taxonID	type	description	language	source
03D541284E087F74FF762EB42216AAE6.taxon	type_taxon	Type-species: Testudobracon niger Quicke, 1986 (original description).	en	Mai, Pham Quynh, Long, Khuat Dang, Hiep, Nguyen Duc, Hoa, Dang Thi, Duong, Tran Dinh (2023): First record of the genus Testudobracon Quicke, 1986 (Hymenoptera: Braconidae: Braconinae) in Vietnam, with description of four new species. Zootaxa 5244 (5): 485-500, DOI: 10.11646/zootaxa.5244.5.5, URL: http://zoobank.org/46962264-5ded-44b6-a42a-66fa97eb5f36
03D541284E087F74FF762EB42216AAE6.taxon	synonymic_list	Testudobracon alius Long, sp. nov.. Oriental: Vietnam Testudobracon asphondyliae Haider & Shujauddin, 2004. Oriental: India Testudobracon athashi Ranjith, 2017. Oriental: India Testudobracon australicolorus Quicke & Ingram, 1993. Australasian: Australia-Queensland Testudobracon flavus Wang, Chen & He, 2003. Eastern Palaearctic: China; Oriental: China, Vietnam Testudobracon descensus Long, sp. nov.. Oriental: Vietnam Testudobracon gibbosa Yang & Chen, 2006. Oriental: China Testudobracon grandiventris Wang, Chen & He, 2003. Oriental: China Testudobracon guangxinensis Wang, Chen & He, 2003. Oriental: China Testudobracon imitator Long, sp. nov.. Oriental: Vietnam Testudobracon longicaudis Maeto, 1991. Eastern Palaearctic: Japan Testudobracon malabaricus Sheeba, 2017. Oriental: India Testudobracon niger Quicke, 1986. Oriental: Indonesia (Java) Testudobracon phiaoacus Long & PQMai, sp. nov.. Oriental: Vietnam Testudobracon pleuralis (Ashmead, 1906). Eastern Palaearctic and Oriental: China, Japan Testudobracon shameeri Ranjith, 2017. Oriental: India Testudobracon tatyanae Quicke & Ingram, 1993. Australasian: Australia-Queensland Testudobracon travencorensis Sheeba, 2017. Oriental: India Testudobracon unicolorus Quicke & Ingram, 1993. Australasian: Australia; Oriental: China Testudobracon watanabei Yang & Chen, 2006. Oriental: China	en	Mai, Pham Quynh, Long, Khuat Dang, Hiep, Nguyen Duc, Hoa, Dang Thi, Duong, Tran Dinh (2023): First record of the genus Testudobracon Quicke, 1986 (Hymenoptera: Braconidae: Braconinae) in Vietnam, with description of four new species. Zootaxa 5244 (5): 485-500, DOI: 10.11646/zootaxa.5244.5.5, URL: http://zoobank.org/46962264-5ded-44b6-a42a-66fa97eb5f36
03D541284E0B7F73FF762D7C2544AD9A.taxon	description	Figures 1 – 2	en	Mai, Pham Quynh, Long, Khuat Dang, Hiep, Nguyen Duc, Hoa, Dang Thi, Duong, Tran Dinh (2023): First record of the genus Testudobracon Quicke, 1986 (Hymenoptera: Braconidae: Braconinae) in Vietnam, with description of four new species. Zootaxa 5244 (5): 485-500, DOI: 10.11646/zootaxa.5244.5.5, URL: http://zoobank.org/46962264-5ded-44b6-a42a-66fa97eb5f36
03D541284E0B7F73FF762D7C2544AD9A.taxon	materials_examined	Material examined. Holotype, ♀, “ Bracn. 1103 ” (IEBR), NC Vietnam: Ha Tinh, Huong Son, Rao An, forest, MT, 18 ° 22 ’ N 106 ° 13 ’ E, 600 m, April 15 – 21.1998, AMNH, K. Long. Paratype, 1 ♀, “ Bracn. 1532 ” (IEBR), NE Vietnam: Tuyen Quang, Na Hang NP, Son Phu, forest, MT, 21 ° 17 ’ 32 ’’ N 105 ° 28 ’ 19 ’’ E, 573 m, 15. vi. 2018, KD Long.	en	Mai, Pham Quynh, Long, Khuat Dang, Hiep, Nguyen Duc, Hoa, Dang Thi, Duong, Tran Dinh (2023): First record of the genus Testudobracon Quicke, 1986 (Hymenoptera: Braconidae: Braconinae) in Vietnam, with description of four new species. Zootaxa 5244 (5): 485-500, DOI: 10.11646/zootaxa.5244.5.5, URL: http://zoobank.org/46962264-5ded-44b6-a42a-66fa97eb5f36
03D541284E0B7F73FF762D7C2544AD9A.taxon	description	Description. Holotype, female, length 2.8 mm; antenna 2.3 mm; fore wing 2.9 mm; ovipositor 1.2 mm (Fig. 1). Head. Antenna with 22 flagellomeres. Length of first flagellomere 1.2 × as long as second flagellomere; first, second and penultimate flagellomeres 4.0, 2.5 and 2.0 × as long as wide; head 1.45 and 1.7 × as wide as long medially in frontal and dorsal view, respectively; frons and vertex granulate; face with mid-longitudinal ridge reaching middle of face; face granulo-rugulose (Fig. 2 b); OOL: OD: POL = 6.0: 2.5: 2.5; length of eye 2.25 × as long as temple in dorsal view; occiput rugose; lateral temples smooth and shiny; height of eye: width of face: width of head = 12: 19: 29; height of clypeus: inter-tentorial distance: tentorio-ocular distance = 2.5: 6.0: 4.0; clypeus with distinct postero-dorsal carina (Fig. 2 b); length of malar space 1.25 × basal width of mandible. Mesosoma. Length of mesosoma 1.2 × its height (Fig. 2 f); mesoscutum shiny, sparsely setose and punctate; middle lobe without medial longitudinal ridge (Fig. 2 a); notauli complete, meeting scutellar sulcus; anteriorly with depressed area, almost smooth; scutellar sulcus deep with 8 carinae (Fig. 2 a); scutellum shiny, smooth; mesopleuron with a deep oval pit medially (Fig. 2 f), smooth ventrally, finely rugo-punctate medially and dorsally; metapleuron shiny, finely punctate; propodeum with a complete midlongitudinal carina, shiny, smooth; in lateral view, metapleuron and propodeum posteriorly densely setose (Fig. 2 d). Wings. Length of pterostigma 2.2 × its maximum width; fore wing vein 1 - SR 0.4 × as long as 1 - M; ration of length of fore wing veins r: 3 - SR: SR 1 = 5: 11: 32. and 2 - SR: 3 SR: r-m = 9: 11: 6; second submarginal cell hardly narrowed distally (Fig. 2 h). Hind wing vein C + SC + R with single straight hamulus and vein R 1 with 3 curved hamuli apically; ratio of length of hind wing veins 1 r-m: SC + R 1 = 6: 8. Legs. Length of fore femur: tibia: tarsus = 16: 18: 13; hind coxa smooth, setose; length of hind femur 4.0 × its maximum width; length of hind femur: tibia: basitarsus = 24: 31: 11. Metasoma. First metasomal tergite rugose, 0 / 7 × as long as wide apically, deeply impressed medio-basally and carinate medially; second metasomal tergite basally 2.3 × wider than long, with sublateral grooves (Fig. 2 g); second tergite 0.8 × as long as third tergite medially; second-third metasomal tergites, rugulose medially (Fig. 2 g); secondfifth tergites rugo-striate laterally; fourth-sixth metasomal tergites faintly rugose medially; sixth metasomal tergite with rounded postero-lateral lobes; medial emargination (incision) deep (Fig. 2 g), 0.6 × as long as tergite medially, and 1.5 × as long as wide; ovipositor 1.0 × metasoma in lateral view, and 0.5 × fore wing. Colour. Body largely yellow (Fig. 1); antennae brown; stemmaticum brown; fore and middle legs yellow; hind leg yellow, except coxa, femur and tibia apically pale brown; propodeum brownish yellow; wing veins yellowish brown; metasomal tergites 1 – 3 yellow, except 2 nd tergite medially and 3 rd tergite basally yellowish brown; 4 th – 6 th tergites pale yellow (Fig. 2 g); postero-lateral lobes of 3 rd- 5 th tergites brown; ovipositor sheath brown; ovipositor yellow, brwonish yellow subapically. Variation. Female (paratype): length of body 2.9 mm; antenna 2.6 mm; fore wing 2.8 mm; ovipositor sheath 1.3 mm; antenna with 23 flagellomeres. Male. Unknown	en	Mai, Pham Quynh, Long, Khuat Dang, Hiep, Nguyen Duc, Hoa, Dang Thi, Duong, Tran Dinh (2023): First record of the genus Testudobracon Quicke, 1986 (Hymenoptera: Braconidae: Braconinae) in Vietnam, with description of four new species. Zootaxa 5244 (5): 485-500, DOI: 10.11646/zootaxa.5244.5.5, URL: http://zoobank.org/46962264-5ded-44b6-a42a-66fa97eb5f36
03D541284E0B7F73FF762D7C2544AD9A.taxon	distribution	Distribution. NE Vietnam (Tuyen Quang); NC Vietnam (Ha Tinh).	en	Mai, Pham Quynh, Long, Khuat Dang, Hiep, Nguyen Duc, Hoa, Dang Thi, Duong, Tran Dinh (2023): First record of the genus Testudobracon Quicke, 1986 (Hymenoptera: Braconidae: Braconinae) in Vietnam, with description of four new species. Zootaxa 5244 (5): 485-500, DOI: 10.11646/zootaxa.5244.5.5, URL: http://zoobank.org/46962264-5ded-44b6-a42a-66fa97eb5f36
03D541284E0B7F73FF762D7C2544AD9A.taxon	biology_ecology	Biology. Unknown.	en	Mai, Pham Quynh, Long, Khuat Dang, Hiep, Nguyen Duc, Hoa, Dang Thi, Duong, Tran Dinh (2023): First record of the genus Testudobracon Quicke, 1986 (Hymenoptera: Braconidae: Braconinae) in Vietnam, with description of four new species. Zootaxa 5244 (5): 485-500, DOI: 10.11646/zootaxa.5244.5.5, URL: http://zoobank.org/46962264-5ded-44b6-a42a-66fa97eb5f36
03D541284E0B7F73FF762D7C2544AD9A.taxon	etymology	Etymology. Named from “ alius ” (Latin for “ another, other ”). Notes. Testudobracon alius sp. nov. is close to T. niger Quicke, from Indonesia, but differs from the latter by the following characters: 1) Second sub-marginal cell of fore wing longer and not narrowed apically; 2) Vein 3 - SR of fore wing 2.1 × vein r (1.8 × in T. niger); and 3) Antenna with 22 flagellomeres and first flagellomere 1.2 × second flagellomere (equal each other in T. niger).	en	Mai, Pham Quynh, Long, Khuat Dang, Hiep, Nguyen Duc, Hoa, Dang Thi, Duong, Tran Dinh (2023): First record of the genus Testudobracon Quicke, 1986 (Hymenoptera: Braconidae: Braconinae) in Vietnam, with description of four new species. Zootaxa 5244 (5): 485-500, DOI: 10.11646/zootaxa.5244.5.5, URL: http://zoobank.org/46962264-5ded-44b6-a42a-66fa97eb5f36
03D541284E0C7F71FF7629E82599A946.taxon	description	Figures 3 – 4	en	Mai, Pham Quynh, Long, Khuat Dang, Hiep, Nguyen Duc, Hoa, Dang Thi, Duong, Tran Dinh (2023): First record of the genus Testudobracon Quicke, 1986 (Hymenoptera: Braconidae: Braconinae) in Vietnam, with description of four new species. Zootaxa 5244 (5): 485-500, DOI: 10.11646/zootaxa.5244.5.5, URL: http://zoobank.org/46962264-5ded-44b6-a42a-66fa97eb5f36
03D541284E0C7F71FF7629E82599A946.taxon	materials_examined	Material examined. Holotype, ♀, “ Bracn. 1561 ” (IEBR), NW Vietnam: Son La, Chieng Sinh, Quyet Tam, orchard, MT, 21 ° 18 ’ 06 ’’ N 103 ° 55 ’ 36 ’’ E, 663 m, 10. v. 2017, KD Long.	en	Mai, Pham Quynh, Long, Khuat Dang, Hiep, Nguyen Duc, Hoa, Dang Thi, Duong, Tran Dinh (2023): First record of the genus Testudobracon Quicke, 1986 (Hymenoptera: Braconidae: Braconinae) in Vietnam, with description of four new species. Zootaxa 5244 (5): 485-500, DOI: 10.11646/zootaxa.5244.5.5, URL: http://zoobank.org/46962264-5ded-44b6-a42a-66fa97eb5f36
03D541284E0C7F71FF7629E82599A946.taxon	description	Description. Holotype, female, length 3.2 mm; antenna 2.6 mm; fore wing 2.9 mm; ovipositor sheath 1.4 mm (Fig. 3). Head. Antenna with 23 flagellomeres; length of first flagellomere subequal to second flagellomere; first, second and penultimate flagellomeres 2.0, 2.0 and 1.2 × as long as wide; head 1.3 and 1.8 × as wide as long medially in frontal and dorsal view, respectively; frons and vertex anteriorly rugo-granulate (Figs 4 a, 4 b); occiput largely rogose; temple laterally coriaceous; malar space rugose; OOL: OD: POL = 7: 3: 3 (Fig. 4 a) ;; oculo-antennal groove indistinct; length of eye 3.25 × as long as temple in dorsal view; lateral temples smooth and shiny; height of eye: width of face: width of head = 16: 16: 33; face rugose (Fig. 4 b); height of clypeus: inter-tentorial distance: tentorio-ocular distance = 2.5: 7.0: 5.0; clypeus with distinct postero-dorsal carina (Fig. 4 b); length of malar space 1.75 × basal width of mandible. Mesosoma. Length of mesosoma 1.2 × its height; mesoscutum with sparse and short setosetae; middle lobe more or less with medial longitudinal ridge (Fig. 4 d); notauli complete, punctate, anteriorly with depressed area, narrowed medially and widened posteriorly, meeting scutelar sulcus; scutellar sulcus deep, with 7 carinae (Fig. 4 d); scutellum convex, smooth; mesopleuron with a short and deep narrow pit medially (Fig. 4 f); precoxal sulcus indistinct; mesopleuron coriaceous ventally and medially, transversely rugose dorsally; metapleuron rugo-punctate; propodeum with a complete mid longitudinal carina, smooth, setose latero-posteriorly. Wings. Length of pterostigma 2.7 × its maximum width; fore wing vein 1 - SR 0.4 × as long as 1 - M; ratio of length of fore wing veins r: 3 - SR: SR 1 = 5: 11: 36, and 2 - SR: 3 - SR: r-m = 10: 11: 6; second submarginal cell slightly narrowed distally (Fig. 4 k). Hind wing vein C + SC + R with single straight hamulus and vein R 1 with 3 curved hamuli apically; ratio of length of hind wing veins 1 r-m: SC + R 1 = 6: 10. Legs. Length of fore femur: tibia: tarsus = 19: 21: 24; hind coxa smooth, setose; length of hind femur: tibia: basitarsus = 28: 33: 11. Metasoma. First metasomal tergite rugose, 0.7 × as long as wide apically apically, deeply impressed mediobasally and carinate medially; second metasomal tergite basally 1.8 × wider than long, with sublateral grooves, rugose, setose (Fig. 4 d); second metasomal tergite 1.1 × as long as third tergite medially; second metasomal tergite areolate-rugulose medially (Fig. 4 c); third tergite confused rugulose; fourth-sixth metasomal tergites finely irregularly rugose medially, rugose laterally (Fig. 4 g); sixth metasomal tergite with rounded postero-lateral lobes; medial emargination (incision) deep (Fig. 4 h), 0.6 × as long as tergite medially, and 0.7 × as long as wide; ovipositor abruptly bent downward (Fig. 4 i), 1.0 × metasoma in lateral view, and 0.6 × as long as fore wing (Fig. 3). Colour. Antenna brown; stemmaticum dark brown; head, pronotum yellow; mesoscutum and mesopleuron ventrally reddish yellow; scutellum, mesopleuron dorsally, mesosternum and propodeum brown to dark brown (Fig. 4 f); fore and middle legs yellow; hind coxa, femur and tibia dark brown, except hind tibia basally, hind trochanters + trochantellus and hind tarsus pale yellow; wing veins yellowish brown; metasomal tergites 1 – 3 yellow, except 1 st tergite apically, 2 nd – 3 rd tergites medially dark brown; 4 th – 6 th tergites pale yellow; postero-lateral lobes of 3 rd – 5 th tergites and 3 rd- 4 th metasomal sutures yellowish brown (Fig. 4 h); ovipositor sheath brown; ovipositor yellow. Male. Unknown.	en	Mai, Pham Quynh, Long, Khuat Dang, Hiep, Nguyen Duc, Hoa, Dang Thi, Duong, Tran Dinh (2023): First record of the genus Testudobracon Quicke, 1986 (Hymenoptera: Braconidae: Braconinae) in Vietnam, with description of four new species. Zootaxa 5244 (5): 485-500, DOI: 10.11646/zootaxa.5244.5.5, URL: http://zoobank.org/46962264-5ded-44b6-a42a-66fa97eb5f36
03D541284E0C7F71FF7629E82599A946.taxon	distribution	Distribution. NW Vietnam (Hoa Binh, Son La).	en	Mai, Pham Quynh, Long, Khuat Dang, Hiep, Nguyen Duc, Hoa, Dang Thi, Duong, Tran Dinh (2023): First record of the genus Testudobracon Quicke, 1986 (Hymenoptera: Braconidae: Braconinae) in Vietnam, with description of four new species. Zootaxa 5244 (5): 485-500, DOI: 10.11646/zootaxa.5244.5.5, URL: http://zoobank.org/46962264-5ded-44b6-a42a-66fa97eb5f36
03D541284E0C7F71FF7629E82599A946.taxon	biology_ecology	Biology. Unknown.	en	Mai, Pham Quynh, Long, Khuat Dang, Hiep, Nguyen Duc, Hoa, Dang Thi, Duong, Tran Dinh (2023): First record of the genus Testudobracon Quicke, 1986 (Hymenoptera: Braconidae: Braconinae) in Vietnam, with description of four new species. Zootaxa 5244 (5): 485-500, DOI: 10.11646/zootaxa.5244.5.5, URL: http://zoobank.org/46962264-5ded-44b6-a42a-66fa97eb5f36
03D541284E0C7F71FF7629E82599A946.taxon	etymology	Etymology. From Latin “ descensus ” meaning “ downward course ”, because ovipositor abruptly bent downward subapically. Notes. Testudobracon descensus sp. nov. is close to T. niger Quicke, from Indonesia, but differs from the latter by the following characters: 1) Vein 3 - SR of fore wing 1.65 × vein r (1.8 × in T. niger); 2) Second sub-marginal cell of fore wing almost not narrowed apically; and 3) Propodeum and medio-dorsal area of 3 rd tergite blackish brown to black; third-fourth metasomal suture brown.	en	Mai, Pham Quynh, Long, Khuat Dang, Hiep, Nguyen Duc, Hoa, Dang Thi, Duong, Tran Dinh (2023): First record of the genus Testudobracon Quicke, 1986 (Hymenoptera: Braconidae: Braconinae) in Vietnam, with description of four new species. Zootaxa 5244 (5): 485-500, DOI: 10.11646/zootaxa.5244.5.5, URL: http://zoobank.org/46962264-5ded-44b6-a42a-66fa97eb5f36
03D541284E0E7F7CFF762DD4223FACDE.taxon	description	Figures 5 – 6	en	Mai, Pham Quynh, Long, Khuat Dang, Hiep, Nguyen Duc, Hoa, Dang Thi, Duong, Tran Dinh (2023): First record of the genus Testudobracon Quicke, 1986 (Hymenoptera: Braconidae: Braconinae) in Vietnam, with description of four new species. Zootaxa 5244 (5): 485-500, DOI: 10.11646/zootaxa.5244.5.5, URL: http://zoobank.org/46962264-5ded-44b6-a42a-66fa97eb5f36
03D541284E0E7F7CFF762DD4223FACDE.taxon	materials_examined	Material examined. “ Bracn. 1034 ” (IEBR), NW Vietnam: Hoa Binh, Mai Chau, Tan Son, orchard, MT, 20 ° 43 ’ 10.3 ’’ N 104 ° 59 ’ 47 ’’ E, 950 m, 10 – 15. vi. 2010, KD Long.	en	Mai, Pham Quynh, Long, Khuat Dang, Hiep, Nguyen Duc, Hoa, Dang Thi, Duong, Tran Dinh (2023): First record of the genus Testudobracon Quicke, 1986 (Hymenoptera: Braconidae: Braconinae) in Vietnam, with description of four new species. Zootaxa 5244 (5): 485-500, DOI: 10.11646/zootaxa.5244.5.5, URL: http://zoobank.org/46962264-5ded-44b6-a42a-66fa97eb5f36
03D541284E0E7F7CFF762DD4223FACDE.taxon	description	Description. Holotype, female, length 3.0 mm; antennae 2.9 mm; fore wing 2.6 mm; ovipositor sheath 1.4 mm (Fig. 5). Head. Antenna with 23 flagellomeres; length of first flagellomere 1.2 × as long as second flagellomere; first, second and penultimate flagellomeres 2.0, 2.0 and 2.0 × as long as wide; head 1.5 and 1.9 × as wide as long medially in anterior and dorsal view, respectively; OOL: OD: POL = 6: 3: 3 (Fig. 6 a); oculo-antennal groove indistinct; length of eye 4.0 × as long as temple in dorsal view; height of eye: width of face: width of head = 15: 15: 34; height of clypeus: inter-tentorial distance: tentorio-ocular distance = 2: 7: 4; clypeus with distinct postero-dorsal carina (Fig. 6 b); length of malar space 1.5 × basal width of mandible. Face, frons and vertex anteriorly finely rugose; occiput irregularly rugose; temple almost smooth. Mesosoma. Length of mesosoma 1.2 × its height (Fig. 6 g); mesoscutum rugo-punctate, middle lobe with indistinct medial longitudinal ridge (Fig. 6 d); notauli complete, anteriorly with depressed area, wide posteriorly, meeting scutellar sulcus scutellar sulcus rather wide, deep, with 6 carinae (Fig. 6 d); scutellum convex, smooth; mesopleuron with a deep oval pit medially (Fig. 6 g); mesopleuron rugo-punctate, but smooth posterior-ventrally; metapleuron rugo-punctate; groove between metapleuron and propodeum wide and deep; propodeum with a complete mid- longitudinal carina, largely smooth medio-posteriorly, rugo-striate antero-laterally. Wings. Length of pterostigma 2.4 × its maximum width; fore wing vein 1 - SR 0.3 × as long as 1 - M; ration of length of fore wing veins r: 3 - SR: SR 1 = 6: 12: 32 and 2 - SR: 3 - SR: r-m = 10: 12: 5; second submarginal cell rather long, slightly narrowed distally (Fig. 6 h). Hind wing vein C + SC + R with straight hamulus and vein R 1 with 3 curved hamuli apically; ratio of length of hind wing veins 1 r-m: SC + R 1 = 6: 9. Legs. Length of fore femur: tibia: tarsus = 19: 18: 21; hind coxa smooth, setose; length of hind femur: tibia: basitarsus = 25: 30: 11. Metasoma. First metasomal tergite rugose, 0.8 × as long as wide apically with a longitudinally impressed area medially; second metasomal tergite basally 2.1 × wider than long with sublateral crenulate grooves, rugose, setose (Fig. 6 f); second metasomal tergite 0.8 × as long as third tergite medially; second metasomal tergite rugulose medially, irregularly rugose laterally; third tergite confused rugulose (Fig. 6 f); fourth – sixth tergites finely rugose medially, irregularly rugose laterally; sixth metasomal tergite with rounded postero-lateral lobes (Fig. 6 e); medial emargination deep, touching suture between T 5 and T 6 (Fig. 6 c) 1.6 × as wide as height medially; ovipositor 0.9 × metasoma in lateral view, and 0.5 × as long as fore wing. Colour. Antenna brown; head yellow, except stemmaticum and occiput dark brown; pronotum mesonotum yellow; mesopleuron ventrally and propodeum brown; fore and middle legs yellow, except telotarsus yellowish brown; hind coxa brown; hind femur and hind tibia apically brown, except hind femur apically and basally, hind tibia basally yellow; hind trochanters and trochantellus, hind tarsus yellow, except telotarsus brown; wing veins yellowish brown; metasomal tergites 1 – 3 yellow, except 2 nd – 3 rd tergites medially dark brown; 4 th – 6 th tergites pale yellow dorsally; postero-lateral lobes of 3 rd – 5 th tergites, 4 th metasomal sutures yellowish brown; ovipositor sheath brown; ovipositor yellow. Male. Unknown.	en	Mai, Pham Quynh, Long, Khuat Dang, Hiep, Nguyen Duc, Hoa, Dang Thi, Duong, Tran Dinh (2023): First record of the genus Testudobracon Quicke, 1986 (Hymenoptera: Braconidae: Braconinae) in Vietnam, with description of four new species. Zootaxa 5244 (5): 485-500, DOI: 10.11646/zootaxa.5244.5.5, URL: http://zoobank.org/46962264-5ded-44b6-a42a-66fa97eb5f36
03D541284E0E7F7CFF762DD4223FACDE.taxon	distribution	Distribution. NW Vietnam (Hoa Binh).	en	Mai, Pham Quynh, Long, Khuat Dang, Hiep, Nguyen Duc, Hoa, Dang Thi, Duong, Tran Dinh (2023): First record of the genus Testudobracon Quicke, 1986 (Hymenoptera: Braconidae: Braconinae) in Vietnam, with description of four new species. Zootaxa 5244 (5): 485-500, DOI: 10.11646/zootaxa.5244.5.5, URL: http://zoobank.org/46962264-5ded-44b6-a42a-66fa97eb5f36
03D541284E0E7F7CFF762DD4223FACDE.taxon	biology_ecology	Biology. Unknown.	en	Mai, Pham Quynh, Long, Khuat Dang, Hiep, Nguyen Duc, Hoa, Dang Thi, Duong, Tran Dinh (2023): First record of the genus Testudobracon Quicke, 1986 (Hymenoptera: Braconidae: Braconinae) in Vietnam, with description of four new species. Zootaxa 5244 (5): 485-500, DOI: 10.11646/zootaxa.5244.5.5, URL: http://zoobank.org/46962264-5ded-44b6-a42a-66fa97eb5f36
03D541284E0E7F7CFF762DD4223FACDE.taxon	etymology	Etymology. From “ imitor ” Latin meaning “ copy, mimic ”, because the new species is similar to Testudobracon descensus, sp. nov. by having ovipositor abruptly bent downward subapically. Notes. Testudobracon imitator sp. nov. is close to T. athashi Ranjith, from India but differs from the latter by following characters: 1) Ovipositor 0.9 × metasoma (0.6 × in T. athashi); 2) Vein 3 - SR of fore wing distinctly longer vein (1.4 ×) vein 2 - SR (in T. athashi vein 3 - SR subequal to vein 2 - SR; and 3) Propodeum, hind coxa and lateral lobes of third – fifth metasomal tergites blackish brown (propodeum, hind coxa yellow in T. athashi); T. imitator, sp. nov. is also close to T. tatyanae Quicke & Ingram, from Australia, but differs from the latter by the following characters: 1) Second-third metasomal tergites with acute postero-lateral lobes (rounded postero-lateral lobes in T. tatyanae); 2) Notauli deep and wide posteriorly, sparsely cranulate; and 3) Third metasomal tergite 2.8 × wider than long (2.25 × in T. tatyanae).	en	Mai, Pham Quynh, Long, Khuat Dang, Hiep, Nguyen Duc, Hoa, Dang Thi, Duong, Tran Dinh (2023): First record of the genus Testudobracon Quicke, 1986 (Hymenoptera: Braconidae: Braconinae) in Vietnam, with description of four new species. Zootaxa 5244 (5): 485-500, DOI: 10.11646/zootaxa.5244.5.5, URL: http://zoobank.org/46962264-5ded-44b6-a42a-66fa97eb5f36
03D541284E037F7DFF762F2C22A7A8A6.taxon	description	Figures 7 – 8	en	Mai, Pham Quynh, Long, Khuat Dang, Hiep, Nguyen Duc, Hoa, Dang Thi, Duong, Tran Dinh (2023): First record of the genus Testudobracon Quicke, 1986 (Hymenoptera: Braconidae: Braconinae) in Vietnam, with description of four new species. Zootaxa 5244 (5): 485-500, DOI: 10.11646/zootaxa.5244.5.5, URL: http://zoobank.org/46962264-5ded-44b6-a42a-66fa97eb5f36
03D541284E037F7DFF762F2C22A7A8A6.taxon	materials_examined	Material examined. Holotype, ♀, “ Bracn. 1570 ” (IEBR), NE Vietnam: Cao Bang, Nguyen Binh, Quang Thanh, soybeans, 22 ° 36 ’ 45.5 N 105 ° 56 ’ 09.4 E, 570 m, sweep, 02. vii. 2022, KDLong.	en	Mai, Pham Quynh, Long, Khuat Dang, Hiep, Nguyen Duc, Hoa, Dang Thi, Duong, Tran Dinh (2023): First record of the genus Testudobracon Quicke, 1986 (Hymenoptera: Braconidae: Braconinae) in Vietnam, with description of four new species. Zootaxa 5244 (5): 485-500, DOI: 10.11646/zootaxa.5244.5.5, URL: http://zoobank.org/46962264-5ded-44b6-a42a-66fa97eb5f36
03D541284E037F7DFF762F2C22A7A8A6.taxon	description	Description. Holotype, female, length 2.4 mm; antenna 2.2 mm; fore wing 2.4 mm; ovipositor sheath 1.3 mm (Fig. 7). Head. Antenna with 22 flagellomeres; first flagellomere as long as second flagellomere; first, second and penultimate flagellomeres 2.0, 2.0 and 1.5 × as long as wide; head 1.35 and 1.6 × as wide as long medially in frontal and dorsal view, respectively; face rugose; frons frons granulo-rugose; vertex granulo-rugose; OOL: OD: POL = 6: 2: 3 (Fig. 8 a); oculo-antennal groove indistinct; length of eye 2.75 × as long as temple in dorsal view; occiput rugose; lateral temple smooth and shiny; eight of eye: width of face: width of head = 12: 14: 27; face granulate with posteriorly widened, transversely striated medial triangular area (Fig. 8 b); height of clypeus: inter-tentorial distance: tentorio-ocular distance = 2: 5: 4; clypeus with distinct postero-dorsal carina; length of malar space 1.25 × basal width of mandible. Mesosoma. Length of mesosoma 1.25 × its height (Fig. 8 f); mesoscutum sparsely punctate; median lobe of mesoscutum with indistinct mid-longitudinal ridge (Fig. 8 d); notauli complete, anteriorly with depressed area, converging posteriorly, hardly fused with scutellar sulcus; scutellar sulcus deep, with 7 carinae (Fig. 8 d); scutellum almost smooth medially, sparsely punctate laterally, with long setae clustered apically; mesopleuron with a deep short oval pit medially, smooth ventrally, rugo-punctate dorsally (Fig. 8 f); metapleuron punctate like anterio-lateral area of propodeum; groove between metapleuron and propodeum deep; propodeum with a complete mid longitudinal carina, smooth medially and posteriorly. Wings. Length of pterostigma 2.7 × its maximum width; fore wing vein 1 - SR 0.4 × as long as 1 - M; ration of length of fore wing veins r: 3 - SR: SR 1 = 5: 10: 29. and 2 - SR: 3 - SR: r-m = 9: 10: 5; second submarginal cell slightly narrowed distally (Fig. 8 k). Hind wing vein C + SC + R with single straight hamulus and vein R 1 with 3 curved hamuli apically; ratio of length of hind wing veins 1 r-m: SC + R 1 = 5: 7. Legs. Length of fore femur: tibia: tarsus = 14: 17: 19; hind coxa smooth, setose; length of hind femur: tibia: basitarsus = 23: 28: 10. Metasoma. First metasomal tergite rugose, 0.55 × as long as wide apically with a longitudinally impressed area medially and posterior carinae; second metasomal tergite basally 1.3 × wider than long, with sublateral grooves (Fig. 8 c); second metasomal tergite as long as third tergite medially; second-third metasomal tergites confused-rugose medially, rugose laterally (Fig. 8 h); fourth-sixth metasomal tergites finely rugose medially, largely rugose laterally; medial emargination deep (Fig. 8 g), 0.7 × as long as tergite medially, and 1.4 × as wide as height; ovipositor 1.0 × metasoma in lateral view, and 0.5 × as long as metasoma and fore wing Colour. Body yellow to reddish yellow (Fig. 7); antenna brown; head and mesonotum reddish yellow, except malar space, mandible, labrum, palpi and lateral side of pronotum yellow; propodeum and metapleuron brown; fore and middle legs yellow, except telotarsi brown; hind leg yellow, except hind coxa and femur medially, tibia apically and hind tarsus brown; wing veins brown but paler basally; first-third metasomal tergites yellow, except third tergite medially and laterally dark brown; fourth – sixth metasomal tergites pale yellow, posterio-lateral protuberances dark brown (Fig. 8 h); ovipositor sheath brown; ovipositor yellow. Male. Unknown.	en	Mai, Pham Quynh, Long, Khuat Dang, Hiep, Nguyen Duc, Hoa, Dang Thi, Duong, Tran Dinh (2023): First record of the genus Testudobracon Quicke, 1986 (Hymenoptera: Braconidae: Braconinae) in Vietnam, with description of four new species. Zootaxa 5244 (5): 485-500, DOI: 10.11646/zootaxa.5244.5.5, URL: http://zoobank.org/46962264-5ded-44b6-a42a-66fa97eb5f36
03D541284E037F7DFF762F2C22A7A8A6.taxon	distribution	Distribution. NE Vietnam (Cao Bang: Phia Oac NP).	en	Mai, Pham Quynh, Long, Khuat Dang, Hiep, Nguyen Duc, Hoa, Dang Thi, Duong, Tran Dinh (2023): First record of the genus Testudobracon Quicke, 1986 (Hymenoptera: Braconidae: Braconinae) in Vietnam, with description of four new species. Zootaxa 5244 (5): 485-500, DOI: 10.11646/zootaxa.5244.5.5, URL: http://zoobank.org/46962264-5ded-44b6-a42a-66fa97eb5f36
03D541284E037F7DFF762F2C22A7A8A6.taxon	biology_ecology	Biology. Unknown.	en	Mai, Pham Quynh, Long, Khuat Dang, Hiep, Nguyen Duc, Hoa, Dang Thi, Duong, Tran Dinh (2023): First record of the genus Testudobracon Quicke, 1986 (Hymenoptera: Braconidae: Braconinae) in Vietnam, with description of four new species. Zootaxa 5244 (5): 485-500, DOI: 10.11646/zootaxa.5244.5.5, URL: http://zoobank.org/46962264-5ded-44b6-a42a-66fa97eb5f36
03D541284E037F7DFF762F2C22A7A8A6.taxon	etymology	Etymology. The new species is named after the locality (Phia Oac NP), where the holotype was collected. Notes. The differences between T. descensus sp. nov. and T. phiaoacus sp. nov. were given in the key; this new species is close to T. guangxinensis Wang, Chen & He, from China, but differs from the latter by the following characters: 1) Propodeum smooth, with a complete median longitudinal carina; 2) Middle lobe of mesoscutum with sparse but discretely punctate, anterio-medially with a weak longitudinal ridge, lateral lobes of mesoscutum with discrete punctures (nearly smooth in T. guangxinensis); and 3) Third – fifth metasomal tergites with oblique posterolateral lobe like protuberances in lateral view (rounded postero-lateral lobe like protuberances in T. guangxinensis)	en	Mai, Pham Quynh, Long, Khuat Dang, Hiep, Nguyen Duc, Hoa, Dang Thi, Duong, Tran Dinh (2023): First record of the genus Testudobracon Quicke, 1986 (Hymenoptera: Braconidae: Braconinae) in Vietnam, with description of four new species. Zootaxa 5244 (5): 485-500, DOI: 10.11646/zootaxa.5244.5.5, URL: http://zoobank.org/46962264-5ded-44b6-a42a-66fa97eb5f36
03D541284E047F7BFF762B8525C2AEE2.taxon	materials_examined	Material examined. Female, “ Bracn. 010 ” (IEBR), N Vietnam: Hai Phong, Cat Ba NP (Is.), forest, sweep, 20 ° 60 ′ 50.00 ″ N 107 ° 20 ′ 25.50 ″ E, 11. vi. 2002, KD Long. Female (Fig. 9 a), antenna with 21 flagellomeres; body length 2.2 mm; antenna 2.0 mm; fore wing 2.3 mm; ovipositor sheath 1.3 mm (Fig. 9 a). Ratio of length of fore wing veins r: 3 - SR: SR 1 = 4: 8: 23. and 2 - SR: 3 - SR: r-m = 7: 8: 5 (Fig. 9 e). Ovipositor 0.9 × metasoma in lateral view, and 0.5 × as long as fore wing.	en	Mai, Pham Quynh, Long, Khuat Dang, Hiep, Nguyen Duc, Hoa, Dang Thi, Duong, Tran Dinh (2023): First record of the genus Testudobracon Quicke, 1986 (Hymenoptera: Braconidae: Braconinae) in Vietnam, with description of four new species. Zootaxa 5244 (5): 485-500, DOI: 10.11646/zootaxa.5244.5.5, URL: http://zoobank.org/46962264-5ded-44b6-a42a-66fa97eb5f36
03D541284E047F7BFF762B8525C2AEE2.taxon	distribution	Distribution. N Vietnam (Hai Phong, Cat Ba NP).	en	Mai, Pham Quynh, Long, Khuat Dang, Hiep, Nguyen Duc, Hoa, Dang Thi, Duong, Tran Dinh (2023): First record of the genus Testudobracon Quicke, 1986 (Hymenoptera: Braconidae: Braconinae) in Vietnam, with description of four new species. Zootaxa 5244 (5): 485-500, DOI: 10.11646/zootaxa.5244.5.5, URL: http://zoobank.org/46962264-5ded-44b6-a42a-66fa97eb5f36
