see C. simplicicornis in Figs. 13–14), wider conjoint dorsal plate and indistinct, undeveloped dorsal processes of median lobe (Figs. 15–16; see C. simplicicornis in Figs. 17–20).</p><p>Remarks. In addition to the distinct color variation noted by Yang et al. (2012), the diverticulum of female genitalia varies among different specimens, with relative length ca. 1.7–1.5X longer than spermathecal duct (Figs. 29–30).</p><p>Distribution. Taiwan (presumed endemic).</p></div>	https://treatment.plazi.org/id/03DE082705458A04FF12FA06FD6CFAFE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hsiao, Yun;Tsai, Cheng-Lung	Hsiao, Yun, Tsai, Cheng-Lung (2019): Cephalomalthinus simplicicornis (Wittmer, 1993) rev. stat. et comb. n.: a resurrected soldier beetle (Coleoptera, Cantharidae) from Taiwan based on morphological and molecular data. Zootaxa 4619 (2): 297-310, DOI: 10.11646/zootaxa.4619.2.6
03DE082705428A07FF12FACAFD6CF82B.text	03DE082705428A07FF12FACAFD6CF82B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cephalomalthinus simplicicornis (Wittmer 1993) Hsiao & Tsai 2019	<div><p>Cephalomalthinus simplicicornis (Wittmer, 1993) rev. stat. et comb. n.</p><p>(Figs. 5–10, 13–14, 17–20, 24–28, 31–32)</p><p>Micropodabrus simplicicornis Wittmer, 1993: 209 .</p><p>Fissocantharis formosana (Pic, 1910); Yang et al. 2012: 47.</p><p>Type material examined. TAIWAN: Holotype ♂ of Micropodabrus simplicicornis Wittmer, 1993 (NHMB): [p] 12-IV-1981 / T. Shimomura leg. // Kao-i, alt. 600m / Taoyuan Pref. / TAIWAN // [h] M. / simplicicornis / Wittmer / [p] det. W. Wittmer. // HOLOTYPUS // CANTHARIDAE / CANTH00000275.</p><p>Other material examined. TAIWAN: 1 ♂ (NMNS), Junghua [in Fushing, Taoyuan], 11.iv.1971, B.-S. Chang leg. // 1282-26814; 1 ♀ (NMNS), Shangbaling, Taoyuan, 18.iv.1990, Sweep &amp; Searching, C.-C. Chiang leg. // NMNS ENT 598-26; 1 ♂ (NMNS), Hotzuhu, Taoyuan, 30.v.1991, Sweeping Net, I.-S. Hsu leg. // NMNS ENT 1636-494; 1 ♀ (NMNS), Taman, Fushing, Taoyuan, 30.iii.1992, Sweeping Net, I.-S. Hsu leg. // NMNS ENT 1638- 43; 1 ♂, 1 ♀ (YHC), Chishanyen, Taipei, 10.iv.2014, S.-F. Yu leg. // In Copulation; 2 ♀ (YHC), Chishanyen, Taipei, 10.iv.2014, S.-F. Yu leg. ; 1 ♀ (YHC), Baling, Fushing, Taoyuan, 9.v.2014, L. Huang leg. ; 1 ♂ (YHC), Chishanyen, Taipei, 9.iv.2015, S.-F. Yu leg. ; 1 ♂ (YHC), Academia Sinica, Nankang, Taipei, 15.iv.2015, L. Huang leg. ; 1 ♂ (YHC), Jingshan-banling, New Taipei, 24.iv.2015, I.-L. Lee leg. ; 2 ♀ (YHC), Chishanyen, Taipei, 8.v.2015, L. Huang leg. ; 1 ♂ (YHC), Fushan Botanical Garden, Ilan, 4.vi.2015, T. Hsieh leg. ; 1 ♂ (YHC), Tai-Jiu-Jia-gonlu 16.5K [= Xinwu Highway, New Taipei], 19.iv.2016, T. Hsieh leg. ; 1 ♂ (YHC), Lalashan, Taoyuan, 27.iv.2016 , L.</p><p>Huang leg.; 1 ♂ (YHC), Baehen-gonlu [=Northern Cross-Island Highway], Fushing, Taoyuan, 23.iv.2017 , L. Huang leg.; 1 ♂ (YHC), Shapaling, Taoyuan, 30.iv.2017 , L. Huang leg.; 10 ♂, 7 ♀ (2 ♂, 2 ♀ in NCHU; 8 ♂, 5 ♀ in YHC), Shienchiyen, Taipei, 10.iv.2018 , Y. Hsiao leg.</p><p>Redescription. Male (Figs. 5–7). Body length: 7.00– 9.50 mm (holotype: 8.00); width: 1.25–2.00 mm (1.80).</p><p>Coloration. Eyes black. Head entirely orange or orange, with vertex black. Antennae entirely black, or orange, with last four to six antennomeres slightly darkened or blackish brown. Mandibles orange, with dark apices. Prothorax orange. Elytra black. Scutellum brownish orange or black. Legs entirely black or orange, or orange, with apical part of femura, tibiae and tarsi slightly or distinctly blackish. Mesoventrites, metaventrites and abdomen black. Body covered with pale pubescence; anterior margin of clypeus, antennae, elytra and legs fringed with bristles.</p><p>Head subquadrate, with vertex moderately (Fig. 13) or distinctly narrowed basally (Fig. 14), nearly as long as wide, vertex flat on surface; surface densely, finely punctate and semilustrous; anterior margin of clypeus arcuate; eyes small, globular and slightly prominent, ratio of eye diameter to interocular space 1: 3.2–4.4 (holotype: 3.4); terminal labial palpomeres rounded ax-shaped; terminal maxillary palpomeres subtriangular, widest medially, apical parts of inner margins acurate, acute at apices; antennae filiform, simple, exceeding elytral midlength; scape, pedicel thick, gradually widened apically, pedicel shortest, antennomeres III–XI subcylindrical, ratio of antennomere lengths: 22.0–32.0 (24.0):10.0:21.0–28.0 (25.0):22.0–30.0 (24.0):25.0–34.0 (27.0):25.0–35.0 (30.0):25.0–36.0 (32.0):25.0–35.0 (33.0):25.0–34.0 (32.0):24.5–32.0 (31.0):28.0–42.0 (36.0).</p><p>Pronotum subquadrate, PW/HW: 0.84–0.91 (0.87), PW/PL: 0.86–0.98 (0.90); anterior and posterior margins moderately arcuate; lateral margins slightly narrowed medially, slightly diverging posteriorly, slightly constricted before posterior angles; anterior angles rounded; posterior angles obtuse; dorsum convex in postero-lateral and hollowed in antero-lateral areas; surface densely, finely punctate and semilustrous. Scutellum triangular, with obtuse apex.</p><p>Elytra elongate, slender, subparallel-sided, EW/PW: 1.37–1.65 (1.50), EL/EW: 2.76–3.27 (3.20); surface densely and rugosely punctate and semilustrous.</p><p>Legs slender; femora mostly straight; tibiae mostly straight, with basal part feebly arcuate; claws bifid.</p><p>Aedeagus elongate (Figs. 17–20); ventral process of each paramere wide, slightly narrowed apically, slightly hooked at apex, curved inwards; conjoint dorsal plate of parameres well-developed medially, forming median lobe, with apical margin subtruncate or slightly rounded and lateral margins fringed with bristles, 0.5 times as long as or slightly shorter than ventral process, distinctly narrower than the width between ventral processes, median lobe somewhat varies in shape (Fig. 20); laterophysis absent; paired angular processes adhered to dorsal side of median lobe apically, protruding from both sides of dorsal plate, distinctly visible from dorsal view (Figs. 17–18). Inner sac enlarged, with basal part as wide as median lobe, slightly shorter than tegmen, elongated posteriorly, swollen apically; ventral surface with rounded, swollen process basally and clustered acuminate setae in apically; dorsal surface with rounded, swollen process basally and horn-like process apically, elongated dorso-anteriorly (Figs. 24–26).</p><p>Female (Figs. 8–9). Body length: 7.00– 8.50 mm; width: 1.25–2.00 mm.</p><p>Very similar to male. Similar to male in coloration. Body wider than male. Eyes smaller than males, ratio of eye diameter to interocular space 1:4.50–4.90. Antennae somewhat shorter than in male, barely extending to elytral midlength, ratio of antennomere lengths: 20.0–22.0:10.0:16.0–20.0:20.0–24.0:20.0–26.0:22.0–25.0:21.0– 26.0:22.0–24.0:21.0–24.0:19.0–24.0:23.0–30.0. PW/HW: 0.93–0.94, PW/PL: 0.91–1.00. EW/PW: 1.50–1.73, EL/ EW: 2.54–2.85. Claws bifid. Abdominal ventrite VII with margins densely pubescent and apical margin nearly straight, or slightly protuberant medially, protuberance triangularly emarginate medially, forming paired median lobes, median lobes rounded apically (Figs. 27–28).</p><p>Female genitalia with vagina stout, abruptly thinned and extended into a slightly long and thick duct ventroapically; diverticulum and spermathecal duct arising from apex of long duct of vagina; diverticulum thin, spiral, distinctly longer than spermathecal duct; spermatheca provided with thin, spiral tube, with length slightly longer than diverticulum; accessory gland thin, slightly shorter than spermatheca; median oviduct situated in middle of vagina (Figs. 31–32).</p><p>Diagnosis. This species closely resembles C. formosanus (Pic, 1910) in color pattern and filiform, simple male antennae and conjoint dorsal plate 0.5 times longer than ventral process. It can be separated from C. formosanus by the smaller, less prominent eyes (Figs. 13–14; see C. formosanus in Figs. 11–12), narrower conjoint dorsal plate and distinctly developed dorsal processes of median lobe, which is clearly visible from dorsal view (Figs. 17–20; see C. formosanus in Figs. 15–16).</p><p>Biological notes. According to the label data, this species is known in northwestern Taiwan. Individuals were observed resting on vegetation during the day.</p><p>Distribution. Taiwan (presumed endemic).</p></div>	https://treatment.plazi.org/id/03DE082705428A07FF12FACAFD6CF82B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hsiao, Yun;Tsai, Cheng-Lung	Hsiao, Yun, Tsai, Cheng-Lung (2019): Cephalomalthinus simplicicornis (Wittmer, 1993) rev. stat. et comb. n.: a resurrected soldier beetle (Coleoptera, Cantharidae) from Taiwan based on morphological and molecular data. Zootaxa 4619 (2): 297-310, DOI: 10.11646/zootaxa.4619.2.6
