taxonID	type	description	language	source
03DE87FCFFE33531FF51FD2CFC84BCCC.taxon	discussion	REMARK The monophyly of the Phalangopsidae cricket family was attested by the molecular phylogenetic analyses of Chintauan-Marquier et al. (2013, 2016), provided it includes part of the Gryllomorphinae crickets (previously classified within the Gryllidae Laicharting, 1781), and excludes the Pteroplistinae Chopard, 1936, which relationships within Grylloidea remain unclear. The resultant topology implied however that the subfamilies usually considered within the Phalangopsidae, at the exception of the Phaloriinae (sensu Desutter-Grandcolas 2015, i. e. including Megacris Desutter-Grandcolas, 2012) which were always recovered monophyletic, were to be critically reviewed: as a consequence, the morphological characters that were traditionaly used to define those subfamilies have to be reevaluated. Phalangopsidae is the most diverse family within Grylloidea and the study of Chintauan-Marquier et al. (2013, 2016), although based on 65 terminals, took into account only a fraction of the phalangopsid clade: for this reason, no classification was derived from the topology proposed by Chintauan-Marquier et al. (2013, 2016). Instead, additional terminals and additional markers were sequenced and are presently under study (see Warren et al. 2019 for preliminary results and discussion). In the present paper, the genera are classified within the subfamilies listed in Cigliano et al. (2020).	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFE33531FC1CFE4CFC46BEEA.taxon	description	(Fig. 2 A-D; Table 1)	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFE33531FC1CFE4CFC46BEEA.taxon	materials_examined	TYPE LOCALITY. — Amérique méridionale [Southern America] after Serville (1831), French Guiana and Suriname after Saussure (1878). TYPE MATERIAL. — Syntype. ‘ Amérique méridionale’ • 1 ♂; MNHN-EO-ENSIF 4009.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFE33531FC1CFE4CFC46BEEA.taxon	discussion	REMARK Serville (1831) described this species on one male and one female, which he considered immature (but see Saussure 1874, 1878). The male type is correctly identified in the MNHN collection. One additional female is identified Phalangopsis longipes Serv. in Serville’s handwriting, but it has posterior locality labels (Guyane française, Saint Laurent du Maroni, Collection Le Moult / septembre). It may have been labelled in a second time by error, as three other females have the same origin. This female may represent the female syntype observed by Serville (MNHN-EO-ENSIF 6828).	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFE33531FC1CFE4CFC46BEEA.taxon	materials_examined	MATERIAL FROM THE MITARAKA. — French Guiana • 1 ♀; Monts Tumuc-Humac, Massif du Mitaraka, layon A; entre 54.4509 O 2.2357 N et 54.4557 O 2.2405 N; alt. entre 280 m et 365 m; 23. II. - 10. III. 2015; F. Legendre & S. Hugel leg.; fn. SH 116, de nuit; MNHN • 2 ♀; Monts Tumuc-Humac, Massif du Mitaraka, GF DZ; 54.450137 O 2.233883 N; alt. 315 m; 23. II. - 10. III. 2015; F. Legendre & S. Hugel leg.; fn. SH 179, SH 180, de nuit; MNHN • 1 ♂; Monts Tumuc-Humac, Massif du Mitaraka, GF DZ; 54.450137 O 2.233883 N; alt. 315 m; 23. II. - 10. III. 2015; F. Legendre & S. Hugel leg.; fn. SH 178, de nuit; MNHN.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFE33531FC1CFE4CFC46BEEA.taxon	description	MEASUREMENTS (IN MM). — See Table 1.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFE33531FC1CFE4CFC46BEEA.taxon	discussion	REMARK The length of the ovipositor of the females collected in Mitaraka is variable (see Table 1), but extends the values given earlier for the species (Desutter-Grandcolas 1992 a, 13.6 - 16.6 mm, mean 14.8, n = 9). The copulatory papilla is however very similar to that reported for that species in the same paper. The number of tergites with yellow bands on posterior edge also varies in the material at hand, until the third tergite for the specimen SH 116 (Fig. 1 A). Male genitalia are illustrated on Fig. 2 B-D.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFE33531FEA0F9C8FD9DBEE6.taxon	description	(Fig. 2)	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFE33531FEA0F9C8FD9DBEE6.taxon	type_taxon	TYPE SPECIES. — Phalangopsis longipes Serville, 1831 by subsequent designation (Kirby 1906).	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFE33531FEA0F9C8FD9DBEE6.taxon	distribution	DISTRIBUTION. — Northern tropical regions of South America (French Guiana, Brazil, Suriname).	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFE33531FEA0F9C8FD9DBEE6.taxon	diagnosis	DIAGNOSIS. — See Desutter-Grandcolas (1992 a).	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFE33531FEA0F9C8FD9DBEE6.taxon	biology_ecology	HABITAT. — Phalangopsis species live in forests, where they are active at night in the leaf litter and hide during the day in cavities at ground level (Desutter-Grandcolas 1995).	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFE23530FF12FE0CFF76BCAC.taxon	description	(Fig. 2 E; Table 2)	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFE23530FF12FE0CFF76BCAC.taxon	materials_examined	TYPE LOCALITY. — French Guiana, Arataye, afflt. Approuague, 8 km NE du saut Pararé, station des Nouragues. TYPE MATERIAL. — Holotype by original designation. French Guiana • 1 ♂; Arataye, afflt. Approuague, 8 km NE du saut Pararé, station des Nouragues; 4. IV. 1988; L. Desutter leg; MNHN-EO-ENSIF 4010. Allotype. French Guiana • 1 ♀; same locality and collector as the holotype; 11. VI. 1988; MNHN-EO-ENSIF 4011. Paratypes: 7 females. French Guiana • 2 ♀; same locality and collector as the holotype; VI. 1988; MNHN-EO-ENSIF 5541, 5542 • 2 ♀; Saül; VIII. 1988; L. Desutter & P. Grandcolas leg.; MNHN-EO-ENSIF 5543, 5544 • 1 ♀; Sinnamary, Paracou, forêt sur sable blanc; IX. 1988; L. Desutter & P. Grandcolas leg.; MNHN-EO-ENSIF 5545 • 2 ♀; Sinnamary, piste de St Elie, PK 15; VII. 1989; P. Grandcolas leg.; MNHN-EO-ENSIF 5546, 5547. MATERIAL FROM THE MITARAKA. — French Guiana • 1 ♀; Monts Tumuc-Humac, Massif du Mitaraka, layon A; 54.451125 O 2.234786 N; alt. 300 m; 23. II. - 10. III. 2015; F. Legendre & S. Hugel leg.; fn. SH 075, de nuit; MNHN.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFE23530FF12FE0CFF76BCAC.taxon	description	MEASUREMENTS (IN MM). — See Table 2.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFE23530FF12FE0CFF76BCAC.taxon	discussion	REMARKS The length of the ovipositor of the Mitaraka female of P. flavilongipes (see Table 2) extends the range previously reported for the species (Desutter-Grandcolas 1992 a, 17.5 – 18 mm, mean value 18 mm, n = 3). The copulatory papilla is similar to that illustrated for the species (Desutter-Grandcolas 1992 a, figure 48).	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFE2353EFF61FA49FF3CBEED.taxon	description	(Figs 3 - 5; Table 3) urn: lsid: zoobank. org: act: 187 D 9 FF 1 - 9 DC 8 - 42 E 0 - BCE 5 - FD 0 BD 7 D 8 D 4 BF	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFE2353EFF61FA49FF3CBEED.taxon	type_taxon	TYPE SPECIES. — Aracopsis hugeli Desutter-Grandcolas, n. sp. by original designation.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFE2353EFF61FA49FF3CBEED.taxon	diagnosis	DIAGNOSIS. — Size small. Mesonotum, metanotum, and tergite 1 well-developed, tergites 2 to 9 short and compressed. Head narrow in side view, very long in front view; eyes small but highly protruding; fastigium slanting from the vertex, with a faint basal transverse furrow; maxillary palpi elongate, article 5 much longer than article 4 and hardly widened in apical third. TI with an inner tympanum; outer tympanum absent. TIII with 3 / 3 apical spurs, the dorsal the longest on each side; 4 / 4 subapical spurs, small and alternate; serrulation small. Male: FWs small, overlapping, rounded distally, with a stridulatory file; tergites 3 to 6 with paired glandular structures. Male genitalia very small, compact; pseudepiphallic sclerite transverse, flat; apical lophi located on a narrow membranous projection between pseudepiphallic sclerite and ectophallic fold; pseudepiphallic arm small with simple and acute apex; ectophallic arc very wide, its width greater than ectophallic apodeme length; basal part of ectophallic apodemes greatly enlarged; endophallic sclerite long and narrow, with impaired lamella-shaped apodeme and a crest-like longitudinal apodeme. No phallic glands associated with pseudepiphallus. Female: apterous. Female genitalia: copulatory papilla elongate, conical, sclerotized. By its external morphology, Aracopsis Desutter-Grandcolas, n. gen. is close to Philippopsis and Phalangopsis, and superficially resembles Aracamby and related genera (Souza-Dias & Desutter-Grandcolas 2014); it can readily be separated from all these genera: – Similar to Philippopsis by its size, body shape, head shape and tibial spurs. Distinguished from Philippopsis by inner tympanum (lacking in Philippopsis), forewings and tergal glands in males (males with short flap-like forewings and without tergal glands in Philippopsis), TIII serrulation (sparse in Philippopsis) and male genitalia (arm long, narrow, curved, with bifid apex; ectophallic arc narrow, with two long evaginations, ectophallic apodemes short and high, in Philippopsis). – Similar to Phalangopsis by maxillary palpi, leg shape and TIII serrulation. Distinguished from Phalangopsis by small size, inner tympanum (lacking in Phalangopsis), shape of maxillary palpi last article (much thinner and longer in Phalangopsis), male FWs (not overlapping in Phalangopsis) and male genitalia (pseudepiphallic lophi much more developed, ectophallic arc narrow, ectophallic apodemes long, almost parallel and thick, in Phalangopsis). – Resembling Aracamby by small size and general body shape. Distinguished by tergites relative size, head shape (wider in Aracamby, with less protruding eyes), last article of maxillary palpi (article 5 strongly curved before mid length in Aracamby), TII apical spurs (four in Aracamby), TIII apical spurs (median spur the longest on inner and outer sides in Aracamby), tibial tympanum (variable in Aracamby), male FWs (long, covering most of abdomen, convex, with a full stridulum in Aracamby) and male genitalia (arm long, thick, almost straight, with bifid apex; ectophallic arc wide and narrow, ectophallic apodemes long and thin, in Aracamby). Females apterous or with pad-like forewings in all four genera; separated by the same morphological characters as males (size, head shape, tympanum, tibial spurs). Female copulatory papilla short and conical in Aracopsis Desutter-Grandcolas, n. gen. and Phalangopsis; elongate and basally membranous in Philippopsis and Aracamby.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFE2353EFF61FA49FF3CBEED.taxon	etymology	ETYMOLOGY. — Taxon named after its phylogenetic relationships to Philippopsis and Phalangopsis, and superficial resemblance to Aracamby.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFE2353EFF61FA49FF3CBEED.taxon	distribution	DISTRIBUTION. — French Guiana.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFE2353EFF61FA49FF3CBEED.taxon	description	DESCRIPTION General morphology Size small. Body (Fig. 3) convex, with well-developed pro-, meso-, metanota, and tergite 1; tergites 2 to 9 reduced and short. Head very long and narrow in side view. Fastigium slightly wider than half scape width, separated from vertex by a faint, transverse furrow (Fig. 4 A). Eyes (Fig. 4 A, B) small, strongly protruding. Ocelli small, set as a triangle; distance between lateral ocelli shorter than distance between median and one lateral ocelli; median ocellus not protruding. Maxillary palpi (Fig. 5 A) very elongate, especially articles 3 to 5; articles 3 to 5 very thin; article 3 slightly shorter than article 4; article 5 the longest, little but regularly widened from mid length toward the tip, truncate at apex. Legs setose, elongate and very thin. TI with a small inner tympanum; outer tympanum absent; two apical, ventral spurs. TII with two long, apical, ventral spurs; no dorsal spurs. TIII with 4 / 4 subapical spurs, short and alternate; 3 / 3 apical spurs (Fig. 5 B, C); inner apical spurs longer than outer spurs; dorsal apical spur the longest on both sides. FIII with a filiform apical part (Fig. 4 C, D). Basitarsomeres all very long and thin. Basitarsomeres III with few small dorsal spines, in addition to apical ones (Fig. 5 B, C). Cerci very long. Coloration (Fig. 3) Contrasting black, light orange, yellow and brown; face yellowish with wide longitudinal, brown bands; scapes yellowish, antennae black brown; pronotum black, more or less mottled with light orange; legs annulated yellow and brown; cerci brown with a wide yellow ring at their base. Male No metanotal gland. FWs (Figs 4 A; 5 D) present, very short, fully overlapping; dorsal field rounded in shape; venation faint; stridulatory file with very small teeth. HWs lacking. Tergites 3 to 6 with a pair of dorsal projections (Fig. 4 E), smaller on tergite 6. Supra anal plate depressed along mid line basally; short and strong setae on basal half, very long ones on distal half. Male genitalia (Fig. 4 F-H) Pseudepiphallic sclerite short and transverse, without extended rami; distal margin V-shaped; apical lobes making a very narrow projection posterior to pseudepiphallic sclerite, well separate the one from the other by a V-shaped indentation; pseudepiphallic arms very short, C-sclerite having the shape of a thin and very long spine. Ectophallic fold very short, hardly distinct between apical lobes and pseudepiphallic parameres; ectophallic apodemes very short, greatly enlarged at the level of the arc. Endophallic sclerite long and narrow, with very short endophallic apodemes (both longitudinal crest and distal lamella). No phallic glands. Female Similar to male, but without FWs and tergal glands. Ovipositor short, about half TIII length, and gently curved upwards (Fig. 4 I); apex slightly widened, without distal teeth. Female genitalia Copulatory papilla (Fig. 5 E-G) short, conical, with acute apex.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFEC353AFCCFFE4CFB4ABA71.taxon	description	(Figs 3 - 5; Table 3) urn: lsid: zoobank. org: act: 155 B 4850 - 5 A 0 F- 4 CD 5 - A 8 EE- 89 F 6 A 81 D 6 E 6 A	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFEC353AFCCFFE4CFB4ABA71.taxon	etymology	ETYMOLOGY. — Species named after our colleague and friend Sylvain Hugel (CNRS), who collected the specimens in the field.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFEC353AFCCFFE4CFB4ABA71.taxon	materials_examined	TYPE LOCALITY. — French Guiana, Mont Tumuc-Humac, Mitaraka. TYPE MATERIAL. — Holotype by present designation. French Guiana • 1 ♂; Monts Tumuc-Humac, Massif du Mitaraka, entre C 100 et C 1000; entre 54.44768 O 2.235494 N et 54.4419 O 2.233664 N; alt. entre 350 m et 415 m; 23. II. - 10. III. 2015; F. Legendre & S. Hugel leg.; fn SH 175, de nuit, sur une souche d’arbre surélevée; MNHN-EO-ENSIF 3984. Allotype. French Guiana • 1 ♀; Monts Tumuc-Humac, Massif du Mitaraka, entre C 1000 et savane roche; entre 54.4419 O 2.23366 N et 54.4346 O 2.238655 N; alt. entre 415 m et 390 m; 23. II. - 10. III. 2015; F. Legendre & S. Hugel leg.; fn SH 221, de nuit, dans un terrrier abandonné; MNHN-EO-ENSIF 3985. Paratypes: 1 male, 3 females. French Guiana • 1 ♂ 1 ♀; Monts Tumuc-Humac, Massif du Mitaraka, entre B 100 et un chablis; entre 54.448813 O 2.236462 N et 54.444953 O 2.243543 N; alt. entre 325 m et 360 m; 28. II. 2015; F. Legendre & S. Hugel leg.; fn SH 283 - 284, de nuit, sur une souche d’arbre renversée; MNHN-EO-ENSIF 3975, 3977 • 1 ♀; same data as holotype; fn SH 176, de nuit, sur une souche d’arbre surélevée; MNHN-EO-ENSIF 3976 • 1 ♀; same data as allotype; fn. SH 223, de nuit, en mauvais état, dans un terrrier abandonné; MNHN-EO-ENSIF 3978. ADDITIONAL MATERIAL EXAMINED. — French Guiana • 1 juvenile; Monts Tumuc-Humac, Massif du Mitaraka, entre C 100 et C 1000; entre 54.44768 O 2.235494 N et 54.4419 O 2.233664 N; alt. entre 350 m et 415 m; 23. II. - 10. III. 2015; F. Legendre & S. Hugel leg.; fn SH 222, de nuit, dans un terrrier abandonné; molecular sample LDG 515, 549; MNHN-EO-ENSIF 3986 • 1 juvenile; Monts Tumuc-Humac, Massif du Mitaraka, vers sommet en Cloche; entre 54.4541 O 2.2349 N et 54.4646 O 2.2329 N; alt. entre 370 m et 470 m; 23. II. - 10. III. 2015; F. Legendre & S. Hugel leg.; fn. SH 420, de nuit; MNHN.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFEC353AFCCFFE4CFB4ABA71.taxon	diagnosis	DIAGNOSIS. — In addition to the characters of the genus, last three articles of maxillary palpi black; face yellowish with a wide brown band under each eye, the area below median ocellus somewhat darker; pronotum deep black with a faint and irregular longitudinal orange line; hindfemur outer side lightly colored in lower half; tergites very dark brown, almost black with dirty-yellow flecks.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFEC353AFCCFFE4CFB4ABA71.taxon	description	DESCRIPTION In addition to the characters of the genus. General morphology TI inner apical spur longer than the outer. TII apical spurs subequal. TIII apical spurs: median spur longer on outer side (Fig. 5 B, C), dorsal spur much longer on inner side, going beyond basitarsomere III mid length (Fig. 5 C). TIII serrulation made of very small teeth, hardly visible among strong and numerous black setae; serrulation lacking between dorsal subapical and apical spurs; on inner side, no spine between isa 1 and isa 2, and between isa 2 and isa 3, one or two (mean 1.5) spines between isa 3 and isa 4, 10 - 14 spines (mean 12) in males, and 15 spines in females above isa 4; on outer side, no spine between osa 1 and osa 2, no spine in males, zero to two (mean 1) spines in females between osa 2 and osa 3, two to three (mean 2.8) spines between osa 3 and osa 4, 9 - 13 spines (mean 11) in males, and 15 - 19 (mean 17) spines in females above osa 4. Tarsi very long; second tarsomeres not widened; basitarsomeres III very long and thin, without dorsal inner spine except one apical, with one to three (mean two) in males, three to four (mean 3.8) in females outer dorsal spines, in addition to distal one. Coloration Face (Fig. 4 B) yellowish, cheeks darker; a wide longitudinal band beneath each eye; area beneath median ocellus also darker; scapes light orange, antennae dark brown. Pronotum (Fig. 4 A) black brown to black, with a faint light orange median longitudinal line. TI, TII dirty yellow with brown rings. FI, FII (Fig. 3 C) dirty yellow with brown rings, and a small, basal brown spot. TIII (Fig. 3 C) dirty yellow with brown dorsal flecks; ventral margin yellow; spurs yellowish with brown tips. FIII (Fig. 4 C, D) dirty yellow marked with brown: in distal half, two brown rings; in basal half, two larger flecks, the first one divided on outer side, simple on inner side, the second simple with additional oblique yellow lines; two additional brown spots on inner side at about one third of FIII length. Basitarsomeres brown with yellow base. Tergites black with dirty-yellow spots (Fig. 3). Male (Fig. 3 B, C) Metanotum without glandular structure. FWs short, not reaching metanotum distal margin (Fig. 4 A); venation (Fig. 5 D) faint, except for a strong M + CuA vein, separating dorsal and lateral fields; file (PCu) extending over half FW width only; harp with three to four widely separated veins; lateral field short and narrow, triangular, without venation; distal margins with short setae, without inflated areas. Stridulatory file with about 40 very small teeth (n = 1). FWs black, with slightly yellowish M + CuA vein. Tergites 3 to 6 with paired transverse projections; each projection with a rounded median conical apex (Fig. 4 E). Supra anal plate as on Fig. 4 E. Subgenital plate short and relatively low; distal margin straight. Male genitalia (Fig. 4 F-H) Very small, about half the size of Philippopsis guianae genitalia. Pseudepiphallic sclerite transverse, without separate rami. Posterior part of pseudepiphallus membranous located below the transverse pseudepiphallic sclerite and narrowed before apex; apical lophi membranous, separate by a V-shaped indentation. B-sclerite very reduced. C-sclerite long and acute, not bifurcate apically as in P. guianae. A-sclerite quadrangular, about twice as long as wide. Pseudepiphallic parameres wellsclerotized, their surface granular; each paramere comprising a flat ventral sclerite (PsP 1 sensu Souza-Dias et al. 2015) and a slightly concave dorsal sclerite (PsP 2 sensu Souza-Dias et al. 2015). Ectophallic fold sclerotized. Ectophallic arc flat and very wide; ectophallic apodemes very short, flat, except for a large dorsal hump at arc level, and not developed ventrally (contra P. guianae). Endophallic sclerite very long and narrow; apodemes very reduced (dorsal crest and anterior lamella). Female (Fig. 3 A) Subgenital plate small, transverse; distal margin bisinuate. Ovipositor small, about half TIII length; dorsal valve apex lanceolate, with many parallel lines, as leaf veins (Fig. 4 I). Female genitalia (Fig. 5 E-G) Copulatory papilla small, conical with a very thin apex; wellsclerotized, except for its very apex. Measurements (in mm) See Table 3.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFEC353AFCCFFE4CFB4ABA71.taxon	biology_ecology	HABITAT AND ECOLOGY Aracopsis hugeli Desutter-Grandcolas, n. gen., n. sp. has been found only by night, foraging in the leaf litter. It has been observed several times on the bare ground beneath the stump of a fallen tree (Fig. 3, S. Hugel, pers. comm.). As Phalangopsis and Philippopsis, it may hide during the day in cavities at ground level (Desutter-Grandcolas 1992 a).	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFEC353AFCCFFE4CFB4ABA71.taxon	description	CALLING SONG The song has been recorded at 23 ° C in rearing conditions with several individuals of different species singing altogether, which precludes detailed analysis and illustration of the call. The song of Aracopsis hugeli Desutter-Grandcolas, n. gen., n. sp. consists in a series of short buzzing trills emitted at length. Each trill has a mean duration of 3.4 s (min 2.6, max 4.1, n = 7), but longer trills have also been recorded (6.1 s and 8.3 s, n = 2); it is made of very short syllables (mean duration: 62 ms, min 48 ms, max 76 ms, n = 30) regularly and rapidly emitted. Dominant and fundamental frequency averages 3.5 kHz, with four additional harmonics of regularly decreasing amplitude.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFEC353AFCCFFE4CFB4ABA71.taxon	discussion	PHYLOGENETIC RELATIONSHIPS Molecular phylogenetic evidence confirm the close relationships of Aracopsis Desutter-Grandcolas, n. gen., Philippopsis and Phalangopsis (Warren et al. 2019).	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFE8353AFCD1FC4FFA2ABC32.taxon	description	(Fig. 6)	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFE8353AFCD1FC4FFA2ABC32.taxon	type_taxon	TYPE SPECIES. — Acantoluzarida nigra Desutter-Grandcolas, 1992 a by original designation.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFE8353AFCD1FC4FFA2ABC32.taxon	distribution	DISTRIBUTION. — French Guiana.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFE8353AFCD1FC4FFA2ABC32.taxon	diagnosis	DIAGNOSIS. — See Desutter-Grandcolas (1992 a: 110).	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFE8353AFCD1FC4FFA2ABC32.taxon	biology_ecology	HABITAT. — Straminicolous species (Desutter-Grandcolas 1995).	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFE83538FCD3FAC9FD3EBD8F.taxon	description	(Fig. 6; Table 4)	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFE83538FCD3FAC9FD3EBD8F.taxon	materials_examined	TYPE LOCALITY. — French Guiana: Arataye, Affluent Approuague, 8 km NE saut Pararé, Nouragues research station. TYPE MATERIAL. — Holotype by original designation. French Guiana • 1 ♂; Arataye, afflt. Approuague, 8 km NE du saut Pararé, station des Nouragues; 19. VI. 1988; L. Desutter & P. Grandcolas leg.; MNHN-EO-ENSIF 3105. Allotype. French Guiana • 1 ♀; same locality and collector as the holotype; 19. VI. 1988; MNHN-EO-ENSIF 3104. Paratypes: 5 males, 14 females. French Guiana • 1 ♂; same locality as the holotype; 31. III. 1988; L. Desutter leg.; MNHN-EO-EN-SIF 5880 • 1 ♂; 1. IV. 1988; L. Desutter leg.; MNHN-EO-ENSIF 5881 • 1 ♀; same locality and collector as the holotype; 31. IV. 1988; MNHN-EO-ENSIF 3104 • 1 ♂ 8 ♀; 3 - 15. VI. 1988; MNHN-EO-ENSIF 5882, 5885 - 5892 • 1 ♂ 1 ♀; 13 - 17. VII. 1988; MNHN-EO-ENSIF 5883, 5893 • 1 ♂; 4 ♀; Sinnamary, piste de St Elie, PK 15; 28. VI. - 24. VII. 1989; P. Grandcolas leg.; MNHN-EO-ENSIF 5884, 5894 - 5897. MATERIAL FROM THE MITARAKA. — French Guiana • 2 ♂; Monts Tumuc-Humac, Massif du Mitaraka, vers sommet en Cloche; entre 54.4541 O 2.2349 N et 54.4646 O 2.2329 N; alt. entre 370 m et 470 m; 23. II. - 10. III. 2015; F. Legendre & S. Hugel leg.; fn. SH 395, SH 456, de nuit; MNHN • 5 ♀; same data as for preceding; fn. SH 396, SH 397, SH 399, SH 503, SH 536, de nuit; MNHN • 1 ♂; same data as for preceding; fn. SH 434, de jour; MNHN • 1 ♀; same data as for preceding; fn. SH 435, de jour; MNHN • 1 ♂; Monts Tumuc-Humac, Massif du Mitaraka, D 2; 54.451125 O 2.234786 N; alt. 300 m; 23. II. - 10. III. 2015; F. Legendre & S. Hugel leg.; fn. SH 077, de nuit; MNHN • 1 ♂; Monts Tumuc-Humac, Massif du Mitaraka, entre C 100 et C 1000; entre 54.44768 O 2.235494 N et 54.4419 O 2.233664 N; alt. entre 350 m et 415 m; 23. II. - 10. III. 2015; F. Legendre & S. Hugel leg.; fn. SH 173, de nuit; MNHN • 1 ♀; same data as for preceding; fn. SH 174, de nuit; MNHN • 1 ♂; Monts Tumuc-Humac, Massif du Mitaraka, Prox Borne 1; entre 54.4365 O 2.2269 N et 54.4355 W 2.2125 N; alt. entre 300 m et 445 m; 23. II. - 10. III. 2015; F. Legendre & S. Hugel leg.; fn. SH 353, de nuit; MNHN • 1 ♀; same data as for preceding; fn. SH 354, de nuit; MNHN • 1 ♀; Monts Tumuc-Humac, Massif du Mitaraka, Layon A; entre 54.4509 O 2.2357 N et 54.4547 O 2.2405 N; alt. entre 280 m et 365 m; 23. II. - 10. III. 2015; F. Legendre & S. Hugel leg.; fn. SH 309, de nuit; MNHN.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFE83538FCD3FAC9FD3EBD8F.taxon	description	COMPLEMENTS OF DESCRIPTION In addition to the characters given by Desutter-Grandcolas (1992 a). Coloration Lateral yellow stripes extended on occiput, vertex, FWs and metanotum, sometimes forked on occiput (Fig. 6); muscular inscriptions interrupting the stripes at pronotum one third. FIII outer side and base of inner side striped with brown; dorsal side and apical part of inner side speckled with brown especially at apex; spurs yellowish, dark at base and apex. Tergites speckled with yellow on lateral margins (Fig. 6 C), this pattern often obscured in males (Fig. 6 D). Female (Fig. 6 E) Bigger than male. Measurements (in mm) See Table 4. Variation Size highly variable. Mouthparts sometimes marked with white. Cerci brown or light brown at base. Yellow rings sometimes present on TI and TII. Females with sometimes five veins on FW lateral field. Females tend to lose their forewings	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFEA3538FF7DFB6EFC82BCEF.taxon	description	(Figs 7 - 11)	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFEA3538FF7DFB6EFC82BCEF.taxon	type_taxon	TYPE SPECIES. — Luzaridella obscura Desutter-Grandcolas, 1992 a by original designation.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFEA3538FF7DFB6EFC82BCEF.taxon	distribution	DISTRIBUTION. — French Guiana and Brazil (Amazonas).	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFEA3538FF7DFB6EFC82BCEF.taxon	diagnosis	DIAGNOSIS. — See Desutter-Grandcolas (1992 a: 103).	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFEA3538FF7DFB6EFC82BCEF.taxon	biology_ecology	HABITAT. — Straminicolous species (Desutter-Grandcolas 1995).	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFEA3525FE87F988FC8BBDAF.taxon	description	(Figs 7, 8, 9 A-E; Table 5) urn: lsid: zoobank. org: act: 9 B 67 BEAE-A 1 A 0 - 4 DE 8 - BB 6 C- 39 FF 8 F 9 F 74 A 0	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFEA3525FE87F988FC8BBDAF.taxon	materials_examined	TYPE LOCALITY. — French Guiana, Monts Tumuc-Humac, Massif du Mitaraka. TYPE MATERIAL. — Holotype by originalpresent designation. French Guiana • 1 ♂; Monts Tumuc-Humac, Massif du Mitaraka vers sommet en Cloche; entre 54.4541 O 2.2349 N et 54.4646 O 2.2329 N; alt. entre 370 m et 470 m; 23. II. - 10. III. 2015; F. Legendre & S. Hugel leg.; fn. SH 412, de nuit; MNHN-EO-ENSIF 9752 (male in bad condition, covered with dry mold). Allotype. French Guiana • 1 ♀; Monts Tumuc-Humac, Massif du Mitaraka vers sommet en Cloche; entre 54.4541 O 2.2349 N et 54.4646 O 2.2329 N; alt. entre 370 m et 470 m; 23. II. - 10. III. 2015; F. Legendre & S. Hugel leg.; fn. SH 398, de nuit; MNHN-EO-ENSIF 9757. Paratypes: 2 females. French Guiana • 1 ♀; Monts Tumuc-Humac, Massif du Mitaraka, Layon A; entre 54.4509 O 2.2357 N et 54.4547 O 2.2405 N; alt. entre 280 m et 365 m; 23. II. - 10. III. 2015; F. Legendre & S. Hugel leg.; fn. SH 111, de nuit; MNHN-EO-ENSIF 9755 • 1 ♀; MontsTumuc-Humac, Massif du Mitaraka, D 2; 54.451125 O 2.234786 N; alt. 300 m; 23. II. - 10. III. 2015; F. Legendre & S. Hugel leg.; fn. SH 080, de nuit; MNHN-EO-ENSIF 9745. OTHER MATERIAL EXAMINED. — French Guiana • 1 ♀; Monts Tumuc-Humac, Massif du Mitaraka, D 2; 54.451125 O 2.234786 N; alt. 300 m; 23. II. - 10. III. 2015; F. Legendre & S. Hugel leg.; fn. SH 027, de nuit; MNHN.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFEA3525FE87F988FC8BBDAF.taxon	etymology	ETYMOLOGY. — Species named after its slender shape compared to other species of the genus.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFEA3525FE87F988FC8BBDAF.taxon	diagnosis	DIAGNOSIS. — Within the genus, species relatively large, with thin PIII at least in males (Fig. 7 A). Male FWs relatively soft, compared to other species of the genus, covering abdomen up to tergite 7; stridulum complete. Male genitalia distinctive (A-sclerite protruding as a thick curved spine; pseudepiphallic apical sclerotization X-shaped, delimiting a pair of distal cavities very lightly sclerotized dorsally, and a free sclerotized process more ventrally; distal twothird of ectophallic apodemes rubbon-like and thin, curved ventrally toward pseudepiphallic parameres, anterior third vertical). Female copulatory papilla oval, with dorsal and ventral sides little sclerotized. Species differing from Luzaridella maculata n. sp. by male and female genitalia (compare Fig. 7 D-G and Fig. 11 E-G), male stridulatory file, a bigger size (compare Table 5 and 6) and sternite coloration (light brown with median part darker). More similar to Luzaridella annulata Desutter-Grandcolas, 1992 from which it can be separated by its larger size, darker coloration (especially for annulated legs), and female copulatory papilla (compare Fig. 9 C-E and Desutter-Grandcolas 1992 a, figure 9).	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFEA3525FE87F988FC8BBDAF.taxon	description	DESCRIPTION In addition to characters of the genus (Desutter-Grandcolas 1992 a): General morphology TIII serrulation lacking between subapical spurs and apical spurs on both sides; on inner side, two to four spines (mean 2.5 in male, 3.5 in females) between isa 1 and isa 2, two to four (mean 2.5 in male, 3.3 in females) spines between isa 2 and isa 3, three to four spines (mean 4 in male, 3.8 in females) between isa 3 and isa 4, 12 - 13 spines in male (mean 12.5) and 11 - 15 spines (mean 13.5) in females above isa 4; on outer side, three to four spines (mean 3 in male, 3.3 in females) between osa 1 and osa 2, three to four spines (mean 3 in male, 3.5 in females) between osa 2 and osa 3, four to six spines (mean 4 in male, 5.3 in females) between osa 3 and osa 4, 11 - 12 spines (mean 11.5) in male, and 12 - 13 (mean 12.5) spines in females above osa 4. Basitarsomere III serrulation: six to seven inner spines (mean 6.5 in male, 6.8 in females), six to nine outer spines (mean 6.5 in male, 7.3 in females) in addition to apical spines. Coloration Head: Face light brown marked with yellow; cheeks darker than face; ocelli marked with black; vertex and occiput dark brown, with four longitudinal yellow lines (Fig. 8 A): two outer lines starting at median ocellus, reaching lateral ocelli, forked at beginning of vertex, and extending to pronotum; two inner lines, shorter than outer stripes. Eyes grey. Antennal pits whitish. Scapes yellow with two brown rings, one at base, one at apex. Antennae yellow at base, then brown with yellow rings. Mouthparts white with light brown spots. Palpi light brown, white on their dorsal and ventral sides. Pronotum DD and LL separated by a wide yellow stripe (Fig. 8 A, B); DD light brown with lighter muscular inscriptions and two yellow spots (close to yellow stripes) on posterior margin; LL ferruginous brown, light brown to yellow close to anterior angle. Legs: FI and FII yellow with three brown rings. TI and TII brown with two and three yellow rings, respectively. Tarsomeres I and II brown, yellow at their base. FIII yellow with brown pattern: outer side and inner base striped with brown; a brown ring near apex; apex brown. TIII brown on dorsal side, lighter on ventral side; with two yellow rings barely visible; spurs yellowish, their base and apex dark; tarsomeres III light brown. Tergites light brown with dark brown and yellow pattern (Fig. 8 C). Sternites light brown with a darker median part. Cerci light brown, with a brown ring close to brown apex. Male Metanotum glandular, covered with many long setae (Fig. 7 C). Forewings (Fig. 7 B) wider than pronotum and abdomen, trapezoidal (wider posteriorly than anteriorly); long for the genus, reaching tergite 7. Venation faint (Fig. 9 A, B); lateral field with four parallel longitudinal veins, the second branched distally; stridulum complete, harp without vein, mirror much wider than long, not crossed by any vein; PCu very oblique, bearing a very short file located very high on the vein and with only five teeth. Male genitalia (Fig. 7 D-G) Genitalia slightly wider than long, with very short rami (Fig. 7 D). Pseudepiphallic arms well-developed; membrane separating them sclerotized apically as two cup-like sclerites connected together and extending toward sclerotized parts of EEI and ectophallic fold as a pair of free sclerotized processes (Fig. 7 D, G); A-sclerite hook-like; B-sclerite (Fig. 7 F) widened before apex, then regularly narrowed toward acute apex; pseudepiphallc parameres simple, not hook-like, between pseudepiphallic apical sclerites and A-sclerites (Fig. 7 D, E, G). Ectophallic apodemes thin and rubbon-like on anterior twothird, as a curved, vertical lamella on posterior third (Fig. 7 D); arc fully sclerotized, sinuous (Fig. 7 D); pseudepiphallic sclerite anterior part vertical, abutting against ectophallic apodeme (Fig. 7 D). Ectophallic fold short, with a trifid apical sclerite (Fig. 7 E). Endophallic sclerite very short, distal apex trifid (Fig. 7 E); endophallic apodeme very short, lamella-like. No dorsal cavity. Female Larger than male. FWs short, not overlapping, reaching tergite 3. Dorsal field with seven to nine longitudinal veins, lateral field with four or five longitudinal veins. Female genitalia Copulatory papilla (Fig. 9 C-E) small (less than 1 mm long), nearly oval in shape, distal margin bisinuate, ventral and dorsal sides less sclerotized. Measurements (in mm) See Table 5. Variation Size highly variable in females. First ring of FI and FII sometimes hardly marked. Dorsal disc and tergites sometimes darker.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFF73520FE9EFB49FDC4BEED.taxon	description	(Figs 9 F-H, 10, 11; Table 6) urn: lsid: zoobank. org: act: ED 281 AA 8 - 746 B- 4 EDC-AD 3 B- 6 E 3 E 32 A 6 AEB 9	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFF73520FE9EFB49FDC4BEED.taxon	materials_examined	TYPE LOCALITY. — French Guiana, Monts Tumuc-Humac, Massif du Mitaraka. TYPE MATERIAL. — Holotype by present designation. French Guiana • 1 ♂; Monts Tumuc-Humac, Massif du Mitaraka, Layon A; entre 54.4509 O 2.2357 N et 54.4547 O 2.2405 N; alt. entre 280 m et 365 m; 23. II. - 10. III. 2015; F. Legendre & S. Hugel leg.; fn. SH 310, de nuit; MNHN-EO-ENSIF 9740). Allotype. French Guiana • 1 ♀; Monts Tumuc-Humac, Massif du Mitaraka, D 2; 54.451125 O 2.234786 N; alt. 300 m; 23. II. - 10. III. 2015; F. Legendre & S. Hugel leg.; fn. SH 076, de nuit; MNHN-EO-ENSIF 9753. Paratypes: 8 males, 9 females. French Guiana • 4 ♂ 2 ♀; Monts Tumuc-Humac, Massif du Mitaraka, same data as the holotype; fn. SH 112, SH 121, SH 562, SH 564, fn. SH 123, SH 266, de nuit; MNHN-EO-ENSIF 9742, 9749, 10765, 10768, 10769, 10771 • 1 ♂ 3 ♀; Monts Tumuc-Humac, Massif du Mitaraka, vers sommet en Cloche; entre 54.4541 O 2.2349 N et 54.4646 O 2.2329 N; alt. entre 370 m et 470 m; 23. II. - 10. III. 2015; F. Legendre & S. Hugel leg.; fn. SH 513, SH 511, SH 512, SH 537, de nuit; MNHN-EO-ENSIF 10767, 10774, 10775, 10776 • 1 ♂ 1 ♀; Monts Tumuc-Humac, Massif du Mitaraka, D 2; 54.451125 O 2.234786 N; alt. 300 m; 23. II. - 10. III. 2015; F. Legendre & S. Hugel leg.; fn. SH 010, SH 011, de jour; MNHN-EO-ENSIF 10766, 10770 • 1 ♂ 3 ♀; same data as preceding; fn. SH 081, SH 042, SH 025, SH 082, de nuit; MNHN-EO-ENSIF 9741, 10773, 10772, 10777 • 1 ♂ Monts Tumuc-Humac, Massif du Mitaraka, Prox Borne 1; entre 54.4365 O 2.2269 N et 54.4355 O 2.2125 N; alt. entre 300 m et 445 m; 23. II. - 10. III. 2015; F. Legendre & S. Hugel leg.; fn. SH 352, de nuit; MNHN-EO-ENSIF 9753. OTHER MATERIAL EXAMINED. — French Guiana • 2 ♂; Monts Tumuc-Humac, Massif du Mitaraka, Layon A; entre 54.4509 O 2.2357 N et 54.4547 O 2.2405 N; alt. entre 280 m et 365 m; 23. II. - 10. III. 2015; F. Legendre & S. Hugel leg.; fn. SH 113, SH 563, de nuit; MNHN • 1 ♂; Monts Tumuc-Humac, Massif du Mitaraka, vers sommet en Cloche; entre 54.4541 O 2.2349 N et 54.4646 O 2.2329 N; alt. entre 370 m et 470 m; 23. II. - 10. III. 2015; F. Legendre & S. Hugel leg.; fn. SH 547, de nuit; MNHN • 1 juvenile; Monts Tumuc-Humac, Massif du Mitaraka, D 2; 54.451125 O 2.234786 N; alt. 300 m; 23. II. - 10. III. 2015; F. Legendre & S. Hugel leg.; fn. SH 002, de jour; MNHN.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFF73520FE9EFB49FDC4BEED.taxon	etymology	ETYMOLOGY. — Species named after its pattern of coloration.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFF73520FE9EFB49FDC4BEED.taxon	diagnosis	DIAGNOSIS. — Within the genus, species close to Luzaridella clara Desutter-Grandcolas, 1992 by its light coloration (Fig. 11), its small and globular male genitalia, and the shape of female copulatory papilla (rectangular and entirely sclerotized); it can be separated from this species by its coloration and length of ovipositor (less than 10 mm in L. clara, i. e., well shorter than FIII and TIII, versus> 13 mm in L. maculata n. sp., longer than TIII). Also resembling Luzaridella miniata n. sp. by general coloration, but differing from that species by sternite coloration (light brown marked with black), male stridulatory file (16 teeth against five in L. miniata n. sp.), male genitalia (compare Figs 7 D-G and 11 E-G), female genitalia (oval and partly desclerotized dorsally and ventrally in L. miniata n. sp.), and a smaller size (compare Tables 5 and 6).	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFF73520FE9EFB49FDC4BEED.taxon	description	DESCRIPTION In addition to the character of the genus (see Desutter-Grandcolas 1992 a). General morphology TIII serrulation lacking between subapical spurs and apical spurs on both sides; on inner side, three to five spines (mean 3.8) in males, four to six spines (mean 4.9) in females between isa 1 and isa 2, four to five spines (mean 4.1) in males, four to six spines (mean 4.8) in females between isa 2 and isa 3, four to six spines (mean 4.5) in males, four to five spines (mean 4.9) in females between isa 3 and isa 4, 9 - 14 spines (mean 11.8) in males, 13 - 15 spines (mean 14.4) in females above isa 4; on outer side, two to four spines (mean 3) in males, three to four spines (mean 3.4) in females between osa 1 and osa 2, four to five spines (mean 4.3) in males, five spines in females between osa 2 and osa 3, four to five spines (mean 4.4) in males, four to six spines (mean 5) in females between osa 3 and osa 4, 9 - 13 spines (mean 10.6) in males, 12 - 14 spines (mean 13.1) in females above osa 4. Barsitarsomere III serrulation: five to nine inner spines (mean 8.6 in males, 7.8 in females) and five to nine outer spines (mean 5.8 in males, 7.5 in females) in addition to apical spines. Phalangopsidae crickets from the Mitaraka Coloration Face yellowish with light brown marks, including a curved line between lower angle of eyes and epistomal suture and sometimes a light yellow, median longitudinal line under median ocellus (Fig. 11 D); clypeus and labrum light yellow, with a pair of brown spots on clypeus; cheeks light to dark brown with an elongate yellowish spot close to the eye more or less prolonged toward mandibula; ocelli partly circled with black. Fastigium and vertex light to dark brown, fastigium sometimes lighter; with five longitudinal yellow lines: two external ones from lateral ocelli to occiput, sometimes forked behind the eyes, extending to pronotum and forewings (Fig. 11 A, C); two inner ones, transparent, shorter than external stripes, joining at vertex mid length; one median and longitudinal, very thin, short and most often barely visible. Scapes yellow marked with brown, especially at base and apex. Antennae yellow at base, then brown with yellow rings. Palpi light brown laterally, lighter on dorsal and ventral margins (Fig. 11 C, D). Pronotum DD and LL separated by a wide yellow stripe (Fig. 11 C); DD light brown with lighter muscular inscriptions; posterior margin largely bordered with black brown, except for two yellow spots close to longitudinal yellow stripes; LL light brown to yellow on lower margin, with a wide black brown band dorsally. Legs (Fig. 11 A): FI and FII yellow with two brown rings. TI and TII brown with two and three yellow rings, respectively. Basitarsomeres I and II brown, yellow at base; tarsomeres 2 and 3 yellow. FIII yellow with brown pattern: outer side and inner base striped with brown; a brown ring near the knee; knee brown. TIII (Fig. 11 A) brown dorsally, lighter ventrally; near the knee, a short black ring and a larger yellow ring; an additional yellow fleck on each side, more or less distinct; spurs yellowish, their base and apex dark. Basitarsomeres III light brown with darker spot near apex; tarsomeres 2 and 3 yellowish. Tergites light brown with dark brown and yellow pattern (Fig. 11 B). Sternites light brown marked with black. Cerci light brown, lighter at base with yellow ring near apex. Male Metanotum glandular, with a wide bunch of long setae along both anterior and posterior margins, separated by a deep transverse furrow. FWs (Fig. 11 A) as wide as pronotum and abdomen, reaching tergite 4. Venation visible but sometimes interrupted; lateral field with five parallel longitudinal veins; stridulum complete, harp without vein, mirror much wider than long, not crossed by any vein; stridulatory file with 16 teeth. Subgenital plate short and high. Male genitalia (Fig. 11 E-H) Very small and globular, with relatively long and thin rami. Pseudepiphallic arms less than half rami length. Dorsal membrane of pseudepiphallus (between arms) present as a small, broadly triangular area, with relatively big apical structures; B-sclerite short, wide over its whole length, not narrowed distally; A-sclerite with a very small apical spine. Pseudepiphallic parameres globular with a non-protruding, small, broad apical tooth. Ectophallic apodemes not extending beyond rami anterior tip; very wide and concave along their whole length, regularly narrowed anteriorly; arc fully sclerotized, narrow compared to ectophallic apodemes. Ectophallic fold very short, sclerotized. Endophallic sclerite with trifid distal margin. Endophallic apodeme very well developed, with a high median longitudinal crest, connected to ectophallic fold, and lamellas along and beyond endophallic sclerite. No dorsal cavity. Female Larger than male. FWs short, little recovering, reaching tergite 1 (Fig. 11 B). Dorsal field with six to eight longitudinal veins, lateral field with four longitudinal veins. Subgenital plate distal margin deeply sinuate. Ovipositor as long or slightly longer than FIII, but longer than TIII. Female genitalia Copulatory papilla (Fig. 9 F-H) entirely sclerotized and short, less than one millimetre long; almost rectangular in shape; upper part with small protuberance. Measurements (in mm) See Table 6. Variation Vertex sometimes brown ferruginous. In darker specimens, tergite coloration sometimes dark brown. In males veins 3 and 4 on FW lateral field sometimes extended near FW lower margin. Some specimens at hand seem in bad condition and present very dark coloration for cheek (light coloration barely visible), vertex (yellow forks sometimes difficult to see), scapes (entirely brownish), base of antennae (brown), palpi (uniform coloration), pronotum (yellow spots not visible), sternites (coloration very variable with black margins or middle part), or metanotum (light brown to yellowish).	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFF23520FC78FF6DFC7ABB82.taxon	description	(Fig. 12)	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFF23520FC78FF6DFC7ABB82.taxon	type_taxon	TYPE SPECIES. — Luzarida annuliger Hebard, 1928 a by original designation.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFF23520FC78FF6DFC7ABB82.taxon	diagnosis	DIAGNOSIS. — See Hebard (1928 a) and Desutter-Grandcolas (1992 a).	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFF23520FC78FF6DFC7ABB82.taxon	distribution	DISTRIBUTION. — Northwest South America (Colombia, Ecuador, Guyana, French Guiana, Brazil, Trinidad).	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFF23520FC78FF6DFC7ABB82.taxon	biology_ecology	HABITAT. — Species most often found at ground level, active at night, or hiding during the day. They may perched at night on diverse low structures, such as fallen dead trunks, raised twigs or leaves, or low plants.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFF2352FFCC9FD6FFE3CBBEE.taxon	description	(Fig. 12; Table 7)	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFF2352FFCC9FD6FFE3CBBEE.taxon	materials_examined	TYPE LOCALITY. — French Guiana, Arataye, Affl. Approuague, 8 km NE pied du Saut Pararé. TYPE MATERIAL. — Holotype by original designation. French Guiana • 1 ♂; Arataye, afflt. Approuague, 8 km NE du saut Pararé, station des Nouragues; 19. V. 1988; L. Desutter leg.; MNHN-EO-ENSIF 3035. Allotype. French Guiana • 1 ♀; same locality and collector as the holotype; 11. V. 1988; MNHN-EO-ENSIF 3032. Paratypes: 1 male, 6 females. French Guiana • 1 ♂ 6 ♀; same locality as the holotype; VI. 1988; L. Desutter & P. Grandcolas leg.; MNHN-EO-ENSIF 5942 - 5948. MATERIAL FROM THE MITARAKA. — French Guiana • 1 ♂; Monts Tumuc-Humac, Massif du Mitaraka, Layon A; entre 54.4509 O 2.2357 N et 54.4547 O 2.2405 N; alt. entre 280 m et 365 m; 23. II. - 10. III. 2015; F. Legendre & S. Hugel leg.; fn. SH 308, de nuit; MNHN.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFF2352FFCC9FD6FFE3CBBEE.taxon	description	COMPLEMENT OF DESCRIPTION In addition to the characters given by Desutter-Grandcolas (1992 a). Coloration Mouthparts dark brown with whitish specks. Four brown specks on posterior margin of pronotum. FIII outer side and base of inner side striped with brown. TIII light brown. Spurs yellowish, with dark base and apex. Cerci whitish dorsally; ventrally black on basal third, with a black ring at second third, and black at apex. Male genitalia As on Fig. 12 D-F. Measurements (in mm) See Table 7. Variation Mirror crossed by two or three veins.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFF2352FFCC9FD6FFE3CBBEE.taxon	discussion	REMARK Two specimens of L. grandis have been photographed in the field: both were found active (Fig. 12 A, B), even singing (Fig. 12 C), in the leaf litter. The photos show a large variation in coloration, especially for head, pronotum and cerci, which can be almost whitish.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFFD352FFE9BFC8FFEB9BD55.taxon	type_taxon	TYPE SPECIES. — Lerneca varipes Walker, 1869 by original monotypy.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFFD352FFE9BFC8FFEB9BD55.taxon	distribution	DISTRIBUTION. — Tropical areas of South and Central America.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFFD352FFE9BFC8FFEB9BD55.taxon	diagnosis	DIAGNOSIS. — See Desutter-Grandcolas (1992 a).	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFFD352FFE9BFC8FFEB9BD55.taxon	biology_ecology	HABITAT. — Diurnal or nocturnal straminicolous species (Desutter-Grandcolas 1995).	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFFD352DFE82FB2EFDE8BEED.taxon	description	(Figs 9 I, J, 13 - 15 A-C; Table 8) urn: lsid: zoobank. org: act: 219361 CD- 0 B 25 - 4 C 29 - A 0 CF-C 09 D 982 F 74 EE	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFFD352DFE82FB2EFDE8BEED.taxon	materials_examined	TYPE LOCALITY. — French Guiana, Monts Tumuc-Humac, Massif du Mitaraka. TYPE MATERIAL. — Holotype by present designation. French Guiana • 1 ♂; Monts Tumuc-Humac, Massif du Mitaraka, Layon A; entre 54.4509 O 2.2357 N et 54.4547 O 2.2405 N; alt. entre 280 m et 365 m; 23. II. - 10. III. 2015; F. Legendre & S. Hugel leg.; fn. SH 281, de nuit; MNHN-EO-ENSIF 10757. Allotype. French Guiana • 1 ♀; Monts Tumuc-Humac, Massif du Mitaraka, D 2; 54.451125 O 2.234786 N; alt. 300 m; 23. II. - 10. III. 2015; F. Legendre & S. Hugel leg.; fn. SH 078, de nuit; MNHN-EO-ENSIF 10761. Paratypes. 3 males, 4 females. French Guiana • 2 ♂ 2 ♀; Monts Tumuc-Humac, Massif du Mitaraka; same data and collectors as the holotype; fn. SH 118, SH 280, fn. SH 119, SH 282; MNHN-EO-ENSIF 10755, 10758, 10759, 10762 • 1 ♂; Monts Tumuc-Humac, Massif du Mitaraka, vers sommet en Cloche; entre 54.4541 O 2.2349 N et 54.4646 O 2.2329 N; alt. entre 370 m et 470 m; 23. II. - 10. III. 2015; F. Legendre & S. Hugel leg.; fn. SH 411, de nuit; MNHN-EO-ENSIF 10756 • 1 ♀; Monts Tumuc-Humac, Massif du Mitaraka, D 2; 54.451125 O 2.234786 N; alt. 300 m; 23. II. - 10. III. 2015; F. Legendre & S. Hugel leg.; fn. SH 013, de jour; MNHN-EO-ENSIF 10760 • 1 ♀; Monts Tumuc-Humac, Massif du Mitaraka, vers Sommet en Cloche; entre 54.4541 O 2.2349 N et 54.4646 O 2.2329 N; alt. entre 370 m et 470 m; 23. II. - 10. III. 2015; F. Legendre & S. Hugel leg.; fn SH 514, de nuit; MNHN-EO-ENSIF 10763. OTHER MATERIAL EXAMINED. — French Guiana • 1 ♂ juvenile; Monts Tumuc-Humac, Massif du Mitaraka, D 2; 54.451125 O 2.234786 N; alt. 300 m; 23. II. - 10. III. 2015; F. Legendre & S. Hugel leg.; fn. SH 003, de jour; MNHN • 1 ♂; Monts Tumuc-Humac, Massif du Mitaraka, Layon A; entre 54.4509 O 2.2357 N et 54.4547 O 2.2405 N; alt. entre 280 m et 365 m; 23. II. - 10. III. 2015; F. Legendre & S. Hugel leg.; fn. SH 120, de nuit; MNHN • 1 ♂; Monts Tumuc-Humac, Massif du Mitaraka, layon A; entre 54.4509 O 2.2357 N et 54.4547 O 2.2405 N; alt. entre 280 m et 365 m; 23. II. - 10. III. 2015; F. Legendre & S. Hugel leg.; fn SH 335, de nuit; MNHN • 1 ♀; same data as for preceding; fn SH 110, de nuit; MNHN • 1 ♀; Monts Tumuc-Humac, Massif du Mitaraka, vers sommet en Cloche; entre 54.4541 O 2.2349 N et 54.4646 O 2.2329 N; alt. entre 370 m et 470 m; 23. II. - 10. III. 2015; F. Legendre & S. Hugel leg.; fn. SH 538, de nuit; MNHN.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFFD352DFE82FB2EFDE8BEED.taxon	etymology	ETYMOLOGY. — Species named after the type locality.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFFD352DFE82FB2EFDE8BEED.taxon	diagnosis	DIAGNOSIS. — Species very close to Lerneca fuscipennis (Saussure, 1874) by male FW venation and genitalia, female FW and genitalia, and general coloration (Fig. 13). It differs from that species by details in coloration (PI and PII, sternites brown except for a wide longitudinal yellow band on each side), male genitalia (C-sclerite bifid, pseudepiphallic parameres as on Fig. 15 C), and a bigger size (in particular lengths of FIII, TIII, pronotum, male FW female ovipositor).	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFFD352DFE82FB2EFDE8BEED.taxon	biology_ecology	HABITAT. — Lerneca mitarakensis n. sp. has been collected during the day and at night in the leaf litter.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFFD352DFE82FB2EFDE8BEED.taxon	description	DESCRIPTION In addition to characters of the genus (Desutter-Grandcolas 1992 a). General morphology TIII serrulation lacking between subapical spurs and apical spurs on both sides; on inner side, one spine between isa 1 and isa 2, one to two spines between isa 2 and isa 3 (mean 1.3), two spines between isa 3 and isa 4, seven to eight spines (mean 7.3) in males, six to seven spines (mean 6.3) in females above isa 4; on outer side, zero to one spine between osa 1 and osa 2 (mean 0.5 in males, mean 0.8 in females), one to two spines between osa 2 and osa 3 (mean 1.8 in males, 1.5 in females), two to three (mean 2.5) spines between osa 3 and osa 4, 12 - 14 spines (mean 13) in males, 12 - 15 (mean 13.3) spines in females above osa 4. Barsitarsomeres III serrulation: one to three inner spines (mean 2) and five to seven outer spines (mean 6 in males, mean 5.8 in females) in addition to apical spines. Coloration. Face black, up to lower margins of antennal pits, and behind eyes; under median ocellus, a short transverse yellow line, with upper and lower margins both straight. Ocelli circled with black. Vertex yellowish brown with six yellow longitudinal lines in addition to a variable, thin median line. Eyes grey brown, usually light yellow dorsally. Mouthparts black. Palpi coloration different in males and females (see below). Antennae brown with yellowish base; scapes yellowish brown, inner margin ivory. Pronotum LL black, anterior angle of lower margin possibly yellow; DD yellowish brown, anterior and posterior margins marked with yellow to light brown flecks, two light yellow stripes along lateral margins. Legs I and II yellow, tibiae and femora brown dorsally. FIII yellow with brown pattern: brown stripes on outer side and on base of inner side, brown spots on dorsal side and on inner apical part. TIII brown, lighter on ventral side; subapical spurs brown, light near apex. Tergites dark brown; sternites brown, with lateral part prolonged ventrally as two longitudinal yellow stripes. Cerci light brown, lighter at base. Male Maxillary palpi light brown. Metanotum brown with two median semicolon projections and an ellipse-shaped projection near posterior margin. FWs brown, speckled with lighter brown. Venation as in L. fuscipennis, especially for the oblique file vein with about 40 teeth close to plectrum; FW membrane longitudinally plicated between stridulatory veins. HWs longer than FWs by less than 1 mm. Tergite I yellow. Supra anal plate with two tufts of large bristles close to posterior margin. Subgenital plate brown. Male genitalia Quite similar to those of L. fuscipennis (compare Fig. 15 A-C and D and Desutter-Grandcolas 1992 a, figures 26 - 27), but spine of C-sclerite bifid, with a shorter branch on outer side; B-sclerite large and concave; outer margin denticulate; subapical spine on ventral margin of pseudepiphallic parameres more narrow than in L. fuscipennis. Female Apterous. Maxillary palpi with articles 4 and 5 lighter. Leg coloration lighter than in males. Tergites brown, speckled with yellow, lateral parts dark brown; metanotum and tergite 1 more spotted than other tergites. Supra anal plate brown. Subgenital plate small, truncated at apex, distal margin concave, not bisinuate. Ovipositor smaller than FIII (see measurements). Female genitalia Similar to those of L. fuscipennis (Fig. 9 I, J). Measurements (in mm) See Table 8. Variation Few specimens with yellow legs I and II, even on top. Tibiae III not always lighter on ventral side, especially in females. Yellow stripes on pronotum usually extended on metanotum and tergites (female), or on forewings (male).	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFFF352CFC21F9C9FBC1BEEA.taxon	description	(Figs 9 K, 14 - 15 E-G, Tables 9, 10)	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFFF352CFC21F9C9FBC1BEEA.taxon	discussion	REMARK The most informative characters given by Saussure (1878) to separate Lerneca inalata (Saussure, 1874) from L. fuscipennis (Saussure, 1878) are the shape of the pronotum (trapezoidal, twice as wide as long, against more quadrangular in L. fuscipennis), the shape of the FWs (very wide, against little widened in L. fuscipennis), the shape of the stridulatory file (transverse and straight, against short and oblique in L. fuscipennis) and the slender shape of the legs. These characters are similarly well-contrasted between L. inalata inalata and Lerneca mitarakensis n. sp. (see Figs 13, 14). All the subspecies recently described in L. inalata differ from the description given by Saussure for pronotum, FW and file shapes, as shown by figures in original papers (Gorochov 2007, 2014; Lima et al. 2016). The taxonomic status and relationships of these taxa will have to be reconsidered with the analysis of the type of L. inalata, and through molecular studies	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFFF352CFC21F9C9FBC1BEEA.taxon	materials_examined	TYPE MATERIAL. — Holotype by original designation. Suriname • 1 ♂ (ZMUH, not examined). MATERIAL EXAMINED. — French Guiana • 1 ♂; Monts Tumuc-Humac, Massif du Mitaraka, Savane roche; 54.4347 O 2.2387 N; alt. 390 m; 23. II. - 10. III. 2015; F. Legendre & S. Hugel leg.; fn. SH 206, de nuit; MNHN • 1 ♂; Roche de Kourou; MNHN. Venezuela • 3 ♂ 9 ♀, identified Diplacustes varipes Walker, 1869 by A. Finot, ex coll. Finot; MNHN.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFFF352CFC21F9C9FBC1BEEA.taxon	diagnosis	DIAGNOSIS. — See Saussure (1874) and Saussure (1878) for general morphology and coloration, and Desutter-Grandcolas (1992 a) for characters of male and female genitalia.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFFF352CFC21F9C9FBC1BEEA.taxon	biology_ecology	HABITAT. — During the Mitaraka expedition, L. inalata has been found de nuit in bedrock grown savannas, i. e., in dryer and more open areas than the surrounding forest.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFFF352CFC21F9C9FBC1BEEA.taxon	description	COMPLEMENT TO DESCRIPTION General morphology TIII serrulation as on Table 9. HWs longer than FWs in both males and females by 4 - 5 mm. Male Stridulatory file with 175 – 200 teeth (see Table 10). Male genitalia As in Desutter-Grandcolas (1992 a, figures 28 - 29) and on Fig. 15 E-G. Female (after specimens from Venezuela) Ovipositor straight, shorter than FIII and TIII. Subgenital plate short and transverse; distal margin almost straight. Female genitalia Copulatory papilla elongate and plicate, as in all the species of the genus; abruptly narrowed distally as in Lerneca inalata beripocone Lima, Martins & Lhano, 2016; widened before apex (Fig. 9 K). Measurements (in mm) See Table 10.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFFF352CFC21F9C9FBC1BEEA.taxon	discussion	REMARK The specimens from French Guiana (Mitaraka, Kourou) and Venezuela correspond to Saussure’s 1874 description of L. inalata except for their larger size (Table 9). The specimens originating from Venezuela are also lighter in coloration and their HWs go well beyond FW tip by a mean length of 4.4 mm. HWs are absent or rudimentary in the type of L. inalata according to Saussure’s description, while they are shorter than FWs in Guianese specimens, showing a possible polymorphism for wing length, as documented in other cricket species. All observed specimens have very similar male genitalia (compare Desutter-Grandcolas 1992 a, figures 28, 29 and Fig. 15 E-G); female genitalia are known for Venezuelan specimens only (Fig. 9 K).	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFF8352AFF73FF6DFC21BC0C.taxon	description	(Figs 16 - 17) urn: lsid: zoobank. org: act: 872 E 35 D 5 - 8 D 0 F- 4 A 60 - 9859 - ACEF 999 D 7361	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFF8352AFF73FF6DFC21BC0C.taxon	type_taxon	TYPE SPECIES. — Mellomima guyanensis Desutter-Grandcolas, n. sp. by present designation.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFF8352AFF73FF6DFC21BC0C.taxon	distribution	DISTRIBUTION. — French Guiana and Brazil (P. Souza Dias, pers. comm.).	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFF8352AFF73FF6DFC21BC0C.taxon	etymology	ETYMOLOGY. — Taxon named after the genera Mellopsis Mews & Sperber, 2010 and Guabamima De Mello, 2010.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFF8352AFF73FF6DFC21BC0C.taxon	diagnosis	DIAGNOSIS. — Genus close to Pizacris Souza-Dias & Desutter-Grandcolas, 2015, Mellopsis Mews & Sperber, 2010 and Guabamima De Mello, 2010 (see Souza-Dias et al. 2015) by its male genitalia (pseudepiphallic sclerite and arms well reduced, pseudepiphallic parameres contrastingly very large and apical, endophallic apodeme well-developed and V shaped) and general morphological traits (tympana, head shape, tibial apical and subapical spurs, and serrulation). Mellomima Desutter-Grandcolas, n. gen. differs from all three genera by its coloration (uniform, except for a longitudinal yellow band from labrum tip to pronotum DD), its shiny appearance (body not at all setose), its pronotum shape (longer than wide, not transverse), its thin and elongate body shape, resembling Amusodes Hebard, 1928, with FIII filiform on half their length, and the presence of two bunches of thick setae on distal margin of male supra anal plate. Mellomima Desutter-Grandcolas, n. gen. also differs from Guabamima by its reduced and rounded FWs and its very small pseudepiphallic arms in males; from Mellopsis by the wide lateral field of its FWs and the stocky shape of pseudepiphallic parameres; and from Pizacris by the coloration of the maxillary palpi (brown with only the upper and lower sides white), its partly overlapping FWs and the stocky shape of pseudepiphallic parameres. Female unknown.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFF8352AFF73FF6DFC21BC0C.taxon	description	DESCRIPTION General morphology Medium size crickets with elongate and thin shape; body shiny and not setose (Fig. 16 A). Fastigium wider than scape at base, thinner at apex, not separated from vertex by a transverse furrow or a shallow depression (Fig. 16 B). Eyes (Fig. 16 C, D) small, not protruding. Ocelli not reduced, set as a triangle; distance between lateral ocelli equal to distance between median and one lateral ocelli; median ocellus vertical, almost apical on vertex. Maxillary palpi (Fig. 16 C) very elongate and extremely thin; article 3 slightly shorter than article 4; article 5 a little longer than article 4, regularly widened toward apex, truncate obliquely at apex. Pronotum longer than wide, not at all transverse (Fig. 16 B). Legs elongate and very thin. TI with a small inner and a small outer tympanum; two apical, ventral spurs, small compared to basitarsomere I. TII with two ventral, apical spurs. TIII with 4 / 4 subapical spurs, very short and alternate, in TIII distal half; with 3 / 3 apical spurs (Fig. 16 E, F): inner spurs longer than outer spurs; median and dorsal inner spurs subequal and very long, dorsal apical spur as long as half basitarsomere III; ventral and dorsal outer spur subequal, median outer spur twice as long as ventral and dorsal outer spurs. TIII serrulation strong between and above subapical spurs. FIII with a long filiform apical part, about half its total length (Fig. 16 A). Basitarsomeres all very long and thin (Fig. 16 A). Basitarsomeres III with small inner and outer dorsal spines, in addition to apical ones. Cerci shorter than abdomen. Coloration Uniform, except for a yellow longitudinal band running from labrum through the face, occiput, pronotum DD and perhaps tergite I. Legs almost completely uniformly colored. Male FWs (Fig. 16 B, C) very short, not reaching tergite I distal margin; not completely overlapping dorsally; without glandular structures; left FW apparently not membranous. Venation: no stridulatory structures; lateral field with four longitudinal parallel veins. HWs lacking. Tergites without glandular structures. Supra anal plate quadrangular; a pair of strong setae bunches on distal margin, the setae regularly arranged according to their size (Fig. 16 G). Subgenital plate short, high, deeply furrowed on apical third. Male genitalia (Fig. 17) Pseudepiphallic sclerite rounded and vertical, without extended rami; inner side of pseudepiphallic sclerite with a pair of projections that come against ectophallic apodemes; pseudepiphallic arms very short, A-sclerite well developed, having the shape of concave lamellas; pseudepiphallic parameres very small, located on inner margin of A-sclerites. Ectophallic fold very short; ectophallic apodemes short and wide, thinner toward apex. Endophallic sclerite short, with well developed endophallic apodemes (both a high longitudinal crest and a distal lamella). No phallic glands. Female Unknown.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFF83528FCEBFAE9FC39BB71.taxon	description	(Figs 16 - 17; Table 11) urn: lsid: zoobank. org: act: C 0814161 - 69 A 9 - 49 CB-B 218 - D 5 A 764 E 9 EA 3 B	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFF83528FCEBFAE9FC39BB71.taxon	materials_examined	TYPE LOCALITY. — French Guiana, Monts Tumuc-Humac, Massif du Mitaraka. TYPE MATERIAL. — Holotype by present designation. French Guiana • 1 ♂; Monts Tumuc-Humac, Massif du Mitaraka vers sommet en Cloche; entre 54.4541 O 2.2349 N et 54.4646 O 2.2329 N; alt. entre 370 m et 470 m; 23. II. - 10. III. 2015; F. Legendre & S. Hugel leg.; fn. SH 546, de nuit; molecular sample LDG 557; MNHN-EO-ENSIF 9754. OTHER MATERIAL EXAMINED. — French Guiana • 1 juvenile; Monts Tumuc-Humac, Massif du Mitaraka, Layon A; entre 54.4509 O 2.2357 N et 54.4547 O 2.2405 N; alt. entre 280 m et 365 m; 23. II. - 10. III. 2015; F. Legendre & S. Hugel leg.; fn. SH 130, de nuit; molecular sample LDG 551; MNHN.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFF83528FCEBFAE9FC39BB71.taxon	etymology	ETYMOLOGY. — Species named after its origin in French Guiana.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFF83528FCEBFAE9FC39BB71.taxon	diagnosis	DIAGNOSIS. — As for the genus.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFF83528FCEBFAE9FC39BB71.taxon	biology_ecology	HABITAT. — The juvenile specimen has been collected by beating the leaf litter during the day, and the male has been found at night, in the leaf litter, in a windfall with dead branches. Mellomima guyanensis, n. gen., n. sp. could be a nocturnal straminicolous species.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFF83528FCEBFAE9FC39BB71.taxon	description	DESCRIPTION In addition to the morphological characters of the genus. General morphology TIII serrulation lacking between subapical spurs and apical spurs on both sides; on inner side, two spines between isa 1 and isa 2, three to four spines between isa 2 and isa 3, five spines between isa 3 and isa 4, 20 - 23 spines above isa 4; on outer side, three to four spines between osa 1 and osa 2, five spines between osa 2 and osa 3, six spines between osa 3 and osa 4, 16 - 19 spines above osa 4. Barsitarsomeres III serrulation: five inner spines and eight outer spines, in addition to apical spines. Coloration Head and pronotum reddish brown, except for yellow longitudinal band (Fig. 16 A, B, D). Maxillary palpi light brown with white lower and upper margins. Antennae yellow at base, light brown with white rings on flagellum length; scapes brown with yellowish ventral side (Fig. 16 D). Abdomen black brown, lighter dorsally without distinct yellow band or spot, except on tergite I. Sternites light brown. Cerci white basally, very light brown otherwise. FI, FII uniformly yellowish brown. TI, TII light reddish brown. FIII reddish brown, lighter at base; few unclear yellow spots on dorsal side (but see juvenile coloration). TIII yellowish brown, darker at base, with a dark ring and an irregular yellow ring near the knee; a small black brown spot at and near the base of each subapical spur; subapical spurs otherwise yellow with black apex; apical spurs yellowish with black apex. Tarsi yellowish to light yellowish brown. Male FWs not reaching tergite I distal margin (Fig. 16 A). FWs as on Figure 16 B, C; dorsal field venation with one faint, longitudinal vein. Supra anal plate as on Figure 16 G. Male genitalia As on Figure 17. Female Unknown. Juveniles Longitudinal yellow band running from labrum to supra anal plate. FIII light yellow on dorsal base; outer side striated with yellow; upper side with many yellow rounded spots. Metanotum and tergite I uniformly brown; tergites 2 – 6 lighter with dark spots and yellow lines near anterior margin. Measurements (in mm) See Table 11.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFFA3516FCCEFD4FFE56BA3C.taxon	type_taxon	TYPE GENUS. — Aclodes Hebard, 1928 b.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFFA3516FCCEFD4FFE56BA3C.taxon	discussion	REMARK Saussure (1874) defined the genus Heterogryllus for one female originating from Brazil, described as Heterogryllus ocellaris. Two other species have been described in the genus, Heterogryllus crassicornis Saussure, 1878 and Heterogryllus bordoni Chopard, 1970, transferred to Aclodes Hebard, 1928 and Paraclodes Desutter-Grandcolas, 1992 n. stat. respectively (see Desutter-Grandcolas 1992 b, 2014). In molecular-based phylogenetic analyses (Chintauan-Marquier et al. 2013, 2016; Warren et al. 2019), Aclodes and Paraclodes n. stat. constitute a well-separate clade within the Paragryllinae. According to the morphological characters used to define the ‘ Aclodae’ group of genera (Desutter-Grandcolas 1992 b), Uvaroviella Chopard, 1923 also belongs to this clade. Among other genera, Heterogryllus, Aclodes, Paraclodes n. stat. and Uvaroviella are presently classified in the subtribe Heterogryllina of the subfamily Phalangopsinae (Cigliano et al. 2020), a relationship not supported by the morphological characteristics of Heterogryllus (Saussure 1874, Desutter-Grandcolas pers. obs.). The tribe Aclodini Desutter-Grandcolas n. tribe is defined here for Aclodes, Paraclodes n. stat. and Uvaroviella.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFFA3516FCCEFD4FFE56BA3C.taxon	diagnosis	DIAGNOSIS. — After Desutter-Grandcolas (1992 b). Size small to medium. General coloration dark brown, marbled with black and light brown; face usually ornated with yellow marks; tibiae and apex of femura circled with yellow, outer side of femora striated with yellow. TI with a small, oval inner tympanum; no outer tympanum. TIII with 4 / 4 subapical spurs; with 2 / 3 apical spurs, inner ventral apical spur regressed. Basitarsomere III with only one row of dorsal spines (lost in Paraclodes bordoni (Chopard, 1970 )). Head small and vertical, longer than wide in front view; fastigium very narrow; article 5 of maxillary palpi little widened toward apex, truncated straight. Wings never developed in both males and females. Cerci and antennae very long. Male genitalia: pseudepiphallic sclerite well-developed but flat, more or less triangular; median lophi flat, prolonging pseudepiphallic sclerite; rami elongated; endophallic sclerite long and narrow; endophallic apodemes (both lateral lamellae and longitudinal median crest) little developed. Females: FWs shorter than in males or absent. Ovipositor straigth, compressed laterally, with short apex. Female genitalia: copulatory papilla very small and little sclerotized, never plicated nor greatly elongate. Aclodini Desutter-Grandcolas n. tribe is a very homogenous clade as far as morphology and male genitalia are concerned. Differences between genera and species relate mainly to male and female genitalia, size, length of FW and ovipositor, number of stridulatory teeth in males and face coloration. Aclodini Desutter-Grandcolas n. tribe can be superficially confused with other genera of Paragryllinae, from which they can be easily separated by the number of TIII subapical and apical spurs, spines of basitarsomere III, FW shape in males, and male and female genitalia.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFFA3516FCCEFD4FFE56BA3C.taxon	biology_ecology	HABITAT. — Aclodini Desutter-Grandcolas n. tribe are nocturnal species, active at night on tree trunks. They hide during the day in cavities such as burrows, hollow trees, or cave entrance when available (see Desutter-Grandcolas 1992 b).	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFFA3516FCCEFD4FFE56BA3C.taxon	discussion	INCLUDED GENERA. — Aclodes Hebard, 1928, Paraclodes Desutter-Grandcolas, 1992 n. stat., Uvaroviella Chopard, 1923 s. str. (see below). Gorochov (2007) subdivided Uvaroviella into nine subgenera (including Aclodes and Paraclodes), defined in an identification key mostly after non-discrete characters of male genitalia added with variable wing characters. These taxonomic entities, which are based on a very incomplete study of the characters of the taxa, are at least inadequate for taxonomic studies and their monophyly cannot be reasonably assessed. In the Orthoptera Species File Online, Aclodes is restored as a genus, but neither the position of all the species described in Aclodes and subsequently transferred in diverse Uvaroviella subgenera by Gorochov (2007 and later papers), nor the status of the other subgenera are reconsidered (see Cigliano et al. 2020). To stabilize the classification and avoid non-monophyletic taxa, we here follow Cigliano et al. for Aclodes and similarly restore Paraclodes as an independent genus, using the original definitions of the two genera: several subgenera are consequently synonymized with Aclodes or Paraclodes n. stat. The analysis of the variation of their morphological characters will have to be really performed to further study species relationships.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFC43516FE9BFCCFFC19B8CB.taxon	description	(Figs 18 - 20)	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFC43516FE9BFCCFFC19B8CB.taxon	type_taxon	TYPE SPECIES. — Aclodes nicuesa Hebard, 1928 b by original designation.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFC43516FE9BFCCFFC19B8CB.taxon	discussion	INCLUDED SPECIES. — Aclodes affinis (Gorochov, 2011), Aclodes andensis (Gorochov, 2007), Aclodes antennalis (Gorochov, 2007), Aclodes babyas (Otte & Perez-Gelabert, 2009), Aclodes bora Desutter-Grandcolas, 1992, Aclodes cavicola Chopard, 1954 (restored), Aclodes chamacoru (Nischk & Otte, 2000), Aclodes crassicornis (Saussure, 1878), Aclodes demissa (Gorochov, 2007), Aclodes dispar (Redtenbacher, 1892), Aclodes feredemissa (Gorochov, 2007), Aclodes finitima (Gorochov, 2007), Aclodes grandis Desutter-Grandcolas, 1992, Aclodes herpon Otte, 2006, Aclodes hypxyros (Nischk & Otte, 2000), Aclodes infuscata Desutter-Grandcolas, 1992, Aclodes iserskyi (Gorochov, 2011), Aclodes leleupae Chopard, 1970, Aclodes maculatum (Caudell, 1912), Aclodes meioptera (Gorochov, 2007), Aclodes mococharu Nischk & Otte, 2000, Aclodes morona (Gorochov, 2011), Aclodes nebulosa (Gorochov, 2007), Aclodes orchestes Otte, 2006, Aclodes parantennalis (Gorochov, 2007), Aclodes pastaza (Gorochov, 2007), Aclodes pequegna Desutter-Grandcolas, 1992, Aclodes pequegnita Desutter-Grandcolas, 1992, Aclodes rumococha Desutter-Grandcolas, 1992, Aclodes scandens Otte, 2006, Aclodes spelaea Desutter-Grandcolas, 1992, Aclodes turbidus Otte, 2006, Aclodes ucayali (Gorochov, 2011), Aclodes vittatum Chopard, 1937. Incertae sedis. Aclodes cryptos (Nischk & Otte, 2000).	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFC43516FE9BFCCFFC19B8CB.taxon	distribution	DISTRIBUTION. — Northern South America, Central America, Trinidad and Saint-Vincent (Cigliano et al. 2020).	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFC43516FE9BFCCFFC19B8CB.taxon	diagnosis	DIAGNOSIS. — See Hebard (1928 b) and Desutter-Grandcolas (1992 a, b).	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFC43514FC36FDACFC86BF6C.taxon	description	(Fig. 18; Table 12)	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFC43514FC36FDACFC86BF6C.taxon	materials_examined	TYPE LOCALITY. — French Guiana, Arataye, tributary to the Approuague river, 8 km NE from the saut Pararé, Nouragues research station. TYPE MATERIAL. — Holotype by original designation. French Guiana • 1 ♂; Arataye, Afflt. Approuague, 8 km NE saut Pararé, station de recherche des Nouragues; 10. VI. 1988; L. Desutter & P. Grandcolas leg.; abri sous roche; MNHN-EO-ENSIF 3061. Allotype. French Guiana • 1 ♀; same locality and collector as the holotype; 1. VI. 1988; abri sous roche; MNHN-EO-ENSIF 3062. Paratypes: 37 males, 25 females. French Guiana • 1 ♂ 3 ♀; same locality as the holotype; IV. 1988; L. Desutter leg.; MNHN-EO-ENSIF 5310, 5263, 5265, 5286 • 11 ♂ 4 ♀; same locality as the holotype; V. 1988; L. Desutter leg.; MNHN-EO-ENSIF 5272, 5264, 5284, 5285, 5289, 5291 - 5293, 5307, 5308, 5311 - 5315 • 5 ♂ 6 ♀; same locality and collector as the holotype; V. 1988; MNHN-EO-ENSIF 5266 - 5271, 5290, 5309, 5316, 3139, 3140 (both males recorded) • 15 ♂ 11 ♀; same locality and collector as the holotype;. VI. 1988; MNHN-EO-ENSIF 5273 - 5283, 5287, 5288, 5294 - 5305, 5306 • 3 ♂; same locality and collector as the holotype; VII. 1988; MNHN-EO-ENSIF 5317, 5318, 5320 • 1 ♂; same locality as the holotype; XI. 1989; P. Grandcolas leg.; MNHN-EO-ENSIF 5319 • 1 ♂ 1 ♀; same locality as the holotype; ZIN. MATERIAL FROM THE MITARAKA. — French Guiana • 2 ♂; Monts Tumuc-Humac, Massif du Mitaraka vers sommet en Cloche; entre 54.4541 O 2.2349 N et 54.4646 O 2.2329 N. alt. entre 370 m et 470 m; 23. II. - 10. III. 2015; F. Legendre & S. Hugel leg.; fn. SH 403, 404, de nuit; MNHN, • 1 juvenile; same data as for preceding; fn SH 405, de nuit; MNHN.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFC43514FC36FDACFC86BF6C.taxon	diagnosis	DIAGNOSIS. — See Desutter-Grandcolas (1992 a: 134, 1992 b: 177).	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFC43514FC36FDACFC86BF6C.taxon	description	COMPLEMENT OF DESCRIPTION General morphology TIII serrulation lacking between subapical spurs 1 and 2 on both sides; on inner side, no spine between isa 2 and isa 3, two to five spines (mean 3.75) between isa 3 and isa 4, 9 - 14 (mean 11.25) spines above isa 4; on outer side, one to two spines (mean 1.75) between osa 2 and osa 3, four spines between osa 3 and osa 4, 11 - 14 (mean 12.25) spines above osa 4. Barsitarsomeres III serrulation: no inner spine and two outer spines, in addition to apical spines. Coloration Face coloration as on Fig. 18 A. Male FWs as on Fig. 18 B. Metanotum with two rounded glandular areas (Fig. 18 C). Male genitalia As on Fig. 18 D-F. Measurements (in mm) See Table 12. Variation The two males from Mitaraka correspond to the description of A. spelaea, except for their larger size (see Table 12). The number of stridulatory teeth of their file (496 teeth, n = 1) fits the upper number found in the type series (385 - 495 teeth, mean 459, n = 11). Their FWs have the particular shape found in the species, but lateral part of dorsal field is much longer (Fig. 18 B). Their male genitalia are also similar, though larger (Fig. 18 D-F). Slight variations in coloration include, in one or both males, entirely dark LL, variously dark cerci, one or two yellow spots near anterior margin of supra anal plate, and entirely dark subgenital plate. CALLING SONG See Desutter-Grandcolas (1992 a) and recordings in the MNHN sound library (data reference: MNHN-SO- 2014 - 359 to 361).	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFC63512FF61F90BFD2ABF2C.taxon	description	(Figs 19, 20; Table 13)	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFC63512FF61F90BFD2ABF2C.taxon	materials_examined	TYPE LOCALITY. — French Guiana, Arataye, tributary to the Approuague river, 8 km NE from the saut Pararé, Nouragues research station. TYPE MATERIAL. — Holotype by original designation. French Guiana • 1 ♂; Arataye, Afflt. Approuague, 8 km NE saut Pararé, station de recherche des Nouragues; 8. VI. 1988; L. Desutter leg.; MNHN-EO-ENSIF 3063. Allotype. French Guiana • 1 ♀; same locality and collector as the holotype; 5. VI. 1988; MNHN-EO-ENSIF 3028. Paratypes: 2 males, 2 females. French Guiana • 1 ♀; same locality and collector as the holotype; V. 1988; MNHN-EO-ENSIF 5262 • 2 ♂ 1 ♀; same locality and collector as the holotype;. VI. 1988; MNHN-EO-ENSIF 5259 - 5261. MATERIAL FROM THE MITARAKA. — French Guiana • 1 ♂; Monts Tumuc-Humac, Massif du Mitaraka, Layon A; entre 54.4509 O 2.2357 N et 54.4547 O 2.2405 N; alt. entre 280 m et 365 m; 23. II. - 10. III. 2015; F. Legendre & S. Hugel leg.; de jour, fn. SH 127; MNHN • 1 ♀; same data as for preceding; de jour, fn SH 128; MNHN.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFC63512FF61F90BFD2ABF2C.taxon	description	COMPLEMENT OF DESCRIPTION General morphology TIII serrulation lacking between subapical spurs 1 and 2 on both sides; on inner side, one spine in male, zero to one spine in female between isa 2 and isa 3, two to three spines in male, two spines in female between isa 3 and isa 4, six spines in male, five to six spines in female above isa 4; on outer side, zero to one spine in male, no spine in female between osa 1 and osa 2, two spines in male, one spine in female between osa 2 and osa 3, three spines in male, two to three spines in female between osa 3 and osa 4, 12 - 13 spines in male, six to seven spines in female above osa 4. Barsitarsomeres III serrulation: zero to two inner spines in male, no inner spine in female; two outer spines in male, one to two outer spines in female, in addition to apical spines. Coloration Face (Fig. 20 A, B) with contrasting yellow pattern; last three articles of maxillary palpi (Fig. 20 C) brown; scapes brown with three yellow lines on basal half of inner side, the innermost with a series of small teeth. Pronotum and FWs as on Fig. 20 B, C: most lateral yellow line of vertex wider than the median one and extending on DD lateral margins. Male FWs (Fig. 20 B, C) reaching tergite 6. Male genitalia As on Fig. 20 D-F (compare with Desutter-Grandcolas 1992 b Fig. 25). Female Distance between FWs longer than FW width, but shorter than FW length. Measurements (in mm) See Table 13. Variation The Mitaraka specimens are larger than the specimens originating from the type locality. They fit however the description of A. pequegna, with the following variation: FW venation (Fig. 20 B, C) with two main veins in the harp, and six shorter ones (five in total in type material); stridulatory file with 282 teeth (304 in dissected paratype); lateral field with five longitudinal parallel veins (four in type material). In the male at hand, the genitalia are somewhat longer and more narrow than type genitalia; the posterior projection of pseudepiphallic distal margin is slightly more narrow and longer than in the Nourague material, and in lateral view, dorsal margin of pseudepiphallic lophi is less angulate (Fig. 20 D-F).	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFC03512FF1BF9C8FAABBBBA.taxon	description	(Figs 21 - 25)	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFC03512FF1BF9C8FAABBBBA.taxon	type_taxon	TYPE SPECIES. — Paraclodes guyanensis Desutter-Grandcolas n. nov. for Paraclodes aptera (Chopard, 1912), with which P. guyanensis Desutter-Grandcolas, 1992, the designated type species of Paraclodes, was synonymized (Desutter-Grandcolas 1994: 324). Homonymy with Paraclodes aptera (Giglio-Tos, 1897).	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFC03512FF1BF9C8FAABBBBA.taxon	discussion	INCLUDED SPECIES. — Paraclodes aptera (Giglio-Tos, 1897), Paraclodes bordoni (Chopard, 1970), Paraclodes cunicula Desutter-Grandcolas, n. sp., Paraclodes furcata Desutter-Grandcolas, n. sp., Paraclodes guyanensis Desutter-Grandcolas, n. stat., Paraclodes nouragui Desutter-Grandcolas, 1992, Paraclodes subaptera Gorochov, 2007.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFC03512FF1BF9C8FAABBBBA.taxon	distribution	DISTRIBUTION. — French Guiana, Venezuela, Panama.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFC03512FF1BF9C8FAABBBBA.taxon	diagnosis	DIAGNOSIS. — See Desutter-Grandcolas (1992 a, b).	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFC0351DFCC2FD4FFE7CBEEA.taxon	description	(Figs 21 - 23 A-C; Table 14) urn: lsid: zoobank. org: act: F 7 F 3 E 68 D- 9 ADB- 4 BAF-ADA 0 - A 8228196 F 61 F	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFC0351DFCC2FD4FFE7CBEEA.taxon	materials_examined	TYPE LOCALITY. — French Guiana, Monts Tumuc-Humac, Massif du Mitaraka. TYPE MATERIAL. — Holotype by present designation. French Guiana • 1 ♂; Monts Tumuc-Humac, Massif du Mitaraka, Layon A; entre 54.4509 O 2.2357 N et 54.4547 O 2.2405 N; alt. entre 280 m et 365 m; 23. II. - 10. III. 2015; F. Legendre & S. Hugel leg.; fn SH 115, de nuit; MNHN-EO-ENSIF 10778. Allotype. French Guiana • 1 ♀; Monts Tumuc-Humac, Massif du Mitaraka; 54.451125 O 2.234786 N; alt. 300 m; 23. II. - 10. III. 2015; F. Legendre & S. Hugel leg.; fn. SH 040, D 2, de nuit; MNHN-EO-ENSIF 10785. Paratypes: 6 males, 4 females. French Guiana • 1 ♂; same data as the holotype; fn SH 264; MNHN-EO-ENSIF 10779 • 1 ♀; same data as the allotype; fn SH 041; MNHN-EO-ENSIF 10786 • 2 ♂; Monts Tumuc-Humac, Massif du Mitaraka, D 2; 54.451125 O 2.234786 N; alt. 300 m; 23. II. - 10. III. 2015; F. Legendre & S. Hugel leg.; fn. SH 071, SH 073, de nuit; MNHN-EO-EN-SIF 10780, 10781 • 2 ♂ 3 ♀; Monts Tumuc-Humac, Massif du Mitaraka, vers sommet en Cloche; entre 54.4541 O 2.2349 N et 54.4646 O 2.2329 N; alt. entre 370 m et 470 m; 23 II. et 10. III. 2015; F. Legendre & S. Hugel leg.; fn. SH 421, SH 462, SH 535, SH 461, SH 460, de nuit; MNHN-EO-ENSIF 10782, 10783, 10787 - 10789 • 1 ♂; Monts Tumuc-Humac, Massif du Mitaraka, entre C 1000 et savane roche; entre 54.4419 O 2.23366 N et 54.4346 O 2.238655 N; alt. entre 415 m et 390 m; 23. II. - 10. III. 2015; F. Legendre & S. Hugel leg.; fn. SH 232, de nuit; MNHN-EO-ENSIF 10784.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFC0351DFCC2FD4FFE7CBEEA.taxon	etymology	ETYMOLOGY. — Species named after the very long size of the pseudepiphallic lobes in male genitalia, making like a wide fork.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFC0351DFCC2FD4FFE7CBEEA.taxon	diagnosis	DIAGNOSIS. — Within the genus, species very close to Paraclodes nouragui Desutter-Grandcolas, 1992 n. stat., and P. subaptera Gorochov, 2007 n. stat., but fully apterous and with distinct male and female genitalia, and coloration. It can be separated from P. nouragui n. stat. by a bigger size, the lack of FWs in males, male genitalia (pseudepiphallic sclerite more narrow, pseudepiphallic apical lobes very long, slightly divergent, not crenulated at base; pseudepiphallic parameres more rounded and vertical), female ovipositor length and copulatory papilla (much bigger, higher, with a smaller aperture dorsally). P. furcata Desutter-Grandcolas, n. sp. can be separated from P. subaptera n. stat. by its male genitalia (pseudepiphallic apical lobes longer, distinctly narrowed and curved toward apex, separated by a very narrow distance at base, but this distance increasing sharply close to apical lobes base (Fig. 21 D, E), pseudepiphallic parameres rounded and vertical, but half the size of P. subaptera n. stat.) and its female genitalia (higher, with a smaller aperture dorsally).	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFC0351DFCC2FD4FFE7CBEEA.taxon	description	DESCRIPTION In addition to the characters of the genus. General morphology Maxillary palpi not very elongated (Fig. 21 A); article 3 shorter than article 4, article 5 the longest. DD posterior margin largely concave (Fig. 21 B). TI with a small inner tympanum; outer tympanum absent. FIII with filiform apical part. TIII lacking spines between subapical spurs and apical spurs on both sides; on inner side, no spine in males and females between isa 1 and isa 2, zero to one spine (mean 0.1) in males, zero to two spines (mean 1) in females between isa 2 and isa 3, two to four spines (mean 2.7) in males, three to four spines (mean 3.6) in females between isa 3 and isa 4, six to eight spines (mean 7.1) in males, nine to ten spines (mean 9.4) in females above isa 4; on outer side, one to two spines (mean 1.9 in males, 1.6 in females) between osa 1 and osa 2, two to four spines (mean 3 in males and females) between osa 2 and osa 3, three to six spines (mean 4.9) in males, three to seven spines (mean 5) in females between osa 3 and osa 4, 12 - 15 spines (mean 13.4) in males, 14 - 15 spines (mean 14.4) in females above osa 4. Barsitarsomere III serrulation: no inner spines in males and females; three to four outer spines (mean 3.3) in males, three to five outer spines (mean 3.4) in females in addition to apical spines. Coloration Face (Fig. 21 A) with a yellow line below median ocellus, bordered with a black line and separated from antennal pit by light brown; face light to dark brown, with two median, wide yellow spots and a small yellow spot below each antennal pit; an additional yellow spot, variable in size and color, close to inner angle of eye. Cheeks yellow, sometimes separated from subgena by light brown; a brown spot behind posterior margin of eye. Fastigium black dorsally, more or less marked with light brown or yellowish laterally (Fig. 21 A). Head dorsum black; vertex with four thin yellow longitudinal lines, the most lateral ones joining a wider yellow band behind the eyes from which it is separated by a triangular brown spot (Fig. 21 B). Scapes yellow, upper margin black brown; inner side darker with more or less delimited yellow spots. Antennae light brown. Maxillary palpi light yellow, last three articles marked with brown on lateral sides (Fig. 21 A). Pronotum: anterior margin yellow; DD (Fig. 21 B) with wide longitudinal yellow band, an irregular yellow band on each lateral margin, two and four black spots near anterior and posterior margins respectively, pyriform inscriptions ochre; LL black, anterior angle largely yellowish (Fig. 21 C). FI, FII (Fig. 21 B, C) yellow, inner and outer lower margins black brown; two black rings in distal half; a large black patch on each side and several small black spots dorsally and laterally. Metanotum with a yellow, longitudinal band, bordered by two lateral black brown spots, prolonged on tergites I and II by a very thin line (Figs 21 B; 22). Legs as on Fig. 22: TI, TII yellow, lower side light ochre; four incomplete black rings. FIII yellow with black brown pattern: outer side with oblique black stripes and a longitudinal black line close to lower margin; two black rings on apical filiform part; inner side less marked with dark brown. TIII yellow, light ochre ventrally; with four incomplete black rings. Spurs and claws yellow with black apex. Tarsomeres 1 brown with yellow base; tarsomeres 2 yellowish; tarsomeres 3 brown. Abdomen variegated yellow, brown and black (Fig. 22). Cerci yellowish, densely covered with black setae. Male Antennae with bunches of long setae. Apterous. Supra anal plate brown with two transverse yellowish spots at base. Subgenital plate brown. Male genitalia (Fig. 21 D-F) Pseudepiphallic sclerite distinctly long and narrow, as long as rami, with abruptly raised base. Pseudepiphallic apical lobes very long, as long as pseudepiphallic sclerite, regularly narrowed toward apex; regularly curved on outer side; inner side abruptly narrowed close to base, the indentation between lobes sharp and of regular width. Pseudepiphallic parameres with large lower part and small upper part; lower part lamella-like, almost vertical in side view, triangular in ventral view. Ectophallic apodemes only slightly longer than endophallic sclerite; apex abruptly widened, as in P. nouragui n. stat .. Endophallic sclerite and apodeme as in P. nouragui n. stat. and P. subaptera n. stat. Female Apterous, without even flap-like FWs laterally. Supra anal plate brown with two transverse yellowish spots at base. Subgenital plate short, transverse; distal margin straight, not sinuate. Ovipositor very long, much longer than FIII and TIII; apex outer sides crenulated. Female genitalia (Fig. 23 A-C) Copulatory papilla broadly triangular in side view, higher laterally than the papilla of P. nouragui n. stat. and P. subaptera n. stat.; dorsal aperture smaller than in the last two species, and not as wide as the papilla over its whole length; papilla more narrow apically than basally in dorsal view (compare Fig. 23 A-C and D-F). Measurements (in mm) See Table 14. Variation Face most often lighter in males than in females (depending on the condition of the specimens?). LL sometimes wholly black, without yellow near lower margin. Fastigium coloration from yellowish to dark brown.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFCF351CFCD1FAA9FDA7BEEB.taxon	description	(Fig. 24; Table 15)	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFCF351CFCD1FAA9FDA7BEEB.taxon	materials_examined	TYPE LOCALITY. — French Guiana, Saint-Laurent-du-Maroni. TYPE MATERIAL OF LARANDA APTERA. — Holotype by original designation. French Guiana • 1 female juvenile; St Laurent du Maroni; collection Le Moult; MNHN-EO-ENSIF 6827. TYPE MATERIAL OF PARACLODES GUYANENSIS. — Holotype by original designation. French Guiana • 1 ♂; Arataye, Afflt. Approuague, 8 km NE saut Pararé, station de recherche des Nouragues; 9. VI. 1988; L. Desutter leg.; MNHN-EO-ENSIF 3042. Allotype. French Guiana • 1 ♀; same locality, date and collector as the holotype; MNHN-EO-ENSIF 5422. Paratypes: 66 males, 46 females. French Guiana • 7 ♂ 5 ♀; same locality and collector as the holotype; IV. 1988; MNHN-EO-ENSIF 5428, 5429, 5434, 5439, 5442, 5476, 5481, 5484, 5486, 5506 - 5508 • 3 ♂ 3 ♀; same locality and collector as the holotype; V. 1988; MNHN-EO-ENSIF 5430, 5431, 5436, 5479, 5492, 5500 • 19 ♂ 8 ♀; same locality as the holotype; VI. 1988; L. Desutter & P. Grandcolas leg.; MNHN-EO-ENSIF 5424, 5426, 5427, 5433, 5435, 5443, 5454, 5469, 5475, 5478 - 5480, 5482, 5483, 5485, 5487 - 5491, 5493, 5495, 5496, 5501, 5502, 5504, 5505 • 4 ♂ 6 ♀; same data as preceding; VII. 1988; MNHN-EO-ENSIF 5425, 5437, 5438, 5440, 5441, 5544, 5494, 5497, 5498, 5503 • 1 ♀; same locality as the holotype; IX. 1989; P. Grandcolas leg.; MNHN-EO-ENSIF 5432 • 16 ♂ 11 ♀; Arataye, Afflt. Approuague, aval du saut Pararé; VII. 1988; L. Desutter & P. Grandcolas leg.; MNHN-EO-ENSIF 5509 - 5524, 5423, 5455 - 5464 • 4 ♂ 1 ♀; Sinnamary, Paracou, forêt sur sable blanc: VIII. 1988; L. Desutter & P. Grandcolas leg.; MNHN-EO-ENSIF 5526 - 5529, 5465 • 2 ♂ 3 ♀; same data as preceding; IX. 1988; MNHN-EO-ENSIF 5525, 5530, 5466 - 5468 • 8 ♂ 1 ♀; Sinnamary, Piste de St Elie, PK 15; VIII. 1988; L. Desutter & P. Grandcolas leg.; MNHN-EO-ENSIF 5531 - 5538, 5474 • 1 ♂; Sinnamary, crique Grégoire; XI. 1968; A. Delplanque & J. Bonfils leg.; MNHN-EO-ENSIF 5539 • 1 ♂ 4 ♀; Saül; VIII. 1988; L. Desutter & P. Grandcolas leg.; MNHN-EO-ENSIF 5540, 5470 - 5473 • 1 ♂ 1 ♀; ZIN. MATERIAL FROM THE MITARAKA. — French Guiana • 1 ♂; Monts Tumuc-Humac, Massif du Mitaraka, Layon A; entre 54.4509 O 2.2357 N et 54.4547 O 2.2405 N; alt. entre 280 m et 365 m; 23. II. - 10. III. 2015; F. Legendre & S. Hugel leg.; SH 265, de nuit; MNHN • 1 ♀; same data as for preceding; fn SH 311, de nuit; MNHN • 1 ♂; Monts Tumuc-Humac, Massif du Mitaraka, D 2; 54.451125 O 2.234786 N; alt. 300 m; 23. II. - 10. III. 2015; F. Legendre & S. Hugel leg.; fn. SH 074, de nuit; MNHN.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFCF351CFCD1FAA9FDA7BEEB.taxon	discussion	REMARK Specimens (see Fig. 24) fitting the description of P. guyanensis for their general shape and male FW size and venation (Desutter-Grandcolas 1992 b, figure 37), face coloration (Desutter-Grandcolas 1992 b, figure 41), the structure of male genitalia (Desutter-Grandcolas 1992 b, figure 38), and female copulatory papilla (Desutter-Grandcolas 1992 b, figure 44). They differ by their larger size (compare measurements) and number of stridulatory teeth (n = 118, n = 1, against 60 - 83 teeth in the specimens from the type locality), and a very short yellow line between median ocellus and the face (in the place of two yellow spots in type material). Male and female genitalia are too similar to P. guyanensis to consider these specimens as a different species, and the lack of samples through the guianese territory impedes a correct appreciation of species variation.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFCF351CFCD1FAA9FDA7BEEB.taxon	description	MEASUREMENTS (IN MM). — See Table 15.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFCE351CFCC9FE6CFB4FBF3D.taxon	description	(Fig. 23 D-F; Table 16)	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFCE351CFCC9FE6CFB4FBF3D.taxon	materials_examined	TYPE LOCALITY. — French Guiana, Arataye, Approuague tributary, Nourragues Research Station. TYPE MATERIAL. — Holotype. French Guiana • 1 ♂; Arataye, Afflt. Approuague, 8 km NE saut Pararé, station de recherche des Nouragues, abri sous roche; 13. IV. 1988; L. Desutter leg.; MNHN-EO-ENSIF 5413. Allotype. French Guiana • 1 ♀; same locality and collector as the holotype; 4. IV. 1988; MNHN-EO-ENSIF 5414. Paratypes: 5 males, 2 females. French Guiana • 4 ♂ 1 ♀; same locality and collector as the holotype; IV. 1988; MNHN-EO-EN-SIF 5415 - 5417, 5419, 5420 • 1 ♀; same locality as the holotype; V. 1988; L. Desutter & P. Grandcolas leg.; MNHN-EO-ENSIF 5421 • 1 ♂; same locality as the holotype; X. 1988; P. Grandcolas leg.; MNHN-EO-ENSIF 5418. MATERIAL FROM THE MITARAKA. — French Guiana • 1 ♀; Monts Tumuc-Humac, Massif du Mitaraka, entre C 100 et C 1000; entre 54.44768 O 2.235494 N et 54.4419 O 2.233664 N; alt. entre 350 m et 415 m; 23. II. - 10. III. 2015; F. Legendre & S. Hugel leg.; fn. SH 177, de nuit; MNHN.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFCE351CFCC9FE6CFB4FBF3D.taxon	discussion	REMARK Specimen fitting the description of P. subaptera n. stat. by its face coloration (Desutter-Grandcolas 1992 b, fig. 43), the presence of very short lobe-like FWs, the relative size of ovipositor and FIII (but ovipositor longer, see Table 16) and the shape of copulatory papilla (compare Fig. 23 D-F and Desutter-Grandcolas 1992 b, figure 46). Its size extends the variation of the species, up to now known by type locality only.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFCE351CFCC9FE6CFB4FBF3D.taxon	description	MEASUREMENTS (IN MM). — See Table 16.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFCE3519FCDBF9C8FE91BD0E.taxon	description	(Fig. 23 G-I, 25; Table 17)	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFCE3519FCDBF9C8FE91BD0E.taxon	materials_examined	TYPE LOCALITY. — French Guiana, Monts Tumuc-Humac, Massif du Mitaraka. TYPE MATERIAL. — Holotype by present designation. French Guiana • 1 ♂; Monts Tumuc-Humac, Massif du Mitaraka, vers sommet en Cloche; entre 54.4541 O 2.2349 N et 54.4646 O 2.2329 N; alt. entre 370 m et 470 m; 23. II. - 10. III. 2015; F. Legendre & S. Hugel leg.; fn. SH 406, de nuit; MNHN-EO-ENSIF 10790. Allotype. French Guiana • 1 ♀; same data as the holotype; fn. SH 409; MNHN-EO-ENSIF 10791. Paratypes: 2 females. French Guiana • 2 ♀; same data as the holotype; fn SH 408, SH 459; MNHN-EO-ENSIF 10792, 10793. Type material more or less covered with fungi, not in good condition.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFCE3519FCDBF9C8FE91BD0E.taxon	etymology	ETYMOLOGY. — Species named after the shape of pseudepiphallic apical lobes in male genitalia, which resembles the ears of a rabbit (cunicula, in Latin). Name in apposition.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFCE3519FCDBF9C8FE91BD0E.taxon	diagnosis	DIAGNOSIS. — Within the genus, small species close to P. subaptera n. stat. by the lack of developed FWs in males and females, and by the general shape of male and female genitalia. It is characterized by its male genitalia (pseudepiphallic sclerite long and very narrow, more so than in P. subaptera n. stat.; pseudepiphallic lobes very long and wide, twisted over their length, resembling rabbit ears; pseudepiphallic parameres short, the lower part narrowed toward apex, the upper part short and broadly squared), its low number of dorsal spines on TIII and basitarsomere III, and its female genitalia (lateral margins rounded in dorsal view, dorsal aperture narrowed distally).	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFCE3519FCDBF9C8FE91BD0E.taxon	description	DESCRIPTION In addition to the characters of the genus (Desutter-Grandcolas 1992 b). General morphology Size small. Maxillary palpi not very elongated (Fig. 25 A); article 3 shorter than article 4, article 5 the longest. DD posterior margin largely concave (Fig. 25 C). TI with a small inner tympanum; outer tympanum absent. FIII with filiform apical part. TIII lacking spines between subapical spurs and apical spurs on both sides; on inner side, no spine in male and females between isa 1 and isa 2, and between isa 2 and isa 3, three spines in male, two spines in females between isa 3 and isa 4, five spines in male, three to five spines (mean 4.2) in females above isa 4; on outer side, one spine in male and females between osa 1 and osa 2, two spines in male and females between osa 2 and osa 3, five spines in male, four to five spines (mean 4.4) in females between osa 3 and osa 4, eight to ten spines (mean 9) in male, nine to 12 spines (mean 10.2) in females above osa 4. Barsitarsomere III serrulation: no inner spines in male and females; two outer spines in male, three outer spines in females, in addition to apical spines. Coloration Face (Fig. 25 A) variably yellow, from almost fully yellow below antennal pits to light brown with large yellow spots and indistinct brownish areas close to epistomal suture and below anterior angles of eye; with a yellow line below median ocellus, bordered with a black line and separated from antennal pit by a short light yellow and light brown coloration. Cheeks light yellowish, with light brown area behind posterior margin of eye. Fastigium black (Fig. 25 C), ocelli ivory. Head dorsum black; vertex with five indistinct yellow longitudinal lines (Fig. 25 C). Anterior side of scapes with a transverse yellow spot, surrounded with black brown. Antennae brown, apical margin of articles yellow. Maxillary palpi very light yellow or brown, last three articles marked with brown on lateral sides (Fig. 25 A). Pronotum: anterior margin light brown; DD (Fig. 25 C) with a wide longitudinal yellow band over its whole length, bordered by two wide black spots close to anterior margin, and an irregular yellow band on each lateral margin; DD otherwise light brown, black brown and black, with yellowish posterior margin; LL black, lower margin bordered with light brown (Fig. 25 B). Metanotum with a yellow, longitudinal band, bordered by two lateral black brown spots, prolonged on tergites I and II by a very thin line. Legs: TI, TII yellow, lower side light ochre; four black rings. FI, FII light yellow, inner and outer lower margins black; one and two (respectively) black rings in distal half; a large black patch on each side and several small black spots dorsally and laterally. FIII yellow with black brown pattern: outer side with many oblique black stripes and a longitudinal black line close to lower margin; two black rings on apical filiform part; inner side less marked with dark brown. TIII black brown with a bright yellow transverse line close to the knee; irregular yellow flecks on the sides, more or less prolonged dorsally. Spurs and claws yellowish with brown apex. Tarsomeres 1 brown with yellow base; tarsomeres 2 and 3 light brown. Cerci light brown. Male Antennae without bunches of strong and long setae. Apterous. Supra anal plate brown with two indistinct transverse yellowish spots at base. Subgenital plate brown; very long, going beyond supra anal plate distal margin by half of its length. Male genitalia (Fig. 25 D-F) Pseudepiphallic sclerite very narrow dorsally, abruptly raised anteriorly. Pseudepiphallic apical lobes very wide and flat, separate by a narrow U-shaped depression. Pseudepiphallic parameres small; lower part wider on its outer side, acute distally; upper part close to midline, broadly squared. Rami wide, slightly concave, widened anteriorly and posteriorly. Ectophallic apodemes reaching only rami mid length; apex abruptly widened, triangular and acute. Female Apterous, without even flap-like FWs laterally. Supra anal plate brown with yellowish base. Subgenital plate short, transverse; distal margin straight, not sinuate. Ovipositor very short, shorter than FIII; apex outer sides very faintly crenulated, almost smooth. Female genitalia (Fig. 23 G-I). Copulatory papilla small, close to that of P. subaptera n. stat., but apical two-third of the papilla more rounded than quadrangular in dorsal view, lower and straighter in side view; dorsal aperture regularly narrowed toward distal margin; apex small, with a distinct sclerotized edge. Measurements (in mm) See Table 17.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFCB3519FC79FF6DFA8CB8D1.taxon	type_taxon	TYPE SPECIES. — Ectecous hedyphonus Saussure, 1878 by original monotypy.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFCB3519FC79FF6DFA8CB8D1.taxon	distribution	DISTRIBUTION. — Northeastern South America (French Guiana, Brazil).	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFCB3519FC79FF6DFA8CB8D1.taxon	diagnosis	DIAGNOSIS. — See Desutter-Grandcolas (1992 a).	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFCB3519FEBAF8EBFDA6BEE9.taxon	type_taxon	TYPE GENUS. — Neoacla Desutter, 1988.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFCB3519FEBAF8EBFDA6BEE9.taxon	diagnosis	DIAGNOSIS. — See Desutter (1987, 1988).	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFCB3519FECEFBEEFDA7BFDD.taxon	description	(Fig. 23 J-L; Table 18)	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFCB3519FECEFBEEFDA7BFDD.taxon	materials_examined	MATERIAL EXAMINED. — French Guiana • 1 ♀; Monts Tumuc-Humac, Massif du Mitaraka, vers sommet en Cloche; entre 54.4541 O 2.2349 N et 54.4646 O 2.2329 N; alt. entre 370 m et 470 m; 23. II. - 10. III. 2015; F. Legendre & S. Hugel leg.; de nuit, fn. SH 407; MNHN.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFCB3519FECEFBEEFDA7BFDD.taxon	discussion	REMARK We separate here one female which differs by its very short ovipositor (less than 16 mm), its female copulatory papilla (short, rounded with a laterally strongly compressed apex: see Fig. 23 J-L), and its size (see Measurements). The specimen is in very bad condition, but clearly belongs to a new species that will have to be described with additional material.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFCB3519FECEFBEEFDA7BFDD.taxon	description	MEASUREMENTS (IN MM). — See Table 18.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFCB3518FCD6FDACFC38BEEA.taxon	description	(Figs 26, 27; Table 19) urn: lsid: zoobank. org: act: BCBABC 04 - B 038 - 4 D 9 F-B 2 E 0 - 8463 E 9 B 6 D 069	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFCB3518FCD6FDACFC38BEEA.taxon	materials_examined	TYPE LOCALITY. — French Guiana, Monts Tumuc-Humac, Massif du Mitaraka. TYPE MATERIAL. — Holotype by present designation. French Guiana • 1 ♂; Monts Tumuc-Humac, Massif du Mitaraka, D 2; 54.451125 O 2.234786 N; alt. 300 m; 23. II. - 10. III. 2015; F. Legendre & S. Hugel leg.; fn. SH 028, de nuit; MNHN-EO-ENSIF 10794. Paratypes: 2 males. French Guiana • 2 ♂; Monts Tumuc-Humac, Massif du Mitaraka, vers sommet en Cloche; entre 54.4541 O 2.2349 N et 54.4646 O 2.2329 N; alt. entre 370 m et 470 m; 23. II. - 10. III. 2015; F. Legendre & S. Hugel leg.; fn. SH 422, SH 423, de nuit; MNHN-EO-ENSIF 10795, 10796.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFCB3518FCD6FDACFC38BEEA.taxon	etymology	ETYMOLOGY. — Species named after the lamella-shape of pseudepiphallic apical lobes in male genitalia.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFCB3518FCD6FDACFC38BEEA.taxon	diagnosis	DIAGNOSIS. — Species very close to E. tenebrosus Desutter-Grandcolas, 1992 by its size, coloration, male FW size and venation, but separated from that species by male genitalia (as in E. tenebrosus, except for lamella-shaped dorsal valves), its slightly more narrow FWs (length / width ratio 1.42, n = 3, against 1.53, n = 6, in E. tenebrosus), with fewer stridulatory teeth (232 against 259 - 264, n = 6, mean 262, in E. tenebrosus).	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFCB3518FCD6FDACFC38BEEA.taxon	description	DESCRIPTION In addition to the character of the genus. General morphology Size only slightly bigger than E. tenebrosus. Fastigium, ocelli and palpi as in E. tenebrosus; with lateral ocelli separate by a distance shorter than their own width. Coloration General coloration as on Fig. 26. Face as in E. tenebrosus: black, with reversed Y-shaped line under median ocellus, a small yellow spot under antennal pit (in an area with many long setae), and a wider yellow area under lower angle of eye; but median yellow line thinner between antennal pits and spot under anterior angle of eye smaller (Fig. 27 A). Maxillary palpi as in E. tenebrosus, but article 5 darker (Fig. 27 A). Vertex (Fig. 27 B) as in E. tenebrosus, with the most lateral yellow lines (behind the eyes) prolonged as a semi-circular line on the occiput (unnoticed in E. tenebrosus description). Pronotum (Fig. 27 B, C) and legs (Fig. 26) as in E. tenebrosus. Male FWs covering abdomen up to half supra anal plate, as in E. tenebrosus. Venation (Fig. 27 D): harp with 6 – 8 veins, mirror crossed by 4 – 5 concentric veins. Stridulatory file with 232 teeth (n = 1). Subgenital plate short, truncated apically. Distal margin of supra anal plate with many long and thick setae (Fig. 27 E). Male genitalia (Fig. 27 F-H) Pseudepiphallic sclerite, rami, ectophallic apodemes, ectophallic fold (short and bifid) and endophallic apodeme as in E. tenebrosus (see Desutter-Grandcolas 1992 b, Figs 59 - 61), but dorsal valves wide and thick, having the shape of nearly vertical lamellas (Fig. 27 H), with sharp, curved denticles on lower margin (Fig. 27 G); median lophi (= apical lobes) entirely membranous (Fig. 27 F, G), convex on outer side, concave with abruptly vertical base on inner side. Female Unknown. Measurements (in mm) See Table 19.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFD43506FF63FE6CFDE4BBBD.taxon	type_taxon	TYPE SPECIES. — Kevanacla orientalis Desutter-Grandcolas, 1992 a: 142 by original designation.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFD43506FF63FE6CFDE4BBBD.taxon	distribution	DISTRIBUTION. — Northeastern South America (French Guiana).	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFD43506FF63FE6CFDE4BBBD.taxon	diagnosis	DIAGNOSIS. — See Desutter-Grandcolas (1992 a).	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFD43506FF70FD4FFC36B8D0.taxon	description	(Figs 28, 29 A, B; Table 20)	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFD43506FF70FD4FFC36B8D0.taxon	materials_examined	TYPE LOCALITY. — French Guiana, Arataye, tributary of Approuague, downstream the saut Pararé. TYPE MATERIAL. — Holotype by original designation. French Guiana • 1 ♂; Arataye, Afflt. Approuague, aval du saut Pararé; 18. VII. 1988; L. Desutter leg.; MNHN-EO-ENSIF 5189. Allotype. French Guiana • 1 ♀; same locality, date and collector as the holotype; MNHN-EO-ENSIF 5190. Paratypes: 1 male, 2 females. French Guiana • 1 ♀; Arataye, Afflt. Approuague, 8 km pied du saut Pararé, station de recherche des Nouragues; VI. 1988; L. Desutter & P. Grandcolas leg.; MNHN-EO-ENSIF 5193 • 1 ♀; same locality and collector; VII. 1988; MNHN-EO-ENSIF 5192. Brazil • 1 ♂; Haut Carsevenne; 1898; F. Geai leg.; MNHN-EO-ENSIF 5191. MATERIAL FROM THE MITARAKA. — French Guiana • 1 ♀; Monts Tumuc-Humac, Massif du Mitaraka, Layon A; entre 54.4509 O 2.2357 N et 54.4547 O 2.2405 N; alt. entre 280 m et 365 m; 23. II. - 10. III. 2015; F. Legendre & S. Hugel leg.; de nuit, SH 129; MNHN.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFD43506FF70FD4FFC36B8D0.taxon	diagnosis	DIAGNOSIS. — See Desutter-Grandcolas (1992 a).	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFD43506FF70FD4FFC36B8D0.taxon	description	COMPLEMENT OF DESCRIPTION General morphology TIII serrulation: spines lacking between subapical spurs and apical spurs, and between subapical spurs 1 and 2 on both sides; on inner side, one spine between isa 2 and isa 3, 15 spines above isa 3; on outer side, three spines between osa 2 and osa 3, two spines between osa 3 and osa 4, 17 spines above osa 4. Barsitarsomere III serrulation: no inner spine and four outer spines, in addition to apical spines. Coloration General coloration as on Figs 28, 29 A. Antennae black brown; yellow ring present every five to seven articles, not on the distal margin of all articles. Measurements (in mm) See Table 20. Variation Maxillary palpi: dorsal part of article 3 lighter than its sides, but not as light as on articles 4 and 5 (Fig. 29 A). LL anterior angle with a small yellowish spot. DD coloration pattern as on Fig. 29 A.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFD43506FC38F968FCFEBEE9.taxon	type_taxon	TYPE SPECIES. — Paragryllus martinii Guérin-Méneville, 1844 by original monotypy.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFD43506FC38F968FCFEBEE9.taxon	diagnosis	DIAGNOSIS. — See Desutter-Grandcolas (1992 a).	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFD43506FC38F968FCFEBEE9.taxon	distribution	DISTRIBUTION. — Tropical areas in Africa and South and Central America, including Trinidad. Only one species known in French Guiana.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFD43506FB97FC2FFBB3BF4D.taxon	materials_examined	MATERIAL FROM THE MITARAKA. — French Guiana • 1 juvenile ♂; Monts Tumuc-Humac, Massif du Mitaraka vers sommet en Cloche; entre 54.4541 O 2.2349 N et 54.4646 O 2.2329 N; alt. entre 370 m et 470 m; 23. II. - 10. III. 2015; F. Legendre & S. Hugel leg.; de nuit, fn. SH 499; MNHN. Specimen in bad condition (fungi).	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFD43506FB97FC2FFBB3BF4D.taxon	discussion	REMARK Unithema is the only strogulomorphine genus recorded today in French Guiana. As the juvenile male at hand present very short FWs, it could belong to this genus. The specimen has a very dark coloration: its pronotum is almost entirely black, without a yellow band along posterior margin, and with only a very thin longitudinal line on DD; its palpi, abdomen and cerci are entirely black. It shows also particular details of coloration, such as alternate black and ivory lines on scape anterior side, a yellow band on posterior margin of metanotum (as in Unithema xanthochosmea Desutter-Grandcolas, 1991), black antennae with yellow articles widely spaced on flagellum, and a thin ivory line under median ocellus, bordered by two yellowish lines along antennal pits. This specimen could belong to a new species.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFD43506FCCCFDECFA33BA53.taxon	type_taxon	TYPE SPECIES. — Unithema guadelupensis Desutter-Grandcolas, 1991: 358 by original designation.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFD43506FCCCFDECFA33BA53.taxon	diagnosis	DIAGNOSIS. — See Desutter-Grandcolas (1991).	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFD43506FCCCFDECFA33BA53.taxon	distribution	DISTRIBUTION. — French Guiana and Lesser Antilles (Guadeloupe).	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFD73503FF78FB69FDA7BA1E.taxon	description	(Figs 23 M-O, 29 C; Table 21)	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFD73503FF78FB69FDA7BA1E.taxon	discussion	REMARK Two subspecies have been separated in P. elapsus (see Desutter-Grandcolas 1992 a) to acknowledge variations in size, coloration, male stridulatory file, female ovipositor length and female copulatory papilla. The only female at hand (Fig. 29 C) does not fully fit neither of the subspecies, and its status will have to be reconsidered with additional specimens.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFD73503FF78FB69FDA7BA1E.taxon	materials_examined	TYPE LOCALITY. — French Guiana, aval du saut Pararé (Paragryllus elapsus elapsus Desutter-Grandcolas, 1992) and Sinnamary (Paragryllus elapsus incertus Desutter-Grandcolas, 1992). TYPE MATERIAL. — Paragryllus elapsus elapsus Desutter-Grandcolas, 1992: Holotype by original designation. French Guiana • 1 ♂; Arataye, aval du saut Pararé; 9. VII. 1988; L. Desutter leg.; MNHN-EO-ENSIF 5138. Allotype. French Guiana • 1 ♀; same locality and collector as the holotype; 12. VII. 1988; MNHN-EO-ENSIF 5139. Paratypes: 1 male, 1 female: French Guiana • 1 ♂ 1 ♀; same locality and collector as the holotype; VII. 1988; MNHN-EO-ENSIF 5140, 5141. Paragryllus elapsus incertus Desutter-Grandcolas, 1992: Holotype by original designation. French Guiana • 1 ♂; Sinnamary; VII. 1977; M. Descamps leg.; MNHN-EO-ENSIF 5142. Allotype. French Guiana • 1 ♀; same locality, date and collector as the holotype; MNHN-EO-ENSIF 5143. Paratype, 1 male: French Guiana • 1 ♂; same locality, date and collector as the holotype; MNHN-EO-ENSIF 5144. MATERIAL FROM THE MITARAKA. — French Guiana • 1 ♀; Monts Tumuc-Humac, Massif du Mitaraka vers sommet en Cloche; entre 54.4541 O 2.2349 N et 54.4646 O 2.2329 N; alt. entre 370 m et 470 m; 23. II. - 10. III. 2015; F. Legendre & S. Hugel leg.; fn. SH 458, de nuit; MNHN.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFD73503FF78FB69FDA7BA1E.taxon	discussion	REMARK Compared to the two subspecies, the female at hand differs by the small size of its ovipositor, well shorter than FIII and TIII (longer in P. elapsus elapsus and P. elapsus incertus), its larger size, the shape of its copulatory papilla (short, not concave laterally, bisinuous distally, Fig. 23 M, N) and details of coloration, such as area between eyes light yellow, this coloration prolonged on occiput; LL black, with a short median yellow line along lower margin; DD yellowish marked with brown and black, with a wide longitudinal, median yellow band bordered distally by a pair of Y-shaped black spots, muscular inscriptions marked with black (Fig. 29 C). TIII serrulation: spines lacking between subapical spurs and apical spurs on both sides; on inner side, one spine between isa 1 and isa 2, one to two spines between isa 2 and isa 3, 20 spines above isa 3; on outer side, one to two spines between osa 1 and osa 2, two spines between osa 2 and osa 3, 18 - 19 spines above osa 3. Barsitarsomeres III serrulation: one to two inner spines and five to six outer spines, in addition to apical spines. Subgenital plate as on Fig. 23 O.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFD73503FF78FB69FDA7BA1E.taxon	description	MEASUREMENTS (IN MM). — See Table 21.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFD13503FF54FCEFFC83BD0C.taxon	type_taxon	TYPE SPECIES. — Rumea gaschei Desutter, 1988: 358 by original designation.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFD13503FF54FCEFFC83BD0C.taxon	diagnosis	DIAGNOSIS. — See Desutter-Grandcolas (1992 a).	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFD13503FF54FCEFFC83BD0C.taxon	distribution	DISTRIBUTION. — Amazon region (Peru, Brazil, French Guiana).	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFD13503FF7CFBEEFB4EBDF5.taxon	description	(Table 22)	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFD13503FF7CFBEEFB4EBDF5.taxon	materials_examined	TYPE LOCALITY. — French Guiana, Sinnamary, Paracou. TYPE MATERIAL. — Holotype by original designation. French Guiana • 1 ♂; Sinnamary, Paracou, forêt sur sables blancs; II. IX. 1988; L. Desutter leg.; MNHN-EO-ENSIF 5102. Paratypes: 2 males. French Guiana • 1 ♂; same locality and collector as the holotype; IX. 1988; MNHN-EO-ENSIF 5103 • 1 ♂; Mana, Acarouany; 11. XI. 1968; J. Bonfils leg.; MNHN-EO-ENSIF 5104. MATERIAL FROM THE MITARAKA. — French Guiana • 2 ♀; Monts Tumuc-Humac, Massif du Mitaraka vers sommet en Cloche; entre 54.4541 O 2.2349 N et 54.4646 O 2.2329 N; alt. entre 370 m et 470 m; 23. II. - 10. III. 2015; F. Legendre & S. Hugel leg.; de nuit, fn. SH 410, SH 457; MNHN. Specimens in bad condition (fungi).	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFD13503FF7CFBEEFB4EBDF5.taxon	diagnosis	DIAGNOSIS. — See Desutter-Grandcolas (1992 b: 162) for males.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFD13503FF7CFBEEFB4EBDF5.taxon	discussion	REMARK Females are not known from the type locality, but only from near the Arataye river, a tributary of Approuague. The females from Mitaraka are close to the types of R. guyanensis as far as size and coloration are concerned, as are the females of Arataye.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
03DE87FCFFD13503FF7CFBEEFB4EBDF5.taxon	description	MEASUREMENTS (IN MM). — See Table 22.	en	Desutter-Grandcolas, Laure, Faberon, Léo (2020): Phalangopsidae crickets (Orthoptera, Grylloidea) from the Mitaraka biological survey, French Guiana. Zoosystema 42 (32): 739-797, DOI: 10.5252/zoosystema2020v42a32
