taxonID	type	description	language	source
03D187BCFFDE480A29A0BF4E2619FB88.taxon	description	(Figs. 1, 25, 39, 65, 107 – 108)	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFDE480A29A0BF4E2619FB88.taxon	description	Description. Because the original description (Péringuey 1896: 160 – 161) is quite detailed, we simply add the following: length 10.8 – 14.0 mm, width 3.0 – 4.2 mm. Apical third of Ant 1 and basal two-thirds of Ant 3 black or dark brown; femora reddish brown with apical third or half black; tibiae nearly black. Head wider than pronotum (Fig. 1), HW / PW 5 1.19 (1.11 – 1.30); Ant 1 / Ant 3 5 3.40 (3.20 – 3.57); labrum with medial lobe slightly protruded anteriorly (Fig. 25); genae long, slightly tumid; eyes large, about 1.8 times as long as genae. Pronotum widest slightly before middle, PL / PW 5 1.56 (1.45 – 1.62). Elytra with lateral margins nearly parallel; outer angles slightly dentate, not sharply pointed at apex (Fig. 39), EW / PW 5 2.27 (2.09 – 2.56), EL / EW 5 1.89 (1.81 – 2.04). Protarsomere 3 strongly asymmetrical in male; tarsal claws with 5 – 6 pectinations on inner side, length of the longest pectination subequal to width of claw base (Fig. 65). Aedeagus stout (Fig. 108), with apical lamella narrow, thickened and rounded at apex (Fig. 107).	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFDE480A29A0BF4E2619FB88.taxon	distribution	Distribution. Angola, Cameroon, Congo, Kenya, Mozambique, South Africa, Senegal, Uganda, Zambia.	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFDE480A29A0BF4E2619FB88.taxon	discussion	Remarks. We did not check the type. Our description is based on specimens determined by Basilewsky. Members of this species are readily distinguished from those of other members of the genus by their red pronota and long tarsal claw pectinations.	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFDD480B28E7B85E24B1FC03.taxon	description	(Figs. 109 – 112) Types. Holotype: a male (CBNH), ‘‘ Malawi: Chitipa District, Jembya Reserve, 18 km SSE Chisenga. 10 - 08 S, 33 - 27 E, 1,870 m. 11 – 20 Dec 1988, J. Rawlins, S. Thomson’ ’ / ‘‘ HOLOTYPE, Dendrocellus bicoloripennis Liang & Kavanaugh n. sp., designated by H. B. Liang & D. H. Kavanaugh 2006 ’’ [red label]. Paratypes (total 4 specimens): 3 females (1 in CAS, 2 in CBNH), same collecting locality and collectors but ‘‘ 1 – 10 Jan 1989 ’’; 1 female (CBNH), same collecting locality and collectors but ‘‘ 11 – 20 Jan 1989 ’’.	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFDD480B28E7B85E24B1FC03.taxon	materials_examined	Type locality. 18 km SSE of Chisenga, Jembya Reserve, Malawi.	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFDD480B28E7B85E24B1FC03.taxon	diagnosis	Diagnosis. Pronotum reddish brown; elytra with bluish-green luster at basal three-fourths, black or dark purple at apical fourth; hind femora with apical half or two-thirds black, tibiae black; tarsal claw pectinations very short, the longest pectination shorter than one-forth width of claw base.	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFDD480B28E7B85E24B1FC03.taxon	description	Description. Length: 9.8 – 11.3 mm, width: 3.2 – 3.4 mm. Head and abdominal sterna with blue luster; neck, pronotum, scutellum, thoracic sterna, episterna, and epimera, Ant 2 – 11, and basal half of femora red or brown; elytra with bluish-green luster at basal three-fourths, black or dark purple at apical fourth with an anteriorly directed triangular extension along suture; apical half or two-thirds of femora, tibiae, tarsomeres 1 – 3, and apical third of Ant 1 black; basal two-thirds of Ant 1, Ant 2 – 11, tarsomeres 4 – 5 brown or yellow. Head densely punctate on vertex; labrum with medial lobe slightly protruded anteriorly; Ant 1 / Ant 3 5 3.22 (3.13 – 3.31); genae developed, strongly tumid; eyes large, convex, 1.8 times as long as genae. Pronotum short, cylindrical, widest slightly before middle, PL / PW 5 1.29 (1.26 – 1.35), HW / PW 5 1.15 (1.14 – 1.16); slightly widened from apical angles to middle, then narrowed posteriad, strongly sinuate before basal angles; basal angles obtuse, strongly protruded laterally. Elytra wide, densely punctate throughout; lateral margins nearly parallel in basal forth, distintly widened posteriad to apical fourth, then narrowed to apex; EL / EW 5 1.69 (1.63 – 1.73), EW / PW 5 2.16 (2.11 – 2.25); striae effaced, almost invisible (only striae 1 – 3 traceable in apical regions); outer apical angles nearly round, only vaguely angulate. Protarsomere 3 strongly asymmetrical in male; tarsal claws with 5 – 6 min pectinations, length of the longest pectination less than one-fifth width of claw base (Fig. 110). Aedeagus stout, with apical lamella moderately long, thin, linguiform, rounded at apex (Figs. 111 – 112).	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFDD480B28E7B85E24B1FC03.taxon	distribution	Distribution. Malawi.	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFDD480B28E7B85E24B1FC03.taxon	etymology	Etymology. This species is named for the pattern of two different colors on the elytra.	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFDD480B28E7B85E24B1FC03.taxon	discussion	Remarks. Members of this new species are similar to those of African Dryptini species Drypta ruficollis Dejean, 1831, Drypta thoracica Boheman, 1848, and Dendrocellus australis Péringuey in having red pronota, but differ from them in having slender tarsal claws with short pectinations (in Drypta ruficollis and Drypta thoracica specimens, the tarsal claws are smooth; and in Dendrocellus australis specimens, the tarsal claws have long pectinations). The bicolored elytral color pattern and effaced elytral striae appear to be unique in the genus (in other members of the genus, the elytra are unicolored, and all striae are distinct).	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFDC481429A6BFCF250DFAD0.taxon	description	(Figs. 2, 3, 28, 69, 77 – 78)	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFDC481429A6BFCF250DFAD0.taxon	description	Specimens examined. Total 23 specimens: Bhutan: 1 female (ISNB), ‘‘ Coll. I. R. Sc. N. B. Inde-British Bootang, Maria Basti 1900, ex Coll. Oberthur’ ’. China: 1 male, ‘‘ Hainan, Tongshi, 340 m’ ’ / ‘‘ 1960. IV. 25, Xuezhong Zhang collector’ ’; 1 male, ‘‘ Hainan, Tongshi, 340 m’ ’ / ‘‘ 1960. IV. 25, Changqing Li collector’ ’; 1 female, ‘‘ Hainan, Qiongzhong, 400 m’ ’ / ‘‘ 1960. VII. 15, Suofu Li collector’ ’; 1 male, ‘‘ Hainan, Jianfengling, 75 m’ ’ / ‘‘ 1980. III. 26, Shuyong Wang colllector’ ’; 1 female, ‘‘ Yunnan, Xishuangbanna, Xiaomengyang, 850 m’ ’, 1958. X. 3, Xuwu Meng collector’ ’; 1 male, ‘‘ Yunnan, Xishuangbanna, Damenglong, 650 m’ ’ / ‘‘ 1958. VIII. 7, Yiran Zhang collector’ ’; 1 male, ‘‘ Yunnan, Hekou, 80 m, 1956. VI. 8, Keren Huang collector’ ’ / ‘‘ Desera geniculata Klug, Kryzhanovsky det. ’’; 1 male, ‘‘ Botanical Garden, Xihuangbanna, Yunnan Prov., 5 - 6 - VII- 2005, Lizhen Li leg. ’’; 1 female, ‘‘ Tibet, Mêdog’ ’ / ‘‘ 1984. VIII’ ’; 1 male, ‘‘ Tibet, Mêdog, 820 m’ ’ / ‘‘ 1983. II. 2, Yinheng Han 16 collector’ ’. India: 1 male (ISNB), ‘‘ geniculata Klug, ex. coll. Bonneull, Inde’ ’ / ‘‘ Coll. R. I. Sc. N. B. Inde’ ’. Indonesia: 1 female (ZMHU), ‘‘ 709 ’’ / ‘‘ Java’ ’ / ‘‘ coelestina Kl. ’’ / ‘‘ Syntypus Drypta coelestina Klug 1834, labeled by MNHUB 2004 ’’ [red label]; 1 male (ISNB) ‘‘ Sud-Celebes, Bantimoeng, C. Ribbe, 1883 ’’, Coll. R. I. Sc. N. B. Indonesia’ ’. Laos: 1 female (ISNB), ‘‘ Laos, Pakneun, 1 - IX- 1918, R. Vitalis de Salvaza’ ’ / ‘‘ 2,074 ’’ / ‘‘ Desera geniculata Kl. Det. R. Vitalis de Salvaza’ ’ / ‘‘ Coll. R. I. Sc. N. B. ’’. Malaysia: 1 female (ZMHU), ‘‘ Malacca, Mniszcch’ ’; 1 male, ‘‘ longicollis 9,953 ’’ / ‘‘ Malacca’ ’ / ‘‘ Soc. Ent. Belg. Coll. Putzeys’ ’, Coll. R. I. Sc. N. B. Malaisie’ ’.	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFDC481429A6BFCF250DFAD0.taxon	materials_examined	Myanmar: 1 male (MCSN), ‘‘ Carin Chebà, 900 – 1,100 m, L. Fea’ ’ / ‘‘ Drypta sp.? ’’ / ‘‘ det. H. Bates 1892 ’’ / ‘‘ Desera geniculata Klug’ ’ / ‘‘ Museo Civico di Genova’ ’. Pakistan: 1 female (CAS), ‘‘ E. Pakistan, Harbang For. 6 mi N. Chiringa, IX- 18 - 61, 50 ft’ ’ / ‘‘ Collectors E. S. Ross, D. Q. Cavagnaro’ ’ / ‘‘ Desera sp. 5, det. D. H. Kavanaugh, 1991 ’’ / ‘‘ Dendrocellus coelestinus Klug 1834, det. Liang H. B. 2005.3 ’’. Thailand: 1 male (ZMHU), ‘‘ Mon Angget, Chiang Mai, Thailand, 31. V. 1990, K. Masumoto’ ’ / ‘‘ Desera geniculata (Klug), det. N. Ito, 1995 ’’; 1 female (CAS), ‘‘ Thailand, Tak, vicinity of Umphang, mid January to Mid April 1991, native collectors, Calif. Acad. Sci. Collection’ ’ / ‘‘ Drypta sp. 19, det. D. H. Kavanaugh 1995 ’’ / ‘‘ Dendrocellus coelestinus Klug 1834, det. Liang H. B. 2005.3 ’’; 1 male (MBC), ‘‘ Thai, 13.5.1993, 19.29 N, 98.18 E, Soppong 750 m, L. Bocák lgt. ’’; 1 female (MBC), ‘‘ Thai, 13.5.1993, 19.29 N, 98.18 E, Soppong 750 m, Vit Kubáň leg. ’’.	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFDC481429A6BFCF250DFAD0.taxon	description	Description. Length 9.5 – 10.4 mm, width 2.4 – 2.9 mm. Black with green and blue luster; apical third of Ant 1 black, basal two-thirds brown; femora yellow or brown with apical sixth or eighth black; tibiae yellow. Head wider than pronotum, HW / PW 5 1.27 (1.18 – 1.33); Ant 1 / Ant 3 5 3.41 (3.21 – 3.62); labrum with medial lobe moderately protruded anteriorly (Fig. 28); genae moderately tumid, eyes large, about 2.3 times as long as genae. Pronotum narrow, PL / PW 5 1.66 (1.57 – 1.74); lateral margins nearly parallel in apical two thirds. Elytra with lateral margins nearly parallel in both sexes; EL / EW 5 2.04 (1.85 – 2.13), EW / PW 5 2.18 (2.00 – 2.29); outer angles slightly dentate; intervals flat, densely punctate. Protarsomere 3 slightly asymmetrical in male; tarsal claws with 3 – 5 short pectinations, length of the longest pectination shorter than half width of claw base (Fig. 69). Aedeagus with apical lamella thin, wide, linguiform (Figs. 77 – 78).	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFDC481429A6BFCF250DFAD0.taxon	distribution	Distribution. Bhutan, China, India, Indonesia (Java, Sumatra, Sulawesi), Laos, Malaysia, Myanmar, Pakistan, Thailand.	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFDC481429A6BFCF250DFAD0.taxon	discussion	Remarks. The syntype of D. coelestinus (Klug) we examined is teneral, with bluish purple luster on pronotum and elytra. D. parallelus Chaudoir was described from Sumatra. When we compared the type of D. parallelus with that of D. coelestinus, we were unable to find any significant difference. Chaudoir (1861: 545) treated D. coelestinus (Klug) as a synonym of D. unidentatus (Macleay). Andrewes (1927: 100) disagreed with this treatment and pointed out that D. coelestinus (Klug) was probably a pale variety of D. geniculatus (Klug). However, adults of D. coelestinus have very much shorter tarsal claw pectinations (Andrewes also noticed this character, 1927: 100) and males have a triangular lamella of the aedeagus, and, therefore, this species is distinct from D. geniculatus (Klug).	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFC34815293CB9822467FAD0.taxon	description	(Figs. 4, 30, 67, 73 – 74)	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFC34815293CB9822467FAD0.taxon	description	1980.6.22, by light trap, Juchang Huang collector’ ’; 1 female, ‘‘ Fujiang, Xiadaoshui’ ’ / ‘‘ 1980. VI. 25, Yuanhong Chen collector’ ’; 1 male, ‘‘ Guangdong, Zhanjiang, 1982.9.23 ’’ / ‘‘ On sugarcane, Jiao Chen collector’ ’; 1 male (BMNH), ‘‘ Guangxi, Jinxiu, Luoxiang, 450 m, 2000. VI. 30, Jian Yao collector’ ’; 1 female, ‘‘ Guangxi, Jinxiu, Linhai Shanzhuang, 1,000 m, 2000. VII. 2, Jian Yao collector’ ’; 1 female, ‘‘ Guangxi, Jinxiu, Shengtangshan, 900 – 1,900 m, 2000. VI. 28, Wenzhu Li collector’ ’; 1 male (IOZ) and 1 female (BMNH), ‘‘ Guangxi, Jinxiu’ ’ / ‘‘ By light trap, 1983. V. 31, Subai Liao collector’ ’. India: 1 female (ISNB), ‘‘ Inde’ ’ / ‘‘ Coll. P. Dupuis’ ’ / ‘‘ Desera geniculata, det. P. Dupuis’ ’ / ‘‘ Desera geniculata Kl. ’’ / ‘‘ Coll. R. I. Sc. N. B. Inde’ ’. Indonesia: 1 female (ISNB), ‘‘ Java’ ’ / ‘‘ Collection P. Dupuis’ ’ / ‘‘ Coll. R. I. Sc. N. B. Java’ ’; 1 female (ZMHU), ‘‘ Java, Boemi Ajoe’ ’ / ‘‘ Dendrocellus geniculatus, Fairm. ’’; 1 male (ZMHU), ‘‘ Java, Baugalan, 1922 ’’ / ‘‘ Drypta coelestina Kl. ’’; 1 male (ZMHU), ‘‘ Deli, auf Sumatra, Hartert’ ’; 1 female (ZMHU), ‘‘ Ober-Langkat, Deli, Sumatra, 1894, M. Ude S. ’’ / ‘‘ 93278 ’’. Japan: 1 male (ISNB), ‘‘ Japon’ ’ / ‘‘ Collect Balls’ ’ / ‘‘ Japan’ ’ / ‘‘ Collection E. Rousseau’ ’ / ‘‘ Coll. R. I. Sc. N. B. Japon’ ’; 1 female (ISNB), ‘‘ Japon’ ’ / ‘‘ Collect Balls’ ’ / ‘‘ Japan’ ’ / ‘‘ Coll. R. I. Sc. N. B. Japon’ ’. Pakistan: 1 female (CAS), ‘‘ E. Pakistan, Ichamati, 28 mi E. Chittagong, IX- 14 - 61, 50 ft’ ’, Collection E. S. Ross, D. Q. Cavagnaro’ ’; 1 male (ZMHU), ‘‘ Süd borneo, Bandjermasin, L. W. Schaufuss G. ’’. Thailand: 1 female (ISNB), ‘‘ Coll. I. R. Sc. N. B. Thailand (Ioei), Na-Haeo (Field res. stat.), 15 - 19. V. 2003, light trap, leg. J. Constant, K. Smets & P. Grootaert’ ’; 1 male (MBC), ‘‘ Thai, 13.5.1993, 19.29 N, 98.18 E, Soppong, 750 m, L. Bocák lgt. ’’; 2 females (MBC), ‘‘ Thai, 13.5.1993, 19.29 N, 98.18 E, Soppong, 750 m, Vit Kubáň leg. ’’. Description. Length 10.2 – 11.2 mm, width 2.7 – 3.0 mm. Black with green luster; Ant 1 with apical fourth blackish brown, basal three fourths yellow; femora yellow, with apical sixth black; tibiae yellow. Head slightly wider than pronotum, HW / PW 5 1.19 (1.12 – 1.24); labrum with medial lobe strongly protruded anteriorly (Fig. 30); Ant 1 / Ant 3 5 2.62 (2.50 – 2.67); genae slightly tumid, eyes large, about 2.3 times as long as genae. Pronotum widest at the middle, slightly narrowed anteriorly, PL / PW 5 1.58 (1.45 – 1.67). Elytra long, lateral margins nearly parallel in both sexes; outer angles slightly dentate, EW / PW 5 2.21 (2.03 – 2.31), EL / EW 5 1.99 (1.93 – 2.05). Protarsomere 3 slightly asymmetrical in male; tarsal claws with 3 or 4 pectinations, length of the longest pectination less than half width of claw base (Fig. 67). Aedeagus with apical lamella thin, triangular (Figs. 73 – 74).	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFC34815293CB9822467FAD0.taxon	distribution	Distribution. China, India, Indonesia (Java, Sumatra, Kalimantan), Japan, Laos, Pakistan, Thailand.	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFC34815293CB9822467FAD0.taxon	discussion	Remarks. We have not examined the type. Members of this species are readily recognized by the strongly protruded medial lobe of the labrum, which is unique in the genus. Dendrocellus confusus is widely distributed in southeastern Asia from Japan and China to Java. Hansen (1967: 406) stated that adults of this species were similar to those of D. geniculatus (Klug) in having short tarsal claw pectinations. However, he wrongly characterized D. geniculatus (Klug) (see remarks under D. geniculatus). Actually, adults of this species are similar to those of D. coelestinus (Klug) in having short claw pectinations, but different from the latter in having a longer medial lobe of the labrum and smaller ratio of Ant 1 / Ant 3.	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFC248162944B98225A2FBE9.taxon	description	(Figs. 5, 45, 48, 50, 53, 56, 83 – 84)	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFC248162944B98225A2FBE9.taxon	description	18 I. R. Sc. N. B. Java – Togoe, 1902, ex. Coll. Oberthur’ ’; 1 female (ISNB), ‘‘ Beokit Gabah, Z. W. K. Sumatra, leg. H. Lucht, I. 1919 ’’ / ‘‘ Coll. I. R. Sc. N. B. Sumatra’ ’; 1 male (ZMHU), ‘‘ W. Sumatra, No. 18, VII. 1991 ’’. Malaysia: 2 females (ZMHU), ‘‘ Near Keningau, Is. Borneo, Sabah, E. Malaysia, 16 II 1989, M. Itoh’ ’ / ‘‘ Desera geniculata (Klug) det. N. Ito, 1993 ’’; 1 male and 1 female (ISNB), ‘‘ Kina Balu, Borneo’ ’ / ‘‘ Collection P. Dupuis’ ’ / ‘‘ Coll. R. I. Sc. N. B. Borneo’ ’; 1 male and 1 female (ISNB), ‘‘ Kina balu, Borneo 1,500 m, H. Hille, Berlin SW 11 ’’ / ‘‘ R. I. Sc. Nat. Belg. I. G. 16.500 ’’ / ‘‘ Coll. R. I. Sc. N. B. Borneo’ ’; 1 female (ISNB), ‘‘ Coll. I. R. Sc. N. B. Nord Borneo, Kina Balu, Fr. Buffat, 1904 ’’; 1 female (ISNB), ‘‘ Coll. I. R. Sc. N. B. Indonesie, N. Borneo, Mont Kina Balu, 5. VIII. 1903, leg. John Waterstradt, ex coll. Oberthur IG. 18.293 ’’. Kalimantan (Malaysia or Indonesia): 1 male (ISNB), ‘‘ Borneo’ ’ / ‘‘ Collection P. Dupuis’ ’ / ‘‘ Coll. R. I. Sc. N. B. Borneo’ ’; 1 female (ISNB), ‘‘ Borneo’ ’ / ‘‘ Collection P. Dupuis’ ’ / ‘‘ Desera unidentata M. L. det. P. Dupuis, 1913 ’’ / ‘‘ Desera unidentata ’’ / ‘‘ Coll. R. I. Sc. N. B. Borneo’ ’; 1 male (ISNB), ‘‘ Borneo’ ’ / ‘‘ Reg. Mus. Hist. Nat., Belg. I. G. 10294 ’’ / ‘‘ longicollis Dj’ ’ / ‘‘ G. Fagel’ ’ / ‘‘ G. Fagel det. Desera longicollis Dej’ ’ / ‘‘ Coll. R. I. Sc. N. B. Borneo’ ’; 1 male and 2 females (CAS), ‘‘ Borneo, M. L. Hauschild’ ’ / ‘‘ Van Dyke collection’ ’ / ‘‘ Dendrocellus crassus Andrewes 1931, det. Liang H. B. 2005.3 ’’. Description. Andrewes (1931) provided a detailed description and illustration. We simply add the following: length 9.4 – 10.4 mm, width 2.9 – 3.3 mm. Apical (not basal as stated in original description) four fifths of Ant 1 black; femora brown, with apical sixth or seventh black; Ant 1 / Ant 3 5 3.45 (3.33 – 3.56); HW / PW 5 1.17 (1.12 – 1.21); PL / PW 5 1.53 (1.48 – 1.57); EW / PW 5 2.36 (2.18 – 2.42); EL / EW 5 1.72 (1.64 – 1.81); labrum with medial lobe slightly protruded anteriorly; genae short, slightly tumid; eyes moderately large, 2.5 times as long as genae; elytral punctation as in Fig. 45; protarsomere 3 symmetrical in male (Fig. 53); tarsal claws with 5 – 6 pectinations, length of the longest pectination subequal to the width of claw base (similar to Fig. 63); last sternum as in Fig. 48 or 50. Aedeagus with apical lamella thin, linguiform (Figs. 83 – 84).	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFC248162944B98225A2FBE9.taxon	distribution	Distribution. Indonesia (Sumatra), Malaysia (N. Kalimantan).	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFC248162944B98225A2FBE9.taxon	discussion	Remarks. We have not examined the type (in BMNH). Members of this species are readily recognized by their elytral intervals sparsely punctate (Fig. 45), femora with apical sixth black, and tibiae completely black.	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFC1481729AAB865240DFB68.taxon	description	(Figs. 6, 42, 58, 95 – 96)	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFC1481729AAB865240DFB68.taxon	description	Description. Because Sloane’s description (1907: 473) is detailed, we simply add the following: length 9.1 – 10.7 mm, width 2.6 – 3.5 mm. Femora brown, with apical ninth or tenth blackish brown; tibiae brown. Ant 1 / Ant 3 5 3.37 (3.13 – 3.50); HW / PW 5 1.16 (1.09 – 1.19); PL / PW 5 1.58 (1.53 – 1.65); EW / PW 5 2.41 (2.22 – 2.58); EL / EW 5 1.74 (1.62 – 1.81); labrum with medial lobe slightly protruded anteriorly; genae short, slightly tumid; eyes large, about 2.5 times as long as genae; elytra with lateral margins nearly parallel in both sexes; elytral outer angles strongly dentate, sharply pointed at apex (Fig. 42); protarsomere 3 slightly asymmetrical in male; tarsal claws with 5 or 6 pectinations, longest pectination slightly longer than width of claw base (Fig. 58). Aedeagus with apical lamella narrow, slightly digitiform (Figs. 95 – 96).	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFC1481729AAB865240DFB68.taxon	distribution	Distribution. New Guinea.	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFC1481729AAB865240DFB68.taxon	discussion	Remarks. We have not examined the type. Members of this species can be recognized by their brown femora with very short dark apices. In most specimens, the elytra possess strongly dentate outer angles as described by Sloane (1907: 473), and in some specimens, the outer angles are sharpened into a short spine (e. g., from Jule Island). We found one specimen (in CAS) of this species with obtuse outer angles; but because it is similar to other specimens of this species in other characters, including shape of the aedeagus, this is probably just an abnormal specimen. This species is the only member of Dendrocellus distributed in New Guinea and appears to be endemic to this island.	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFC0481229A4B8EA241EFB54.taxon	description	(Figs. 8 – 9, 26, 40, 44, 47, 49, 52, 55, 64, 87 – 88)	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFC0481229A4B8EA241EFB54.taxon	description	20 ‘‘ Takao Formosa, Sauter XI- 07 ’’ / ‘‘ Type’ ’ [red label], ‘‘ W. Hansen vid., 1966, Desera gilsoni Dup. ’’; 1 male and 1 female (ISNB), ‘‘ Desera gilsoni Dup., P. Dupuis det. 1913 ’’ / ‘‘ Type’ ’ / ‘‘ Formosa, Polisha, X. 09, H. Sauter S. G. ’’ / ‘‘ Paraype’ ’ / ‘‘ W. Hansen vid., 1966, Desera gilsoni Dup. ’’; 1 female (ISNB), ‘‘ Desera gilsoni Dup., P. Dupuis det. 1913 ’’ / ‘‘ Formosa, Hoozan, I. 10, H. Sauter S. G. ’’ / ‘‘ Paratype’ ’ / ‘‘ W. Hansen vid., 1966, Desera gilsoni Dupuis’ ’; 1 female (ISNB), ‘‘ Desera gilsoni Dup., P. Dupuis det. 1913 ’’ / ‘‘ Formosa, Hoozan, XII. 09, H. Sauter S. G. ’’ / ‘‘ Paratype’ ’ / ‘‘ W. Hansen vid., 1966, Desera gilsoni Dupuis’ ’; 1 male (ZMHU), ‘‘ China’ ’ / ‘‘ Dendrocellus geniculatus Klug, det. Ing. Jedlicka’ ’; 1 female (ZMHU), ‘‘ S. Formosa, Fuhosho, 4.09, H. Sauter S. V. ’’; 1 female (ZMHU), ‘‘ Formosa, Hoozan, 1.10, H. Sauter S. G. ’’; 1 female (ZMHU), ‘‘ Formosa, Hoozan, XII. 09, H. Sauter S. G. ’’; 1 male (ZMHU), ‘‘ Formosa, Polisha, X- 09, H. Sauter S. G. ’’; 1 female (ZMHU), ‘‘ Formosa, Hoozan, I. 10, H. Sauter S. G’ ’; 1 male (CAS), ‘‘ Chung Kon, Hainan Id, VII- 22 - 35 ’’ / ‘‘ L. Gressitt Collector’ ’ / ‘‘ L. Gressitt Collection’ ’; 1 male (CAS), ‘‘ Dwa Bi, Hainan Id, VII- 22 - 35 ’’ / ‘‘ L. Gressitt Collector’ ’ / ‘‘ L. Gressitt Collection’ ’; 1 male (CAS), ‘‘ Kuraru, VI- 14 - 34, Formosa’ ’ / ‘‘ L. Gressitt Collector’ ’ / ‘‘ L. Gressitt Collection’ ’; 1 female (CAS), ‘‘ Rokki, Formosa, V- 17 - 34 ’’ / ‘‘ L. Gressitt collector’ ’ / ‘‘ L. Gressitt Collection’ ’; 1 female (CAS), ‘‘ Rokki, Formosa, V- 15 - 34 ’’ / ‘‘ L. Gressitt collector’ ’ / ‘‘ Van Dyke Collection’ ’; 1 male, ‘‘ Hainan, Baoting, 80 m’ ’, 1960. V. 14, Suofu Li collector’ ’; 1 male, ‘‘ Hainan, Baoting, 80 m’ ’ / ‘‘ 1960. VII. 25, Xuezhong Zhang collector’ ’; 1 female, ‘‘ Hainan, Tongshi, 340 m’ ’ / ‘‘ 1960. VIII. 5 ’’, Suofu Li collector’ ’; 1 female, ‘‘ Hainan, Qiongzhong, 400 m’ ’ / ‘‘ 1960. VII. 15 ’’, Suofu Li collector’ ’; 1 male, ‘‘ Hainan, Jianfengling, 70 m’ ’ / ‘‘ 1980. III. 26, light trap, Fuji Pu collector’ ’; 1 male, ‘‘ Fujian, Jiangle, Longqishan, 500 m’ ’ / ‘‘ 1991. VIII. 13, Shimei Song collector’ ’; 1 female, ‘‘ Fujian, Jiangyang, Huangkeng, 270 – 360 m’ ’ / ‘‘ 1960. III. 26, Yiran Zhang collector’ ’; 1 male, ‘‘ Fujian, Jianyang, Tongyou, 170 m’ ’ / ‘‘ 1978, Shixiong Shen collector’ ’; 1 male, ‘‘ Fujian, Fuzhou’ ’ / ‘‘ Light trap, 1979. VIII. 12 ’’; 1 male, ‘‘ Shanghai, Jiangwan, 6.8.1942 ’’; 1 female, ‘‘ Zi-ka-wei, 19.5.20 ’’ [Shanghai]; 1 male, ‘‘ Guangxi, Jinxiu, Shengtangshan, 900 m, 1999. V. 17, Fusheng Huang collector’ ’; 1 female, ‘‘ Guangxi, Napo, Defu, 1,350 m, 2000. VI. 18, Wenzhu Li collector’ ’; 1 female, ‘‘ Guangxi, Napo, Defu, 1,350 m, 2000. VI. 18, Chaodong Zhu collector’ ’; 1 female, ‘‘ Guangxi, Napo, Defu, 1,350 m, 2000. VI. 19, Jian Yao collector’ ’; 1 female, ‘‘ Guangxi, Tianlin, Yaojiawan, 1,200 – 1,400 m, 2002. VI. 1, Xiujuan Yang collector’ ’; 1 female, ‘‘ Guangxi, Tianlin, Laoshan Nongchang, 1,200 – 1,600 m, 2002. VI. 4, Xiujuan Yang collector’ ’; 2 males and 1 female, ‘‘ Guangxi, Jinxiu County’ ’ / ‘‘ Light trap, 1983. V. 30 - 31, Subai Liao collector’ ’; 1 male, ‘‘ Guangxi, Longshen’ ’ / ‘‘ 1980. VI. 16 ’’; 1 male, ‘‘ Guangxi, Longshen’ ’ / ‘‘ 1980. VI. 16, Shimei Song collector’ ’; 1 female, ‘‘ Guangxi, Longshen, 300 m’ ’ / ‘‘ 1963. V. 23, Wang Chunguang collector’ ’, 2 males and 1 female, ‘‘ Guangxi, Guilin, Yanshan, 200 m, light trap’ ’ / ‘‘ 1963. V. 12 - 13, Wang Chunguang collector’ ’; 1 female, ‘‘ Guangxi, Guilin, Yanshan, 1952.10.3 ’’; 1 female, ‘‘ Guangxi, Guilin, Yanshan, 1953.6.5 ’’; 1 male, ‘‘ Guangxi, Guilin, Yanshan, 1953.8.18 ’’; 1 male, ‘‘ China, Jiangxi, Longnan, Jiulianshan’ ’ / ‘‘ 1975. VI. 8, light trap, Youwei Zhang collector’ ’; 1 male, ‘‘ Jiangsi, Dayu, Lannijing, Sanjiangkou, 400 m’ ’ / ‘‘ 1985. VIII. 11, Subai Liao collector’ ’; 3 males, ‘‘ Guizhou, Chishui, Jinsha, 500 m’ ’ / ‘‘ 2000. IX. 21 - 22, Liang H. B. ’’; 2 females, ‘‘ Guizhou, Luodian’ ’ / ‘‘ 1980. VI- VII’ ’; 1 male, ‘‘ Guangdong, Shenzhen, 1980.5, Shaohua Du collector’ ’; 1 male and 1 female, ‘‘ Guangdong, Shipai, 1964.5, Shoujian Chen collector’ ’; 1 female, ‘‘ Yunnan, Xishuangbanna, Yunjinghong, 650 m’ ’ / ‘‘ 1958. VII. 13, Xuwu Meng collector’ ’; 1 male, ‘‘ Yunnan, Xishuangbanna, Damenglong, 650 m’ ’ / ‘‘ 1958. VII. 17, Yiran Zhang collector’ ’; 1 female, ‘‘ China, Yunnan, Fugong, Shangpa, Guquan, roadside, 26.88897 ° N, 98.86558 ° E’ ’ / ‘‘ 1,230 m, 2004.4.21, Liang H. B., Li X. Y. coll., California Academy & IOZ, Chinese Acad. Sci. ’’; 1 female, ‘‘ China, Yunnan, Fugong, Lumadeng, Yaping, Yaping Bridge’ ’ / ‘‘ 2004.4.30 day, Guangxu Peng coll., Institute of Zool., CAS and California Acad. Sciences’ ’. 1 female, ‘‘ China, Yunnan, Gongshan, Cikai, Dashaba, along Nujiang, 27.73268 ° N, 98.67099 ° E’ ’, 1,435 m, 2004.11.10, night, Hongbin Liang collector, California Academy & IOZ, Chinese Acad. Sci. ’’; 1 female, ‘‘ Sichuan, Beibei, 15.10.1940 ’’; 21 males and 8 females, ‘‘ Tibet, Mêdog, 1,000 – 1,100 m’ ’ / ‘‘ 1983. I. 23, Zai Lin collector’ ’; 1 male and 3 females, ‘‘ Tibet, Mêdog, 800 m’ ’ / ‘‘ 1983. II. 7, Zai Lin collector’ ’; 2 females, ‘‘ Tibet, Mêdog, 820 m’ ’ / ‘‘ 1982. II. 2, Yinheng Han collector’ ’; 1 female, ‘‘ Tibet, Mêdog, Baibung, 850 m’ ’ / ‘‘ 1983. III. 7, Yinheng Han collector’ ’. India: 1 female (ZMHU), ‘‘ Ind. Ori. Sal. ’’ / ‘‘ geniculata Kl. ’’ / ‘‘ Hist. - Coll. (Coleoptera), Nr. 709, Drypta geniculata Klug, India Orient., Salinger, Zool. Mus. Berlin’ ’ / ‘‘ Holotypus, Drypta geniculata Klug 1834, Labeled by MNHUB 2004 ’’ [red label]; 1 male (ZMHU), ‘‘ S. India, Karikal, Pondicherry State, Febr. 1966, P. S. N. ’’; 1 female (ISNB), ‘‘ Pedong’ ’ / ‘‘ Pedong (Desgodins), ‘‘ Coll. R. I. Sc. N. B. Malaisie? ’’ [India]; 1 male (ISNB), ‘‘ Pedon, A. Desgodins’ ’ / ‘‘ Pedong’ ’ / ‘‘ Coll. R. I. Sc. N. B. Malaisie? ’’ [India]. Indonesia: 1 male (ZMHU), ‘‘ Batoe Doelang, Batoe Lanteh, Zeb. 800 – 1,200 m, Bensch 10 - 15.5.27 ’’; 1 female (ZMHU), ‘‘ Sumatra, Dryrolle’ ’ / ‘‘ 40538 ’’; 1 female (ZMHU), ‘‘ W. Sumatra ’’ / ‘‘ No. 18, VII 1991 ’’; 3 males (ISNB), ‘‘ Jadraboera, Sumatra W. C., E. Wegas’ ’ / ‘‘ Collection E. Rousseau’ ’ / ‘‘ Coll. R. I. Sc. N. B. Sumatra’ ’; 1 male and 1 female (ISNB), ‘‘ Sumatra’ ’ / ‘‘ Collection P. Dupuis’ ’ / ‘‘ Desera ’’ / ‘‘ Coll. R. I. Sc. N. B. Sumatra’ ’; 1 female (ISNB), ‘‘ Java, Koller’ ’ / ‘‘ Coll. R. I. Sc. N. B. Java’ ’; 1 male (ISNB), ‘‘ Sumatra’ ’ / ‘‘ Coll. Ogier de Baulny’ ’ / ‘‘ 6,180 ’’ / ‘‘ Coll. R. I. Sc. N. B. Sumatra’ ’. Malaysia: 1 female (ZMHU), ‘‘ Malacca, Tengah Gebirge, P. Zobris V. ’’; 1 male (ISNB), ‘‘ D. geniculatus Malacca (DL) Kl. ’’ / ‘‘ Soc. Ent. Belg. Coll. Putzeys’ ’ / ‘‘ Coll. R. Sc. N. B. Malaisie’ ’. Pakistan: 3 males and 8 females (CAS), ‘‘ Pakistan, Ichamati 28 mi E. Chittagong, IX- 14 - 61, 50 ft’ ’ / ‘‘ Collectors E. S. Ross, D. Q. Cavagnaro’ ’. Philippines: 1 male (ISNB), ‘‘ Ile Basilan, Philippines’ ’ / ‘‘ R. Mus. Illist. Nat. Belg. I. G. 12.595 ’’ / ‘‘ Coll. R. I. Sc. N. B. Philippines’ ’; 1 male (CAS), ‘‘ Mt. Makiling, Laguna P. I., I- 1 - 1931 ’’ / ‘‘ F. C. Hadden Collection’ ’; 1 male (CAS), ‘‘ Mt. Makiling, Laguna P. I., VIII- 9 - 1931 ’’ / ‘‘ V. Madrid collector’ ’ / ‘‘ F. C. Hadden Collection’ ’; 1 female (CAS), ‘‘ Mt. Makiling, Laguna P. I., VIII- 1 - 1931 ’’ / ‘‘ F. C. Hadden Collection’ ’ / ‘‘ Elevation 3,000 ft’ ’; 1 female (CMNH), ‘‘ Philippines: Mt. Baco, Mindoro, Sept 1987 ’’ / ‘‘ Desera geniculata Klug’ ’; 1 male (CAS), ‘‘ San Jose, Mindoro P. I., 1945, E. S. Ross’ ’. Sikkim: 1 male (ZMHU), ‘‘ Himalaya Sikkim’ ’ / ‘‘ Desera geniculata Klug, H. E. Andrewes’ ’; 12 males and 9 females (ZMHU), ‘‘ Himalaya Sikkim, Coll. Hauser’ ’. Thailand: 1 female (CMNH), ‘‘ Thailand: Chiang Mai Prov., San Sai, 6 – 8 May 1986 ’’; 1 male and 2 females (ZMHU), ‘‘ Thailand, Wang Takrai-Park, Nakhon Nayok Prov. ’’ / ‘‘ 4 - 7.9.1978 ’’, gekätschert, leg. H. J. Bremer’ ’; 1 female (ZMHU), ‘‘ Nordost-Thailand, Khon Kaen ad Iucem’ ’ / ‘‘ Dr. Sastri, Saowakontha leg. 2.9.1980 ’’; 1 male (CAS), ‘‘ Thailand, Chonburi, February, 1982, G. A. Shook, coll. ’’. Vietnam: 6 males and 4 females (ZMHU), ‘‘ Annam, Phuc-Son, Nov. Dez. H. Fruhstorfer’ ’; 1 female (ISNB), ‘‘ Coll. R. I. Sc. N. B. ’’ / ‘‘ Tonkin Hoabinh (North Vietnam), Ex coll. Oberthur, IG. 18.293 ’’; 1 female (ISNB), ‘‘ Annam Cua Thung’ ’ / ‘‘ R. I. Sc. N. B. 21042 ’’ / ‘‘ Coll. R. I. Sc. N. B. North Vietnam’ ’; 1 female (ISNB), ‘‘ Annam Phuison’ ’ / ‘‘ Coll. R. I. Sc. N. B. North Vietnam’ ’; 1 male and 2 females (ISNB), ‘‘ Annam Phuc-Son Nov. Dez, H. Fruhstorfer’ ’ / ‘‘ Coll. R. I. Sc. N. B North 22 Vietnam’ ’; 1 male (CAS), ‘‘ Quang Nam Province, Kim Lien, Republic of Vietnam, Bruce W. Miller, VII. 26.1968 ’’ / ‘‘ D. H. Kavanaugh Collection’ ’; 1 male, ‘‘ Tonkin, Hoa-Binh, III. 1937, leg A. de Cooman’ ’; 1 male, ‘‘ Tonkin, Hoa-Binh, leg A. de Cooman’ ’. Description. Length 8.0 – 10.2 mm, width 2.4 – 3.0 mm. Black with green or blue luster, often mixed with coppery luster; Ant 1 with apical third dark brown, Ant 3 with basal two-thirds slightly darker than apical third; femora yellow, with apical sixth or seventh blackish brown; tibiae yellow. Head wider than pronotum, HW / PW 5 1.24 (1.18 – 1.31); Ant 1 / Ant 3 5 3.63 (3.46 – 3.76); labrum (Fig. 26) with medial lobe slightly protruded anteriorly; genae short, tumid; eyes large, about 2.6 times as long as genae. Pronotum widest at middle, slightly narrowed anteriorly, PL / PW 5 1.51 (1.45 – 1.63). Elytra with lateral margins slightly widened posteriad to apical third, then narrowed gradually to outer apical angles in female (Fig. 8), nearly parallel in male; EL / EW 5 1.82 (1.69 – 1.89), EW / PW 5 2.29 (2.10 – 2.47); outer angles obtuse or slightly dentate (Fig. 40), punctation of intervals as in Fig. 44. Protarsomere 3 slightly asymmetrical in male (Fig. 52); length of the longest tarsal claw pectinations slightly shorter than width of claw base (Fig. 64). Last sternum as in Fig. 47 or 49. Aedeagus with apical lamella narrow, digitiform, apex thickened (Figs. 87 – 88).	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFC0481229A4B8EA241EFB54.taxon	distribution	Distribution. Bangladesh, Bhutan, Cambodia, China, India, Indonesia, Japan, Laos, Malaysia, Pakistan, Philippines, Sikkim, Thailand, Vietnam.	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFC0481229A4B8EA241EFB54.taxon	discussion	Remarks. We compared the types of Dendrocellus gilsoni (Dupuis) with that of D. geniculatus (Klug) and no significant difference was found. Hansen (1967: 405) described D. gilsoni continentalis from Laos, stating that it differs from the nominate subspecies in having a shorter testaceous portion of Ant 1 at base, longer black portion of femora at apex, and more convex elytral intervals. We examined many specimens with these features from China, Vietnam, Thailand, and Philippines, and no significant difference was found except for the femora black at apical fifth and Ant 1 brown at apical two-fifths. In our opinion, these are just variant forms of D. geniculatus (Klug). Moreover, Hansen (1967: 401) stated D. geniculatus (Klug) has short tarsal claw pectinations. Actually, D. geniculatus (Klug) has long and fine tarsal claw pectinations (Fig. 64). D. geniculatus (Klug) is widely distributed in southeastern Asia, from India east to Japan and south to Java. Its members can be recognized by the following combination of character states: small size (usually less than 10 mm), large ratio of Ant 1 / Ant 3 (3.46 – 3.76), elytra often with coppery luster, elytral outer angles obtuse or slightly dentate, tarsal claw pectinations fine and long, lamella of aedeagus very narrow and thickened at apex. Adults of several species (i. e., D. coelestinus, D. confusus, D. crassus; see ‘‘ Specimens examined’ ’ for each species) often have been wrongly determined as D. geniculatus (Klug).	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFC5481329A3B92D27FDFB91.taxon	description	(Fig. 10)	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFC5481329A3B92D27FDFB91.taxon	materials_examined	Type. Lectotype, here designated, a female (MCSN), labeled ‘‘ Carin Chebà, 900 – 1,100 m, L. Fea, V XII- 88 ’’ / ‘‘ Typus’ ’ / ‘‘ Dendrocellus gestroi Bates’ ’ / ‘‘ Gestroi Bates’ ’ / ‘‘ Dendrocellus Gestroi Typus Bates’ ’ / ‘‘ Syntypus, Dendrocellus gestroi Bates 1892 ’’ [red label] /, ‘‘ Museo Civico di Genova’ ’ / ‘‘ LECTOTYPE, Dendrocellus gestroi Bates, designated by H. B. Liang & D. H. Kavanaugh 2006 ’’ [red label].	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFC5481329A3B92D27FDFB91.taxon	description	Description. Length 10.0 mm, width 3.2 mm. Black with blue luster, apical twothirds of Ant 1, Ant 2 and basal two-thirds of Ant 3 blackish brown, basal onethird of Ant 1, apical one-third of Ant 3 and Ant 4 – 11 brown; mandibles and palpi brown; tibiae and apical one-third of femora black, basal two-thirds of femora dark red, tarsi dark brown. Head with vertex slightly convex, densely punctate; Ant 1 / Ant 3 5 3.33; labrum with medial lobe slightly protruded anteriorly; genae long, distinctly tumid; eyes small, 1.9 times as long as genae. Pronotum densely punctate, PL / PW 5 1.36, HW / PW 5 1.06, widest slightly before middle; apical angles round, lateral margins straight and widened from apical angles to widest point, then narrowed posteriorly and sinuate to basal angles; basal angles acute, pointed laterally; lateral furrow distinct; basal foveae deep; middle line fine. Elytra short and wide, lateral margins widened from base to apical third, then narrowed to apex; EL / EW 5 1.56, EW / PW 5 2.22; intervals slightly convex, moderately punctate, interspace between two adjacent punctures 1.5 – 2.0 times diameter of punctures; outer angles strongly dentate, apex pointed (similar to Fig. 41). Tarsal claws with 5 – 6 long pectinations, length of individual pectination subequal to width of claw base (similar to Fig. 60).	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFC5481329A3B92D27FDFB91.taxon	distribution	Distribution. Myanmar.	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFC5481329A3B92D27FDFB91.taxon	discussion	Remarks. In his original description of D. gestroi, Bates gave no indication of the number of specimens in his type series. This means that, although only a single female specimen is to be found in the MCSN collection, bearing both ‘‘ typus’ ’ and ‘‘ syntypus’ ’ labels (see above), it must, according to the International Code of Zoological Nomenclature (Article 73.2), be considered a syntype and designated as a lectotype. The single known specimen of this species is similar to those of D. unidentatus (Macleay) in having acute elytral outer angles, long tarsal claw pectinations, and black tibiae, but differs from the latter in having a short pronotum (PL / PW 5 1.36, but 1.50 – 1.67 in D. unidentatus) and wider elytra (EL / EW 5 1.56, but 1.67 – 1.93 in D. unidentatus). Its acute elytral outer angles and long tarsal claw pectinations are different from those of D. nepalensis (Hope) and D. kulti (Jedlicka) adults. The dark red base of the femora and small eyes of this specimen (and presumably all members of this species) are apparently unique within the genus.	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFC4481C292AB85D2447FB92.taxon	description	(Figs. 11, 61, 99 – 100) Types. Holotype: a male (ISNB), ‘‘ Barway, P. Cardon’ ’ / ‘‘ Coll. R. I. Sc. N. B. Inde’ ’ / ‘‘ HOLOTYPE, Dendrocellus inexpectus Liang & Kavanaugh n. sp., designated by H. B. Liang & D. H. Kavanaugh 2006 ’’ [red label]. Paratypes (total 9 specimens): 1 male and 3 females (ISNB), ‘‘ Barway, P. Cardon’ ’ / ‘‘ Coll. R. I. Sc. N. B. Inde’ ’ / ’’ PARATYPE, Dendrocellus inexpectus Liang & Kavanaugh n. sp., designated by H. B. Liang & D. H. Kavanaugh 2006 ’’ [yellow label]; 2 males (ISNB), ‘‘ Konbir’ ’ / ‘‘ Coll. R. I. Sc. N. B. Inde’ ’ / ’’ PARATYPE, Dendrocellus inexpectus Liang & Kavanaugh n. sp., designated by H. B. Liang & D. H. Kavanaugh 2006 ’’ [yellow label]; 2 males (ISNB), ‘‘ Belgaum Bombay, H. E. Andrewes’ ’ / ‘‘ Desera geniculata Klug, det. H. E. Andrewes, 1920 ’’ / ‘‘ W. Hansen vid. 1966, Desera geniculata Klug’ ’ / ‘‘ Coll. R. I. Sc. N. B. Inde’ ’ / ’’ PARATYPE, Dendrocellus inexpectus Liang & Kavanaugh n. sp., designated by H. B. Liang & D. H. Kavanaugh 2006 ’’ [yellow label]; 1 male (ISNB), ‘‘ Barnai, 1892 ’’ / ‘‘ Collection E. Candeze’ ’ / ‘‘ Dendrocellus geniculatus Klug, M. Maindron det. 1910 ’’ / ‘‘ W. Hansen vid. 1966, Desera geniculata Klug’ ’ / ‘‘ Coll. R. I. Sc. N. B. 24 Inde’ ’ / ’’ PARATYPE, Dendrocellus inexpectus Liang & Kavanaugh n. sp., designated by H. B. Liang & D. H. Kavanaugh 2006 ’’ [yellow label].	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFC4481C292AB85D2447FB92.taxon	materials_examined	Type locality. Barway (near Chota Nagpur), India.	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFC4481C292AB85D2447FB92.taxon	diagnosis	Diagnosis. Femora brown with apical sixth or seventh dark brown; tibia brown; apical two thirds of Ant 1 black; elytral outer angles slightly dentate; aedeagus with apical lamella short, round at apex (Fig. 99).	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFC4481C292AB85D2447FB92.taxon	description	Description. Length 9.2 – 10.5 mm, width 2.6 – 3.1 mm. Black with green luster, mixed with some copper luster, apical third of Ant 1 black, basal two thirds of Ant 1, Ant 2 – 11 brown; mandibles and palpi brown; tibia and apical sixth or seventh of femora dark brown, basal two-thirds of femora brown. Head with vertex slightly convex, densely punctate; Ant 1 / Ant 3 5 3.54 (3.31 – 3.69); labrum with medial lobe slightly protruded anteriorly; genae short, distinctly tumid; eyes large, 2.7 times as long as genae. Pronotum densely punctate, PL / PW 5 1.48 (1.43 – 1.57), HW / PW 5 1.21 (1.16 – 1.27), widest slightly before middle; apical angles round; lateral margins slightly widened from apical angles to widest point, strongly narrowed behind, then sinuate to basal angles; basal angles obtuse, pointed laterally; lateral furrow indistinct; basal foveae shallow; middle line fine. Elytra short and wide, EL / EW 5 1.76 (1.69 – 1.85), EW / PW 5 2.35 (2.28 – 2.50); intervals nearly flat, densely punctate; outer angles slightly dentate. Protarsomere 3 slightly asymmetrical in male; tarsal claws with 5 – 6 long pectinations, length of individual pectination subequal to width of claw base (Fig. 61). Aedeagus with apical lamella thin, very short, apex round (Figs. 99 – 100).	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFC4481C292AB85D2447FB92.taxon	distribution	Distribution. Barway and Bombay in India.	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFC4481C292AB85D2447FB92.taxon	etymology	Etymology. This species is named for its discovery, which was unexpected.	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFC4481C292AB85D2447FB92.taxon	discussion	Remarks. Members of this species are very similar to those of D. geniculatus (Klug). In fact, we had not distinguished this species from D. geniculatus (Klug) until the specimens were dissected for genitalia comparison. Males can be distinguished from those of D. geniculatus (Klug) by the short and round lamella of the aedeagus. Moreover, femora of both sexes of this new species are brown with dark brown apex, whereas the femora of most D. geniculatus (Klug) specimens, including those of the holotype, are typically yellow with a black apex. Some specimens of D. geniculatus (Klug) do possess brown femora (e. g., from Vietnam and Sumatra), but the apex is still black, not dark brown.	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFCB481D291CB85C24CFFC5F.taxon	description	(Figs. 12, 34, 62, 89 – 90) Types. Holotype: a male (ISNB), ‘‘ Java, Seokaboemi’ ’ / ‘‘ R. I. Sc. N. B. 19.458, Coll. Le Moult’ ’ / ‘‘ Coll. R. I. Sc. N. B. Java’ ’ / ’’ HOLOTYPE, Dendrocellus javanus Liang & Kavanaugh n. sp., designated by H. B. Liang & D. H. Kavanaugh 2006 ’’ [red label]. Paratypes (total 5 specimens): 1 female (ISNB), ‘‘ Java, Soc. Ent. Belg., Coll. Putzeys’ ’ / ‘‘ Coll. R. I. Sc. N. B. Java’ ’ / ’’ PARATYPE, Dendrocellus javanus Liang & Kavanaugh n. sp., designated by H. B. Liang & D. H. Kavanaugh 2006 ’’ [yellow label]; 1 male and 1 female (ISNB), ‘‘ Java’ ’ / ‘‘ R. Mus. Hist. Nat. Belg. I. G. 12.595 ’’ / ‘‘ Coll. R. I. Sc. N. B. Java’ ’ / ’’ PARATYPE, Dendrocellus javanus Liang & Kavanaugh n. sp., designated by H. B. Liang & D. H. Kavanaugh 2006 ’’ [yellow label]; 1 female (ISNB), ‘‘ Java’ ’ / ‘‘ Soc. Ent. Belg. Coll. Putzeys’ ’ / ‘‘ Coll. R. I. Sc. N. B. Java’ ’ / ’’ PARATYPE, Dendrocellus javanus Liang & Kavanaugh n. sp., designated by H. B. Liang & D. H. Kavanaugh 2006 ’’ [yellow label]; 1 male (ZMHU), ‘‘ Java Occid. Dr. Standgr. ’’ / ‘‘ Dendrocellus geniculatus Klug’ ’ / ’’ PARA- TYPE, Dendrocellus javanus Liang & Kavanaugh n. sp., designated by H. B. Liang & D. H. Kavanaugh 2006 ’’ [yellow label].	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFCB481D291CB85C24CFFC5F.taxon	materials_examined	Type locality. Java, Indonesia.	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFCB481D291CB85C24CFFC5F.taxon	diagnosis	Diagnosis. Femora brown with apical fifth or sixth black; tarsal claws (Fig. 62) with longest pectination subequal to width of tarsal claw base; elytra with outer angles slightly dentate; aedeagus with apical lamella moderately long, slightly digitiform, not thickened at apex (Figs. 89 – 90).	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFCB481D291CB85C24CFFC5F.taxon	description	Description. Length 9.2 – 10.2 mm, width 2.7 – 2.9 mm. Black, head, pronotum, and elytra with green luster, mixed with slight blue luster; apical third of Ant 1 and basal two thirds of Ant 3 black, basal two thirds of Ant 1, apical third of Ant 3, and Ant 4 – 11 brown or yellow; apical fifth or sixth of femora black, remainder brown or yellow; tibiae and tarsi brown or yellow. Head (Fig. 34) wider than pronotum, HW / PW 5 1.18 (1.13 – 1.23); vertex convex, coarsely punctate; Ant 1 / Ant 3 5 3.35 (3.24 – 3.46); labrum with medial lobe slightly protruded anteriorly; genae moderately tumid; eyes large, about 2.2 times as long as genae. Pronotum densely punctate, narrow, PL / PW 5 1.46 (1.40 – 1.50); lateral margins gradually widened from apical angles to middle, strongly sinuate before basal angles; basal angles protruded laterally; basal foveae quite deep; middle line shallow. Elytra long, slightly widened from base to apical third, then narrowed to apex in both sexes (more so in female); EL / EW 5 1.82 (1.70 – 1.86), EW / PW 5 2.23 (2.12 – 2.32); outer angles slightly dentate (similar to Fig. 40); interval flat, densely punctate. Protarsomere 3 slightly asymmetrical in male; tarsal claw with 5 – 6 long pectinations, length of the longest pectination subequal to width of claw base (Fig. 62). Aedeagus with apical lamella moderately long, slightly digitiform, unthickened at apex (Figs. 89 – 90).	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFCB481D291CB85C24CFFC5F.taxon	distribution	Distribution. Presently known only from type locality.	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFCB481D291CB85C24CFFC5F.taxon	etymology	Etymology. The new species is named for Java, its type locality.	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFCB481D291CB85C24CFFC5F.taxon	discussion	Remarks. Members of this new species are similar to those of D. geniculatus (Klug) in having tumid genae, green body luster, and similar femoral and tibial color, but differ in having the ratio of Ant 1 / Ant 3 smaller and the lamella of aedeagus not thickened (Fig. 90). They are also similar to adults of D. ternatensis Chaudoir in the form of male genitalia, but differ from the latter in femoral color and in having more tumid genae (Fig. 34) and smaller ratio of Ant 1 / Ant 3.	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFCA481E296AB81B2632FD73.taxon	description	(Figs. 13 – 14, 35, 81 – 82)	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFCA481E296AB81B2632FD73.taxon	description	Description. Length 9.6 – 11.6 mm, width 3.2 – 3.9 mm. Black with blue luster; Ant 1 pale yellow or with apical third brown; femora black with blue luster, tibiae yellow (dark brown in specimen from Hubei). Head almost equal in width to 26 pronotum, HW / PW 5 1.03 (0.97 – 1.10), Ant 1 / Ant 3 5 3.39 (3.13 – 3.62); labrum with medial lobe slightly or moderately protruded anteriorly; genae short, distinctly tumid (less so in males); eyes large, slightly protruded laterally, 2.6 times as long as genae. Pronotum short, PL / PW 5 1.29 (1.19 – 1.35), widest at apical third. Elytra wide and short, El / EW 5 1.58 (1.51 – 1.66), EW / PW 5 2.14 (2.07 – 2.24), outer angles slightly round (Fig. 35); intervals slightly convex, sparsely punctate, punctures small, mixed with some larger ones. Protarsomere 3 slightly asymmetrical in male; tarsal claws with 3 – 5 short pectinations, length of individual pectination less than half width of claw base (similar to Fig. 64). Aedeagus with apical lamella thin, wide, linguiform (Figs. 81 – 82).	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFCA481E296AB81B2632FD73.taxon	distribution	Distribution. China (Hubei, Taiwan, Tibet, Yunnan).	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFCA481E296AB81B2632FD73.taxon	discussion	Remarks. We compared type specimens of Dendrocellus kulti kulti (Jedlicka) and D. kulti reconditus (Hansen) and found no significant difference, except that the former (length 9.6 mm) is slightly smaller than the latter (length 11.4 – 11.6 mm). Members of this species can be recognized by their short pronota, blue elytra and black femora. This seemed to us at first to be just a blue color variation of D. nepalensis (Hope); but the color is very stable in all specimens we examined, and we treat it as a distinct species. Dendrocellus kulti (Jedlicka) is distributed only in China, from Tibet to Taiwan, its range partly overlapping with that of D. nepalensis Hope.	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFC9481F28E2BEFF24E1FD50.taxon	description	(Figs. 15, 38, 70, 105 – 106) Types. Holotype: a male (ISNB), ‘‘ Rhodésie du Nord, Abercorn, 19 - V- 1943, H. - J. Brédo’ ’ / ‘‘ R. I. Sc. N. B. I. G. 15.472 ’’ / ‘‘ Coll. R. I. Sc. N. B. Rhodésie du Nord’ ’ / ‘‘ HOLOTYPE, Dendrocellus micropectinatus Liang & Kavanaugh n. sp., designated by H. B. Liang & D. H. Kavanaugh 2006 ’’ [red label]. Paratypes (total 12 specimens): 1 female (ISNB), same collecting data as holotype; 1 female (ISNB), ‘‘ Rhodésia du Nord, Abercorn, XI- 1943, H. - J. Brédo’ ’ / ‘‘ R. Mus. Hist. Nat. Belg. I. G. 15.472 ’’ / ‘‘ Coll. R. I. Sc. N. B. Rhodésie du Nord’ ’ / ’’ PARATYPE, Dendrocellus micropectinatus Liang & Kavanaugh n. sp., designated by H. B. Liang & D. H. Kavanaugh 2006 ’’ [yellow label]; 1 male and 6 females (ISNB), ‘‘ Coll. R. I. Sc. N. B. Congo belg. Kafakumba, coll. Le Moult’ ’ / ’’ PARATYPE, Dendrocellus micropectinatus Liang & Kavanaugh n. sp., designated by H. B. Liang & D. H. Kavanaugh 2006 ’’ [yellow label]; 2 males (MBC), ‘‘ Zambia, 8.11.02, Mwinilunga, ca 20 km N-W, leg. Wachtel’ ’ / ’’ PARATYPE, Dendrocellus micropectinatus Liang & Kavanaugh n. sp., designated by H. B. Liang & D. H. Kavanaugh 2006 ’’ [yellow label]; 1 female (MBC), ‘‘ leg. Wachtel, Zambia, Mwinilunga, 20 km NW, 8.11.2002 ’’ / ’’ PARATYPE, Dendrocellus micropectinatus Liang & Kavanaugh n. sp., designated by H. B. Liang & D. H. Kavanaugh 2006 ’’ [yellow label].	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFC9481F28E2BEFF24E1FD50.taxon	materials_examined	Type locality. Abercorn, Zambia.	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFC9481F28E2BEFF24E1FD50.taxon	diagnosis	Diagnosis. Pronotum red with purple luster; femora with apical half black, tibiae black; tarsal claw pectinations extremely short, the longest pectination shorter than one-fifth width of claw base (Fig. 70).	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFC9481F28E2BEFF24E1FD50.taxon	description	Description. Length: 9.8 – 10.9 mm, width: 2.9 – 3.4 mm. Head and abdominal sterna with blue luster; neck, pronotum, scutellum, thoracic sterna, episterna, and epimera, Ant 4 – 11, and basal half of femora red or brown; pronotum with purple luster; elytra with green luster; apical one-third or apical half of hind femora, tibiae, and apical half of Ant 1 black; tarsomeres, Ant 2, Ant 3, and basal half of Ant 1 dark brown. Head densely punctate on vertex; labrum with medial lobe slightly protruded anteriorly; Ant 1 / Ant 3 5 3.02 (2.88 – 3.14); genae oblique, hardly tumid; eyes large, convex, 2.5 times as long as genae. Pronotum short, cylindrical, widest at apical third, PL / PW 5 1.38 (1.34 – 1.43), HW / PW 5 1.19 (1.11 – 1.26); slightly widened from apical angles to anterior one-third, then narrowed posteriad, slightly sinuate before basal angles; basal angles obtuse, weakly protruded laterally. Elytra nearly parallel in basal half, slightly widened posteriad to apical fourth, then narrowed to apex; EL / EW 5 1.85 (1.79 – 1.99), EW / PW 5 2.16 (1.95 – 2.26); striae shallow, punctate; intervals flat, densely punctate; outer apical angles nearly rectangular, not pointed (Fig. 38). Protarsomere 3 strongly asymmetrical in male; tarsal claws with 3 – 5 min pectinations, length of the longest pectination less than one-fifth width of claw base (Fig. 70). Aedeagus stout, with apical lamella moderately long, thin, rounded at apex (Figs. 105 – 106).	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFC9481F28E2BEFF24E1FD50.taxon	distribution	Distribution. Congo, Zambia.	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFC9481F28E2BEFF24E1FD50.taxon	etymology	Etymology. This species is named for the very short pectinations on its tarsal claws.	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFC9481F28E2BEFF24E1FD50.taxon	discussion	Remarks. Members of this new species are similar to those of another African Dendrocellus species, D. bicoloripennis n. sp., but differ from them in having unicolorous green elytra with distinct humeri and nearly parallel lateral margins. In D. bicoloripennis adults, the elytra are bicolored bluish green and dark purple or black with indistinct humeri and lateral margins distinctly widened posteriorly.	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFC8481829A7BF0226E1F986.taxon	description	(Figs. 16, 36, 68, 79 – 80)	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFC8481829A7BF0226E1F986.taxon	description	Description. Length 9.8 – 11.7 mm, width 3.2 – 3.8 mm. Black, head and pronotum with blue luster; elytra with purple luster, and in some specimens mixed with slight blue or green luster; Ant 1 yellow or with apical fourth brown; femora black with blue luster, tibiae yellow. Head equal to or slightly wider than pronotum, HW / PW 5 1.06 (1.00 – 1.11); Ant 1 / Ant 3 5 3.31 (3.07 – 3.48); labrum with medial lobe slightly or moderately protruded anteriorly; genae short, moderately tumid; eyes large, convex, 2.7 times as long as genae. Pronotum short, PL / PW 5 1.30 (1.25 – 1.39), widest at apical third. Elytra short, EW / PW 5 2.18 (2.00 – 2.31), EL / EW 5 1.63 (1.53 – 1.70), widened posteriad to apical fourth, then slightly narrowed to apex; outer angles broadly rounded (Fig. 36); intervals slightly convex, sparsely punctate, punctures small, mixed with some larger ones. Protarsomere 3 slightly asymmetrical in male; tarsal claws with 4 – 5 short pectinations, length of the longest pectination less than half width of claw base (Fig. 68). Aedeagus with apical lamella wide, thin, linguiform (Figs. 79 – 80).	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFC8481829A7BF0226E1F986.taxon	distribution	Distribution. Bangadesh, Bhutan, China, India, Myanmar, Nepal, Sikkim, Vietnam.	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFC8481829A7BF0226E1F986.taxon	discussion	Remarks. We have not examined the type. Members of this species are readily recognized by their short pronota and purple elytra. They are very similar to D. kulti Jedlicka members, but differ in having purple elytra and more rounded elytral outer angles (Figs. 35 – 36).	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFCE48192928BCCF2504FB7E.taxon	description	(Figs. 17, 46, 63, 85 – 86) Types. Holotype: a male (ISNB), ‘‘ Kina Balu, Borneo’ ’ / ‘‘ Collection P. Dupuis’ ’ / ‘‘ Coll. R. I. Sc. N. B. ’’ / ‘‘ HOLOTYPE, Dendrocellus nigripennis Liang & Kavanaugh n. sp., designated by H. B. Liang & D. H. Kavanaugh 2006 ’’ [red label]. Paratypes (total 2 specimens): 1 male and 1 female (ISNB), ‘‘ Bornéo’ ’ / ‘‘ Collection P. Dupuis’ ’ / ‘‘ Coll. R. I. Sc. N. B. ’’ / ‘‘ PARATYPE, Dendrocellus nigripennis Liang & Kavanaugh n. sp., designated by H. B. Liang & D. H. Kavanaugh 2006 ’’ [yellow label].	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFCE48192928BCCF2504FB7E.taxon	materials_examined	Type locality. Mount Kinabalu, Pulau Kalimantan.	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFCE48192928BCCF2504FB7E.taxon	diagnosis	Diagnosis. Elytra black with very slight blue luster, intervals sparsely punctate (Fig. 46); femora with apical half black, basal half brown; tibiae and tarsi black; protarsomere 3 symmetrical in male; tarsal claws with 4 – 5 pectinations, length of the longest pectination nearly half width of claw base (Fig. 63); aedeagus with lamella wide, apex truncate (Figs. 85 – 86).	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFCE48192928BCCF2504FB7E.taxon	description	Description. Length 9.2 – 11.0 mm, width 2.8 – 3.4 mm. Black, head and pronotum with blue luster; elytra with very slight blue luster, restricted to outer intervals in many specimens; tibiae, tarsi, apical half of femora, and apical fourfifths of Ant 1 black, Ant 2, Ant 3, and basal fifth of Ant 1 dark brown; Ant 4 – 11 and palpi brown. Head wider than pronotum, HW / PW 5 1.22 (1.21 – 1.23); vertex convex, coarsely punctate; labrum with medial lobe slightly protruded anteriorly; Ant 1 / Ant 3 5 3.48 (3.40 – 3.56); genae slightly tumid; eyes large, about 2.4 times as long as genae. Pronotum coarsely punctate, PL / PW 5 1.65 (1.63 – 1.70); lateral margins strongly sinuate before basal angles; basal angles protruded laterally; basal foveae deep. Elytra short, EL / EW 5 1.66 (1.62 – 1.68), EW / PW 5 2.41 (2.33 – 2.58); lateral margins slightly emarginate in the basal fourth, widened posteriad to apical third, then narrowed to apex; outer angles slightly dentate, pointed; intervals convex, sparsely punctate, punctures small, mixed with some larger ones, interspace between two adjacent punctures 1.5 – 2.0 times diameter of punctures (Fig. 46). Protarsomere 3 symmetrical in male and female; tarsal claws with 4 – 5 pectinations, length of the longest pectination subequal to the width of claw base (Fig. 63). Aedeagus with apical lamella wide, apex truncate (Figs. 85 – 86).	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFCE48192928BCCF2504FB7E.taxon	distribution	Distribution. Known only from the type locality.	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFCE48192928BCCF2504FB7E.taxon	etymology	Etymology. The new species is named for its black elytra.	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFCE48192928BCCF2504FB7E.taxon	discussion	Remarks. Initially, we assumed this to be just a black variety of D. crassus (Andrewes). However, the color of the femora and shape of the apex of the aedeagus (Fig. 85) are both very different from those of D. crassus members, so we treat this as a distinct species.	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFCE481A2905B8F92491FB69.taxon	description	(Figs. 18, 31, 66, 91 – 92) Types. Holotype: a male (ISNB), ‘‘ Queensland’ ’ / ‘‘ Collection P. Dupuis’ ’ / ‘‘ Coll. R. I. Sc. N. B., Australia-Queensl. ’’ / ‘‘ Desera smaragdina Ch., dét. P. Dupuis’ ’ / ‘‘ Desera smaragdina Ch. ’’ / ‘‘ HOLOTYPE, Dendrocellus queenslandicus Liang & Kavanaugh n. sp., designated by H. B. Liang & D. H. Kavanaugh 2006 ’’ [red label]. Paratypes (total 4 specimens): 1 male (ISNB), ‘‘ Queensland’ ’ / ‘‘ Collection P. Dupuis’ ’ / ‘‘ Coll. R. I. Sc. N. B., Australia-Queensl. ’’ / ‘‘ Desera smaragdina Ch., dét. P. Dupuis’ ’ / ‘‘ Desera smaragdina Ch. ’’ / ‘‘ PARATYPE, Dendrocellus queenslandicus Liang & Kavanaugh n. sp., designated by H. B. Liang & D. H. Kavanaugh 2006 ’’ [yellow label]; 1 male (ISNB), ‘‘ Queensland’ ’ / ‘‘ Collection P. Dupuis’ ’ / ‘‘ Coll. R. I. 30 Sc. N. B. ’’ / ‘‘ PARATYPE, Dendrocellus queenslandicus Liang & Kavanaugh n. sp., designated by H. B. Liang & D. H. Kavanaugh 2006 ’’ [yellow label]; 1 male (CMNH), ‘‘ Mainoru, ENE of Katherine, N. T., 14 Dec. 1982, A. Walford- Huggins’ ’ / ‘‘ Walford-Huggins collection, Carnegie Museum Accession 35338 ’’ / ‘‘ Desera smaragdina Chd’ ’ / Desera smaragdina (Chaudoir) det. R. Davidson’ ’ / ’’ PARATYPE, Dendrocellus queenslandicus Liang & Kavanaugh n. sp., designated by H. B. Liang & D. H. Kavanaugh 2006 ’’ [yellow label]; 1 male (CMNH), ‘‘ Australia QLD. 7 km. NE / Tolga, May 8, 1989, R. I Storey, at light’ ’ / ‘‘ Desera smaragdina (Chaudoir) det. R. Davidson’ ’ / ’’ PARATYPE, Dendrocellus queenslandicus Liang & Kavanaugh n. sp., designated by H. B. Liang & D. H. Kavanaugh 2006 ’’ [yellow label].	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFCE481A2905B8F92491FB69.taxon	materials_examined	Type locality. Queensland, Australia.	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFCE481A2905B8F92491FB69.taxon	diagnosis	Diagnosis. Elytra with parallel sides, intervals densely punctate; femora with apical third black, basal two-thirds brown; tibiae and tarsi yellow; protarsomere 3 asymmetrical in male; tarsal claws with 5 pectinations, length of the longest pectination subequal to half width of claw base (Fig. 66); aedeagus with apical lamella thin, digitiform, unthickened at apex (Fig. 91).	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFCE481A2905B8F92491FB69.taxon	description	Description. Length 9.0 – 9.9 mm, width 2.5 – 2.8 mm. Black, head and pronotum with blue luster mixed with slight green luster; elytra with green luster mixed with slight blue luster; apical two thirds of Ant 1 black, basal third dark brown, Ant 2 and Ant 3 dark brown, Ant 4 – 11 brown; apical third of femora black, basal twothirds brown; tibiae and tarsi yellow. Head much wider than pronotum, HW / PW 5 1.34 (1.27 – 1.37); vertex convex, coarsely punctate; Ant 1 / Ant 3 5 3.31 (3.23 – 3.38); labrum with medial lobe slightly protruded anteriorly; genae short, oblique; eyes large, about 2.8 times as long as genae (Fig. 31). Pronotum coarsely punctate, narrow, PL / PW 5 1.64 (1.60 – 1.71); lateral margins slightly sinuate before basal angles; basal angles almost rectangular; basal foveae deep. Elytra long, lateral margins almost parallel, EL / EW 5 2.04 (2.00 – 2.06), EW / PW 5 2.21 (2.16 – 2.30); outer angles slightly dentate, pointed; intervals convex, densely punctate, interspace between two adjacent punctures narrower than diameter of punctures. Protarsomere 3 slightly asymmetrical in male; tarsal claws with 5 pectinations, length of the longest pectination subequal to half width of claw base (Fig. 66). Aedeagus with apical lamella thin, narrow, not thickened at apex (Figs. 91 – 92).	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFCE481A2905B8F92491FB69.taxon	distribution	Distribution. Known from Queensland and Northern Territory, Australia.	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFCE481A2905B8F92491FB69.taxon	etymology	Etymology. The new species is named for its type locality.	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFCE481A2905B8F92491FB69.taxon	discussion	Remarks. This new species at first appears to be just a variety of D. ternatensis Chaudoir, based on the similarity in genitalia. However, the longer black portion of the femora and longer, parallel-sided elytra of its members are quite different from those of D. ternatensis members, so we treat this as a distinct species.	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFCD481B2982B8E52460FA07.taxon	description	(Figs. 7, 29, 33, 37, 57, 75 – 76)	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFCD481B2982B8E52460FA07.taxon	materials_examined	Other specimens examined. Total 6 specimens: China: 1 male, ‘‘ Guangxi, Tianlin, Lidongshan, 1,300 – 1,400 m, 2002. V. 28, Jianwen Liu collector’ ’; 1 male, ‘‘ Guangxi, Tianlin, Langpin, 1,300 m, 2002. VI. 1, Jianwen Liu collector’ ’; 1 female, ‘‘ Guangxi, Tianlin, Yaojiawan, 1,200 – 1,400 m, 2002. VI. 1, Xiujuan Yang collector’ ’. India: 1 female (MNHN), ‘‘ Ex Musaeo Chaudoir’ ’ / ‘‘ Museum Paris’ ’ / ‘‘ rugicollis Chaudoir, flavipes Schmidt-Göbel, Indes. Bor. Or. C. Laferte´ ’’; 1 male (MNHN), ‘‘ nepalensis Hope’ ’ / ‘‘ Ex Musaeo Chaudoir’ ’ / ‘‘ Museum Paris’ ’ / ‘‘ rugicollis Chaudoir, flavipes Schmidt-Göbel, Indes. Bor. Or. C. Laferte´ ’’; 1 male (MNHN), ‘‘ Sabatoa, Boucard’ ’ / ‘‘ Ex Musaeo Chaudoir’ ’ / ‘‘ rugicollis Chaudoir, flavipes Schmidt-Göbel, Indes. Bor. Or. C. Laferte´ ’’.	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFCD481B2982B8E52460FA07.taxon	description	Description. Length 9.4 – 10.0 mm, width 2.8 – 3.2 mm. Black with blue and green luster (head and pronotum with blue luster only in the paralectotype specimen); palpi, tibia, basal two-thirds of Ant 1, Ant 2 – 11, basal six sevenths of femora yellow; apical third of Ant 1, apical seventh of femora brown or black. Head (Fig. 33) with vertex slightly convex; Ant 1 / Ant 3 5 3.16 (2.93 – 3.31); labrum with medial lobe moderately protruded anteriorly (Fig. 29); genae short, slightly tumid; eyes large, about 2.5 times as long as genae. Pronotum cylindrical, HW / PW 5 1.14 (1.11 – 1.19), PL / PW 5 1.40 (1.34 – 1.48); lateral margins widened from apical angles to middle, then narrowed posteriad, sinuate before basal angles; basal angles nearly rectangular; middle line very shallow. Elytra with lateral margins slightly widened posteriad from base to apical third, then narrowed to apex (both sexes); outer angles obtuse, not pointed (Fig. 37); EL / EW 5 1.83 (1.71 – 1.92), EW / PW 5 2.20 (2.06 – 2.38); striae shallow; intervals densely punctate. Protarsomere 3 slightly asymmetrical in male; tarsal claw slender, with 5 – 6 long pectinations, length of individual pectination longer width of claw base (Fig. 57). Aedeagus stout, with apical lamella wide, linguiform (Figs. 75 – 76) Variation. Apical seventh of femora and apical third of Ant 1 are paler (brown) in lectotype and paralectotype than in others (black).	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFCD481B2982B8E52460FA07.taxon	distribution	Distribution. China, India, Nepal.	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFCD481B2982B8E52460FA07.taxon	discussion	Remarks. Schmidt-Göbel has been recognized as the author of this species, but this is incorrect. When he described his new genus, Dendrocellus (Schmidt-Göbel 1846), he included Drypta flavipes Wiedemann in that genus and redescribed that species, based on specimens that he had misidentified. He clearly did not intend to describe a new species at that time. Chaudoir (1861: 546) recognized that Schmidt- Göbel’s description of D. flavipes Wiedemann actually represented a distinct and new species, for which he provided the name, Dendrocellus rugicollis. According to Article 72.4.2 of the International Code of Zoological Nomenclature, when a new nominal species group taxon is based on a published misidentification by an earlier author, the type series consists of or includes the specimens that had been misidentified. Consequently, the specimens studied by Schmidt-Göbel constitute the type series for D. rugicollis Chaudoir; and we have selected one of two Schmidt- Göbel specimens in NMPC as lectotype, the other as paralectotype. Andrewes (1921: 173 [in a footnote], and again in 1923: 8) treated the name D. flavipes Schmidt-Göbel as a synonym of Dendrocellus geniculatus (Klug). When we examined Schmidt-Göbel’s specimens, we found that they possess quite long tarsal claw pectinations (Fig. 57) and an aedeagus with linguiform lamella; and this species, therefore, is clearly distinct from D. geniculatus (Klug).	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFCC481B2950B9B0259EFA4B.taxon	description	(Figs. 19, 59)	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFF348252910B8AC2462FA94.taxon	description	(Figs. 20, 27, 41, 60, 101 – 102) Types. Holotype: a male (CAS), ‘‘ Wang Sa Shui, S. Kiangsi, S. China, 7 - 11 - 36 ’’ / ‘‘ L. Gressitt Collector’ ’ / ‘‘ HOLOTYPE, Dendrocellus sinicus Liang & Kavanaugh n. sp., designated by H. B. Liang & D. H. Kavanaugh 2006 ’’ [red label] / ‘‘ California Academy of Sciences Type No. 18233 ’’. Paratypes (total 6 specimens): 1 male (CAS), ‘‘ Tai Au Hong, S. Kiangsi, ’’ / ‘‘ L. Gressitt Collector’ ’ / ‘‘ L. Gressitt Collection’ ’ / ‘‘ PARATYPE, Dendrocellus sinicus Liang & Kavanaugh n. sp., designated by H. B. Liang & D. H. Kavanaugh 2006 ’’ [yellow label]; 1 female (CAS), ‘‘ Gang-keu, SW. Fukien, S. China’ ’ / ’’ L. Gressitt Collector’ ’ / ‘‘ L. Gressitt Collection’ ’ / ‘‘ PARATYPE, Dendrocellus sinicus Liang & Kavanaugh n. sp., designated by H. B. Liang & D. H. Kavanaugh 2006 ’’ [yellow label]; 2 females, ‘‘ Guangdong, Xinfeng, September-October 1983, Wenlian Huang collector’ ’ / ‘‘ PARATYPE, Dendrocellus sinicus Liang & Kavanaugh n. sp., designated by H. B. Liang & D. H. Kavanaugh 2006 ’’ [yellow label]; 1 female, ‘‘ Guangdong, Xinfeng, August 1983, Minlian Huang collector’ ’ / ‘‘ PARATYPE, Dendrocellus sinicus Liang & Kavanaugh n. sp., designated by H. B. Liang & D. H. Kavanaugh 2006 ’’ [yellow label]; 1 female, ‘‘ Guangdong, Xinfeng, September 1983 ’’ / ‘‘ PARATYPE, Dendrocellus sinicus Liang & Kavanaugh n. sp., designated by H. B. Liang & D. H. Kavanaugh 2006 ’’ [yellow label].	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFF348252910B8AC2462FA94.taxon	materials_examined	Type locality. Jiangxi, China.	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFF348252910B8AC2462FA94.taxon	diagnosis	Diagnosis. Elytra with sharp outer angles (Fig. 41), intervals densely punctate; femora with apical third black, basal-two thirds brown; tibiae and tarsi yellow, protarsomere 3 asymmetrical in male; tarsal claws with 5 – 6 long pectinations, length of the longest pectination equal to width of claw base (Fig. 60).	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFF348252910B8AC2462FA94.taxon	description	Description. Length 10.6 – 11.0 mm, width 3.1 – 3.4 mm. Black, head and pronotum with blue luster mixed with slight green luster; elytra with green luster mixed with slight coppery luster; apical third of Ant 1 dark brown, basal twothirds brown, Ant 2 and Ant 3 brown, Ant 4 – 11 yellow; apical third of femora black, basal two thirds brown; tibiae and tarsi yellow. Head wider than pronotum, HW / PW 5 1.22 (1.18 – 1.27); vertex convex; Ant 1 / Ant 3 5 3.53 (3.47 – 3.64); labrum (Fig. 27) coarsely punctate, with medial lobe slightly protruded anteriorly; genae long, moderately tumid; eyes large, 2.5 times as long as genae. Pronotum densely punctate, narrow, PL / PW 5 1.53 (1.47 – 1.60); lateral margins gradually widened from apical angles to middle, sinuate before basal angles; basal angles protruded laterally, nearly rectangular; basal foveae very deep; middle line shallow. Elytra short, EL / EW 5 1.75 (1.69 – 1.79), EW / PW 5 2.19 (2.11 – 2.33); lateral margins widened posteriad to apical third, then narrowed to apex; outer angles strongly dentate, pointed (Fig. 41); intervals convex, densely punctate, interspace between two adjacent punctures narrower than diameter of punctures. Protarsomere 3 slightly asymmetrical in male; tarsal claw with 6 – 7 pectinations, the longest pectination subequal to the width of claw base (Fig. 60). Aedeagus with apical lamella short, round at apex (Figs. 101 – 102). Variation. The pronotum in the holotype is shorter (PL / PW 5 1.47), and lateral margins are more rounded than in the paratypes (PL / PW 5 1.52 – 1.60).	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFF348252910B8AC2462FA94.taxon	distribution	Distribution. Presently known from three provinces of China: Jiangxi, Fujian, and Guangdong.	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFF348252910B8AC2462FA94.taxon	etymology	Etymology. This new species is named for China, the country of its type locality.	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFF348252910B8AC2462FA94.taxon	discussion	Remarks. Members of this new species are similar to those of D. unidentatus (Macleay). However, the lateral margins of the pronota are more rounded before the middle (Fig. 20), the femora have a longer black portion, the tibiae are yellow, and the genae are more rounded than in D. unidentatus members. They are similar also to D. schultzei (Heller) members in shape of the pronotum and elytral outer angles, but differ from the latter in having green and coppery elytra and their tibiae and bases of femora all pale yellow (all black in D. schultzei).	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFF24826299EB9552618FBB3.taxon	description	(Fig. 21)	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFF24826299EB9552618FBB3.taxon	description	34 Description. Length 10.0 mm, width 3.0 mm. Black with green luster, apical half of Ant 1 and basal two-thirds of Ant 3 black, basal half of Ant 1, apical third of Ant 3, and Ant 4 – 11 brown; mandibles and palpi brown; apical sixth of femora black, basal two-thirds of femora brown; tibiae and tarsi nearly black (especially hind ones). Head with vertex convex, densely punctate; Ant 1 / Ant 3 5 3.60; labrum with medial lobe slightly protruded anteriorly; genae distinctly tumid; eyes large, 2.5 times as long as genae. Pronotum densely punctate, PL / PW 5 1.46, HW / PW 5 1.14, widest slightly before the middle; apical angles round, lateral margins slightly widened from apical angles to widest point, gradually narrowed behind, then strongly sinuate to basal angles; basal angles obtuse, pointed laterally; lateral furrow indistinct; basal foveae shallow; middle line fine. Elytra long, lateral margins slightly widened to apical third, then narrowed to apex; EL / EW 5 1.87, EW / PW 5 2.35 (2.28 – 2.50); intervals nearly flat, densely punctate; outer angles slightly dentate. Tarsal claws with 5 long pectinations, length of individual pectination subequal to width of claw base.	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFF24826299EB9552618FBB3.taxon	distribution	Distribution. Australia.	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFF24826299EB9552618FBB3.taxon	discussion	Remarks. The type locality is questionable. Moore et al. (1987: 311) stated the following: ‘‘ … Melbourne, Vic., this locality seems to be an error as the species distribution is limited to northern Qld. ’’. The actual distribution of D. smaragdinus needs further clarification. The description of D. smaragdinus was based on a single female specimen. It may be simply a dark variation of D. geniculatus (Klug), with blackish tibia and tarsi. Comparison of male genitalia is required to resolve the status of this nominal species. Since its original description, D. smaragdinus has been compared only by Sloane (1907: 473, based on his specimen, not the type) and only with his D. elegans. However, we have found that many of Sloane’s comparisons were inaccurate. For example, he claimed that D. elegans members have (1) pronota with lateral margins more deeply sinuate posteriorly and more strongly reflexed laterally toward the base, and (2) elytra wider, more ampliate, and more declivous basally than D. smaragdinus members. This is incorrect, and we suspect that his comparison was based on a D. queenslandicus specimen rather than one of D. smaragdinus.	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFF14827298AB83F2629FBB7.taxon	description	(Figs. 22, 32, 93 – 94)	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFF14827298AB83F2629FBB7.taxon	materials_examined	Type. Lectotype, here designated, a female (MHNP), labeled ‘‘ Ex Musaeo, Chaudoir’ ’ / ‘‘ Paris Museum’ ’ / ‘‘ ternatensis Chaud. Ternate, Deyrolle’ ’ / ‘‘ LECTO- TYPE, Dendrocellus ternatensis Chaudoir, designated by H. B. Liang & D. H. Kavanaugh 2006 ’’ [red label]. Other specimens examined. Total 10 specimens: Indonesia: 1 female (holotype, MNHN), ‘‘ Ex Musaeo, Chaudoir’ ’ / ‘‘ Paris Museum’ ’ / ‘‘ ternatensis Chaud. Ternate, Deyrolle’ ’; 1 male, (ZMHU), ‘‘ Batchian, Schaum’ ’ [Bacan, Indonesia] / ‘‘ 41699 ’’ / ‘‘ No. 95, examined by Prof. Thaxter for Laboulbeniaceae’ ’ / ‘‘ tenatensis ’’; 2 females (CAS), ‘‘ Tondano, N. Celebis, VI- 1 - 32 ’’ / ‘‘ Van Braekel Collector’ ’ / F. C. Hadden Collection’ ’; 1 male and 1 female (CAS), ‘‘ Indonesia, Moluccas, Ambon Island, 70 m, 21. V. 1960, A. M. R. Wegner, Coll. at light’ ’; 1 male (CAS), ‘‘ Indonesia, Moluccas, Ambon Island, 70 m, 18. VI. 1960, A. M. R. Wegner, Coll. at light’ ’; 1 female (CAS), ‘‘ Indonesia, Moluccas, Ambon Island, 70 m, 13. V. 1960, A. M. R. Wegner, Calif. Academy of Scc. 1972 ’’; 1 female (CAS), ‘‘ Indonesia, Moluccas, Ambon Island, 70 m, 6. V. 1961, A. M. R. Wegner, Coll. at light’ ’; 1 male (CAS), ‘‘ Indonesia, Moluccas, Ambon Island, 70 m, 17. I. 1962, A. M. R. Wegner, Coll. at light’ ’; 1 male (CMNH), ‘‘ Indonesia, Sulawesi Utara, Dumoga-Song N. P., 23 Aug. 1985, Coll. Chen Young’ ’ / ‘‘ Base Camo 211 m, Project Wallace’ ’.	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFF14827298AB83F2629FBB7.taxon	description	Description. Length 8.6 – 10.3 mm, width 2.7 – 3.2 mm. Black with green luster, apical third of Ant 1 and basal two-thirds of Ant 3 black or dark brown, basal third of Ant 1, apical third of Ant 3, and Ant 4 – 11 brown; mandibles and palpi brown; apical fourth of femora black, basal three fourths yellow; tibiae and tarsi yellow. Head (Fig. 32) with vertex convex, densely punctate; Ant 1 / Ant 3 5 3.60 (3.46 – 3.86); labrum with medial lobe slightly protruded anteriorly; genae short, slightly tumid; eyes large, 2.9 times as long as genae. Pronotum densely punctate, PL / PW 5 1.52 (1.48 – 1.59), HW / PW 5 1.21 (1.13 – 1.28), widest slightly before the middle; apical angles round, lateral margins slightly widened from apical angles to widest point, gently narrowed behind, then strongly sinuate to basal angles; basal angles obtuse, pointed laterally; lateral furrow indistinct; basal foveae shallow; middle line fine. Elytra long, lateral margins slightly widened to apical third, then narrowed to apex; EL / EW 5 1.79 (1.73 – 1.91), EW / PW 5 2.33 (2.23 – 2.40); intervals nearly flat, densely punctate; outer angles slightly dentate, pointed. Protarsomere 3 slightly asymmetrical in male; tarsal claw with 5 long pectinations, length of individual pectination subequal to width of claw base. Aedeagus with apical lamella thin, slightly digitiform, not thickened at apex (Figs. 93 – 94).	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFF14827298AB83F2629FBB7.taxon	distribution	Distribution. Indonesia (Ambon, N. Sulawesi, Ternate).	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFF14827298AB83F2629FBB7.taxon	discussion	Remarks. This species was described based on a single female specimen. Specimens from Ambon, northern Sulawesi, and Batchian are similar to the type from Ternate, based on color of femora (apical fourth black) and ratio of Ant 1 / Ant 3 (3.46 – 3.86), but slightly smaller (length 8.2 – 9.5 mm) than the type (length 10.3 mm). The apical black portions of the femora in the specimen from Sulawesi are slightly shorter than in other specimens of this species, nearly as in specimens of D. javanaus in this regard, but different from the latter in having oblique, only slightly tumid genae (genae less oblique and moderately tumid in D. javanus specimens).	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFF048202991B82326CEFD27.taxon	description	(Figs. 23, 43, 97 – 98)	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFF048202991B82326CEFD27.taxon	description	Description. Labrum with medial lobe slightly protruded anteriorly; genae long, distinctly tumid; eyes large, about 2.3 times as long as genae; lateral margins of pronotum nearly parallel before middle (Fig. 23); outer apical angles of elytra strongly dentate, pointed (Fig. 43); protarsomere 3 slightly asymmetrical in male; aedeagus with apical lamella short, round at apex (Figs. 97 – 98). For a more comprehensive description, see Liang et al. (2004: 381) and Andrewes (1919: 167).	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFF048202991B82326CEFD27.taxon	distribution	Distribution. China, India, Indonesia (Java, Sumatra), Malaysia, Myanmar, Pakistan, Thailand, Vietnam.	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFF748212938BE9025FEFB0B.taxon	description	(Figs. 24, 51, 54, 103 – 104)	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFF748212938BE9025FEFB0B.taxon	description	Description. Length 11.6 – 12.8 mm, width 3.4 – 3.8 mm. Black with green and blue luster; apical third of Ant 1 black, basal two-thirds red or brown; femora brown with apical fifth or sixth black; tibiae brown; scutellum, and meso- and metasternum brown, prosternum and metepisterna also brown in some specimens. Head wider than pronotum, HW / PW 5 1.19 (1.15 – 1.23); Ant 1 / Ant 3 5 3.94 (3.69 – 4.08); medial lobe of labrum slightly protruded anteriorly; genae very long, slightly tumid; eyes large, about 2.3 times as long as genae. Pronotum narrow, widest slightly before the middle; PL / PW 5 1.52 (1.47 – 1.55); lateral margins slightly widened from apical angles to widest point, gently narrowed behind, then slightly sinuate to basal angles; basal angles obtuse, pointed laterally; lateral furrow indistinct; basal foveae shallow; middle line fine. Elytra with lateral margins slightly widened to apical fourth, then narrowed to apex; outer angles acute, very slightly dentate; EL / EW 5 1.85 (1.70 – 1.94), EW / PW 5 2.33 (2.24 – 2.36); intervals flat, densely punctate. Protarsomere 3 strongly asymmetrical in male (Fig. 51); tarsal claws with 5 – 6 long pectinations, length of the longest pectination subequal to width of claw base (similar to Fig. 65). Aedeagus stout, with apical lamella narrow, digitiform, thickened at apex (Figs. 103 – 104).	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFF748212938BE9025FEFB0B.taxon	distribution	Distribution. Angola, Cameroon, Congo, Gabon, Gambia, Senegal, Sierra Leone, Tanzania.	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFF748212938BE9025FEFB0B.taxon	discussion	Remarks. We have not examined the types and have simply followed Basilewsky’s (1960) interpretation of the names and synonymies. All members of this species have brown meso- and metasterna and some also have the prosternum brown. In these and other features, they are very similar to another African species, D. australis; but they differ from the latter in having blue pronota, longer ratio of Ant 1 / Ant 3, and brown tibiae.	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFD0480728A7BCCF256BFB5F.taxon	description	1967: 277.	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFD0480728A7BCCF256BFB5F.taxon	type_taxon	Type species. Dendrocellus discolor Schmidt-Göbel 1846 (5 Desera nepalensis Hope 1831), designated by Andrewes 1939.	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFD0480728A7BCCF256BFB5F.taxon	description	Description. Body narrow, elongate (Figs. 1 – 24), length 8.0 – 14.0 mm, width 2.5 – 4.2 mm. Black with blue or green luster, in some species mixed with coppery or purple luster; in some species, the pronotum red or brown, sternum brown with blue luster. Microsculpture distinct, forming isodiametric meshes. Head punctate and pubescent; Ant 1 elongate, 2.5 – 3.8 times as long as Ant 3; mandibles about 3.0 times as long as labrum, hooked at apex; maxillary and labial apical palpomeres securiform; labrum with medial lobe slightly (Figs. 25 – 27), moderately (Figs. 28, 29), or strongly (Fig. 30) protruded anteriorly; genae short and oblique (Figs. 31, 32), or long and distinctly tumid (Figs. 33, 34). Pronotum cylindrical, about 1.3 – 1.8 times as long as wide, punctate and pubescent, punctures larger than those on head and elytra. Elytra truncate at apex, outer angles more or less round (Figs. 35, 36), obtuse (Figs. 37, 38), slightly dentate (Figs. 39, 40), or strongly dentate and sharply pointed at apex (Figs. 41 – 43); striae shallow, punctate; intervals flat or slightly convex, punctate and pubescent; in some species, punctures nearly equal in size, quite dense, interspace between two adjacent punctures narrower than diameter of punctures (Fig. 44), in other species, punctures unequal in size, sparse, interspace between two adjacent punctures often wider than diameter of punctures (Figs. 45, 46). Hindwings fullsized, functional. Sterna densely punctate and pubescent; apical margin of terminal sternum slightly protruded posteriad medially in male (Figs. 47, 48), almost straight or widely rounded in female (Figs. 49, 50). Legs slender, protarsomere 3 strongly (Fig. 51) or slightly (Figs. 52) asymmetrical or symmetrical (Figs. 53) in male, symmetrical in female (Figs. 54 – 56), tarsomere 4 bilobed (Fig. 51 – 56); tarsal claws slender, more or less pectinate on inner side (Figs. 57 – 70), very different from those of Drypta (claws thick, strongly curved, and smooth on inner side; e. g., Figs. 71, 72). In Asian species, aedeagus generally thin and long (Fig. 73, 74, 77 – 102), with apical lamella wide and long, round or truncate at apex (Figs. 73 – 86), narrow, digitiform (Figs. 87 – 96), or short and round at apex (Figs. 97 – 102); in African species, aedeagus stout, with lamella long or short (Figs. 103 – 108), apex round, thickened (Figs. 104, 108) or thin (Fig. 106).	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFD0480728A7BCCF256BFB5F.taxon	biology_ecology	Biology. These beetles live on vegetation, but adults of some species (e. g., D. nepalensis (Hope )) also hide under shrub debris or inside decayed tree trunks. Adults are attracted to light traps.	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFD0480728A7BCCF256BFB5F.taxon	distribution	Distribution. Asia, Australia and Africa.	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFCC482428A7BA0024B7FB22.taxon	materials_examined	Other specimens examined. Total 1 specimen: 1 female (MBC), ‘‘ Bali, 8 km e. Cekik, W. Bali NP, 8 ° 11 9 S, 114 ° 30 9 E, 13 - 16.12.2000, leg. Gerstmeier’ ’, ‘‘ Desera schultzei Heller, det. M. Baehr ‘ 02 ’’.	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFCC482428A7BA0024B7FB22.taxon	description	Description. Length 10.8 – 11.4 mm, width 3.4 – 3.6 mm. Black, head and pronotum with slight blue luster; elytra with very slight green luster; apical two-thirds of Ant 1 black, basal third brown, Ant 2 and Ant 3 dark brown, Ant 4 – 11 brown; femora and tibiae black, tarsi dark brown. Head slightly wider than pronotum, HW / PW 5 1.11 (1.08 – 1.13); vertex convex, coarsely punctate; Ant 1 / Ant 3 5 3.63 (3.50 – 3.72); labrum with medial lobe slightly protruded anteriorly; genae long, distinctly tumid; eyes large, 2.5 times as long as genae. Pronotum densely punctate, wide, PL / PW 5 1.47 (1.45 – 1.49); lateral margins gradually widened from apical angles to middle, sinuate before basal angles; basal angles protruded laterally, nearly rectangular; basal foveae shallow; middle line fine. Elytra short, EL / EW 5 1.69 (1.67 – 1.72), EW / PW 5 2.35 (2.30 – 2.38); lateral margins widened posteriad to apical third, then narrowed to apex; outer angles sharp, with a short tooth; intervals flat, densely punctate, interspace between two adjacent punctures narrower than diameter of punctures. Tarsal claws with 6 long pectinations, the longest pectination subequal to the width of claw base (Fig. 59).	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFCC482428A7BA0024B7FB22.taxon	distribution	Distribution. Philippines, Indonesia.	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFCC482428A7BA0024B7FB22.taxon	discussion	Remarks. Members of this species are similar to those of D. unidentatus (Macleay) in having long claw pectinations and a large ratio of Ant 1 / Ant 3, but differ in having a shorter pronotum, nearly black elytra, and black femora. They are also similar to members of D. gestroi Bates, as stated by Heller (1921: 303), but differ in having a longer pronota than D. gestroi members. The femora and tibiae of adults of D. schultzei are all black, which is unique within the genus.	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFF6482129C4B8C424E1FA0E.taxon	distribution	Distribution. Sumatra.	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
03D187BCFFF6482129C4B8C424E1FA0E.taxon	discussion	Remark. Lorenz (1998: 477; 2005: 503) moved Drypta dimidiata Putzeys into genus Desera. Dr. Rienk de Jong in Leiden Museum (Netherlands) kindly helped us examine the type specimen. It does not possess any trace of pectination in the inner side of tarsal claw, and therefore is a true member of Drypta as originally described.	en	Liang, Hongbin, Kavanaugh, David H. (2007): Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species. The Coleopterists Bulletin (Oxford, England) 61 (1): 1-39, DOI: 10.1649/934.1, URL: http://dx.doi.org/10.1649/934.1
