identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03D487B8C47ECD44FF22FD5E13E5FEE3.text	03D487B8C47ECD44FF22FD5E13E5FEE3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus Ashmead 1893	<div><p>Key to Dissomphalus males of the Cerrado, Caatinga and Northern relicts of the Atlantic Forest, Brazil</p><p>1. Tergal process absent ( amplus species-group)............................................................... 2</p><p>- Tergal process present (Figs. 13, 18, 22).................................................................... 3</p><p>2. Mandible bidentate; posterior hypopygeal margin straight; aedeagal ventral ramus shorter than dorsal body (Figs 5–8).......................................................................... D. boitata Colombo &amp; Azevedo sp. nov.</p><p>- Mandible tridentate; posterior hypopygeal margin weakly concave; aedeagal ventral ramus longer than dorsal body (Figs 9– 11)................................................................... D. boto Colombo &amp; Azevedo sp. nov.</p><p>3. Tergal process located medially (Fig. 61)................................................................... 4</p><p>- Tergal process located laterally (Figs. 13, 18, 22)............................................................. 9</p><p>4. Tergal process with one pair of conspicuous depressions, transversally elliptical, tubercle present, directed each other (vallensis species-group) (Fig. 61)................................................................................ 5</p><p>- Tergal process with one pair of conspicuous depressions, never transversally elliptical, tubercle present, never directed each other................................................................................................ 6</p><p>5. Parameral apex convex and oblique; aedeagal ventral ramus as long as aedeagal dorsal body; inner lobe of aedeagal dorsal body never setose............................................................... D. bifurcatus Azevedo, 1999</p><p>- Parameral apex never convex and oblique; aedeagal ventral ramus shorter than aedeagal dorsal body; inner lobe of aedeagal dorsal body setose (Figs 60–65)........................................... D. torem Colombo &amp; Azevedo sp. nov.</p><p>6. Tufts of tergal process linear and horizontal ( linearis species-group)......................... D. linearis Azevedo, 1999</p><p>- Tufts of tergal process neither linear nor horizontal........................................................... 7</p><p>7. Tergal process with large tubercles; apex of aedeagal dorsal body concave; apex of aedeagal ventral ramus with row of small ventral teeth (longiclypeus species-group)............................................ D. scamatus Azevedo, 1999</p><p>- Tergal process without tubercles; apex of aedeagal dorsal body never concave; apex of aedeagal ventral ramus without row of small ventral teeth..................................................................................... 8</p><p>8. Tergal process with depression shallow, tufts large; aedeagal dorsal body nearly as long as ventral ramus, apex with two large teeth (apertus species-group)..................................................... D. subpilosus Azevedo, 1999</p><p>- Tergal process with depression deep; aedeagal dorsal body much longer than ventral ramus, apex without two large teeth (altivolans species-group)............................................................ D. megomphalus Evans, 1969</p><p>9. Aedeagal ventral ramus very longer than dorsal body, tubular, wide, rounded apex, without apical filament (brasiliensis species-group).......................................................... D. tubulatus Redighieri &amp; Azevedo, 2006</p><p>- Aedeagal ventral ramus never tubular (Figs. 15, 20, 24)...................................................... 10</p><p>10. Aedeagal ventral ramus with apical filament ( microstictus species-group)................... D. microstictus Evans, 1969</p><p>- Aedeagal ventral ramus without apical filament (Figs. 15, 20, 24).............................................. 11</p><p>11. Tergal process with callus; volsella with small basal projection (Figs 80–84)....................................................................................................... D. matintaperera Colombo &amp; Azevedo sp. nov.</p><p>- Tergal process without callus; volsella without such projection................................................. 12</p><p>12. Tergal process with one pair of conspicuous depressions, tubercle present, pit on top, lateral margin setose (Figs 71–75)......................................................................... D. caipora Colombo &amp; Azevedo sp. nov.</p><p>- Tergal process not as above............................................................................. 13</p><p>13. Tergal process with depression deeply excavated, depressions partially covered by flap........................................................................................................ D. ubracus Brito &amp; Azevedo, 2017</p><p>- Tergal process not deeply excavated, depressions not covered by flap........................................... 14</p><p>14. Pits of tergal process drop-shaped, sharp anteriorly and rounded posteriorly (guttus species-group).................................................................................................... D. latimerus Azevedo, 2003</p><p>- Pits of tergal process never drop-shaped, usually circular..................................................... 15</p><p>15. Tuft of tergal process with one distinct long seta ( gilvipes species-group) (Fig. 27)................................. 16</p><p>- Tuft of tergal process without such seta................................................................... 18</p><p>16. Median clypeal lobe duckbill-shaped; gena with one pair spines (Figs 25–29)........ D. irupe Colombo &amp; Azevedo sp. nov.</p><p>- Median clypeal lobe not duckbill-shaped; gena without spines................................................. 17</p><p>17. Notauli complete; apex of dorsal body narrowed in lateral view............................... D. gilvipes Evans, 1979</p><p>- Notauli incomplete or absent; apex of dorsal body wide in lateral view..................... D. krombeini Azevedo, 1999</p><p>18. Depressions of tergal process inclined and linear (rufipalpis species-group)...................... D. infissus Evans, 1969</p><p>- Tergal processes without depressions or if present never inclined and linear....................................... 19</p><p>19. Depressions of tergal process ovoid, deep and large, about 0.5 × as long as tergite, rim usually setose (bicavatus species-group)................................................................................................... 20</p><p>- Depressions of tergal process not as above................................................................. 22</p><p>20. Aedeagal basal process without median tooth........................................ D. geniculatus Azevedo, 2001</p><p>- Aedeagal basal process with median tooth................................................................ 21</p><p>21. Aedeagal basal process wide.................................................... D. lobisserratus Azevedo, 2001</p><p>- Aedeagal basal process narrow and sharpened....................................... D. curvilongus Azevedo, 2001</p><p>22. Tubercles of tergal process large with high rim and tuft densely setose ( conicus species-group) (Figs. 13, 18, 22)......... 23</p><p>- Tubercles of tergal process small........................................................................ 29</p><p>23. Basivolsela with thick, conspicuous bristles................................ D. laminaris Redighieri &amp; Azevedo, 2004</p><p>- Basivolsela without such bristles....................................................................... 24</p><p>24. Outer lobe of aedeagal dorsal body with warts............................. D. verrucosus Redighieri &amp; Azevedo, 2004</p><p>- Outer lobe of aedeagal dorsal body without warts.......................................................... 25</p><p>25. Aedeagal ventral ramus with apical half translucent, narrow, apex with two horn-shaped expansions................................................................................................... D. cervoides Azevedo, 2003</p><p>- Aedeagal ventral ramus without such expansions............................................................ 26</p><p>26. Basivolsella with large, wide, rounded expansion directed upward at inner margin; ventral margin of aedeagal ventral ramus with deep median emargination; dorsal margin of outer lobe of aedeagal dorsal body strongly conical in lateral view........................................................................................ D. conicus Azevedo, 2003</p><p>- Basivolsella without such expansion; aedeagal ventral ramus without emargination; dorsal margin of outer lobe of aedeagal dorsal body not conical in lateral view.................................................................... 27</p><p>27. Parameral apex bifurcated; aedeagal ventral ramus longer than dorsal body, with small basal extension (Figs 12–16)........................................................................... D. curupira Colombo &amp; Azevedo sp. nov.</p><p>- Parameral apex not bifurcated; aedeagal ventral ramus equal or shorter than dorsal body, without basal extension......... 28</p><p>28. Mandible with three teeth; parameral apex serrated; aedeagal ventral ramus as long as the dorsal body (Figs 21–24)............................................................................... D. saci Colombo &amp; Azevedo sp. nov.</p><p>- Mandible with two teeth; parameral apex rounded; aedeagal ventral ramus shorter than dorsal body (Figs 17–20).................................................................................. D. iara Colombo &amp; Azevedo sp. nov.</p><p>29. Depression of tergal process with edge without thick and conspicuous bristles (punctatus species-group) (Figs. 31, 35, 39)...................................................................................................... 30</p><p>- Depression of tergal process with edge with thick and conspicuous bristles....................................... 33</p><p>30. Median clypeal lobe trapezoidal; inner margin of basivolsella with large rounded callus.............. D. napo Evans, 1979</p><p>- Median clypeal lobe tridentate; inner margin of basivolsella without callus....................................... 31</p><p>31. Mandible with two teeth; parameral apex straight (Figs 38–41)................ D. jurupari Colombo &amp; Azevedo sp. nov.</p><p>- Mandible with three teeth; parameral apex rounded.......................................................... 32</p><p>32. Posterior hypopygeal margin weakly concave; parameral apex bifurcated in ventral view (Figs 30–33)..................................................................................... D. acutipupu Colombo &amp; Azevedo sp. nov.</p><p>- Posterior hypopygeal margin straight; parameral apex not bifurcated in ventral view (Figs 34–37)....... D. cuca Colombo &amp; Azevedo sp. nov.</p><p>33. Tubercles of tergal process slightly directed mesad (tuberculatus species-group)............... D. plaumanni Evans, 1964</p><p>- Tubercles of tergal process never directed mesad............................................................ 34</p><p>34. Paramere never peanut-shaped or S-shaped (setosus species-group)............................................. 35</p><p>- Paramere peanut-shaped or S-shaped (Figs. 68, 69).......................................................... 39</p><p>35. Dorsal margin of paramere with apical spine; aedeagal ventral ramus with apical half divided into two long tubular rami.......................................................................... D. magnus Redighieri &amp; Azevedo, 2006</p><p>- Dorsal margin of paramere without spine; aedeagal ventral ramus without such rami............................... 36</p><p>36. Mandible with four teeth............................................................................... 37</p><p>- Mandible with two or three teeth........................................................................ 38</p><p>37. Median clypeal lobe trapezoidal, median tooth straight, higher in lateral view; outer lobe of aedeagal dorsal body leaf-shaped (Figs 47–50)........................................................ D. capelobo Colombo &amp; Azevedo sp. nov.</p><p>- Median clypeal lobe tridentate, median tooth rounded, with posterior and lateral sides high and lower anterior and central regions; outer lobe of aedeagal dorsal body never leaf-shaped (Figs 51–55)....... D. iracema Colombo &amp; Azevedo sp. nov.</p><p>38. Mandible two teeth; median clypeal lobe undefined; posterior hypopygeal margin concave (Figs 42–46).................................................................................... D. boibumba Colombo &amp; Azevedo sp. nov.</p><p>- Mandible with three teeth; median clypeal lobe trapezoidal; posterior hypopygeal margin straight (Figs 56–59).............................................................................. D. joaquinae Colombo &amp; Azevedo sp. nov. 39. Median clypeal lobe trilobate; posterior hypopygeal margin weakly concave; paramere peanut-shaped (Figs 66–70)............................................................................ D. boiuna Colombo &amp; Azevedo sp. nov.</p><p>- Median clypeal lobe undefined; posterior hypopygeal margin straight; paramere S-shaped (Figs 76–79)................................................................................... D. mapinguari Colombo &amp; Azevedo sp. nov.</p></div>	https://treatment.plazi.org/id/03D487B8C47ECD44FF22FD5E13E5FEE3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colombo, Wesley D.;Alencar, Isabel D. C. C.;Limeira-De-Oliveira, Francisco;Azevedo, Celso O.	Colombo, Wesley D., Alencar, Isabel D. C. C., Limeira-De-Oliveira, Francisco, Azevedo, Celso O. (2018): New species and records of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Cerrado, Caatinga and relicts of the Atlantic Forest from northeastern Brazil. Zootaxa 4462 (1): 1-40, DOI: 10.11646/zootaxa.4462.1.1
03D487B8C478CD44FF22FE09148BFBB6.text	03D487B8C478CD44FF22FE09148BFBB6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus megomphalus Evans 1979	<div><p>Dissomphalus megomphalus Evans, 1979</p><p>Dissomphalus megomphalus Evans, 1979: 277, 279 (♂, holotype from Ecuador).</p><p>Diagnosis. Male. Dark castaneous. Mandible with three apical teeth. Median clypeal lobe tridentate. Front coriaceous. Vertex slightly convex. Propodeum strongly striate (rugulose). T2 vertical, large, deep concavity, its width 0.5 × width of tergite, two elliptical, large tufts densely short-haired, nearly joined. Posterior hypopygeal margin slightly concave. Genitalia: cuspis thin, much arched; aedeagal ventral ramus much smaller than dorsal body, wide, apical margin oblique, truncate; aedeagal dorsal body with two pairs apical lobes, outer pair laminar, surface subvertical, concave internally, basal half wide, nearly touching each other; inner pair laminar, membranous, hairy, ventral margin strongly denticulate. Female unknown.</p><p>Variations. The main variations found in this series are in the metasomal tergal processes, which have the depression semicircular or circular.</p><p>Remarks. This species was previously known from Ecuador (Evans 1979) and Brazil (Pará, Ceará) (Azevedo 1999a). Now it is recorded for the first time from Piauí and Maranhão.</p><p><a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-41.709446&amp;materialsCitation.latitude=-4.099167" title="Search Plazi for locations around (long -41.709446/lat -4.099167)">Material</a> examined. BRAZIL, PI[auí], 1♂ Piracuruca, PN de Sete Cidades, Posto do ICMBio, 04°05'57''S 41°42'34''W, 01–14.II.2013, Armadilha Suspensa, F Limeira-de-Oliveira, JS Pinto Júnior cols. (CZMA) ; MA[ranhão], 1♂ Mirador, Parq [ue] Estadual Mirador, Base da Geraldina, 03–07.V.2008, Armadilha luminosa, JC Silva, F Limeira-de-Oliveira cols. (CZMA) ; 1♂ Riachão, Fazenda Altos, 22.VIII.2009, Armadilha Malaise, F Limeira-de-Oliveira, MB Aguiar Neto cols. (CZMA) .</p></div>	https://treatment.plazi.org/id/03D487B8C478CD44FF22FE09148BFBB6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colombo, Wesley D.;Alencar, Isabel D. C. C.;Limeira-De-Oliveira, Francisco;Azevedo, Celso O.	Colombo, Wesley D., Alencar, Isabel D. C. C., Limeira-De-Oliveira, Francisco, Azevedo, Celso O. (2018): New species and records of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Cerrado, Caatinga and relicts of the Atlantic Forest from northeastern Brazil. Zootaxa 4462 (1): 1-40, DOI: 10.11646/zootaxa.4462.1.1
03D487B8C478CD49FF22FABB1447FB2D.text	03D487B8C478CD49FF22FABB1447FB2D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus boitata Colombo & Alencar & Limeira-De-Oliveira & Azevedo 2018	<div><p>Dissomphalus boitata Colombo &amp; Azevedo sp. nov.</p><p>(Figs 5–8)</p><p>Description. Male. Head and mesosoma black; metasoma dark castaneous. Mandible with two apical teeth; median clypeal lobe undefined, wide, median tooth weakly rounded; frons strongly coriaceous; eyes setose. Notauli complete; pronotal disc coriaceous. Metasomal tergite II without tergal process. Posterior hypopygeal margin straight. Genitalia: paramere with undefined apex, slightly arched inward, constricted, wide, specially basally, higher than the basiparamere; volsella very wide; aedeagal ventral ramus shorter than dorsal body, apex weakly rounded, convergent, laminar; aedeagal dorsal body with two pairs of apical lobes, dorsal margin folded across its length, apical and ventral margins serrated, inner pair membranous, basal process absent; apodeme extending beyond genital ring. Female unknown.</p><p>Variations. Paramere with narrow apex.</p><p>Remarks. This species is allocated in the amplus species-group because it lacks the tergal processes. It is similar to Dissomphalus clovisi Colombo &amp; Azevedo by having the mandibles bidentate, the median clypeal lobe undefined, and the aedeagus without the basal process. However, D. boitata sp. nov. has the paramere with the apex undefined, higher than the basiparamere, and the aedeagal ventral ramus with apex weakly rounded, whereas D. clovisi has the paramere with the apex serrate, smaller than basiparamere, and the aedeagal ventral ramus with the apex narrow.</p><p>Material examined. Holotype ♂, BRAZIL, MA[ranhão], Carolina, PN Chapada das Mesas, Riacho Cancela, 07°06'44.2''S 47°17'56.8''W, 01–10.VIII.2013, Armadilha suspensa, JA Rafael, F Limeira-de-Oliveira, TTA Silva cols. (CZMA). Paratypes, 1♂, same data of holotype (CZMA).</p><p>Etymology. The epithet boitata from Tupi-Guarani refers to an indigenous myth and means a snake of fire. Treat as a noun in apposition.</p><p>Distribution. Brazil (Maranhão).</p></div>	https://treatment.plazi.org/id/03D487B8C478CD49FF22FABB1447FB2D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colombo, Wesley D.;Alencar, Isabel D. C. C.;Limeira-De-Oliveira, Francisco;Azevedo, Celso O.	Colombo, Wesley D., Alencar, Isabel D. C. C., Limeira-De-Oliveira, Francisco, Azevedo, Celso O. (2018): New species and records of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Cerrado, Caatinga and relicts of the Atlantic Forest from northeastern Brazil. Zootaxa 4462 (1): 1-40, DOI: 10.11646/zootaxa.4462.1.1
03D487B8C475CD48FF22FA8A1477FF39.text	03D487B8C475CD48FF22FA8A1477FF39.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus boto Colombo & Alencar & Limeira-De-Oliveira & Azevedo 2018	<div><p>Dissomphalus boto Colombo &amp; Azevedo sp. nov.</p><p>(Figs 9–11)</p><p>Description. Male. Head and mesosoma black; metasoma light castaneous. Mandible with three apical teeth; median clypeal lobe undefined, wide, median tooth rounded; frons strongly coriaceous; eyes setose. Notauli incomplete. Pronotal disc strongly coriaceous. Metasomal tergite II without tergal process. Posterior hypopygeal margin weakly concave. Genitalia: paramere with weakly rounded apex, slightly arched inward, as long as basiparamere; aedeagal ventral ramus longer than dorsal body, apex rounded, curved dorsally, laminar, very narrow in apical half; aedeagal dorsal body with two pair of apical lobes, dorsal margin folded across its length, apical and ventral margins serrated, inner pair membranous, basal process absent; apodeme extending beyond genital ring. Female unknown.</p><p>Variations. Apical lobes of aedeagal dorsal body with apical and ventral margins weakly serrated.</p><p>Remarks. This species belongs to amplus species-group because it lacks the tergal processes. It is similar to Dissomphalus amplus Redighieri &amp; Azevedo by having the mandibles tridentate, the median clypeal lobe undefined, and the aedeagus without a basal process. However, D. boto sp. nov. has the aedeagal ventral ramus very narrow and longer than the dorsal body, whereas D. amplus has the aedeagal ventral ramus wide and shorter than the dorsal body.</p><p>Material examined. Holotype ♂, BRAZIL, CE[ará], Ubajara, PN Ubajara, Cachoeira do Cafundó, 03°50'13''S 40°54'35''W, 16–31.XII.2012, Armadilha Malaise, JS Pinto Junior, F Limeira-de-Oliveira cols. (CZMA). Paratypes, 1♂, same data of holotype (CZMA).</p><p>Etymology. The epithet boto from Portuguese refers to a Brazilian folk tale in which women explain their outof-wedlock pregnancy by boto (dolphin-like animal) in the Amazon region. Treat as a noun in apposition. Distribution. Brazil (Ceará).</p></div>	https://treatment.plazi.org/id/03D487B8C475CD48FF22FA8A1477FF39	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colombo, Wesley D.;Alencar, Isabel D. C. C.;Limeira-De-Oliveira, Francisco;Azevedo, Celso O.	Colombo, Wesley D., Alencar, Isabel D. C. C., Limeira-De-Oliveira, Francisco, Azevedo, Celso O. (2018): New species and records of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Cerrado, Caatinga and relicts of the Atlantic Forest from northeastern Brazil. Zootaxa 4462 (1): 1-40, DOI: 10.11646/zootaxa.4462.1.1
03D487B8C474CD4BFF22FB48143FFE49.text	03D487B8C474CD4BFF22FB48143FFE49.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus curvilongus Azevedo 2001	<div><p>Dissomphalus curvilongus Azevedo, 2001</p><p>Dissomphalus curvilongus Azevedo, 2001: 354 –355 (♂, holotype from Brazil).</p><p>Diagnosis. Male. Black. Mandible with two apical teeth. Median clypeal lobe trapezoidal. Frons strongly coriaceous. Genitalia: volsella with long apical tooth; aedeagal ventral ramus as long as dorsal body, laminar, surface subhorizontal, wide, apex converging, bidentate; aedeagal dorsal body with two pairs apical lobes, outer pair divided in 2 rami, dorsal ramus laminar, progressively thinner, outer margin conspicuously serrated, apical area setose serrated, ventral ramus thin, not serrated; inner pair membranous, stout, inner surface setose; basal process with median tooth, pair of lateral expansions somewhat wide, long, more than 0.5 × Length paramere, curved downward, nearly reaching ramus aedeagus, setose serrated.</p><p>Variations. The main variation found in this series is the development of the aedeagal basal process, which can be longer than the apical lobes, equally as long, or shorter than the apical lobes.</p><p>Remarks. This species was previously known from Trinidad &amp; Tobago and Brazil (Amapá, Amazonas, Pará, Acre, Rondônia, Ceará and Maranhão) (Azevedo 2001). Now it is recorded for the first time from Piauí and one more male was found in Maranhão.</p><p><a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-41.709446&amp;materialsCitation.latitude=-4.099167" title="Search Plazi for locations around (long -41.709446/lat -4.099167)">Material</a> examined. BRAZIL, PI[auí], 1♂ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-41.709446&amp;materialsCitation.latitude=-4.099167" title="Search Plazi for locations around (long -41.709446/lat -4.099167)">Piracuruca</a>, PN de Sete Cidades, Posto do ICMBio, 04°05'57''S 41°42'34''W, 15–30.VI.2013, Armadilha Malaise, JA Rafael, F Limeira-de-Oliveira, TTA Silva cols. (CZMA) ; 4♂ 01–27.III.2013, 1♂ 09–20.V.2013, 3♂ 19-28.II.2013, F Limeira-de-Oliveira, TTA Silva cols. (CMZA); 1♂ Varredura, JA Rafael, F Limeira-de-Oliveira, TTA Silva cols. (CZMA); 2♂ 16.II–10.III.2013, F Limeira-de- Oliveira, JS Pinto Júnior cols. (CZMA); MA[ranhão], 6♂ Mirador, Parq[ue] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.86889&amp;materialsCitation.latitude=-6.6236115" title="Search Plazi for locations around (long -45.86889/lat -6.6236115)">Estadual Mirador</a>, Base da Geraldina, 06°37'25''S 45°52'08''W, 08.III–16.V.2013, Armadilha Malaise, LS Santos, F Limeira-de-Oliveira, LLM Santos cols. (CZMA) ; 1♂ 01–10.XII.2013, TL Rocha, F Limeira-de-Oliveira, LLM Santos cols. (CZMA); 1♂ 29.V.2014, Arm[adilha] suspensa (CZMA); 1♂ 12–20.VIII.2013, Armadilha suspensa, F Limeira-de-Oliveira, LLM Santos, LS Santos cols. (CZMA); 1♂ C[ândido] Mendes, Fazenda Sete Irmãos, 01°51'37''S 45°46'10''W, 06.X.2008, Armadilha Malaise, F Limeira-de-Oliveira, JA Rafael, PAM Moraes cols. (CZMA); 1♂ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-47.308777&amp;materialsCitation.latitude=-7.118222" title="Search Plazi for locations around (long -47.308777/lat -7.118222)">Carolina</a>, PN <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-47.308777&amp;materialsCitation.latitude=-7.118222" title="Search Plazi for locations around (long -47.308777/lat -7.118222)">Chapada</a> das Mesas, Riacho Sucuruiu, 07°07'05.6''S 47°18'31.6''W, 01–15.VII.2013, Armadilha Malaise, JA Rafael, F Limeirade-Oliveira, TTA Silva cols. (CZMA) .</p></div>	https://treatment.plazi.org/id/03D487B8C474CD4BFF22FB48143FFE49	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colombo, Wesley D.;Alencar, Isabel D. C. C.;Limeira-De-Oliveira, Francisco;Azevedo, Celso O.	Colombo, Wesley D., Alencar, Isabel D. C. C., Limeira-De-Oliveira, Francisco, Azevedo, Celso O. (2018): New species and records of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Cerrado, Caatinga and relicts of the Atlantic Forest from northeastern Brazil. Zootaxa 4462 (1): 1-40, DOI: 10.11646/zootaxa.4462.1.1
03D487B8C474CD48FF22FE381357FBA9.text	03D487B8C474CD48FF22FE381357FBA9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus subpilosus Azevedo. Of 1999	<div><p>Dissomphalus subpilosus Azevedo, 1999</p><p>Dissomphalus subpilosus Azevedo, 1999a: 354 –355 (♂, holotype from Brazil).</p><p>Diagnosis. Male. Dark castaneous. Mandible with three apical teeth. Clypeus flat, tridentate, median carina incomplete. Front coriaceous. Vertex slightly convex. T2 vertical, very shallow depression, occupying median third, width and of length of tergite, distant from anterior margin, pair rounded, very large tufts densely covered by short hairs, far from each other 0.75 × own diameter, surrounded by some hairs inward, longer than those, tufts, tergite with median longitudinal band of hairs posteriorly tuft. Genitalia: paramere with apex oblique-truncate; aedeagal ventral ramus conspicuous, smaller than dorsal body, margins smooth, parallel, apex oblique-truncate, corner rounded, laminar, surfaces subhorizontal, divergent laterally, downward; aedeagal dorsal body with two pairs of lobes, outer large, ventral margin smooth, narrowing apically, dorsal margin much produced inward basally, surfaces subvertical, somewhat divergent ventrally, very concave internally, inner lobe small, inner margin serrated up to apex, dorsal smooth, inner surface densely hairy, somewhat convex. Female unknown.</p><p>Variations. The main variation found in this series is the metasomal tergal process with the depression semicircular or circular.</p><p>Remarks. This species was previously known from Brazil (Ceará) and Bolivia (Azevedo 1999a). Now, one more male has been found in Ceará. In this series the specimens are quite similar to those previously studied, without any variation that is worthy of mention.</p><p><a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-40.909725&amp;materialsCitation.latitude=-3.8369443" title="Search Plazi for locations around (long -40.909725/lat -3.8369443)">Material</a> examined. BRAZIL, CE[ará], 1♂ Ubajara, PN Ubajara, Cachoeira do Cafundó, 03°50'13''S 40°54'35''W, 16–31.XII.2012, Armadilha Malaise, F Limeira-de-Oliveira, JS Pinto Junior cols. (CZMA).</p></div>	https://treatment.plazi.org/id/03D487B8C474CD48FF22FE381357FBA9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colombo, Wesley D.;Alencar, Isabel D. C. C.;Limeira-De-Oliveira, Francisco;Azevedo, Celso O.	Colombo, Wesley D., Alencar, Isabel D. C. C., Limeira-De-Oliveira, Francisco, Azevedo, Celso O. (2018): New species and records of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Cerrado, Caatinga and relicts of the Atlantic Forest from northeastern Brazil. Zootaxa 4462 (1): 1-40, DOI: 10.11646/zootaxa.4462.1.1
03D487B8C477CD4BFF22FE6F177BFB86.text	03D487B8C477CD4BFF22FE6F177BFB86.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus geniculatus Azevedo 2001	<div><p>Dissomphalus geniculatus Azevedo, 2001</p><p>Dissomphalus geniculatus Azevedo, 2001: 185 (♂, holotype from Costa Rica).</p><p>Diagnosis. Male. Black. Mandible with two apical teeth. Median clypeal lobe trapezoidal. Frons strongly coriaceous. Genitalia: paramere with concave apex, corner produced; basivolsella with sharpened and large apical tooth, base digitus produced; aedeagal ventral ramus as long as dorsal body, laminar, surface subvertical, apical half convergent, apex bidentate, inner tooth larger; aedeagal dorsal body with three pairs apical lobes, outer pair bifurcated in two arms, dorsal arm with series of sharp teeth and ventral arm smaller and sharp; two inner pairs membranous, stout, median one small, setose; basal process with pair arched teeth.</p><p>Variations. The main variations found in this series are in the apex of the paramere, which can be weakly or strongly bifurcated.</p><p>Remarks. This species was previously known from Costa Rica and Panama (Azevedo 2001). Now it is recorded for the first time from Brazil (Maranhão).</p><p><a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.86889&amp;materialsCitation.latitude=-6.6236115" title="Search Plazi for locations around (long -45.86889/lat -6.6236115)">Material</a> examined. BRAZIL, MA[ranhão], 1♂ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.86889&amp;materialsCitation.latitude=-6.6236115" title="Search Plazi for locations around (long -45.86889/lat -6.6236115)">Mirador</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.86889&amp;materialsCitation.latitude=-6.6236115" title="Search Plazi for locations around (long -45.86889/lat -6.6236115)">Parq</a> [ue] Estadual Mirador, Base da Geraldina, 06°37'25''S 45°52'08''W, 14–18.VIII.2012, Armadilha Malaise, DWA Marques, F Limeira-de-Oliveira, JSP Júnior cols. (CZMA) ; 1♂ 06°37'06''S 45°51'51''W, 01–12.VIII.2013, LS Santos, F Limeira-de-Oliveira, LLM Santos cols. (CZMA).</p></div>	https://treatment.plazi.org/id/03D487B8C477CD4BFF22FE6F177BFB86	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colombo, Wesley D.;Alencar, Isabel D. C. C.;Limeira-De-Oliveira, Francisco;Azevedo, Celso O.	Colombo, Wesley D., Alencar, Isabel D. C. C., Limeira-De-Oliveira, Francisco, Azevedo, Celso O. (2018): New species and records of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Cerrado, Caatinga and relicts of the Atlantic Forest from northeastern Brazil. Zootaxa 4462 (1): 1-40, DOI: 10.11646/zootaxa.4462.1.1
03D487B8C477CD4BFF22FB311546F923.text	03D487B8C477CD4BFF22FB311546F923.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus lobisserratus Azevedo 2001	<div><p>Dissomphalus lobisserratus Azevedo, 2001</p><p>Dissomphalus lobisserratus Azevedo, 2001: 188, 190 (♂, holotype from Brazil).</p><p>Diagnosis. Male. Black. Mandible with two apical teeth. Median clypeal lobe trapezoidal. Frons somewhat strongly coriaceous, punctures shallow. Genitalia: basivolsella with large sharp apical tooth; aedeagal ventral ramus as long as dorsal body, laminar, surface horizontal, wide, parallel, outer margin narrowing into narrow apex; aedeagal dorsal body with three pairs apical lobes, outer pair thin, evenly convex, apex thin directed each other; dorsal inner pair short, laminar, surface horizontal, parallel, wide, apex broad; ventral pair laminar, apex thin, border strongly serrated, inner surface setose; basal process with pair lateral wide teeth, serrated apical margin, wide median tooth.</p><p>Variations. In this series the specimens are quite similar to those previously studied.</p><p>Remarks. This species was previously known from Brazil (Ceará) (Azevedo 2001). Now it is recorded for the first time from Maranhão.</p><p><a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.880276&amp;materialsCitation.latitude=-6.63" title="Search Plazi for locations around (long -45.880276/lat -6.63)">Material</a> examined. BRAZIL, MA[ranhão], 2♂ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.880276&amp;materialsCitation.latitude=-6.63" title="Search Plazi for locations around (long -45.880276/lat -6.63)">Mirador</a>, Parque Estadual Mirador, Base da Geraldina, 416m, 06°37'48''S 45°52'49''W, Armadilha Malaise, 10–20.XI.2013, F Limeira-de-Oliveira, LLM Santos, TL Rocha cols. (CZMA) ; 1♂ Armadilha suspensa, 01–10.XI.2013 (CZMA) .</p></div>	https://treatment.plazi.org/id/03D487B8C477CD4BFF22FB311546F923	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colombo, Wesley D.;Alencar, Isabel D. C. C.;Limeira-De-Oliveira, Francisco;Azevedo, Celso O.	Colombo, Wesley D., Alencar, Isabel D. C. C., Limeira-De-Oliveira, Francisco, Azevedo, Celso O. (2018): New species and records of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Cerrado, Caatinga and relicts of the Atlantic Forest from northeastern Brazil. Zootaxa 4462 (1): 1-40, DOI: 10.11646/zootaxa.4462.1.1
03D487B8C477CD4AFF22F8CF148CFBBD.text	03D487B8C477CD4AFF22F8CF148CFBBD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus tubulatus Redighieri & Azevedo 2006	<div><p>Dissomphalus tubulatus Redighieri &amp; Azevedo, 2006</p><p>Dissomphalus tubulatus Redighieri &amp; Azevedo, 2006: 320 (♂, holotype from Brazil).</p><p>Diagnosis. Male. Black. Mandible with three apical teeth. Clypeus broadly projected forward, median lobe not defined, median tooth conspicuous, sharpened. Frons weakly coriaceous, punctures large, shallow. Vertex straight. T2 with pair hairy pitted median tubercles, separated each other by slightly more than their own diameters. Posterior hypopygeal margin straight. Genitalia: paramere with conspicuous basal setae in ventral margin; digitus with huge basal projection; aedeagal ventral ramus longer than dorsal body, broad, tubular, wide, apex rounded, apical surface partially microspinose; aedeagal dorsal body outer pair very short, wide; inner pair filamentous, arched downward.</p><p>Variations. In this series the specimens are quite similar to those previously studied.</p><p>Remarks. This species was previously known from Brazil (Paraíba) (Redighieri &amp; Azevedo 2006). Now it is recorded for the first time from Maranhão and Piauí.</p><p><a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-43.561665&amp;materialsCitation.latitude=-9.141055" title="Search Plazi for locations around (long -43.561665/lat -9.141055)">Material</a> examined. BRAZIL, PI[auí], 2♂ Guaribas, PN Serra das Confusões, Andorinhas, 515 m, 09°08'27.8''S 43°33'42''W, 01–10.XI.2013, Armadilha Malaise, JA Rafael, F Limeira-de-Oliveira, TTA Silva cols. (CZMA) ; 1♂ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-41.709446&amp;materialsCitation.latitude=-4.099167" title="Search Plazi for locations around (long -41.709446/lat -4.099167)">Piracuruca</a>, PN de Sete Cidades, Posto do ICMBio, 04°05'57''S 41°42'34''W, 01–14.II.2013, Armadilha suspensa, F Limeira-de-Oliveira, JS Pinto Júnior cols. (CZMA); MA[ranhão] , 3♂ Mirador, Parq[ue] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.86889&amp;materialsCitation.latitude=-6.6236115" title="Search Plazi for locations around (long -45.86889/lat -6.6236115)">Estadual Mirador</a>, Base da Geraldina, 06°37'25''S 45°52'08''W, 08–14.III.2013, Armadilha Malaise, LS Santos, F Limeira-de-Oliveira, LLM Santos cols. (CZMA) ; 5♂ 10–16.V.2013, LS Santos, F Limeira-de-Oliveira, LLM Santos cols. (CZMA); 5♂ 06°37'06''S 45°51'51''W, 411 m, 02–12.VIII.2013, LS Santos, F Limeira-de-Oliveira, LLM Santos cols. (CZMA); 1♂ 03–06. X.2 0 0 8 (CZMA); 1♂ 1–10.XI.2013 (CZMA); 1♂ 06°35'58''S 45°50'49''W, 402m, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.846943&amp;materialsCitation.latitude=-6.5994444" title="Search Plazi for locations around (long -45.846943/lat -6.5994444)">Armadilha de Malaise</a>, 10–20.X.2013, F Limeira-de-Oliveira, LLM Santos, TL Rocha cols. (CZMA) ; 1♂ 01–10.I.2014, F Limeira-de-Oliveira, LLM Santos, LS Santos cols. (CZMA); 1♂ 06°37'25''S 45°52'08''W, 409 m, 01–10.X.2013, F Limeira-de-Oliveira, LLM Santos, TL Rocha cols. (CZMA); 2♂ 06°37'48''S 45°52'49''W, 416m, 01–10.XII.2013, F Limeira-de-Oliveira, LLM Santos, LS Santos cols. (CZMA); 1♂ Armadilha Suspensa, 10– 20.XI.2013 (CZMA) ; 3♂ C[ândido] Mendes, Fazenda Sete Irmãos, 01°51'37''S 45°46'10''W, 03–06.X.2008, Armadilha Malaise, F Limeira-de-Oliveira, JA Rafael, PAM Moraes cols. (CZMA); 6♂ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-47.29911&amp;materialsCitation.latitude=-7.1122775" title="Search Plazi for locations around (long -47.29911/lat -7.1122775)">Carolina</a>, PN <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-47.29911&amp;materialsCitation.latitude=-7.1122775" title="Search Plazi for locations around (long -47.29911/lat -7.1122775)">Chapada</a> das Mesas, Riacho Cancela, 07°06'44.2''S 47°17'56.8''W, 255m, 01.VII–31.VIII.2013, Armadilha Malaise, JA Rafael, F Limeira-de-Oliveira, TTA Silva cols. (CZMA) ; 1♂ Riacho Sucuruiu, 01–10.IX.2013 (CZMA) ; 1♂ 07°06'44.6''S 47°17'56.8''W, 240m, 11–14.VI.2013, JA Rafael, F Limeira-de-Oliveira, TTA Silva cols. (CZMA); 1♂ 07°07'05.6''S 47°18'31.6''W, 14–30.VI.2013 (CZMA).</p></div>	https://treatment.plazi.org/id/03D487B8C477CD4AFF22F8CF148CFBBD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colombo, Wesley D.;Alencar, Isabel D. C. C.;Limeira-De-Oliveira, Francisco;Azevedo, Celso O.	Colombo, Wesley D., Alencar, Isabel D. C. C., Limeira-De-Oliveira, Francisco, Azevedo, Celso O. (2018): New species and records of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Cerrado, Caatinga and relicts of the Atlantic Forest from northeastern Brazil. Zootaxa 4462 (1): 1-40, DOI: 10.11646/zootaxa.4462.1.1
03D487B8C476CD4DFF22FAAB1545FF15.text	03D487B8C476CD4DFF22FAAB1545FF15.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus cervoides Azevedo 2003	<div><p>Dissomphalus cervoides Azevedo, 2003</p><p>Dissomphalus cervoides Azevedo, 2003: 30 –32 (♂, holotype from Brazil).</p><p>Diagnosis. Male. Black. Mandible with two apical teeth. Median clypeal lobe trapezoidal. Frons coriaceous. Vertex slightly convex. T2 small, shallow lateral depression, tubercle pit-topped, dense setae directed backward. Posterior hypopygeal margin straight, pair small conical median protuberances. Genitalia: paramere with dorsal margin with basal callus, ventral margin concave above; basivolsella with scales below; aedeagal ventral ramus smaller than dorsal body, laminar, surface horizontal, translucent, apex short-horned at inner, outer corners, inner margin expanded, serrated below apex; aedeagal dorsal body with three pairs apical lobes, dorsal pair thick, base with outer projection, surface vertical, parallel, wide in lateral view, arched downward, dorsal margin angled, ventral margin serrated, ventral pair membranous, stout, rounded, much smaller than dorsal one, median pair less stout, less membranous, smaller than median one; basal process with pair lateral rounded expansions outward; ventral side genital ring wide.</p><p>Variations. In this series the specimens are quite similar to those previously studied.</p><p>Remarks. This species was previously known from Panama, Trinidad &amp; Tobago, Ecuador, Peru, Brazil (Amazonas, Pará, Rondônia, Mato Grosso, Goiás) and Bolivia (Azevedo 2003). Now it is recorded for the first time from Maranhão.</p><p><a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-47.29911&amp;materialsCitation.latitude=-7.1122775" title="Search Plazi for locations around (long -47.29911/lat -7.1122775)">Material</a> examined. BRAZIL, MA[ranhão], 1♂ Carolina, PN Chapada das Mesas, Riacho Cancela, 07°06'44.2''S 47°17'56.8''W, 255m, 01–10.VIII.2013, Armadilha suspensa, JA Rafael, F Limeira-de-Oliveira, TTA Silva cols. (CZMA) ; 1♂ 07°06'44.2''S 47°17'58.3''W, 01–15.VII.2013 (CZMA); 1♂ 28–31.X.2008, F Limeira-de-</p><p>Oliveira, PAM Moraes cols. (CZMA); 1♂ 07°06'44.2''S 47°17'56.8''W, 255m, 01–10.IX.2013, Armadilha Malaise, JA Rafael, F Limeira-de-Oliveira, TTA Silva cols. (CZMA).</p></div>	https://treatment.plazi.org/id/03D487B8C476CD4DFF22FAAB1545FF15	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colombo, Wesley D.;Alencar, Isabel D. C. C.;Limeira-De-Oliveira, Francisco;Azevedo, Celso O.	Colombo, Wesley D., Alencar, Isabel D. C. C., Limeira-De-Oliveira, Francisco, Azevedo, Celso O. (2018): New species and records of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Cerrado, Caatinga and relicts of the Atlantic Forest from northeastern Brazil. Zootaxa 4462 (1): 1-40, DOI: 10.11646/zootaxa.4462.1.1
03D487B8C471CD4DFF22FE991515FC03.text	03D487B8C471CD4DFF22FE991515FC03.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus conicus Azevedo 2003	<div><p>Dissomphalus conicus Azevedo, 2003</p><p>Dissomphalus conicus Azevedo, 2003: 32 (♂, holotype from Brazil).</p><p>Diagnosis. Male. Black. Mandible with two apical teeth. Median clypeal lobe trapezoidal. Frons strongly coriaceous. Vertex badly convex. T2 small, shallow lateral depression, tubercle wide pit on top, dense tuft setae directed backward. Posterior hypopygeal margin straight. Genitalia: basivolsella with large, wide, rounded expansion directed upward in inner margin; aedeagal ventral ramus as long as dorsal body, laminar, surface vertical, wide, dorsal median emargination in lateral view; aedeagal dorsal body with three pairs apical lobes; outer pair parallel, dorsal margin conical, angled medially upward in lateral view, ventral margin with two large, long teeth, apical margin apical tooth serrated; dorsal and ventral inner pairs stout, membranous, dorsal one smaller, colliculate, dorsal margin serrated; aedeagal base long central tooth, slightly inclined upward.</p><p>Variations. In this series the specimens are quite similar to those previously studied.</p><p>Remarks. This species was previously known from Brazil (Ceará, Pernambuco, Distrito Federal, Minas Gerais, Espírito Santo, Rio de Janeiro, São Paulo, Paraná) (Azevedo 2003), Brazil (Alagoas, Bahia) (Redighieri &amp; Azevedo 2006). Now some more males have been found in Ceará.</p><p><a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-40.909725&amp;materialsCitation.latitude=-3.8369443" title="Search Plazi for locations around (long -40.909725/lat -3.8369443)">Material</a> examined. BRAZIL, CE[ará], 1♂ Ubajara, PN Ubajara, Cachoeira do Cafundó, 03°50'13''S 40°54'35''W, 13–17.XI.2013, Armadilha Malaise, DWA Marques, F Limeira-de-Oliveira cols. (CZMA) ; 1♂ 01– 15.I.2013, JS Pinto Junior, F Limeira-de-Oliveira cols. (CZMA).</p></div>	https://treatment.plazi.org/id/03D487B8C471CD4DFF22FE991515FC03	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colombo, Wesley D.;Alencar, Isabel D. C. C.;Limeira-De-Oliveira, Francisco;Azevedo, Celso O.	Colombo, Wesley D., Alencar, Isabel D. C. C., Limeira-De-Oliveira, Francisco, Azevedo, Celso O. (2018): New species and records of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Cerrado, Caatinga and relicts of the Atlantic Forest from northeastern Brazil. Zootaxa 4462 (1): 1-40, DOI: 10.11646/zootaxa.4462.1.1
03D487B8C471CD4CFF22FBAB14D0FE6D.text	03D487B8C471CD4CFF22FBAB14D0FE6D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus laminaris Redighieri & Azevedo 2004	<div><p>Dissomphalus laminaris Redighieri &amp; Azevedo, 2004</p><p>Dissomphalus laminaris Redighieri &amp; Azevedo, 2004: 331 (♂, holotype from Brazil).</p><p>Diagnosis. Male. Black. Mandible bidentate or tridentate. Median clypeal lobe trapezoidal. Frons strongly coriaceous, very punctate. T2 with pair of shallow or deep, rounded lateral depressions, tubercle present, flattopped, small tuft of setae more or less dense. Posterior hypopygeal margin weakly concave. Genitalia: paramere with apex rounded, barely arched inward, series of small rounded teeth in ventral side; ventral ramus longer than dorsal body, straight, laminar, surface horizontal, apical half divided into two arms, outer one with apex rounded and inner one short and connected to dorsal body; dorsal body with two pairs apical lobes, outer pair laminar, straight, apex with two sharpened teeth, one directed upward or directed downward, several small teeth around; inner pair stout, membranous; basal process laminar, slightly arched upward; apodeme extending beyond elliptical genital ring. Female unknown.</p><p>Variations. The main variations found in this series are the paramere wider than the basiparamere and the external lobe of the aedeagal dorsal body longer than the basivolsella.</p><p>Remarks. This species was previously known from Brazil (Espírito Santo) (Redighieri &amp; Azevedo 2004), Brazil (Paraíba, Pernambuco, Alagoas, Sergipe, Bahia, São Paulo) (Redighieri &amp; Azevedo 2006). Now it is recorded for the first time from Ceará, Piauí, and Maranhão.</p><p><a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-40.909725&amp;materialsCitation.latitude=-3.8369443" title="Search Plazi for locations around (long -40.909725/lat -3.8369443)">Material</a> examined. BRAZIL, CE[ará], 1♂ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-40.909725&amp;materialsCitation.latitude=-3.8369443" title="Search Plazi for locations around (long -40.909725/lat -3.8369443)">Ubajara</a>, PN Ubajara, Cachoeira do Cafundó, 03°50'13''S 40°54'35''W, (CZMA) ; 1♂ 16–31.XII.2012, Armadilha suspensa, JS Pinto Junior, F Limeira-de-Oliveira cols. (CZMA) ; 1♂ 15–28.II.2013, (CZMA); PI[auí], 3♂ Caracol, PN Serra das Confusões, 15–31.VII.2013 (1♂) , 1– 10.IX.2013 (1♂), 20–30.IX.2013 (1♂), Armadilha Malaise, JA Rafael, F Limeira-de-Oliveira, TTA Silva cols. (CZMA) ; 5♂ 01–11.VIII.2013, (CZMA); 16♂ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-43.490612&amp;materialsCitation.latitude=-9.219972" title="Search Plazi for locations around (long -43.490612/lat -9.219972)">Riacho</a> dos <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-43.490612&amp;materialsCitation.latitude=-9.219972" title="Search Plazi for locations around (long -43.490612/lat -9.219972)">Bois</a>, 575m, 09°13'11.9''S 43°29'26.2''W, 15.VII– 30.XI.2013, Armadilha Malaise, JA Rafael, F Limeira-de-Oliveira, TTA Silva cols. (CZMA) ; 2♂ 20–31.I.2014, JA Rafael, F Limeira-de-Oliveira, TL Rocha, S Pereira cols. (CZMA) ; 2♂ PN <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-43.561695&amp;materialsCitation.latitude=-9.141055" title="Search Plazi for locations around (long -43.561695/lat -9.141055)">Andorinhas</a>, 09°08'27.8''S 43°33'42.1''W, 515m, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-43.561695&amp;materialsCitation.latitude=-9.141055" title="Search Plazi for locations around (long -43.561695/lat -9.141055)">Armadilha Malaise</a>, JA <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-43.561695&amp;materialsCitation.latitude=-9.141055" title="Search Plazi for locations around (long -43.561695/lat -9.141055)">Rafael</a>, F Limeira-de-Oliveira, TTA Silva cols. (CZMA) ; 1♂ 03.VII.2013 (CZMA); 2♂ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-43.561695&amp;materialsCitation.latitude=-9.219972" title="Search Plazi for locations around (long -43.561695/lat -9.219972)">Guaribas</a>, PAR [QUE]NA[CIONAL] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-43.561695&amp;materialsCitation.latitude=-9.219972" title="Search Plazi for locations around (long -43.561695/lat -9.219972)">Serra</a> das <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-43.561695&amp;materialsCitation.latitude=-9.219972" title="Search Plazi for locations around (long -43.561695/lat -9.219972)">Confusões</a>, Andorinhas, 09°13'11.9''S 43°33'42.1''W, 515m, 15.II–10.XI.2013, Armadilha Malaise, JA Rafael, F Limeira-de-Oliveira, TTA Silva cols. (CZMA) ; 2♂ 11–20.VIII.2013, (CZMA); 2♂ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-41.709446&amp;materialsCitation.latitude=-4.099167" title="Search Plazi for locations around (long -41.709446/lat -4.099167)">Piracuruca</a>, PN de <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-41.709446&amp;materialsCitation.latitude=-4.099167" title="Search Plazi for locations around (long -41.709446/lat -4.099167)">Sete Cidades</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-41.709446&amp;materialsCitation.latitude=-4.099167" title="Search Plazi for locations around (long -41.709446/lat -4.099167)">Posto</a> do ICMBio, 04°05'57''S 41°42'34''W, 01–10.III.2013, Armadilha suspensa, F Limeira-de-Oliveira, TTA Silva cols. (CZMA) ; 5♂ 01.II– 15.XII.2013, F Limeira-de-Oliveira, JS Pinto Júnior cols. (CZMA) ; 1♂ 12–27.III.2013, F Limeira-de-Oliveira, TTA Silva cols. (CZMA) ; 5♂ 01.II–27.III.2013, Armadilha Malaise, (CZMA) ; 3♂ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-43.501167&amp;materialsCitation.latitude=-9.211861" title="Search Plazi for locations around (long -43.501167/lat -9.211861)">Saco da Cachorra</a>, 522m, 09°12'42.7''S 43°30'04.2''W, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-43.501167&amp;materialsCitation.latitude=-9.211861" title="Search Plazi for locations around (long -43.501167/lat -9.211861)">Armadilha Malaise</a>, 01.IX–10.I.2014, JA Rafael, F Limeira-de-Oliveira, TL Rocha, S Pereira cols. (CZMA) ; 1♂ [ Armadilha] Suspensa, 15.I.2014, [CZMA] ; MA[ranhão], 1♂ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-47.308777&amp;materialsCitation.latitude=-7.118222" title="Search Plazi for locations around (long -47.308777/lat -7.118222)">Carolina</a>, PN <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-47.308777&amp;materialsCitation.latitude=-7.118222" title="Search Plazi for locations around (long -47.308777/lat -7.118222)">Chapada</a> das <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-47.308777&amp;materialsCitation.latitude=-7.118222" title="Search Plazi for locations around (long -47.308777/lat -7.118222)">Mesas</a>, Riacho Sucuruiu, 07°07'05.6''S 47°18'31.6''W, 240m, 14–30.VI.2013, Armadilha Malaise, JA Rafael, F Limeira-de-Oliveira, TTA Silva cols. (CZMA) .</p></div>	https://treatment.plazi.org/id/03D487B8C471CD4CFF22FBAB14D0FE6D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colombo, Wesley D.;Alencar, Isabel D. C. C.;Limeira-De-Oliveira, Francisco;Azevedo, Celso O.	Colombo, Wesley D., Alencar, Isabel D. C. C., Limeira-De-Oliveira, Francisco, Azevedo, Celso O. (2018): New species and records of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Cerrado, Caatinga and relicts of the Atlantic Forest from northeastern Brazil. Zootaxa 4462 (1): 1-40, DOI: 10.11646/zootaxa.4462.1.1
03D487B8C470CD4CFF22FE411549FB7F.text	03D487B8C470CD4CFF22FE411549FB7F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus verrucosus Redighieri & Azevedo. Eighteen 2004	<div><p>Dissomphalus verrucosus Redighieri &amp; Azevedo, 2004</p><p>Dissomphalus verrucosus Redighieri &amp; Azevedo, 2004: 330 (♂, holotype from Brazil).</p><p>Diagnosis. Male. Black. Mandible bidentate. Median clypeal lobe trapezoidal. Frons strongly coriaceous, very punctate. T2 with pair of shallow, elliptical lateral depressions, close to margin of T1, each depression with prominent flat-topped tubercle, pit on top, bearing tuft of setae inclined outward. Posterior hypopygeal margin straight. Genitalia: paramere with apex rounded, slightly arched inward, very wide, specially basally; ventral ramus smaller than dorsal body, laminar, surface horizontal, basal half wider, apex narrow, rounded, barely arched outward; dorsal body with two pairs apical lobes, outer pair wide, laminar, surface horizontal, apex rounded and arched upward, apical half of outer side with laminar projection directed upward with numerous warts in basal surface; inner pair stout, membranous and setae; apodeme extending beyond elliptical genital ring. Female unknown.</p><p>Variations. In this series the specimens are quite similar to those previously studied.</p><p>Remarks. This species was previously known from Brazil (Espírito Santo) (Redighieri &amp; Azevedo 2004) and Brazil (São Paulo) (Redighieri &amp; Azevedo 2006). Now it is recorded for the first time from Piauí and Maranhão.</p><p><a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-43.561695&amp;materialsCitation.latitude=-9.141055" title="Search Plazi for locations around (long -43.561695/lat -9.141055)">Material</a> examined. BRAZIL, PI[auí], 1♂ Guaribas, PN Serra das Confusões, Andorinha, 09°08'27.8''S 43°33'42.1''W, 515m, JA Rafael, F Limeira-de-Oliveira, TTA Silva cols. (CZMA) ; MA[ranhão], 2♂, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.86889&amp;materialsCitation.latitude=-6.6236115" title="Search Plazi for locations around (long -45.86889/lat -6.6236115)">Mirador</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.86889&amp;materialsCitation.latitude=-6.6236115" title="Search Plazi for locations around (long -45.86889/lat -6.6236115)">Parq</a> [ue] Estadual Mirador, Base da Geraldina, 06°37'25''S 45°52'08''W, 10–16.V.2013, Armadilha Malaise, LS Santos, F Limeira-de-Oliveira, LLM Santos cols. (CZMA) .</p></div>	https://treatment.plazi.org/id/03D487B8C470CD4CFF22FE411549FB7F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colombo, Wesley D.;Alencar, Isabel D. C. C.;Limeira-De-Oliveira, Francisco;Azevedo, Celso O.	Colombo, Wesley D., Alencar, Isabel D. C. C., Limeira-De-Oliveira, Francisco, Azevedo, Celso O. (2018): New species and records of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Cerrado, Caatinga and relicts of the Atlantic Forest from northeastern Brazil. Zootaxa 4462 (1): 1-40, DOI: 10.11646/zootaxa.4462.1.1
03D487B8C470CD4FFF22FAAF1447FE49.text	03D487B8C470CD4FFF22FAAF1447FE49.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus curupira Colombo & Alencar & Limeira-De-Oliveira & Azevedo 2018	<div><p>Dissomphalus curupira Colombo &amp; Azevedo sp. nov.</p><p>(Figs 12–16)</p><p>Description. Male. Head and mesosoma black; metasoma light castaneous. Mandible with three apical teeth; median clypeal lobe trapezoidal, median tooth weakly rounded; frons coriaceous, punctate; eyes weakly setose. Notauli complete; pronotal disc strongly coriaceous, weakly punctate. Metasomal tergite II with tergal process with non-conspicuous lateral depressions, tubercle present, pit on top, tuft of setae present. Posterior hypopygeal margin weakly concave. Genitalia: paramere with bifurcated rounded apex, very wide, higher than the basiparamere; aedeagal ventral ramus longer than dorsal body, apex rounded, laminar, small basal extension present, very narrow; aedeagal dorsal body with four pair of apical lobes, outer lobe narrow than others, apex rounded, slightly arched inward, median pair with apex rounded, slightly arched outward, ventral inner pair membranous, setose, wide, dorsal inner pair with apex bifurcated forming two teeth; basal process absent; apodeme extending beyond genital ring. Female unknown.</p><p>Remarks. This species belongs to conicus species-group by having the median clypeal lobe trapezoidal and metasomal tergite II with one pair of lateral depressions, with tubercle, with pit on the top and tuft of setae. This species is not comparable to any other in the species-group. It is mainly different because it has the paramere with a bifurcated rounded apex, the aedeagal ventral ramus with a basal small extension, and the aedeagal dorsal body with four apical lobes.</p><p><a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-47.29911&amp;materialsCitation.latitude=-7.1122775" title="Search Plazi for locations around (long -47.29911/lat -7.1122775)">Material</a> examined. Holotype ♂, BRAZIL, MA[ranhão], Carolina, PN Chapada das Mesas, Riacho Cancela, 07°06'44.2''S 47°17'56.8''W, 255m, 15–31.VII.2013, Armadilha Malaise, JA Rafael, F Limeira-de-Oliveira, TTA Silva cols. (CZMA) . Paratypes: MA[ranhão], 2♂, same data of holotype, except 01–10.VIII.2013, [Armadilha] suspensa lâmina d'água; 1♂ 14–30.VI.2013; 4♂ 01–15.VII.2013, 1♂ 10–20.IX.2013; 2♂ 14.VI–10.VIII.2013, Armadilha suspensa; 1♂ Mirador, Parq[ue] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.86889&amp;materialsCitation.latitude=-6.6236115" title="Search Plazi for locations around (long -45.86889/lat -6.6236115)">Estadual Mirador</a>, Base da Geraldina, 06°37'25''S 45°52'08''W, 14– 18.VIII.2012, Armadilha Malaise, JS Pinto Junior, DWA Marques, F Limeira-de-Oliveira cols. (CZMA) .</p><p>Etymology. The epithet curupira from Tupi refers to a Brazilian folk tale in which a boy uses his feet in reverse to confuse hunters in order to protect the trees and the animals. Treat as a noun in apposition.</p><p>Distribution. Brazil (Maranhão).</p></div>	https://treatment.plazi.org/id/03D487B8C470CD4FFF22FAAF1447FE49	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colombo, Wesley D.;Alencar, Isabel D. C. C.;Limeira-De-Oliveira, Francisco;Azevedo, Celso O.	Colombo, Wesley D., Alencar, Isabel D. C. C., Limeira-De-Oliveira, Francisco, Azevedo, Celso O. (2018): New species and records of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Cerrado, Caatinga and relicts of the Atlantic Forest from northeastern Brazil. Zootaxa 4462 (1): 1-40, DOI: 10.11646/zootaxa.4462.1.1
03D487B8C473CD4FFF22FE621477FACA.text	03D487B8C473CD4FFF22FE621477FACA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus iara Colombo & Alencar & Limeira-De-Oliveira & Azevedo 2018	<div><p>Dissomphalus iara Colombo &amp; Azevedo sp. nov.</p><p>(Figs 17–20)</p><p>Description. Male. Head and mesosoma black; metasoma dark castaneous. Mandible with two apical teeth; median clypeal lobe undefined, wide, median tooth weakly pointed; frons coriaceous, punctate; eyes weakly setose. Notauli complete; pronotal disc coriaceous. Metasomal tergite II with tergal process with weakly conspicuous lateral depressions, tubercle present, pit on top, tuft of setae present. Posterior hypopygeal margin straight. Genitalia: paramere with rounded apex, slightly arched inward, shorter than the basiparamere; aedeagal ventral ramus shorter than dorsal body, apex rounded, laminar, convergent; aedeagal dorsal body with two pair of apical lobes, outer pair wide, narrow, ventral margin serrated, inner pair membranous, basal process absent; apodeme extending beyond genital ring. Female unknown.</p><p>Remarks. This species belongs to conicus species-group by having metasomal tergite II with one pair of lateral depressions, with tubercle, with pit on the top and tuft of setae. This species is not similar to any other member of this species-group. It is mainly different because it has the paramere with a rounded apex, which is slightly arched inward.</p><p><a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-40.909725&amp;materialsCitation.latitude=-3.8369443" title="Search Plazi for locations around (long -40.909725/lat -3.8369443)">Material</a> examined. Holotype ♂, BRAZIL, CE[ará], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-40.909725&amp;materialsCitation.latitude=-3.8369443" title="Search Plazi for locations around (long -40.909725/lat -3.8369443)">Ubajara</a>, PN Ubajara, Cachoeira do Cafundó, 03°50'13''S, 40°54'35''W, 20–22.VII.2013, Armadilha Malaise, JA Rafael, F Limeira-de-Oliveira, TTA Silva cols. (CZMA) . Paratypes: 4♂ same data of holotype; 1♂ same data of holotype, except 01–14.II.2013; 4♂ 18.XI– 15.I.2013, JS Pinto Junior, F Limeira-de-Oliveira cols.; 2♂ 13–17.XI.2013, DWA Marques, F Limeira-de-Oliveira cols.; 2♂ 01–14.II.2013, F Limeira-de-Oliveira, JA Rafael cols. (CZMA).</p><p>Etymology. The epithet iara from Tupi refers to a Brazilian folk tale, in which a beautiful mermaid lives on the Amazon River, with brown skin, long green hair, and brown eyes. Treat as a noun in apposition.</p><p>Distribution. Brazil (Ceará).</p></div>	https://treatment.plazi.org/id/03D487B8C473CD4FFF22FE621477FACA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colombo, Wesley D.;Alencar, Isabel D. C. C.;Limeira-De-Oliveira, Francisco;Azevedo, Celso O.	Colombo, Wesley D., Alencar, Isabel D. C. C., Limeira-De-Oliveira, Francisco, Azevedo, Celso O. (2018): New species and records of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Cerrado, Caatinga and relicts of the Atlantic Forest from northeastern Brazil. Zootaxa 4462 (1): 1-40, DOI: 10.11646/zootaxa.4462.1.1
03D487B8C473CD4EFF22FAE01447FE11.text	03D487B8C473CD4EFF22FAE01447FE11.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus saci Colombo & Alencar & Limeira-De-Oliveira & Azevedo 2018	<div><p>Dissomphalus saci Colombo &amp; Azevedo sp. nov.</p><p>(Figs 21–24)</p><p>Description. Male. Head and mesosoma black; metasoma light castaneous. Mandible with three apical teeth; median clypeal lobe tridentate, median tooth pointed; frons strongly coriaceous; eyes glabrous. Notauli complete; pronotal disc strongly coriaceous. Metasomal tergite II with tergal process with non-conspicuous lateral depressions, tubercle present, pit on top, tuft of setae present. Posterior hypopygeal margin straight. Genitalia: paramere with serrated apex, slightly arched inward, wide, higher than the basiparamere; aedeagal ventral ramus as long as dorsal body, apex inverted, laminar, medium constriction; aedeagal dorsal body with two pairs of apical lobes, dorsal margin wide, apical and ventral margins weakly serrated, inner pair membranous, weakly setose, apex rounded in ventral view, basal process present, tooth-shaped; apodeme not extending beyond genital ring. Female unknown.</p><p>Remarks. This species belongs to conicus species-group by having metasomal tergite II with one pair of lateral depressions, with tubercle, with pit on top and tuft of setae. It is similar to D. umbilicus Azevedo by having the posterior hypopygeal margin straight and metasomal tergite II with tubercle pit-topped, tuft of setae. However, D. saci sp. nov. has the mandible tridentate, the median clypeal lobe tridentate, and the aedeagal ventral ramus as long as the dorsal body, whereas D. umbilicus has the mandible bidentate, the median clypeal lobe trapezoidal, and the aedeagal ventral ramus longer than the dorsal body.</p><p><a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.86889&amp;materialsCitation.latitude=-6.6236115" title="Search Plazi for locations around (long -45.86889/lat -6.6236115)">Material</a> examined. Holotype ♂, BRAZIL, MA[ranhão], Mirador, Parq[ue] Estadual Mirador, Base da Geraldina, 06°37'25''S 45°52'08''W, 14–18.VIII.2012, Armadilha Malaise, DWA Marques, F Limeira-de-Oliveira, JSP Junior cols. (CZMA) . Paratypes, 1♂ Carolina, PN Chapada das Mesas, Riacho Sucuruiu, 01–10.IX.2013, Arm [adilha] Malaise (CZMA) ; 1♂ 07°07'05.6''S 47°18'31.6''W, 10–20.X.2013, Arm[adilha] Malaise, JA Rafael, F Limeira-de-Oliveira, TTA Silva cols. (CZMA); 1♂ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-47.29911&amp;materialsCitation.latitude=-7.1122775" title="Search Plazi for locations around (long -47.29911/lat -7.1122775)">Riacho Cancela</a>, 07°06'44.2''S 47°17'56.8''W, 225m, 20– 31.VIII.2013, Armadilha Malaise, JA Rafael, F Limeira-de-Oliveira, TTA Silva cols. (CZMA) .</p><p>Etymology. The epithet saci from Tupi refers to a young black man of the Brazilian folklore with a single leg, who carries a carapace over his head that gives him magical powers. Treat as a noun in apposition.</p><p>Distribution. Brazil (Maranhão).</p></div>	https://treatment.plazi.org/id/03D487B8C473CD4EFF22FAE01447FE11	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colombo, Wesley D.;Alencar, Isabel D. C. C.;Limeira-De-Oliveira, Francisco;Azevedo, Celso O.	Colombo, Wesley D., Alencar, Isabel D. C. C., Limeira-De-Oliveira, Francisco, Azevedo, Celso O. (2018): New species and records of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Cerrado, Caatinga and relicts of the Atlantic Forest from northeastern Brazil. Zootaxa 4462 (1): 1-40, DOI: 10.11646/zootaxa.4462.1.1
03D487B8C472CD4EFF22FDD715C8F987.text	03D487B8C472CD4EFF22FDD715C8F987.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus gilvipes Evans 1979	<div><p>Dissomphalus gilvipes Evans, 1979</p><p>Dissomphalus gilvipes Evans, 1979: 278, 279, 283 (♂, holotype from Colombia).</p><p>Diagnosis. Male. Dark castaneous. Mandible with four apical teeth. Clypeus wholly broadly projected toward, small tooth, median carina slightly high. Frons coriaceous. Vertex straight. Notaulus and parapsidal furrow complete, former very convergent posteriorly. T2 with pair of subcircular lateral depressions, each one with linear tuft of setae, being lateral one especially longer than others. Posterior hypopygeal margin straight, with small median callus. Genitalia: paramere very wide, apex as conical projection; aedeagal ventral ramus as long as dorsal body, laminar, apex blunt, oblique with outer corner sharpened; dorsal body with two pairs apical lobes, outer pair narrowing apically in dorsal and lateral view; inner pair somewhat stout, membranous setae. Female unknown.</p><p>Variations. The main variation found in this series is the metasomal tergal process with the depressions closer each other.</p><p>Remarks. This species was previously known from Panama (Azevedo 2003), Colombia (Evans 1979), Ecuador, Peru (Azevedo 1999c), Brazil (Amazonas, Pará, Rondônia, Mato Grosso, Distrito Federal São Paulo and Paraná) (Azevedo 1999c), Brazil (Acre, Pernambuco, Minhas Gerais and Espírito Santo (Azevedo 2003), Brazil (Alagoas and Sergipe) (Redighieri &amp; Azevedo 2006) and Bolivia (Azevedo 1999c). Now it is recorded for the first time from Ceará and Maranhão.</p><p><a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-40.909725&amp;materialsCitation.latitude=-3.8369443" title="Search Plazi for locations around (long -40.909725/lat -3.8369443)">Material</a> examined. BRAZIL, CE[ará], 1♂ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-40.909725&amp;materialsCitation.latitude=-3.8369443" title="Search Plazi for locations around (long -40.909725/lat -3.8369443)">Ubajara</a>, PN Ubajara, Cachoeira do Cafundó, 03°50'13''S 40°54'35''W, 16–31.XII.2012, Armadilha suspensa, JS Pinto Junior, F Limeira-de-Oliveira cols. (CZMA) ; MA[ranhão], 1♂ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-47.299526&amp;materialsCitation.latitude=-7.1122775" title="Search Plazi for locations around (long -47.299526/lat -7.1122775)">Carolina</a>, PN <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-47.299526&amp;materialsCitation.latitude=-7.1122775" title="Search Plazi for locations around (long -47.299526/lat -7.1122775)">Chapada</a> das Mesas, Riacho Cancela, 07°06'44.2''S 47°17'58.3''W, 255m, 01– 15.VII.2013, Armadilha Malaise, JA Rafael, F Limeira-de-Oliveira, TTA Silva cols. (CZMA) ; 1♂ 20– 31.VIII.2013, 1♂ 1 0– 20.XI.2013, Suspensa lâmina d'água (CZMA); 1♂ 20–31.VIII. 2013,1♂ 07°06'44.2''S 47°17'56.8''W (CZMA); 1♂ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-47.308777&amp;materialsCitation.latitude=-7.118222" title="Search Plazi for locations around (long -47.308777/lat -7.118222)">Riacho Sucuruiu</a>, 07°07'05.6''S 47°18'31.6''W, 10–20.X.2013, Armadilha Malaise, JA Rafael, F Limeira-de-Oliveira, TTA Silva cols. (CZMA) . 1♂ Mirador, Parq[ue] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.846943&amp;materialsCitation.latitude=-6.5994444" title="Search Plazi for locations around (long -45.846943/lat -6.5994444)">Estadual Mirador</a>, Base da Geraldina, 06°35'58''S 45°50'49''W, 402m, 15–13.IX.2013, Armadilha Malaise, LS Santos, F Limeira-de-Oliveira, LLM Santos cols. (CZMA) ; 2♂ 06°37'06''S 45°51'51''W, 411m, 02–12.VIII.2013, Armadilha Malaise, F Limeirade-Oliveira, LLM Santos, TL Rocha cols. (CZMA); 1♂ 1–10.XI.2013.</p></div>	https://treatment.plazi.org/id/03D487B8C472CD4EFF22FDD715C8F987	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colombo, Wesley D.;Alencar, Isabel D. C. C.;Limeira-De-Oliveira, Francisco;Azevedo, Celso O.	Colombo, Wesley D., Alencar, Isabel D. C. C., Limeira-De-Oliveira, Francisco, Azevedo, Celso O. (2018): New species and records of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Cerrado, Caatinga and relicts of the Atlantic Forest from northeastern Brazil. Zootaxa 4462 (1): 1-40, DOI: 10.11646/zootaxa.4462.1.1
03D487B8C472CD51FF22F9271402FDD9.text	03D487B8C472CD51FF22F9271402FDD9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus krombeini Azevedo 1999	<div><p>Dissomphalus krombeini Azevedo, 1999</p><p>Dissomphalus krombeini Azevedo, 1999b: 52 –54 (♂, holotype from U.S.A.).</p><p>Diagnosis. Male. Dark castaneous; wings hyaline. Mandible with four apical teeth. Clypeus broadly projected forward, median tooth minute, median carina slightly high, angled in profile, lateral lobes undefined. Frons weakly coriaceous. Vertex slightly convex. Notaulus short, very weak. Parapsidal furrow incomplete anteriorly. T2 with pair of shallow subcircular lateral depressions, small tubercle present, giving rise to linear tuft of setae, lateral one distinctly longer than others. Hypopygium short, wide, posterior margin slightly broadly concave. Genitalia: paramere with wide apex, truncate, arched inward, corners somewhat sharp, base dorsal margin much projected in- and downward; cuspis long and much arched; aedeagal ventral ramus nearly as long as dorsal body, basal three fourths wide with inner margin parallel and straight and outer margin slightly convex, apical fourth narrow with apex truncate and inclined, outer corner sharp; dorsal body with pair of apical lobe, outer margin bulging, inner margin weakly setose, apex narrow, basal process with median spine. Female unknown.</p><p>Variations. In this series the specimens are quite similar to those previously studied.</p><p>Remarks. This species was previously known from U.S.A. (Azevedo 1999b), Dominican Republic, Saint Vincent, Dominica, Costa Rica, Ecuador (Azevedo 2003), Brazil (Distrito Federal, Espírito Santo, São Paulo, Paraná) (Azevedo 2003), Brazil (Santa Catarina) (Redighieri &amp; Azevedo 2006) and Argentina (Azevedo 2003). Now it is recorded for the first time from Maranhão.</p><p>Material examined. BRAZIL, MA[ranhão], 1♂, Carolina, PN Chapada das Mesas, Riacho Sucuruiu, 08.V.2014, Arm [adilha] Malaise, (CZMA).</p></div>	https://treatment.plazi.org/id/03D487B8C472CD51FF22F9271402FDD9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colombo, Wesley D.;Alencar, Isabel D. C. C.;Limeira-De-Oliveira, Francisco;Azevedo, Celso O.	Colombo, Wesley D., Alencar, Isabel D. C. C., Limeira-De-Oliveira, Francisco, Azevedo, Celso O. (2018): New species and records of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Cerrado, Caatinga and relicts of the Atlantic Forest from northeastern Brazil. Zootaxa 4462 (1): 1-40, DOI: 10.11646/zootaxa.4462.1.1
03D487B8C46DCD51FF22FDD21447F98E.text	03D487B8C46DCD51FF22FDD21447F98E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus irupe Colombo & Alencar & Limeira-De-Oliveira & Azevedo 2018	<div><p>Dissomphalus irupe Colombo &amp; Azevedo sp. nov.</p><p>(Figs 25–29)</p><p>Description. Male. Head and mesosoma black; metasoma dark castaneous. Mandible with four apical teeth; median clypeal lobe trapezoidal, broadly like duckbill, high in lateral view, median tooth rounded; frons coriaceous, punctate; eyes glabrous; antenna with moniliform flagellomeres; gena with one pair of spines. Notauli absent; pronotal disc weakly coriaceous; metasomal tergite II with tergal process with subrounded lateral depression, tuft of setae, most lateral one directed outward, distinctly longer than others. Posterior hypopygeal margin straight. Genitalia: paramere with rounded apex, strongly arched inward, very constricted, very wide basally, higher than the basiparamere, few, long bristles present; aedeagal ventral ramus shorter than dorsal body, apex narrow, laminar; aedeagal dorsal body with one pair of apical lobe, long, weakly curved dorsally, ventral margin weakly serrated, membranous; basal process absent; apodeme not extending beyond genital ring. Female unknown.</p><p>Remarks. This species belongs to the gilvipes species-group by having metasomal tergite II with a lateral tergal process, the tufts of this tergal process have most lateral setae distinctly longer than the others. It is not similar to other species of its species-group. This species is mainly different because it has the median clypeal lobe trapezoidal, broadly like a duckbill, the gena with one pair of spines, and the paramere with a rounded apex, that is strongly arched inward and very constricted.</p><p><a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.86889&amp;materialsCitation.latitude=-6.6236115" title="Search Plazi for locations around (long -45.86889/lat -6.6236115)">Material</a> examined. Holotype ♂, BRAZIL, MA[ranhão], Mirador, Parq[ue] Estadual Mirador, Base da Geraldina, 06°37'25''S 45°52'08''W, 11–16.I.2013, Armadilha Malaise, LS Santos, JT Câmara, LLM Santos cols. (CZMA) . Paratypes, 1♂ same data of holotype, except 06°37'48''S 45°52'49''W, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.880276&amp;materialsCitation.latitude=-6.63" title="Search Plazi for locations around (long -45.880276/lat -6.63)">Armadilha Suspensa</a>, 01– 10.XI.2013, F Limeira-de-Oliveira, LLM Santos, TL Rocha cols. (CZMA) ; 1♂ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-47.29911&amp;materialsCitation.latitude=-7.1122775" title="Search Plazi for locations around (long -47.29911/lat -7.1122775)">Carolina</a>, PN <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-47.29911&amp;materialsCitation.latitude=-7.1122775" title="Search Plazi for locations around (long -47.29911/lat -7.1122775)">Chapada</a> das Mesas, Riacho Sucuruiu, 07°06'44.2''S 47°17'56.8''W, 240m, 01–15.VII.2013, Armadilha Malaise, JA Rafael, F Limeirade-Oliveira, TTA Silva cols. (CZMA) ; 1♂ 225m, 01–10.VIII.2013, Suspensa lâmina d'água (CZMA).</p><p>Etymology. The epithet irupe from Guarani refers to a Brazilian indigenous myth, in which an indigenous woman becomes the aquatic plant victoria regia (common name) when she falls in the river. Treat as a noun in apposition.</p><p>Distribution. Brazil (Maranhão).</p></div>	https://treatment.plazi.org/id/03D487B8C46DCD51FF22FDD21447F98E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colombo, Wesley D.;Alencar, Isabel D. C. C.;Limeira-De-Oliveira, Francisco;Azevedo, Celso O.	Colombo, Wesley D., Alencar, Isabel D. C. C., Limeira-De-Oliveira, Francisco, Azevedo, Celso O. (2018): New species and records of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Cerrado, Caatinga and relicts of the Atlantic Forest from northeastern Brazil. Zootaxa 4462 (1): 1-40, DOI: 10.11646/zootaxa.4462.1.1
03D487B8C46DCD50FF22F9591744FDD9.text	03D487B8C46DCD50FF22F9591744FDD9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus latimerus Azevedo 2003	<div><p>Dissomphalus latimerus Azevedo, 2003</p><p>Dissomphalus latimerus Azevedo, 2003: 48 –49 (♂, holotype from Brazil).</p><p>Diagnosis. Male. Black. Forewing subhyaline. Mandible with three apical teeth. Median clypeal lobe trapezoidal. Frons somewhat strongly coriaceous. Vertex nearly straight. T2 with lateral drop-shaped tubercle, sharpened anteriorly, rounded behind, pit-topped, tuft of setae directed backward, surrounded by rounded depression. Posterior hypopygeal margin straight. Genitalia: paramere very wide, digitus with large basal projection; aedeagal ventral ramus smaller than dorsal body, laminar, surface subhorizontal, apex sharp, outer margin convex, inner margin straight, base apparently raising at middle of aedeagus; aedeagal dorsal body with two pairs apical lobes, outer pair laminar, surface convex, apex rounded, inner pair membranous, stout, setose; aedeagal base with many warts. Female unknown.</p><p>Variations. The main variations found in this series are the aedeagal dorsal body with wide and small warts.</p><p>Remarks. This species was previously known from Brazil (Amazonas) (Azevedo 2003). Now it is recorded for the first time from Maranhão.</p><p><a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.864166&amp;materialsCitation.latitude=-6.6183333" title="Search Plazi for locations around (long -45.864166/lat -6.6183333)">Material</a> examined. BRAZIL, MA[ranhão], 1♂ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.864166&amp;materialsCitation.latitude=-6.6183333" title="Search Plazi for locations around (long -45.864166/lat -6.6183333)">Mirador</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.864166&amp;materialsCitation.latitude=-6.6183333" title="Search Plazi for locations around (long -45.864166/lat -6.6183333)">Parq</a> [ue] Estadual Mirador, Base da Geraldina, 06°37'06''S 45°51'51''W, 411m, 20–31.XII.2013, Armadilha suspensa, TL Rocha, F Limeira-de-Oliveira, LLM Santos cols. (CZMA) ; 1♂ 01–12.VIII.2013, Armadilha Malaise, LS Santos, F Limeira-de-Oliveira, LLM Santos cols. (CZMA).</p></div>	https://treatment.plazi.org/id/03D487B8C46DCD50FF22F9591744FDD9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colombo, Wesley D.;Alencar, Isabel D. C. C.;Limeira-De-Oliveira, Francisco;Azevedo, Celso O.	Colombo, Wesley D., Alencar, Isabel D. C. C., Limeira-De-Oliveira, Francisco, Azevedo, Celso O. (2018): New species and records of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Cerrado, Caatinga and relicts of the Atlantic Forest from northeastern Brazil. Zootaxa 4462 (1): 1-40, DOI: 10.11646/zootaxa.4462.1.1
03D487B8C46CCD50FF22FD0F17F2F9E3.text	03D487B8C46CCD50FF22FD0F17F2F9E3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus linearis Azevedo 1999	<div><p>Dissomphalus linearis Azevedo, 1999</p><p>Dissomphalus linearis Azevedo, 1999a: 337 –338 (♂, holotype from Brazil).</p><p>Diagnosis. Male. Body dark castaneous, head and mesosoma darker. Mandible with two apical teeth. Median clypeal lobe tridentate. Frons coriaceous, punctures shallow, large. Frontal impression short. Vertex straight. Tergal process vertical, depression large, not so deep, width 0.4 × tergite width, small emargination in upper margin forming two convex lobes, their lateral margins parallel, dentiform process inward above level of tuft, depression broadly in touch posterior margin T1, two linear and horizontal lateral hairy tufts, slightly directed toward each other, far from each other by their lengths, T2 with additional pair of light spots. Hypopygeal slender stalk, widening apicad, posterior margin slightly concave. Genitalia: paramere wide, cuspis long, much arched, apex truncate; aedeagal ventral ramus smaller than dorsal body, convergent apicad, laminar, wide, inclined, surface horizontal, apex sharpened, inner margin somewhat concave medially; aedeagal dorsal body with two pairs apical lobes, outer pair laminar, vertical, outer surface convex, wide with rounded apex in lateral view; inner pair stout, hairy, membranous. Female unknown.</p><p>Variations. In this series the specimens are quite like those previously studied.</p><p>Remarks. This species was previously known from Brazil (Amazonas, Pará) (Azevedo 1999a). Now is recorded for the first time from Ceará, Piauí and Maranhão.</p><p><a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-40.909725&amp;materialsCitation.latitude=-3.8369443" title="Search Plazi for locations around (long -40.909725/lat -3.8369443)">Material</a> examined. BRAZIL, CE[ará], 1♂ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-40.909725&amp;materialsCitation.latitude=-3.8369443" title="Search Plazi for locations around (long -40.909725/lat -3.8369443)">Ubajara</a>, PN Ubajara, Cachoeira do Cafundó, 03°50'13''S 40°54'35''W, 16–31.XII.2012, Armadilha Malaise, JS Pinto Junior, F Limeira-de-Oliveira cols. (CZMA) ; 1♂ 01– 15.I.2013 (CZMA); PI[auí], 1♂ Caracol, PN Serra das Confusões, Riacho dos Bois, 2014, Armadilha suspensa (CZMA); 3♂ 09°13'11.9''S 43°29'26.2''W, 575m, 01.VIII–30.XI.2013, Armadilha Malaise, JA Rafael, F Limeirade-Oliveira, TTA Silva cols. (CZMA); MA[ranhão], 1♂ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.880276&amp;materialsCitation.latitude=-6.63" title="Search Plazi for locations around (long -45.880276/lat -6.63)">Mirador</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.880276&amp;materialsCitation.latitude=-6.63" title="Search Plazi for locations around (long -45.880276/lat -6.63)">Parque Estadual do Mirador</a>, Base da Geraldina, 416m, 06°37'48''S 45°52'49''W, Armadilha de Malaise, 20–30.XI.2013, F Limeira-de-Oliveira, LLM Santos, TL Rocha cols. (CZMA) .</p></div>	https://treatment.plazi.org/id/03D487B8C46CCD50FF22FD0F17F2F9E3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colombo, Wesley D.;Alencar, Isabel D. C. C.;Limeira-De-Oliveira, Francisco;Azevedo, Celso O.	Colombo, Wesley D., Alencar, Isabel D. C. C., Limeira-De-Oliveira, Francisco, Azevedo, Celso O. (2018): New species and records of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Cerrado, Caatinga and relicts of the Atlantic Forest from northeastern Brazil. Zootaxa 4462 (1): 1-40, DOI: 10.11646/zootaxa.4462.1.1
03D487B8C46CCD53FF22F90414AEFDD9.text	03D487B8C46CCD53FF22F90414AEFDD9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus scamatus Azevedo 1999	<div><p>Dissomphalus scamatus Azevedo, 1999</p><p>Dissomphalus scamatus Azevedo, 1999a: 343 (♂, holotype from Brazil).</p><p>Diagnosis. Male. Black. Mandible forming oblique cutting-edge, angulate medially. Median clypeal lobe very large, as long as half-length clypeus medially. Front strongly coriaceous. Vertex straight. T2 with shallow, elliptical median depression, posterior margin concave medially, pair of conspicuous, elliptical and inclined tubercles, far from each other their smallest diameters, densely covered by setae. Genitalia: paramere wide; aedeagal ventral ramus as long as dorsal body, apical area laminar, surface vertical, ventral margin with row of 5–6 small ventral teeth, median area of ventral surface micro-scaled; dorsal body of aedeagus narrowing apically, apex denticulate in lateral view; inner surface densely setose. Female unknown.</p><p>Variations. In this series the specimens are quite similar to those previously studied.</p><p>Remarks. This species was previously known from Brazil (Paraná) (Azevedo 1999a) and Brazil (Espírito Santo and São Paulo) (Redighieri &amp; Azevedo 2006). Now it is recorded for the first time from Ceará and Maranhão.</p><p><a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-40.909725&amp;materialsCitation.latitude=-3.8369443" title="Search Plazi for locations around (long -40.909725/lat -3.8369443)">Material</a> examined. BRAZIL, CE[ará], 3♂ Ubajara, PN Ubajara, Cachoeira do Cafundó, 03°50'13''S 40°54'35''W, 16–31.XII.2012, Armadilha Malaise, JS Pinto Junior, F Limeira-de-Oliveira cols. (CZMA) ; MA[ranhão], 1♂ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.86889&amp;materialsCitation.latitude=-6.6236115" title="Search Plazi for locations around (long -45.86889/lat -6.6236115)">Mirador</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.86889&amp;materialsCitation.latitude=-6.6236115" title="Search Plazi for locations around (long -45.86889/lat -6.6236115)">Parq</a> [ue] Estadual Mirador, Base da Geraldina, 06°37'25''S 45°52'08''W, 13–19.X.2012, Armadilha Malaise, LS Santos, F Limeira-de-Oliveira, LLM Santos cols. (CZMA) ; 1♂ 08–14.III.2013, LS Santos, F Limeira-de-Oliveira, LLM Santos cols. (CZMA).</p></div>	https://treatment.plazi.org/id/03D487B8C46CCD53FF22F90414AEFDD9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colombo, Wesley D.;Alencar, Isabel D. C. C.;Limeira-De-Oliveira, Francisco;Azevedo, Celso O.	Colombo, Wesley D., Alencar, Isabel D. C. C., Limeira-De-Oliveira, Francisco, Azevedo, Celso O. (2018): New species and records of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Cerrado, Caatinga and relicts of the Atlantic Forest from northeastern Brazil. Zootaxa 4462 (1): 1-40, DOI: 10.11646/zootaxa.4462.1.1
03D487B8C46FCD52FF22FD0F147BFD0D.text	03D487B8C46FCD52FF22FD0F147BFD0D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus microstictus Evans 1969	<div><p>Dissomphalus microstictus Evans, 1969</p><p>Dissomphalus microstictus Evans, 1969: 17 (♂, holotype from Argentina).</p><p>Diagnosis. Male. Mandible with two apical teeth. Median clypeal lobe trapezoidal. Frons strongly coriaceous, densely punctate, punctures shallow, large. Vertex straight, corner somewhat rounded. Ocelli large. Head developed posteriorly eye. Metapectal-propodeal complex with only median carina. T2 with two depressions shallow, large, subcircular, nearly reaching posterior margin of T1, each one with minute setae median tubercle, lateral area of depression with setae. Hypopygeal slender stalk, posterior margin straight, corner rounded. Genitalia: paramere very wide, dorsal margin very evenly developed, apex produced and sharpened; volsella setae, digitus smooth in upper margin; aedeagal ventral ramus extremely short, slightly higher than base of paramere, laminar, wide, surfaces horizontal, inner margin parallel, outer margin convergent, apex narrow and rounded, outer margin with an apical filament very long, nearly with same size of ramus, slightly arched, dorsal body large, laminar, surfaces vertical, base wider in dorsal view, body is very wide in lateral view, with rounded apex directed downward, ventral margin complex, serrated with rounded teeth, inner surface setae. Female unknown.</p><p>Variations. The main variations found in this series are the metasomal tergal process with the depression having various shapes, such as circular, oval, or semicircular.</p><p>Remarks. This species was previously known from Brazil (Amapá, Amazonas, Pará, Pernambuco, Distrito Federal, São Paulo, Paraná and Santa Catarina) (Azevedo 1999c), Brazil (Alagoas, Sergipe, Bahia, Espírito Santo and Rio de Janeiro) (Redighieri &amp; Azevedo 2006), Argentina (Evans 1969) and Bolivia (Azevedo 1999c). Now it is recorded for the first time from Piauí and Maranhão.</p><p>Material examined. BRAZIL, 3♂ [locality missing], JA Rafael, F Limeira-de-Oliveira, TTA Silva cols. (CZMA) ; PI[auí], 29♂ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-41.709446&amp;materialsCitation.latitude=-4.099167" title="Search Plazi for locations around (long -41.709446/lat -4.099167)">Piracuruca</a>, PN de Sete Cidades, Posto do ICMBio, 04°05'57''S 41°42'34''W, 12– 27.III.2013, Armadilha Malaise, F Limeira-de-Oliveira, TTA Silva cols. (CZMA) ; 1♂ 15–30.VI.2013; 17♂ 01– 10.III.2013, F Limeira-de-Oliveira, JS Pinto Júnior cols. (CZMA); 9♂ 15–28.II.2013; 1♂ 01–15.XII.2012, F Limeira-de-Oliveira, JS Pinto Júnior cols. (CZMA); 2♂ 01–14.II.2013, JA Rafael, F Limeira-de-Oliveira, TTA Silva cols. (CZMA); 3♂ 19–28.II.2013; 2♂ 01–10.III.2013, Armadilha Suspensa (CZMA); 3♂ 13–26.IV.2013; 3♂ Caracol, PN Serra das Confusões, 15–31.VII.2013, Armadilha Malaise, JA Rafael, F Limeira-de-Oliveira, TTA Silva cols. (CZMA) ; 1♂ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-43.501167&amp;materialsCitation.latitude=-9.211861" title="Search Plazi for locations around (long -43.501167/lat -9.211861)">Saco</a> das Cachorras, 09°12'42.7''S 43°30'04.2''W, 522m, Armadilha de Malaise, 01– 10.IX.2013, JA Rafael, F Limeira-de-Oliveira, TTA Silva cols. (CZMA) ; 2♂ 10–30.XI.2013, JA Rafael, F Limeira-de-Oliveira, TTA Silva cols. (CMZA); 7♂ 01–31.I.2014, JA Rafael, F Limeira-de-Oliveira, TL Rocha, S Pereira cols. (CZMA); 1♂ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-43.490612&amp;materialsCitation.latitude=-9.219972" title="Search Plazi for locations around (long -43.490612/lat -9.219972)">Riacho</a> dos Bois, 09°13'11.9''S 43°29'26.2''W, 575m, 10–20.X.2013, Armadilha Malaise, JA Rafael, F Limeira-de-Oliveira, TTA Silva cols. (CZMA) ; 1♂ 20–31.X.2013; 1♂ 09°13'11.9''S 43°29'26.2''W, 01–11.VIII.2013 (CZMA); 1♂ Guaribas, PN Serra das Confusões, 11–20.VIII.2013, Armadilha Malaise, JA Rafael, F Limeira-de-Oliveira, TTA Silva cols. (CZMA) ; 2♂ 09°08'27.8''S 43°33'42.1''W, 515m, 20– 31.X.2013, Armadilha Malaise, JA Rafael, F Limeira-de-Oliveira, TTA Silva cols. (CZMA); 1♂ 01–10.XI.2013; MA[ranhão], 1♂ Mirador, Parque Estadual Mirador, 18–25.III.2012, Armadilha Malaise, F Limeira-de-Oliveira, DWA Marques cols. (CZMA); 1♂ Mirador, Parq[ue] Estadual Mirador, Base da Geraldina, 22.II–01.III.2009, Armadilha luminosa, MB Aguiar-Neto, MJA Holanda cols. (CZMA) ; 2♂ 06°35'58''S 45°50'49''W, 402m, 10– 20.X.2013; 5♂ 06°48'25''S 45°52'08''W, 10–16.V.2013, Armadilha Malaise, LS Santos, F Limeira-de-Oliveira, LLM Santos cols. (CZMA); 1♂ 01–10.I.2014; 1♂ 13–19.X.2012; 2♂ 08–14.III.2013; 1♂ 06°37'06''S 45°51'51''W, 411m, 02–12.VIII.2013, LS Santos, F Limeira-de-Oliveira, LLM Santos cols. (CZMA); 1♂ 06°37'48''S 45°52'49''W, 416m, 20–31.VIII.2013 {date probably mistranscribed}, Armadilha Malaise, F Limeira-de-Oliveira, LLM Santos cols. (CZMA); 1♂ 10–20.XI.2013; 3♂ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.109444&amp;materialsCitation.latitude=-6.808056" title="Search Plazi for locations around (long -45.109444/lat -6.808056)">Base da Cágados</a>, 06°48'29''S 45°06'34''W, 27.IX–02.X.2011, Armadilha Malaise, DWA Marques, F Limeira-de-Oliveira cols. (CZMA) ; 1♂ 06°48'29''S 45°06'28''W, 26.XI– 03.XII.2001, (CZMA); 2♂ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-47.308777&amp;materialsCitation.latitude=-7.118222" title="Search Plazi for locations around (long -47.308777/lat -7.118222)">Carolina</a>, PN Chapada das Mesas, Riacho Sucuruiu, 07°07'05.6''S 47°18'31.6''W, 240m, 10–20.X.2013, Armadilha Malaise, JA Rafael, F Limeira-de-Oliveira, TTA Silva cols. (CZMA) ; 2♂ 01.IX– 10.X.2013; 1♂ 14–30.VI.2013; 1♂ 20–30.IX.2013; 1♂ 20–31.VIII.2013 (CZMA); 1♂ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-47.090332&amp;materialsCitation.latitude=-7.073389" title="Search Plazi for locations around (long -47.090332/lat -7.073389)">Riacho Corrente</a>, 07°04'24.2''S 47°05'25.2''W, 288m, 01–10.I.2014, Armadilha Malaise, JA Rafael, F Limeira-de-Oliveira, TTA Silva cols. (CZMA) ; 4♂ 20–31.VIII.2013; 1♂ 07°06'44.2''S 47°17'56.8''W, 225m, 2 0– 31.VIII.2013; 2♂ 07°04'24.2''S 47°05'42.2''W, 298m, 01–10.VIII.2013 (CZMA); 1♂ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-47.29911&amp;materialsCitation.latitude=-7.1122775" title="Search Plazi for locations around (long -47.29911/lat -7.1122775)">Riacho Cancela</a>, 07°06'44.2''S 47°17'56.8''W, 225m, 01–10.X.2013, Suspensa lâmina d´agua, JA Rafael, F Limeira-de-Oliveira, TTA Silva cols. (CZMA) ; 3♂ [data missing] (CZMA); 1♂ Riachão, Fazenda Altos, 18–22.VIII.2009, Armadilha Malaise, F Limeira-de-Oliveira, MB Aguiar Neto cols. (CZMA).</p></div>	https://treatment.plazi.org/id/03D487B8C46FCD52FF22FD0F147BFD0D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colombo, Wesley D.;Alencar, Isabel D. C. C.;Limeira-De-Oliveira, Francisco;Azevedo, Celso O.	Colombo, Wesley D., Alencar, Isabel D. C. C., Limeira-De-Oliveira, Francisco, Azevedo, Celso O. (2018): New species and records of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Cerrado, Caatinga and relicts of the Atlantic Forest from northeastern Brazil. Zootaxa 4462 (1): 1-40, DOI: 10.11646/zootaxa.4462.1.1
03D487B8C46ECD55FF22FCDB15C7FE11.text	03D487B8C46ECD55FF22FCDB15C7FE11.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus napo Evans 1979	<div><p>Dissomphalus napo Evans, 1979</p><p>Dissomphalus napo Evans, 1979: 282 (♂, holotype from Ecuador).</p><p>Diagnosis. Male. Mandible with two apical teeth. Median clypeal lobe trapezoidal. Frons strongly coriaceous, densely punctate, punctures shallow, large. Vertex straight, corner somewhat rounded. Ocelli large. Head developed posteriorly eye. Metapectal-propodeal complex with only median carina. T2 with two depressions shallow, large, subcircular, nearly reaching posterior margin of T1, each one with minute setae median tubercle, lateral area of depression with setae. Hypopygeal slender stalk, posterior margin straight, corner rounded. Genitalia: paramere very wide, dorsal margin very evenly developed, apex produced and sharpened; volsella setae, digitus smooth in upper margin; aedeagal ventral ramus extremely short, slightly higher than base of paramere, laminar, wide, surfaces horizontal, inner margin parallel, outer margin convergent, apex narrow and rounded, outer margin with an apical filament very long, nearly with same size of ramus, slightly arched, dorsal body large, laminar, surfaces vertical, base wider in dorsal view, body wide in lateral view, with rounded apex directed downward, ventral margin complex, serrated with rounded teeth, inner surface setae. Female unknown.</p><p>Variations. The main variations found in this series are the basivolsella with a small projection and the paramere very wide.</p><p>Remarks. This species was previously known from Ecuador (Evans 1979), Peru, Paraguay (Azevedo 1999c), Brazil (Amazonas) (Azevedo 1999c), Brazil (Acre and Mato Grosso), Brazil (Sergipe, Alagoas, Pernambuco and Paraíba) (Redighieri &amp; Azevedo 2006). Now it is recorded for the first time from Piauí and Maranhão.</p><p>Material examined. BRAZIL, 2♂ [locality missing], JA Rafael, F Limeira-de-Oliveira, TTA Silva cols. (CZMA); PI[auí], 3♂ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-43.561695&amp;materialsCitation.latitude=-9.141055" title="Search Plazi for locations around (long -43.561695/lat -9.141055)">Guaribas</a>, PN <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-43.561695&amp;materialsCitation.latitude=-9.141055" title="Search Plazi for locations around (long -43.561695/lat -9.141055)">Serra</a> das Confusões, Andorinha, 09°08'27.8''S 43°33'42.1''W, 515m, 10.XI– 31.XII.2013, Armadilha Malaise, JA Rafael, F Limeira-de-Oliveira, TTA Silva cols. (CZMA) ; 2♂ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-43.490612&amp;materialsCitation.latitude=-9.219972" title="Search Plazi for locations around (long -43.490612/lat -9.219972)">Riacho</a> dos Bois, 09°13'11.9''S 43°29'26.2''W, 575m, 20–30.XI.2013, Armadilha Malaise, JA Rafael, F Limeira-de-Oliveira, TTA Silva cols. (CZMA) ; 1♂ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-43.490612&amp;materialsCitation.latitude=-9.219972" title="Search Plazi for locations around (long -43.490612/lat -9.219972)">Caracol</a>, PN Serra das Confusões, Riacho dos Bois, 09°13'11.9''S 43°29'26.2''W, 575m, JA Rafael, F Limeira-de-Oliveira, TTA Silva cols. (CZMA) ; 1♂ 01–10.XI.2013; MA[ranhão], 3♂ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-47.308777&amp;materialsCitation.latitude=-7.118222" title="Search Plazi for locations around (long -47.308777/lat -7.118222)">Carolina</a>, PN <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-47.308777&amp;materialsCitation.latitude=-7.118222" title="Search Plazi for locations around (long -47.308777/lat -7.118222)">Chapada</a> das Mesas, Riacho Sucuruiu, 07°07'05.6''S 47°18'31.6''W, 240m, 01.X–20.IX.2013, Armadilha Malaise, JA Rafael, F Limeira-de-Oliveira, TTA Silva cols. (CZMA) ; 3♂ 9–14.II.2014; 1♂ 07°06'44.2''S 47°17'56.8''W, 15–31.VII.2013; 1♂ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-47.35583&amp;materialsCitation.latitude=-7.116611" title="Search Plazi for locations around (long -47.35583/lat -7.116611)">Riacho Estiva</a>, 07°06'59.8''S 47°21'21.0''W, 265m, Armadilha Suspensa, 20– 31.X.2013, JA Rafael, F Limeira-de-Oliveira, TTA Silva cols. (CZMA) ; 1♂ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-47.29911&amp;materialsCitation.latitude=-7.1122775" title="Search Plazi for locations around (long -47.29911/lat -7.1122775)">Riacho Cancela</a>, 07°06'44.2''S 47°17'56.8''W, 255m, 01–10.VIII.2013, Armadilha suspensa, JA Rafael, F Limeira-de-Oliveira, TTA Silva cols. (CZMA) ; 1♂ 225m, 01–15.VII.2013; 1♂ Pov [oado] Canto Grande, Balneário Urupuxete, 28–31.X.2008, Armadilha suspensa, F Limeira-de-Oliveira, PAM Moraes Silva cols. (CZMA) ; 1♂ Caxias, Res[erva] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-43.425003&amp;materialsCitation.latitude=-4.9119444" title="Search Plazi for locations around (long -43.425003/lat -4.9119444)">Ecol</a> [ógica] Inhamum, Povoado Coités, 04°54'43''S 43°25'30''W, 27–29.XII.2012, Armadilha Malaise, F Limeira-de-Oliveira, LS Santos cols. (CZMA) ; 1♂ Mirador, Parque Estadual Mirador, Armadilha de Malaise, 01–10.XII.2013 ; 3♂ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.86889&amp;materialsCitation.latitude=-6.6236115" title="Search Plazi for locations around (long -45.86889/lat -6.6236115)">Base da Geraldina</a>, 06°37'25''S 45°52'08''W, 416m, 08.III–20.X.2013, Armadilha Malaise, TL Rocha, F Limeira-de- Oliveira, LLM Santos cols. (CZMA) ; 1♂ 06°35'58''S 45°50'49''W, 402m, F Limeira-de-Oliveira, LLM Santos, TL Rocha cols. (CZMA); 1♂ 06°37'48''S 45°52'46''W, 428m, 20–31.VIII.2013, F Limeira-de-Oliveira, LLM Santos, LS Santos cols. (CZMA); 8♂ 06°37'48''S 45°52'49''W, 416m, 20.VIII–10.XII.2013, Armadilha Malaise, TL Rocha, F Limeira-de-Oliveira, LLM Santos cols. (CZMA); 1♂ 20–26.II.2014; 1♂ 22.III.2009, Arm [adilha] luminosa, MB Aguiar-Neto, MJA Holanda cols. (CZMA); 2♂ 06°37'06''S 45°51'51''W, 411m, 01–12.VIII.2013, Armadilha Malaise, LS Santos, F Limeira-de-Oliveira, LLM Santos cols. (CZMA).</p></div>	https://treatment.plazi.org/id/03D487B8C46ECD55FF22FCDB15C7FE11	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colombo, Wesley D.;Alencar, Isabel D. C. C.;Limeira-De-Oliveira, Francisco;Azevedo, Celso O.	Colombo, Wesley D., Alencar, Isabel D. C. C., Limeira-De-Oliveira, Francisco, Azevedo, Celso O. (2018): New species and records of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Cerrado, Caatinga and relicts of the Atlantic Forest from northeastern Brazil. Zootaxa 4462 (1): 1-40, DOI: 10.11646/zootaxa.4462.1.1
03D487B8C469CD55FF22FD9A1447F956.text	03D487B8C469CD55FF22FD9A1447F956.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus acutipupu Colombo & Alencar & Limeira-De-Oliveira & Azevedo 2018	<div><p>Dissomphalus acutipupu Colombo &amp; Azevedo sp. nov.</p><p>(Figs 30–33)</p><p>Description. Male. Head, mesosoma and metasoma black. Mandible with three apical teeth; median clypeal lobe tridentate, median tooth pointed; frons strongly coriaceous, punctate; eyes glabrous. Notauli complete; pronotal disc strongly coriaceous, punctate. Tergal process with circular depression, margins not angulate, dense tufts setae directed backward, tubercle absent. Posterior hypopygeal margin weakly concave. Genitalia: paramere with rounded apex, arched inward, constricted, very wide, specially basally, higher than the basiparamere, bifurcated in ventral view; basivolsella with projection, wide; digitus wide; volsella sickle-shaped; aedeagal ventral ramus longer than dorsal body, apex strongly narrow, laminar, medium rounded expansion; aedeagal dorsal body with two pairs of apical lobes, outer lobe with dorsal margin folded across its length, apical margin weakly serrated, ventral margin weakly serrated, inner projection present, inner lobe membranous; basal process absent; apodeme extending beyond genital ring. Female unknown.</p><p>Remarks. This species belongs to the punctatus species-group by having the tergal process with tufts of setae directed backward and the tubercles absent. It is similar to Dissomphalus jurupari sp. nov., by having the median clypeal lobe tridentate, the paramere very wide, especially basally, higher than the basiparamere, and the basal process absent. However, D. acutipupu sp. nov., has the mandible tridentate and the posterior hypopygeal margin weakly concave, whereas D. jurupari sp. nov. has the mandible bidentate and the posterior hypopygeal margin straight.</p><p><a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.864166&amp;materialsCitation.latitude=-6.6183333" title="Search Plazi for locations around (long -45.864166/lat -6.6183333)">Material</a> examined. Holotype ♂, BRAZIL, MA[ranhão], Mirador, Parq[ue] Estadual Mirador, Base da Geraldina, 06°37'06''S 45°51'51''W, 411m, 02–12.VIII.2013, Armadilha Malaise, F Limeira-de-Oliveira, LLM Santos, TL Rocha cols. (CZMA) . Paratypes, MA [ranhão], same data of holotype, except: 1♂ 07.XI.2013; 1♂ 06°35'58''S 45°50'49''W, 402m, F <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.846943&amp;materialsCitation.latitude=-6.5994444" title="Search Plazi for locations around (long -45.846943/lat -6.5994444)">Limeira-de-Oliveira</a>, LLM Santos, LS Santos cols. (CZMA) ; 3♂ 10–31.X.2013; 1♂ 06°37'58''S 45°52'49''W, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.880276&amp;materialsCitation.latitude=-6.6327777" title="Search Plazi for locations around (long -45.880276/lat -6.6327777)">Armadilha Suspensa</a>, 20–28.II.2014 ; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-47.29911&amp;materialsCitation.latitude=-7.1122775" title="Search Plazi for locations around (long -47.29911/lat -7.1122775)">Carolina</a>, PN <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-47.29911&amp;materialsCitation.latitude=-7.1122775" title="Search Plazi for locations around (long -47.29911/lat -7.1122775)">Chapada</a> das Mesas, Riacho Cancela, 07°06'44.2''S 47°17'56.8''W, 225m, Suspensa lâmina d'água, JA Rafael, F Limeira-de-Oliveira, TTA Silva cols., 1♂ 15–31.VII.2013, 1♂ 01–10.XI.2013 (CZMA) ; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-47.308777&amp;materialsCitation.latitude=-7.118222" title="Search Plazi for locations around (long -47.308777/lat -7.118222)">Riacho Sucuruiu</a>, 07°07'05.6''S 47°18'31.6''W, 240m, Armadilha de Malaise, JA Rafael, F Limeira-de-Oliveira, TTA Silva cols. (CZMA) ; 1♂ 10–20.X.2013, 1♂ 20– 31.VIII.2013 (CZMA); 1♂ Caxias, Res[erva] Ecol [ógica] Inhamum, 09–12.I.2006, Armadilha Malaise, GA Cunha cols. (CZMA) .</p><p>Etymology. The epithet acutipupu from Tupi refers to a woman-man (hermaphrodite) from Brazilian folklore. Treat as a noun in apposition.</p><p>Distribution. Brazil (Maranhão).</p></div>	https://treatment.plazi.org/id/03D487B8C469CD55FF22FD9A1447F956	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colombo, Wesley D.;Alencar, Isabel D. C. C.;Limeira-De-Oliveira, Francisco;Azevedo, Celso O.	Colombo, Wesley D., Alencar, Isabel D. C. C., Limeira-De-Oliveira, Francisco, Azevedo, Celso O. (2018): New species and records of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Cerrado, Caatinga and relicts of the Atlantic Forest from northeastern Brazil. Zootaxa 4462 (1): 1-40, DOI: 10.11646/zootaxa.4462.1.1
03D487B8C469CD54FF22F9541484FB61.text	03D487B8C469CD54FF22F9541484FB61.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus cuca Colombo & Alencar & Limeira-De-Oliveira & Azevedo 2018	<div><p>Dissomphalus cuca Colombo &amp; Azevedo sp. nov.</p><p>(Figs 34–37)</p><p>Description. Male. Head and mesosoma black; metasoma dark castaneous. Mandible with three apical teeth; median clypeal lobe tridentate, median tooth rounded; frons strongly coriaceous, punctate; eyes glabrous. Notauli complete; pronotal disc strongly coriaceous, punctate. Tergal process with non-conspicuous depressions, dense tufts few setae weakly directed backward, tubercle absent. Posterior hypopygeal margin straight. Genitalia: paramere with rounded apex, slightly arched inward, higher than the basiparamere; aedeagal ventral ramus as long as dorsal body, apex very narrow, laminar, constricted; aedeagal dorsal body with two pairs of apical lobes, outer lobe very long, wide, curved ventrally, inner pair membranous, basal process absent; apodeme not extending beyond genital ring. Female unknown.</p><p>Remarks. This species belongs to the punctatus species-group by having the tergal process with its tufts of setae weakly directed backward and the tubercles absent. It is important to note, however, that the tergal process is not the standard found in punctatus species-group. Nevertheless, it fits into the definition of the group, even with fewer bristles. This species is not similar to any other species of its species-group, differing mainly by having the tergal process with few bristles, the aedeagal ventral ramus with the apex very narrow, and the aedeagal dorsal body with the outer lobe very long.</p><p><a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-47.308777&amp;materialsCitation.latitude=-7.118222" title="Search Plazi for locations around (long -47.308777/lat -7.118222)">Material</a> examined. Holotype ♂, BRAZIL, MA[ranhão], Carolina, PN Chapada das Mesas, Riacho Sucuruiu, 07°07'05.6''S 47°18'31.6''W, 10–20.X.2013, Armadilha Malaise, JA Rafael, F Limeira-de-Oliveira, TTA Silva cols. (CZMA) . Paratypes, MA[ranhão], Carolina, PN Chapada das Mesas, 1♂ 31.VIII.2013, Armadilha suspensa, F Limeira-de-Oliveira, DWA Marques cols. (CZMA) ; 1♂ same data of holotype, except: 01–10.VIII.2013; 2♂ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-47.29911&amp;materialsCitation.latitude=-7.1122775" title="Search Plazi for locations around (long -47.29911/lat -7.1122775)">Riacho Cancela</a>, 07°06'44.2''S 47°17'56.8''W, 225m, 15.VII–10.X.2013, Armadilha Malaise, JA Rafael, F Limeirade-Oliveira, TTA Silva cols. (CZMA) ; Mirador, Parque Estadual do Mirador, 1♂ 25.III.2012, Armadilha Malaise, F Limeira-de-Oliveira, DWA Marques cols. (CZMA) ; 1♂ Mirador, Parque Estadual Mirador, Base da Geraldina, Armadilha Malaise, 01–10.XII.2013 ; 1♂ 07.XI.2013, F Limeira-de-Oliveira, LLM Santos, TL Rocha cols. (CZMA); 2♂ 06°37'25''S 45°52'08''W, 13–19.X.2012, Armadilha Malaise, LS Santos, F Limeira-de-Oliveira, LLM Santos cols. (CZMA); 1♂ 06°35'58''S,45°50'49''W, 402m, 02–12.VIII.2013, Armadilha Malaise, LS Santos, F Limeira-de-Oliveira, LLM Santos cols. (CZMA); 1♂ 06°37'06''S 45°51'51''W, 411m, F Limeira-de-Oliveira, LLM Santos, TL Rocha cols. (CZMA); 1♂ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.109444&amp;materialsCitation.latitude=-6.808056" title="Search Plazi for locations around (long -45.109444/lat -6.808056)">Base da Cágados</a>, 06°48'29''S 45°06'34''W, 29.IX–02.X.2011, Armadilha Malaise, DWA Marques, F Limeira-de-Oliveira cols. (CZMA) ; CE[ará], 1♂ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-40.909725&amp;materialsCitation.latitude=-3.8369443" title="Search Plazi for locations around (long -40.909725/lat -3.8369443)">Ubajara</a>, PN Ubajara, Cachoeira do Cafundó, 03°50'13''S 40°54'35''W, 01–14.II.2013, Armadilha Malaise, F Limeira-de-Oliveira, JA Rafael cols. (CZMA) ; PI[auí], 1♂ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-43.561695&amp;materialsCitation.latitude=-9.141055" title="Search Plazi for locations around (long -43.561695/lat -9.141055)">Guaribas</a>, PN <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-43.561695&amp;materialsCitation.latitude=-9.141055" title="Search Plazi for locations around (long -43.561695/lat -9.141055)">Serra</a> das Confusões, Andorinha, 09°08'27.8''S 43°33'42.1''W, 515m, 01– 10.XI.2013, Armadilha Malaise, JA Rafael, F Limeira-de-Oliveira, TTA Silva cols. (CZMA) .</p><p>Etymology. The epithet cuca from Tupi refers to a Brazilian folk tale in which an old woman with an alligator head scares the children on their parents. Treat as a noun in apposition.</p><p>Distribution. Brazil (Ceará, Piauí and Maranhão).</p></div>	https://treatment.plazi.org/id/03D487B8C469CD54FF22F9541484FB61	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colombo, Wesley D.;Alencar, Isabel D. C. C.;Limeira-De-Oliveira, Francisco;Azevedo, Celso O.	Colombo, Wesley D., Alencar, Isabel D. C. C., Limeira-De-Oliveira, Francisco, Azevedo, Celso O. (2018): New species and records of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Cerrado, Caatinga and relicts of the Atlantic Forest from northeastern Brazil. Zootaxa 4462 (1): 1-40, DOI: 10.11646/zootaxa.4462.1.1
03D487B8C468CD57FF22FB581447FF39.text	03D487B8C468CD57FF22FB581447FF39.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus jurupari Colombo & Alencar & Limeira-De-Oliveira & Azevedo 2018	<div><p>Dissomphalus jurupari Colombo &amp; Azevedo sp. nov.</p><p>(Figs 38–41)</p><p>Description. Male. Head and mesosoma black; metasoma dark castaneous. Mandible with two apical teeth; median clypeal lobe tridentate, median tooth rounded; frons coriaceous, punctate; eyes glabrous. Notauli complete; pronotal disc coriaceous. Tergal process with non-conspicuous depressions, margins not angulate, dense tufts few setae weakly directed backward, tubercle absent. Posterior hypopygeal margin straight. Genitalia: paramere with straight apex, arched inward, wide basally, higher than the basiparamere, large bristles present in ventral view; aedeagal ventral ramus shorter than dorsal body, apex very narrow, laminar, translucid; aedeagal dorsal body with three pair of apical lobes, outer pair long, curved ventrally, dorsal pair smaller than outer pair, curved ventrally, inner pair membranous, basal process absent; apodeme extending beyond genital ring. Female unknown.</p><p>Remarks. This species belongs to the punctatus species-group by having the tergal process with weak tufts of setae directed backward, and the tubercles absent. It is important to note that the tergal process is not the standard found in punctatus species-group. However, it fits into the definition of the group, even with few bristles. This species is similar to Dissomphalus acutipupu sp. nov., by having the median clypeal lobe tridentate, the paramere very wide, especially basally, higher than the basiparamere, the basal process absent. However, D. jurupari sp. nov. has the mandible bidentate and the posterior hypopygeal margin straight, whereas D. acutipupu sp. nov., has the mandible tridentate and the posterior hypopygeal margin weakly concave.</p><p>Material examined. Holotype ♂, BRAZIL, MA[ranhão], Carolina, PN Chapada das Mesas, Riacho Cancela, 07°06'44.2''S 47°17'58.3''W, 255m, 01–15.VII.2013, Armadilha Malaise, JA Rafael, F Limeira-de-Oliveira, TTA Silva cols. (CZMA). Paratypes: 2♂ same data of holotype, except: 01.VI–10.VIII.2013, Armadilha Suspensa (CZMA).</p><p>Etymology. The epithet jurupari from Tupi refers to a character in Brazilian folklore known as the messenger of the sun on Earth. Treat as a noun in apposition. Distribution. Brazil (Maranhão).</p></div>	https://treatment.plazi.org/id/03D487B8C468CD57FF22FB581447FF39	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colombo, Wesley D.;Alencar, Isabel D. C. C.;Limeira-De-Oliveira, Francisco;Azevedo, Celso O.	Colombo, Wesley D., Alencar, Isabel D. C. C., Limeira-De-Oliveira, Francisco, Azevedo, Celso O. (2018): New species and records of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Cerrado, Caatinga and relicts of the Atlantic Forest from northeastern Brazil. Zootaxa 4462 (1): 1-40, DOI: 10.11646/zootaxa.4462.1.1
03D487B8C46BCD57FF22FE3A15A2FAFD.text	03D487B8C46BCD57FF22FE3A15A2FAFD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus infissus Evans 1969	<div><p>Dissomphalus infissus Evans, 1969</p><p>Dissomphalus infissus Evans, 1969: 14, 19, 22 (♂, holotype from Argentina).</p><p>Diagnosis. Male. Mandible with three apical teeth. Median clypeal lobe trapezoidal. Frons coriaceous, punctures small, shallow. Ocellar triangle distant from vertex. Vertex straight, small median depression, corner rounded. T2 with pair of very shallow and circular depressions, not reaching anterior margin, each depression with setae linear tubercle, inclined, posterior part directed inward, latero-anterior area depression sparse-setae. Hypopygium with slender stalk, posterior margin straight, corner rounded. Genitalia: paramere short or wide, dorsal margin evenly very developed, apex concave, very produced dorsally; volsella with stout protuberance near digitus, base volsella with species-group of teeth; aedeagal ventral ramus very short, base wide, apical half narrow, slightly convergent, apex very sharpened, inner margin of base sinuous, outer margin expanded laterally, so that ramus surrounds dorsal body laterally, this margin with small tooth; aedeagal dorsal body very wide with apex in lateral view, laminar, lamina completely folded longitudinally in ventral side, inner lamina membranous, sinuous, setae inside, outer lamina with series of grooves; in dorsal view body is slightly wider in base; inner ventral margin base of genitalia angulate. Female unknown.</p><p>Variations. In this series the specimens are quite similar to those previously studied.</p><p>Remarks. This species was previously known from Brazil (São Paulo and Paraná) (Azevedo 1999c), Brazil (Espírito Santo) (Azevedo 2003), Brazil (Paraíba, Pernambuco, Alagoas and Santa Catarina) (Redighieri &amp; Azevedo 2006) and Argentina (Evans 1969). Now it is recorded for the first time from Piauí and Maranhão.</p><p>Material examined. BRAZIL, PI[auí], 1♂ Caracol, PN Serra das Confusões, 20–30.IX.2013, Armadilha Malaise, JA Rafael, F Limeira-de-Oliveira, TTA Silva cols. (CZMA) ; 1♂ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-43.561695&amp;materialsCitation.latitude=-9.141055" title="Search Plazi for locations around (long -43.561695/lat -9.141055)">Andorinha</a>, 09°08'27.8''S 43°33'42.1''W, 515m, (CZMA) ; 1♂ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-43.490612&amp;materialsCitation.latitude=-9.219972" title="Search Plazi for locations around (long -43.490612/lat -9.219972)">Riacho</a> dos Bois, 09°13'11.9''S 43°29'26.2''W, 575m, 01–11.VIII.2013 ; MA[ranhão], 1♂ Mirador, Parq[ue] Estadual Mirador, Base da Geraldina, 22–26.VIII.2006, Arm [adilha] Malaise, F L [imeira-de-]O[liveira] col. (CZMA) ; 1♂ 06°37'48''S 45°52'49''W, 416m, 10–20.X.2013, Arm[adilha] Malaise, TL Rocha, F Limeira-de-Oliveira, LLM Santos cols. (CZMA); 1♂ 20–30.XI.2013; 1♂ 06°37'58''S 45°50'49''W, 402m, 01– 11.VIII.2013, JA Rafael, F Limeira-de-Oliveira, TTA Silva cols. (CZMA).</p></div>	https://treatment.plazi.org/id/03D487B8C46BCD57FF22FE3A15A2FAFD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colombo, Wesley D.;Alencar, Isabel D. C. C.;Limeira-De-Oliveira, Francisco;Azevedo, Celso O.	Colombo, Wesley D., Alencar, Isabel D. C. C., Limeira-De-Oliveira, Francisco, Azevedo, Celso O. (2018): New species and records of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Cerrado, Caatinga and relicts of the Atlantic Forest from northeastern Brazil. Zootaxa 4462 (1): 1-40, DOI: 10.11646/zootaxa.4462.1.1
03D487B8C46BCD56FF22FA791528FC9D.text	03D487B8C46BCD56FF22FA791528FC9D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus magnus Redighieri & Azevedo 2006	<div><p>Dissomphalus magnus Redighieri &amp; Azevedo, 2006</p><p>Dissomphalus magnus Redighieri &amp; Azevedo, 2006: 324 –326 (♂, holotype from Brazil).</p><p>Diagnosis. Male. Mandible with three apical teeth. Median clypeal lobe trapezoidal. Frons coriaceous, punctures small, shallow. Ocellar triangle distant from vertex. Vertex straight, small median depression, corner rounded. T2 with pair of very shallow and circular depressions, not reaching anterior margin, each depression with setae linear tubercle, inclined, posterior part directed inward, latero-anterior area depression sparse-setae. Hypopygium with slender stalk, posterior margin straight, corner rounded. Genitalia: paramere short or wide, dorsal margin evenly very developed, apex concave, very produced dorsally; volsella with stout protuberance near digitus, base volsella with species-group of teeth; aedeagal ventral ramus very short, base wide, apical half narrow, slightly convergent, apex very sharpened, inner margin of base sinuous, outer margin expanded laterally, so that ramus surrounds dorsal body laterally, this margin with small tooth; aedeagal dorsal body very wide with apex in lateral view, laminar, lamina completely folded longitudinally in ventral side, inner lamina membranous, sinuous, setae inside, outer lamina with series of grooves; in dorsal view body is slightly wider in base; inner ventral margin base of genitalia angulate. Female unknown.</p><p>Variations. The main variations found in this series are the pronotal disc strongly setose and the aedeagal ventral ramus strongly bifurcated.</p><p>Remarks. This species was previously known from Brazil (Paraíba) (Redighieri &amp; Azevedo 2006). Now it is recorded for the first time from Ceará and Maranhão.</p><p>This species has the tergal process with the lateral depression with one pair of tubercles and the lateral margin setose. Such characteristics are diagnostic of the setosus species-group. Thus, we transfer it into this species-group.</p><p><a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-40.909725&amp;materialsCitation.latitude=-3.8369443" title="Search Plazi for locations around (long -40.909725/lat -3.8369443)">Material</a> examined. BRAZIL, CE[ará], 1♂ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-40.909725&amp;materialsCitation.latitude=-3.8369443" title="Search Plazi for locations around (long -40.909725/lat -3.8369443)">Ubajara</a>, PN Ubajara, Cachoeira do Cafundó, 03°50'13''S 40°54'35''W, 16–31.XII.2012, Armadilha Malaise, F Limeira-de-Oliveira, JS Pinto Júnior cols. (CZMA) ; 1♂ 13– 17.XI.2012, DWA Marques, F Limeira-de-Oliveira cols. (CZMA); 2♂ 15–28.II.2013, Armadilha suspensa, JS Pinto Júnior, F Limeira-de-Oliveira cols. (CZMA); 7♂ 01.XII–15.I.2013, Armadilha Malaise; MA[ranhão], 4♂ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-47.29911&amp;materialsCitation.latitude=-7.1122775" title="Search Plazi for locations around (long -47.29911/lat -7.1122775)">Carolina</a>, PN <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-47.29911&amp;materialsCitation.latitude=-7.1122775" title="Search Plazi for locations around (long -47.29911/lat -7.1122775)">Chapada</a> das Mesas, Riacho Cancela, 07°06'44.2''S 47°17'56.8''W, 255m, 01.VII–10.VIII.2013, Armadilha Malaise, JA Rafael, F Limeira-de-Oliveira, TTA Silva cols. (CZMA) ; 1♂ 07°06'44.2''S 47°17'58.3''W, 225m, 10–20.IX.2013, (CZMA); 2♂ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-47.308777&amp;materialsCitation.latitude=-7.118222" title="Search Plazi for locations around (long -47.308777/lat -7.118222)">Riacho Sucuruiu</a>, 07°07'05.6''S 47°18'31.6''W, 240m, 15.VII–10.IX.2013, Armadilha Malaise, JA Rafael, F Limeira-de-Oliveira, TTA Silva cols. (CZMA) ; 1♂ Mirador, Parq[ue] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.864166&amp;materialsCitation.latitude=-6.6183333" title="Search Plazi for locations around (long -45.864166/lat -6.6183333)">Estadual Mirador</a>, Base da Geraldina, 06°37'06''S 45°51'51''W, 411m, 02–12.VIII.2013, Armadilha Malaise, F Limeira-de- Oliveira LLM Santos, TL Rocha cols. (CZMA) ; 1♂ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.00611&amp;materialsCitation.latitude=-6.73" title="Search Plazi for locations around (long -45.00611/lat -6.73)">Posto Avançado do Mel</a>, 06°43'48''S 45°00'22''W, 18– 25.III.2012, Armadilha suspensa, DWA Marques, F Limeira-de-Oliveira cols. (CZMA) ; 6♂ 06°37'48''S 45°52'49''W, 416m, 01–10.XII.2013, Armadilha Malaise, F Limeira-de-Oliveira, LLM Santos, LS Santos cols. (CZMA); 1♂ 06°35'58''S 45°50'49''W, 402m, 20–31.XII.2013, Armadilha Malaise, F Limeira-de-Oliveira, LLM Santos, TL Rocha cols. (CZMA); 1♂ 10–20.X.2013 (CZMA).</p></div>	https://treatment.plazi.org/id/03D487B8C46BCD56FF22FA791528FC9D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colombo, Wesley D.;Alencar, Isabel D. C. C.;Limeira-De-Oliveira, Francisco;Azevedo, Celso O.	Colombo, Wesley D., Alencar, Isabel D. C. C., Limeira-De-Oliveira, Francisco, Azevedo, Celso O. (2018): New species and records of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Cerrado, Caatinga and relicts of the Atlantic Forest from northeastern Brazil. Zootaxa 4462 (1): 1-40, DOI: 10.11646/zootaxa.4462.1.1
03D487B8C46ACD56FF22FC111447F8C2.text	03D487B8C46ACD56FF22FC111447F8C2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus boibumba Colombo & Alencar & Limeira-De-Oliveira & Azevedo 2018	<div><p>Dissomphalus boibumba Colombo &amp; Azevedo sp. nov.</p><p>(Figs 42–46)</p><p>Description. Male. Head, mesosoma and metasoma dark castaneous. Mandible with two apical teeth; median clypeal lobe undefined, median tooth undefined; frons coriaceous. Notauli complete; pronotal disc coriaceous. Metasomal tergite II with lateral depression composed of tubercle with wide fovea, thick and conspicuous bristles on the border of each depression. Posterior hypopygeal margin concave. Genitalia: paramere with rounded apex, slightly arched inward, bifurcated in ventral view, wide, specially basally, higher than basiparamere; aedeagal ventral ramus as long as dorsal body, apex weakly rounded, laminar, narrow, strongly constricted; aedeagal dorsal body with three pair of apical lobes, outer lobe long, narrow, apex strongly serrated with bristles, medium pair narrow, apex pointed, slightly arched inward, inner pair wide, apex rounded, strongly slightly arched outward, basal process absent; apodeme extending beyond genital ring. Female unknown.</p><p>Remarks. This species belongs to the setosus species-group by having the tergal process with lateral depression with one pair of tubercles and the lateral margin setose. It is not similar to any other species of its species-group. It is different mainly by having the paramere bifurcated in ventral view and the aedeagal ventral ramus strongly constricted.</p><p><a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.864166&amp;materialsCitation.latitude=-6.6183333" title="Search Plazi for locations around (long -45.864166/lat -6.6183333)">Material</a> examined. Holotype ♂, BRAZIL, MA[ranhão], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.864166&amp;materialsCitation.latitude=-6.6183333" title="Search Plazi for locations around (long -45.864166/lat -6.6183333)">Mirador</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.864166&amp;materialsCitation.latitude=-6.6183333" title="Search Plazi for locations around (long -45.864166/lat -6.6183333)">Parq</a> [ue] Estadual Mirador, Base da Geraldina, 06°37'06''S 45°51'51''W, 411m, 02–12.VIII.2013, Armadilha Malaise, F Limeira-de-Oliveira LLM Santos, TL Rocha cols. (CZMA) . Paratypes, MA[ranhão], Carolina, PN Chapada das Mesas, 1♂ Riacho Sucuruiu, 08.V.2014, Arm [adilha] Malaise, (CZMA) ; 1♂ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-47.29911&amp;materialsCitation.latitude=-7.1122775" title="Search Plazi for locations around (long -47.29911/lat -7.1122775)">Riacho Cancela</a>, 07°06'44.2''S 47°17'56.8''W, 255m, 15– 31.VII.2013, Armadilha Malaise, JA Rafael, F Limeira-de-Oliveira, TTA Silva cols. (CZMA) .</p><p>Etymology. The epithet boibumba refers to Boi-bumbá, it is a popular dance in Maranhian folklore. Treat as a noun in apposition.</p><p>Distribution. Brazil (Maranhão).</p></div>	https://treatment.plazi.org/id/03D487B8C46ACD56FF22FC111447F8C2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colombo, Wesley D.;Alencar, Isabel D. C. C.;Limeira-De-Oliveira, Francisco;Azevedo, Celso O.	Colombo, Wesley D., Alencar, Isabel D. C. C., Limeira-De-Oliveira, Francisco, Azevedo, Celso O. (2018): New species and records of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Cerrado, Caatinga and relicts of the Atlantic Forest from northeastern Brazil. Zootaxa 4462 (1): 1-40, DOI: 10.11646/zootaxa.4462.1.1
03D487B8C46ACD59FF22F8E81447FBBC.text	03D487B8C46ACD59FF22F8E81447FBBC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus capelobo Colombo & Alencar & Limeira-De-Oliveira & Azevedo 2018	<div><p>Dissomphalus capelobo Colombo &amp; Azevedo sp. nov.</p><p>(Figs 47–50)</p><p>Description. Male. Head and mesosoma black; metasoma dark castaneous. Mandible with four apical teeth;</p><p>median clypeal lobe trapezoidal, median tooth straight, high in lateral view; frons coriaceous, punctate; eyes glabrous. Notauli complete; pronotal disc coriaceous, punctate. Metasomal tergite II with lateral depression composed of tubercle with wide fovea, thick and conspicuous bristles on the border of each depression. Posterior hypopygeal margin straight. Genitalia: paramere with straight apex, wide, as long as basiparamere; aedeagal ventral ramus as long as dorsal body, apex pointed, laminar, divergent, narrow; aedeagal dorsal body with two pair of apical lobes, outer lobe leaf-shaped, median folded region, dorsal pair narrow, apex rounded, slightly arched outward; basal process absent; apodeme extending beyond genital ring. Female unknown.</p><p>Variations. Apical lobes of aedeagal dorsal body very short, short or wide.</p><p>Remarks. This species belongs to the setosus species-group by having the tergal process with lateral depression with one pair of tubercles, and the lateral margin setose. This species is not similar to other species of its species-group. It is different mainly because has the mandible with four apical teeth and the paramere not arched, the aedeagal ventral ramus with the apex pointed, and the aedeagal dorsum with the outer lobe leaf-shaped.</p><p>Material examined. Holotype ♂, BRAZIL, MA[ranhão], Carolina, PN Chapada das Mesas, Riacho Cancela, 07°06'44.2''S 47°17'58.3''W, 255m, 01–10.VIII.2013, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-47.299526&amp;materialsCitation.latitude=-7.1122775" title="Search Plazi for locations around (long -47.299526/lat -7.1122775)">Armadilha</a> suspensa, JA <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-47.299526&amp;materialsCitation.latitude=-7.1122775" title="Search Plazi for locations around (long -47.299526/lat -7.1122775)">Rafael</a>, F Limeira-de-Oliveira, TTA Silva cols. (CZMA) . Paratypes: MA[ranhão], 5♂ same data of holotype, except: 01–15.VII.2013, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-47.308777&amp;materialsCitation.latitude=-7.118222" title="Search Plazi for locations around (long -47.308777/lat -7.118222)">Armadilha Malaise</a>; 2♂ 07°06'44.2''S 47°17'56.8''W, 11–14.VI.2013, Armadilha Malaise; 1♂ 225m, 07°06'44.2''S 47°17'56.8''W, 14–30.VI.2013, 2♂ Riacho Sucuruiu, 07°07'05.6''S 47°18'31.6''W , 240m, 14–30.VI.2013, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.846943&amp;materialsCitation.latitude=-6.5994444" title="Search Plazi for locations around (long -45.846943/lat -6.5994444)">Armadilha Malaise</a> (CZMA); 1♂ 20–31.VIII.2013; 1♂ 07°06'44.2''S 47°17'56.8''W, 11–14.VI.2013; 2♂ 01– 10.IX.2013; 1♂ 08.V.2014; 1♂ Mirador, Parq[ue] Estadual Mirador, Base da Geraldina, 06°35'58''S 45°50'49''W , 402m, 01–15.IX.2013, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.880276&amp;materialsCitation.latitude=-6.63" title="Search Plazi for locations around (long -45.880276/lat -6.63)">Armadilha Malaise</a>, LS Santos, F Limeira-de-Oliveira, LLM <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.880276&amp;materialsCitation.latitude=-6.63" title="Search Plazi for locations around (long -45.880276/lat -6.63)">Santos</a> cols. (CZMA); 1♂ 01– 10.XII.2013, F <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.880276&amp;materialsCitation.latitude=-6.63" title="Search Plazi for locations around (long -45.880276/lat -6.63)">Limeira-de-Oliveira</a>, LLM Santos, TL Rocha cols. (CZMA); 2♂ 06°37'48''S 45°52'49''W , 416m, 01–20.XI.2013, Armadilha Suspensa, F Limeira-de-Oliveira, LLM Santos, TL Rocha cols. (CZMA); 3♂ 06°37'25''S 45°52'08''W, 14–18.VIII.2012, Armadilha Malaise, DWA Marques, F Limeira-de-Oliveira, JSP Júnior cols. (CZMA); 1♂ 13–19.X.2012, Armadilha Malaise, LS Santos, F Limeira-de-Oliveira, LLM Santos cols. (CZMA); 1♂ Caxias, Res[erva] Ecol[ógica] Inhamum, 25–28.XI.2005, GA Cunha col., (CZMA).</p><p>Etymology. The epithet capelobo, cape from Tupi and lobo from Portuguese refers to a a monster-like character in Brazilian folklore. Treat as a noun in apposition.</p><p>Distribution. Brazil (Maranhão).</p></div>	https://treatment.plazi.org/id/03D487B8C46ACD59FF22F8E81447FBBC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colombo, Wesley D.;Alencar, Isabel D. C. C.;Limeira-De-Oliveira, Francisco;Azevedo, Celso O.	Colombo, Wesley D., Alencar, Isabel D. C. C., Limeira-De-Oliveira, Francisco, Azevedo, Celso O. (2018): New species and records of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Cerrado, Caatinga and relicts of the Atlantic Forest from northeastern Brazil. Zootaxa 4462 (1): 1-40, DOI: 10.11646/zootaxa.4462.1.1
03D487B8C465CD58FF22FB7114CFFE11.text	03D487B8C465CD58FF22FB7114CFFE11.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus iracema Colombo & Alencar & Limeira-De-Oliveira & Azevedo 2018	<div><p>Dissomphalus iracema Colombo &amp; Azevedo sp. nov.</p><p>(Figs 51–55)</p><p>Description. Male. Head and mesosoma black; metasoma dark castaneous. Mandible with four apical teeth; median clypeal lobe tridentate, median tooth rounded, with the posterior and lateral sides high and the lower anterior and central regions, dustpan-shaped; frons strongly coriaceous, punctate, eyes glabrous with few bristles. Notauli complete; pronotal disc strongly coriaceous. Metasomal tergite II with lateral depression composed of tuber with wide fovea, thick and conspicuous bristles directed backwards on the border of each depression. Posterior hypopygeal margin straight. Genitalia: paramere with weakly rounded apex, slightly arched inward, very wide, specially basally, higher than basiparamere; basivolsella very wide, straight; aedeagal ventral ramus as long as dorsal body, apex rounded, laminar, curved dorsally; aedeagal dorsal body with three pair of apical lobes, outer pair sickle-shaped, small, curved dorsally, inner pair wide, arched inward, dorsal pair wide, apex rounded, curved dorsally; basal process absent; apodeme extending beyond genital ring. Female unknown.</p><p>Remarks. This species belongs to the setosus species-group by having the tergal process with lateral depression with one pair of tubercles, and the lateral margin setose. It is not similar to any other species of its species-group. It is different mainly by having the mandible with four apical teeth, the basivolsella very wide, the aedeagal dorsal body with the outer pair of the apical lobe sickle-shaped, and the inner pair curved dorsally.</p><p><a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-47.308777&amp;materialsCitation.latitude=-7.118222" title="Search Plazi for locations around (long -47.308777/lat -7.118222)">Material</a> examined. Holotype ♂, BRAZIL, MA[ranhão], Carolina, PN Chapada das Mesas, Riacho Sucuruiu, 07°07'05.6''S 47°18'31.6''W, 240m, 20–30.IX.2013, Armadilha Malaise, JA Rafael, F Limeira-de-Oliveira, TTA Silva cols. (CZMA) . Paratypes, 1♂ CE[ará], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-40.909725&amp;materialsCitation.latitude=-3.8369443" title="Search Plazi for locations around (long -40.909725/lat -3.8369443)">Ubajara</a>, PN Ubajara, Cachoeira do Cafundó, 03°50'13''S 40°54'35''W, 18–30.XI.2012, Armadilha Malaise, JS Pinto Júnior, F Limeira-de-Oliveira cols. (CZMA) ; 1♂ same data of holotype, except: 07°06'44.2''S 47°17'56.8''W, 15–31.VII.2013; 1♂ same data of holotype, except: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-47.299526&amp;materialsCitation.latitude=-7.1122775" title="Search Plazi for locations around (long -47.299526/lat -7.1122775)">Riacho Cancela</a>, 07°06'44.2''S 47°17'58.3''W, 225m, 20–31.VIII.2013, Suspensa lâmina d'água, JA Rafael, F Limeira-de- Oliveira, TTA Silva cols. (CZMA) ; 1♂ 07°06'44.2''S 47°17'56.8''W, 255m, 11–14.VI.2013, Armadilha Malaise, (CZMA); MA[ranhão], Mirador, Parq[ue] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.864166&amp;materialsCitation.latitude=-6.6183333" title="Search Plazi for locations around (long -45.864166/lat -6.6183333)">Estadual Mirador</a>, Base da Geraldina, 1♂ 06°37'06''S 45°51'51''W, 411m, 02–12.VIII.2013, Armadilha Malaise, F Limeira-de-Oliveira LLM Santos, TL Rocha cols. (CZMA) ; 1♂ 06°35'58''S 45°50'49''W, 402m, 01–15.IX.2013, Armadilha Malaise, LS Santos, F Limeira-de-Oliveira, LLM Santos cols. (CZMA).</p><p>Etymology. The epithet iracema from Tupi refers to a beautiful indigenous woman of the Brazilian romance by José de Alencar published in 1865 and performed in Ceará. Treat as a noun in apposition.</p><p>Distribution. Brazil (Ceará and Maranhão).</p></div>	https://treatment.plazi.org/id/03D487B8C465CD58FF22FB7114CFFE11	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colombo, Wesley D.;Alencar, Isabel D. C. C.;Limeira-De-Oliveira, Francisco;Azevedo, Celso O.	Colombo, Wesley D., Alencar, Isabel D. C. C., Limeira-De-Oliveira, Francisco, Azevedo, Celso O. (2018): New species and records of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Cerrado, Caatinga and relicts of the Atlantic Forest from northeastern Brazil. Zootaxa 4462 (1): 1-40, DOI: 10.11646/zootaxa.4462.1.1
03D487B8C464CD58FF22FD9A1447FA7E.text	03D487B8C464CD58FF22FD9A1447FA7E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus joaquinae Colombo & Alencar & Limeira-De-Oliveira & Azevedo 2018	<div><p>Dissomphalus joaquinae Colombo &amp; Azevedo sp. nov.</p><p>(Figs 56–59)</p><p>Description. Male. Head and mesosoma black; metasoma dark castaneous. Mandible with three apical teeth; median clypeal lobe trapezoidal, median tooth undefined, high in lateral view; frons coriaceous, punctate, eyes setose. Notauli complete; pronotal disc coriaceous, punctate. Metasomal tergite II with lateral depression composed of tubercle with wide fovea, thick and conspicuous bristles on the border of each depression. Posterior hypopygeal margin straight. Genitalia: paramere with weakly rounded apex, slightly arched inward, wide, as long as basiparamere, pointed projection in ventral view; aedeagal ventral ramus as long as dorsal body, apex bifurcated forming two teeth, pointed, laminar, curved dorsally; aedeagal dorsal body with two pair of apical lobes, dorsal margin folded across its length, wide, apical and ventral margins weakly serrated, inner pair membranous, spineshaped, basal process absent; apodeme extending beyond genital ring. Female unknown.</p><p>Remarks. This species belongs to the setosus species-group by having the tergal process with the lateral depression with one pair of tubercles and the lateral margin setose. It is similar to D. magnus Redighieri &amp; Azevedo by having the mandible with three apical teeth, the posterior hypopygeal margin straight, and the aedeagal dorsal body with two pairs of apical lobes. However, D. joaquinae sp. nov. has the median clypeal lobe trapezoidal, the paramere without dorsal margin with the apical spine, the aedeagal ventral ramus with the apex bifurcated, whereas D. magnus, has the median clypeal lobe tridentate, the paramere with the dorsal margin with an apical spine, the aedeagal ventral ramus with the apical half divided in two long tubular rami.</p><p>Material examined. Holotype ♂, BRAZIL, MA[ranhão], Carolina, PN Chapada das Mesas, Riacho Cancela, 07°06'44.2''S 47°17'56.8''W, 255m, 01–10.VIII.2013, Armadilha suspensa, JA Rafael, F Limeira-de-Oliveira, TTA Silva cols. (CZMA). Paratypes: 3♂ same data of holotype, except: 01–31.VII.2013 (CZMA).</p><p>Etymology. The epithet joaquinae is named after the folklore of a ghost, Ana Joaquina Jansen Pereira, a rich woman from São Luís (Maranhão), who was forced to perpetually wander in the streets of the city, because she used to mistreat her slaves. Treat as a noun in apposition.</p><p>Distribution. Brazil (Maranhão).</p></div>	https://treatment.plazi.org/id/03D487B8C464CD58FF22FD9A1447FA7E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colombo, Wesley D.;Alencar, Isabel D. C. C.;Limeira-De-Oliveira, Francisco;Azevedo, Celso O.	Colombo, Wesley D., Alencar, Isabel D. C. C., Limeira-De-Oliveira, Francisco, Azevedo, Celso O. (2018): New species and records of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Cerrado, Caatinga and relicts of the Atlantic Forest from northeastern Brazil. Zootaxa 4462 (1): 1-40, DOI: 10.11646/zootaxa.4462.1.1
03D487B8C464CD5BFF22F9E9133DFDD9.text	03D487B8C464CD5BFF22F9E9133DFDD9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus plaumanni Evans 1964	<div><p>Dissomphalus plaumanni Evans, 1964</p><p>Dissomphalus plaumanni Evans, 1964: 47, 52, 58–59 (♂, holotype from Brazil).</p><p>Diagnosis. Male. Head black, mesosoma nearly black. Mandible with four apical teeth. Median clypeal lobe trapezoidal. Frons coriaceous, punctures large, shallow. Vertex slightly convex, rounded corner. T2 with pair of rounded lateral depressions, each one with tubercles, tuft of hairs directed back- and inward, anterior margin of depression, some hairs. Genitalia: paramere very wide; cuspis thick; digitus rounded dorsal expansion; aedeagal ventral ramus laminar, surface horizontal, very short, as short as basiparamere, wide, apex acute; aedeagal dorsal body with pair of apical lobes, laminar, wide, rounded apex in lateral view, compressed in dorsal view, inner surface stout, membranous and setose; genital triangular. Female unknown.</p><p>Variations. In this series the specimens are quite similar to those previously studied.</p><p>Remarks. This species was previously known from Mexico, Belize, Guatemala, Costa Rica, Panama, Trinidad &amp; Tobago, Ecuador, Peru (Azevedo, 1999c), Brazil (Evans, 1964), Brazil (Amazonas, Pará, Maranhão, Acre, Rondônia, Rio de Janeiro, São Paulo, Paraná, and Santa Catarina) (Azevedo 1999c), Brazil (Espírito Santo) (Redighieri &amp; Azevedo 2004), Brazil (Sergipe, Alagoas, and Bahia) (Redighieri &amp; Azevedo 2006), and Bolivia (Azevedo 1999c). Now it is recorded for the first time from Piauí and more males are found in Maranhão.</p><p><a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-47.29911&amp;materialsCitation.latitude=-7.1122775" title="Search Plazi for locations around (long -47.29911/lat -7.1122775)">Material</a> examined. BRAZIL, PI[auí], 1♂ Piracuruca, PN de Sete Cidades, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-47.29911&amp;materialsCitation.latitude=-7.1122775" title="Search Plazi for locations around (long -47.29911/lat -7.1122775)">Posto</a> do ICMBio, 04°05'57''S 41°42'34''W, 15–28.II.2013, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-47.29911&amp;materialsCitation.latitude=-7.1122775" title="Search Plazi for locations around (long -47.29911/lat -7.1122775)">Armadilha</a> suspensa, F Limeira-de-Oliveira, JS Pinto Júnior cols. (CZMA); 1♂ 12– 27.III.2013, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-47.29911&amp;materialsCitation.latitude=-7.1122775" title="Search Plazi for locations around (long -47.29911/lat -7.1122775)">Armadilha Malaise</a>, F Limeira-de-Oliveira, TTA <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-47.29911&amp;materialsCitation.latitude=-7.1122775" title="Search Plazi for locations around (long -47.29911/lat -7.1122775)">Silva</a> cols. (CZMA); MA[ranhão], 4♂ Carolina, PN Chapada das Mesas, Riacho Cancela, 07°06'44.2''S 47°17'56.8''W, 255m, 15.VII–31.XII.2013, Armadilha Malaise, JA Rafael, F Limeira-de-Oliveira, TTA Silva cols. (CZMA); 1♂ Riacho Sucuruiu, 08.V.2014, Arm [adilha] Malaise, (CZMA); 1♂ Caxias, Res [erva] Ecol [ógica] Inhamum, 25–28.XI.2005, GA Cunha col., (CZMA).</p></div>	https://treatment.plazi.org/id/03D487B8C464CD5BFF22F9E9133DFDD9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colombo, Wesley D.;Alencar, Isabel D. C. C.;Limeira-De-Oliveira, Francisco;Azevedo, Celso O.	Colombo, Wesley D., Alencar, Isabel D. C. C., Limeira-De-Oliveira, Francisco, Azevedo, Celso O. (2018): New species and records of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Cerrado, Caatinga and relicts of the Atlantic Forest from northeastern Brazil. Zootaxa 4462 (1): 1-40, DOI: 10.11646/zootaxa.4462.1.1
03D487B8C467CD5BFF22FD0F143FF972.text	03D487B8C467CD5BFF22FD0F143FF972.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus bifurcatus Azevedo 1999	<div><p>Dissomphalus bifurcatus Azevedo, 1999</p><p>Dissomphalus bifurcatus Azevedo, 1999a: 322, 323 (♂, holotype from Brazil).</p><p>Diagnosis. Male. Black. Mandible tridentate. Clypeus with sharp tooth. Frons weakly coriaceous, punctures very small. Vertex nearly straight, weak median depression. T2 with median depression elliptical, deep, rather large, pair of lateral hairy tubercles, so that depression becomes biconcave in dorsal view, tubercles with hole and tuft of hairs, slightly directed toward each other, distant each other 2× their diameters, lateral margins of depression with hairs. Genitalia: aedeagal ventral ramus as long as dorsal body, apical half thin, somewhat dilated below apex, basal half wider, inner margin straight basally, excavated medially, outer margin broadly convex basally; aedeagal dorsal body with two pairs apical lobe, outer lobe with pair of slender teeth dorsad, apex bifurcated, outer arm slightly wider than inner arm; inner pair membranous and hairy.</p><p>Female (in copula). Dark castaneous. Head: very weakly coriaceous, densely punctate, punctures large, somewhat shallow, separated by 0.2–1.0 × their diameters, but absent in narrow median band it frons. Mandible tetradentate, lower larger, apical margin very oblique. Median clypeal lobe trapezoidal, apical margin angularly concave, median carina very high, angled before apical margin. Eye subcircular, with about 7 facets. Malar space 1.4× HE. LH 1.36 × WH, head with sides parallel, slightly convex. Vertex straight, occipital carina visible in dorsal view. Pronotal disc with lateral margin slightly converging posterad. Propodeal disc with maximum width 1.13 × minimum width. Mid tibia not spinose. Metasoma with long petiole.</p><p>Variations. In this series the specimens are quite similar to those previously studied.</p><p>Remarks. This species was previously known from Brazil (Pará) (Azevedo 1999a), and Brazil (Paraíba, Alagoas and Sergipe) (Redighieri &amp; Azevedo 2006). Now it is recorded for the first time from Piauí, and Maranhão.</p><p><a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-43.561695&amp;materialsCitation.latitude=-9.141055" title="Search Plazi for locations around (long -43.561695/lat -9.141055)">Material</a> examined. BRAZIL, PI[auí], 1♂ Guaribas, PN Serra das Confusões, Andorinhas, 09°08'27.8''S 43°33'42.1''W, 515m, 01–10.XI.2013, Armadilha Malaise, JA Rafael, F Limeira-de-Oliveira, TTA Silva cols. (CZMA) ; MA[ranhão], 1♂ Mirador, Parq[ue] <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.86889&amp;materialsCitation.latitude=-6.6236115" title="Search Plazi for locations around (long -45.86889/lat -6.6236115)">Estadual Mirador</a>, Base da Geraldina, 06°37'25''S 45°52'08''W, 08- 14.III.2013, Armadilha Malaise, LS Santos, F Limeira-de-Oliveira, LLM Santos cols. (CZMA) ; 1♂ 06°37'48''S 45°52'49''W, 416m, 20–31.XII.2013; 1♂ 09°08'27.8''S 43°33'42.1''W, 515m, 01–10.XI.2013, JA Rafael, F Limeirade-Oliveira, TTA Silva cols. (CZMA).</p></div>	https://treatment.plazi.org/id/03D487B8C467CD5BFF22FD0F143FF972	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colombo, Wesley D.;Alencar, Isabel D. C. C.;Limeira-De-Oliveira, Francisco;Azevedo, Celso O.	Colombo, Wesley D., Alencar, Isabel D. C. C., Limeira-De-Oliveira, Francisco, Azevedo, Celso O. (2018): New species and records of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Cerrado, Caatinga and relicts of the Atlantic Forest from northeastern Brazil. Zootaxa 4462 (1): 1-40, DOI: 10.11646/zootaxa.4462.1.1
03D487B8C467CD5AFF22F8B81447FC9C.text	03D487B8C467CD5AFF22F8B81447FC9C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus torem Colombo & Alencar & Limeira-De-Oliveira & Azevedo 2018	<div><p>Dissomphalus torem Colombo &amp; Azevedo sp. nov.</p><p>(Figs 60–65)</p><p>Description. Male. Head and mesosoma black; metasoma dark castaneous. Mandible with three apical teeth; median clypeal lobe trilobate, median tooth pointed; frons coriaceous, punctate; eyes glabrous. Notauli complete; pronotal disc coriaceous, punctate. Metasomal tergite II with tergal process with one pair of conspicuous median depressions, transversally elliptical, tubercle present, directed each other. Posterior hypopygeal margin straight. Genitalia: paramere with weakly rounded apex, shorter than the basiparamere, slightly arched inward; aedeagal ventral ramus shorter than dorsal body, apex rounded, laminar, basal projection present; aedeagal dorsal body with two pair of apical lobes, outer lobe very wide, curved ventrally, bifurcated apex forming two teeth, inner pair with apex setose and half basal membranous, basal process absent; apodeme not extending beyond genital ring. Female unknown.</p><p>Remarks. This species belongs to the vallensis species-group by having the tergal process with one pair of conspicuous median depressions, transversally elliptical, tubercle present, directed toward each other. It is similar to Dissomphalus extrarramis Azevedo by having the mandible with three apical teeth, the aedeagal dorsal body with two pairs of apical lobes, outer lobe arched downward apically with two sharpened teeth at the edge, and paramere smaller than basiparamere. However, D. torem sp. nov. has the aedeagal ventral ramus with basal projection present, inner lobe with apex very setose, and two teeth in the basal part of the outer lobe, whereas D. extrarramis, has the aedeagal ventral ramus without a basal projection present, inner lobe without apex densely setose, and never with two teeth in the basal part of the outer lobe.</p><p>Material examined. Holotype ♂, BRAZIL, MA[ranhão], Carolina, PN Chapada das Mesas, Riacho Sucuruiu, 240 m, 07°07'05.6''S 47°18'31.6''W, 11–14.VI.2013, JA Rafael, F Limeira-de-Oliveira, TTA Silva cols. (CZMA). Paratypes: 1♂ same data of holotype, except: Mirador, Par[que] Estadual Mirador, Base da Geraldina, 06°37'25''S 45°52'52.08''W, 10–16.V.2013, F Limeira-de-Oliveira, LLM Santos, TL Rocha cols. (CZMA).</p><p>Etymology. The epithet torem refers to a ritual dance of the indigenous tradition, whose participants imitate animals, very common in the state of Ceará. Treat as a noun in apposition.</p><p>Distribution. Brazil (Maranhão).</p></div>	https://treatment.plazi.org/id/03D487B8C467CD5AFF22F8B81447FC9C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colombo, Wesley D.;Alencar, Isabel D. C. C.;Limeira-De-Oliveira, Francisco;Azevedo, Celso O.	Colombo, Wesley D., Alencar, Isabel D. C. C., Limeira-De-Oliveira, Francisco, Azevedo, Celso O. (2018): New species and records of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Cerrado, Caatinga and relicts of the Atlantic Forest from northeastern Brazil. Zootaxa 4462 (1): 1-40, DOI: 10.11646/zootaxa.4462.1.1
03D487B8C466CD5AFF22FC4B1794F8A7.text	03D487B8C466CD5AFF22FC4B1794F8A7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus ubracus Brito & Azevedo 2017	<div><p>Dissomphalus ubracus Brito &amp; Azevedo, 2017</p><p>Dissomphalus ubracus Brito &amp; Azevedo, 2017: 65 (♂, holotype from Panama).</p><p>Diagnosis. Male. Body castaneous. Mandible with three distal teeth. Median clypeal lobe ill defined, with one angulate tooth; median clypeal carina high in profile, incomplete apically, straight in profile or nearly so. Frons weakly coriaceous and punctures small. Vertex crest weakly concave. Pronotal disc weakly coriaceous; anterior margin ecarinate. Metasomal tergite II with tergal process lateral, with deeply excavated, subcircular and large depression covered by tergite, without tubercle, pit absent. Posterior hypopygeal margin strongly concave. Genitalia: paramere with dorsal margin wide basally; apical margin truncate. Aedeagal ventral ramus shorter than dorsal body, wide, abruptly narrowing apicad; cross section laminar; surface horizontal; inner margin straight; outer margin slightly sinuous; apex short, simple, slightly curved outward; additional inner ramus absent; basal stub absent. Aedeagal dorsal body with two pairs of apical lobes; outer lobe short, horizontal and wide, with apical margin abruptly acute and dorsad; inner pair stout, membranous and setose. Apodeme not extending beyond genital ring. Female unknown.</p><p>Variations. The main variations found in this series are the metasomal tergal process with an invagination that varies from short to long.</p><p>Remarks. This species was previously known from Panama (Brito &amp; Azevedo 2017). Now it is recorded for the first time from South America (Brazil: Ceará and Maranhão).</p><p><a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-40.909725&amp;materialsCitation.latitude=-3.8369443" title="Search Plazi for locations around (long -40.909725/lat -3.8369443)">Material</a> examined. BRAZIL, CE[ará], 2♂ Ubajara, PN Ubajara, Cachoeira do Cafundó, 03°50'13''S 40°54'35''W, 01–22.III.2013, Armadilha Malaise, TTA Silva, F Limeira-de-Oliveira cols. (CZMA) ; MA[ranhão], 1♂ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-47.29911&amp;materialsCitation.latitude=-7.1122775" title="Search Plazi for locations around (long -47.29911/lat -7.1122775)">Carolina</a>, PN Chapada das Mesas, Riacho Cancela, 07°06'44.2''S 47°17'56.8''W, 255m, 01–10.VIII.2013, Armadilha Malaise, JA Rafael, F Limeira-de-Oliveira, TTA Silva cols. (CZMA) ; 1♂ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.109444&amp;materialsCitation.latitude=-6.808056" title="Search Plazi for locations around (long -45.109444/lat -6.808056)">Mirador</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.109444&amp;materialsCitation.latitude=-6.808056" title="Search Plazi for locations around (long -45.109444/lat -6.808056)">Parq</a> [ue] Estadual Mirador, Base da Cágados, 06°48'29''S 45°06'34''W, 27.IX–02.X.2011, Armadilha Malaise, F Limeira-de-Oliveira, DWA Marques cols. (CZMA) .</p></div>	https://treatment.plazi.org/id/03D487B8C466CD5AFF22FC4B1794F8A7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colombo, Wesley D.;Alencar, Isabel D. C. C.;Limeira-De-Oliveira, Francisco;Azevedo, Celso O.	Colombo, Wesley D., Alencar, Isabel D. C. C., Limeira-De-Oliveira, Francisco, Azevedo, Celso O. (2018): New species and records of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Cerrado, Caatinga and relicts of the Atlantic Forest from northeastern Brazil. Zootaxa 4462 (1): 1-40, DOI: 10.11646/zootaxa.4462.1.1
03D487B8C461CD5DFF22FF291447FC2A.text	03D487B8C461CD5DFF22FF291447FC2A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus boiuna Colombo & Alencar & Limeira-De-Oliveira & Azevedo 2018	<div><p>Dissomphalus boiuna Colombo &amp; Azevedo sp. nov.</p><p>(Figs 66–70)</p><p>Description. Male. Head and mesosoma black; metasoma dark castaneous. Mandible with two apical teeth; median clypeal lobe trilobate, median tooth pointed; frons coriaceous, punctate; eyes glabrous. Notauli complete; pronotal disc coriaceous, punctate. Metasomal tergite II with tergal process with one pair of conspicuous lateral depressions, tubercle absent, tuft of setae, thick and conspicuous bristles on the border of each depression, directed into depressions. Posterior hypopygeal margin weakly concave. Genitalia: paramere with rounded apex, narrow, peanut-shaped, higher than the basiparamere; aedeagal ventral ramus longer than dorsal body, apex strongly bifurcated, laminar, very wide; aedeagal dorsal body with two pair of apical lobes, outer lobe very wide, curved ventrally, rounded apex, ventral margin strongly serrated, inner pair membranous; basal process absent; apodeme extending beyond genital ring. Female unknown.</p><p>Remarks. This species does not fit into any currently defined species-group by having the aedeagal ventral ramus with the apex strongly bifurcated, laminar, very wide, the aedeagal dorsal body with the outer apical lobe very wide and curved ventrally.</p><p><a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-47.29911&amp;materialsCitation.latitude=-7.1122775" title="Search Plazi for locations around (long -47.29911/lat -7.1122775)">Material</a> examined. Holotype ♂, BRAZIL, MA[ranhão], Carolina, PN Chapada das Mesas, Riacho Cancela, 07°06'44.2''S 47°17'56.8''W, 255m, 01–10.VIII.2013, Armadilha Malaise, JA Rafael, F Limeira-de-Oliveira, TTA Silva cols. (CZMA) . Paratypes: 2♂ same data of holotype, except: 225m, 01–10.XI.2013; 1♂ 07°06'44.2''S 47°17'58.3''W, 20–30.IX.2013; 2♂ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.880276&amp;materialsCitation.latitude=-6.63" title="Search Plazi for locations around (long -45.880276/lat -6.63)">Mirador</a>, Parque Estadual Mirador, Base da Geraldina, 06°37'48''S 45°52'49''W, 10–30.XI.2013, F Limeira-de-Oliveira, LLM Santos, TL Rocha cols. (CZMA) ; 1♂ Armadilha Suspensa, 01– 10.XI.2013, (CZMA).</p><p>Etymology. The epithet boiuna from Tupi refers to an Amazon myth and means a huge dark snake is able to flip boats. Treat as a noun in apposition.</p><p>Distribution. Brazil (Maranhão).</p></div>	https://treatment.plazi.org/id/03D487B8C461CD5DFF22FF291447FC2A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colombo, Wesley D.;Alencar, Isabel D. C. C.;Limeira-De-Oliveira, Francisco;Azevedo, Celso O.	Colombo, Wesley D., Alencar, Isabel D. C. C., Limeira-De-Oliveira, Francisco, Azevedo, Celso O. (2018): New species and records of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Cerrado, Caatinga and relicts of the Atlantic Forest from northeastern Brazil. Zootaxa 4462 (1): 1-40, DOI: 10.11646/zootaxa.4462.1.1
03D487B8C461CD5CFF22FB801447FF39.text	03D487B8C461CD5CFF22FB801447FF39.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus caipora Colombo & Alencar & Limeira-De-Oliveira & Azevedo 2018	<div><p>Dissomphalus caipora Colombo &amp; Azevedo sp. nov.</p><p>(Figs 71–75)</p><p>Description. Male. Head and mesosoma black; metasoma dark castaneous. Mandible with three apical teeth; median clypeal lobe undefined, median tooth pointed; frons strongly coriaceous, punctate; eyes glabrous. Notauli complete; pronotal disc strongly coriaceous. Metasomal tergite II with tergal process with one pair of strongly conspicuous lateral depressions, tubercle present, pit on top, tuft of setae. Posterior hypopygeal margin weakly concave. Genitalia: paramere with apex curved, pointed, wide, as long as basiparamere; aedeagal ventral ramus shorter than dorsal body, rounded apical extension, laminar, convergent; aedeagal dorsal body with three pairs of apical lobes, outer lobe with dorsal margin folded across its length, weakly serrated, apical margin divergent, curved ventrally, ventral margin wide projection, dorsal pair membranous, inner pair membranous, translucid; basal process absent; apodeme extending beyond genital ring. Female unknown.</p><p>Remarks. This species does not fit any of the other species-groups because it has metasomal tergite II with the tergal process with one pair of strongly conspicuous lateral depressions, the tubercle present, with a pit on top and a tuft of setae, and the paramere with the apex weakly curved and pointed with one pair of apical lobes with the ventral margin with a wide projection.</p><p><a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.86889&amp;materialsCitation.latitude=-6.6236115" title="Search Plazi for locations around (long -45.86889/lat -6.6236115)">Material</a> examined. Holotype ♂, BRAZIL, MA[ranhão], Mirador, Parq[ue] Estadual Mirador, Base da Geraldina, 06°37'25''S 45°52'08''W, 13–19.X.2012, Armadilha Malaise, LS Santos, F Limeira-de-Oliveira, LLM Santos cols. (CZMA) . Paratypes, 1♂ same data of holotype, except: 06°37'06''S 45°51'51''W, 411m, 01– 12.VIII.2013, Armadilha Malaise, LS Santos, F Limeira-de-Oliveira, LLM Santos cols. (CZMA); 1♂ 02– 12.VIII.2013, F Limeira-de-Oliveira LLM Santos, TL Rocha cols. (CZMA); 1♂ 06°37'48''S 45°52'49''W, 416m, 01–10.XII.2013; 1♂ 06°35'58''S 45°50'49''W, 402m, 10–20.X.2013; 1♂ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-47.29911&amp;materialsCitation.latitude=-7.1122775" title="Search Plazi for locations around (long -47.29911/lat -7.1122775)">Carolina</a>, PN Chapada das Mesas, Riacho Cancela, 07°06'44.2''S 47°17'56.8''W, 225m, 20–31.VIII.2013, Armadilha Malaise, JA Rafael, F Limeira-de- Oliveira, TTA Silva cols. (CZMA) ; 1♂ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-47.308777&amp;materialsCitation.latitude=-7.118222" title="Search Plazi for locations around (long -47.308777/lat -7.118222)">Riacho Sucuruiu</a>, 07°07'05.6''S, 47°18'31.6''W, 10–20.X.2013, Armadilha Malaise, JA Rafael, F Limeira-de-Oliveira, TTA Silva cols. (CZMA).</p><p>Etymology. The epithet caipora from Tupi refers to an inhabitant of the forests, who reigns over all the</p><p>animals and destroys the hunters who do not fulfill the hunting agreement made with him. Treat as a noun in apposition.</p><p>Distribution. Brazil (Maranhão).</p></div>	https://treatment.plazi.org/id/03D487B8C461CD5CFF22FB801447FF39	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colombo, Wesley D.;Alencar, Isabel D. C. C.;Limeira-De-Oliveira, Francisco;Azevedo, Celso O.	Colombo, Wesley D., Alencar, Isabel D. C. C., Limeira-De-Oliveira, Francisco, Azevedo, Celso O. (2018): New species and records of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Cerrado, Caatinga and relicts of the Atlantic Forest from northeastern Brazil. Zootaxa 4462 (1): 1-40, DOI: 10.11646/zootaxa.4462.1.1
03D487B8C460CD5CFF22FEF214D7F98E.text	03D487B8C460CD5CFF22FEF214D7F98E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus mapinguari Colombo & Alencar & Limeira-De-Oliveira & Azevedo 2018	<div><p>Dissomphalus mapinguari Colombo &amp; Azevedo sp. nov.</p><p>(Figs 76–79)</p><p>Description. Male. Head and mesosoma black; metasoma dark castaneous. Mandible with two apical teeth; median clypeal lobe undefined, median tooth rounded; frons coriaceous, punctate; eyes weakly setose. Notauli complete; pronotal disc strongly coriaceous. Metasomal tergite II with tergal process with one pair of conspicuous lateral depressions, tubercle present, pit on top and tuft of setae, thick and conspicuous bristles on the border of each depression. Posterior hypopygeal margin straight. Genitalia: paramere with pointed apex, wide, S-shaped, higher than the basiparamere; aedeagal ventral ramus shorter than dorsal body, apex weakly rounded, small projection, laminar; aedeagal dorsal body with two pair of apical lobes, outer lobe long, divergent, apex weakly bifurcated, inner pair membranous; basal process absent; apodeme extending beyond genital ring. Female unknown.</p><p>Variations. Paramere very arched or slightly arched.</p><p>Remarks. This species does not fit any other species-group by having the metasomal tergite II with tergal process with one pair of conspicuous lateral depressions, the tubercle present, with a pit on top and tuft of setae, thick and conspicuous bristles on the border of each depression, and the paramere with pointed apex, wide, Sshaped.</p><p><a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-43.561695&amp;materialsCitation.latitude=-9.141055" title="Search Plazi for locations around (long -43.561695/lat -9.141055)">Material</a> examined. Holotype ♂, BRAZIL, PI[auí], Guaribas, PN Serra das Confusões, Andorinhas, 09°08'27.8''S 43°33'42.1''W, 515m, 01–10.XI.2013, Armadilha Malaise, JA Rafael, F Limeira-de-Oliveira, TTA Silva cols. (CZMA) . Paratypes: MA[ranhão], 1♂ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.846943&amp;materialsCitation.latitude=-6.5994444" title="Search Plazi for locations around (long -45.846943/lat -6.5994444)">Mirador</a>, Parque Estadual Mirador, Base da Geraldina, 07.XI.2013, F Limeira-de-Oliveira, LLM santos, TL Rocha cols. (CZMA); 1♂ 06°37'25''S 45°52'08''W, 14– 18.VIII.2012, Armadilha Malaise, JS Pinto Júnior, F L[imeira]-de-Oliveira, DWA Marques cols. (CZMA); 1♂ 01– 10.X.2013, F Limeira-de-Oliveira, LLM Santos, TL Rocha cols. (CZMA); 2♂ 13–19.X.2012, LS Santos, F Limeira-de-Oliveira, LLM Santos cols. (CZMA); 1♂ 14–18.VIII.2012, DWA Marques, F Limeira-de-Oliveira, JSP Júnior cols. (CZMA); 1♂ 10–16.V.2013, LS Santos, F Limeira-de-Oliveira, LLM Santos cols. (CZMA); 1♂ 30.VII–06.VIII.2011, DWA Marques, F Limeira-de-Oliveira cols. (CZMA); 2♂ 06°37'06''S 45°51'51''W, 411m, 02–12.VIII.2013, F Limeira-de-Oliveira, LLM Santos, TL Rocha cols. (CZMA); 1♂ 06°37'48''S 45°52'49''W, 416m, Armadilha Suspensa, 01–10.XI.2013, F Limeira-de-Oliveira, LLM Santos, TL Rocha cols. (CZMA); 1♂ 06°35'58''S 45°50'49''W, 402m, Armadilha Malaise, 01–10.XII.2013, F Limeira-de-Oliveira, LLM Santos, TL Rocha cols. (CZMA); 1♂ Carolina, PN Chapada das Mesas, Riacho Cancela , 07°06'44.2''S 47°17'58.3''W, 255m, 01–15.VII.2013, Armadilha Malaise, JA Rafael, F Limeira-de-Oliveira, TTA Silva cols. (CZMA); 2♂ 07°06'44.2''S 47°17'56.8''W, 225m, 20–31.VIII.2013, JA Rafael, F Limeira-de-Oliveira, TTA Silva cols. (CZMA); 1♂ Caxias, Res[erva] Ecol[ógica] Inhamum, 26–29.XII.2005, Armadilha Malaise, GA Cunha cols. (CZMA).</p><p>Etymology. The epithet mapinguari from Tupi refers to a legendary creature described as being covered in a long hair red, living in the Amazon forest of Brazil and Bolivia. Treat as a noun in apposition.</p><p>Distribution. Brazil (Piauí and Maranhão).</p></div>	https://treatment.plazi.org/id/03D487B8C460CD5CFF22FEF214D7F98E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colombo, Wesley D.;Alencar, Isabel D. C. C.;Limeira-De-Oliveira, Francisco;Azevedo, Celso O.	Colombo, Wesley D., Alencar, Isabel D. C. C., Limeira-De-Oliveira, Francisco, Azevedo, Celso O. (2018): New species and records of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Cerrado, Caatinga and relicts of the Atlantic Forest from northeastern Brazil. Zootaxa 4462 (1): 1-40, DOI: 10.11646/zootaxa.4462.1.1
03D487B8C460CD5FFF22F92B1447FD69.text	03D487B8C460CD5FFF22F92B1447FD69.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dissomphalus matintaperera Colombo & Alencar & Limeira-De-Oliveira & Azevedo 2018	<div><p>Dissomphalus matintaperera Colombo &amp; Azevedo sp. nov.</p><p>(Figs 80–84)</p><p>Description. Male. Head and mesosoma black; metasoma dark castaneous. Mandible with two apical teeth; median clypeal lobe trapezoidal, broadly like duckbill in dorsal view, high in lateral view, median tooth straight; frons strongly coriaceous, punctate; eyes weakly setose. Notauli complete; pronotal disc coriaceous, punctate. Metasomal tergite II with tergal process with one pair of non-conspicuous lateral depressions, tubercle absent, callus present, tuft of setae. Posterior hypopygeal margin straight. Genitalia: paramere with weakly rounded apex, wide; as long as basiparamere; volsella with small basal projection; aedeagal ventral ramus shorter than dorsal body, apex rounded, weakly serrated, laminar, curved ventrally; aedeagal dorsal body with two pair of apical lobes, outer lobe wide, curved ventrally, dorsal margin weakly serrated, ventral margin strongly setose, inner pair membranous; basal process absent; apodeme extending beyond genital ring. Female unknown.</p><p>Remarks. This species does not fit other species-groups because it has the clypeus broadly duckbill-like in dorsal view, the metasomal tergite II with the tergal process with one pair of non-conspicuous lateral depressions, without tubercle, but with a callus and tuft of setae.</p><p><a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.864166&amp;materialsCitation.latitude=-6.6183333" title="Search Plazi for locations around (long -45.864166/lat -6.6183333)">Material</a> examined. Holotype ♂, BRAZIL, MA[ranhão], Mirador, Parq[ue] Estadual Mirador, Base da Geraldina, 06°37'06''S 45°51'51''W, 411m, 02–12.VIII.2013, Armadilha Malaise, F Limeira-de-Oliveira LLM Santos, TL Rocha cols. (CZMA) . Paratypes, 1♂ same data of holotype, except: 20–30.X.2013 (CZMA); 1♂ same data of holotype, except: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-47.090332&amp;materialsCitation.latitude=-7.073389" title="Search Plazi for locations around (long -47.090332/lat -7.073389)">Riacho Corrente</a>, 07°04'24.2''S 47°05'25.2''W, 288m, 01–10.I.2014, JA Rafael, F Limeirade-Oliveira, TL Rocha, S Pereira cols. (CZMA) ; 3♂ same data of holotype, except: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-47.29911&amp;materialsCitation.latitude=-7.1122775" title="Search Plazi for locations around (long -47.29911/lat -7.1122775)">Carolina</a>, PN <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-47.29911&amp;materialsCitation.latitude=-7.1122775" title="Search Plazi for locations around (long -47.29911/lat -7.1122775)">Chapada</a> das Mesas, Riacho Cancela, 07°06'44.2''S 47°17'56.8''W, 225m, 15.VII–31.VIII.2013, Armadilha Malaise, JA Rafael, F Limeira-de-Oliveira, TTA Silva cols. (CZMA).</p><p>Etymology. The epithet matintaperera from Tupi refers to a Brazilian folkloric woman who wears old, dark and scary clothes. Treat as a noun in apposition.</p><p>Distribution. Brazil (Maranhão).</p></div>	https://treatment.plazi.org/id/03D487B8C460CD5FFF22F92B1447FD69	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Colombo, Wesley D.;Alencar, Isabel D. C. C.;Limeira-De-Oliveira, Francisco;Azevedo, Celso O.	Colombo, Wesley D., Alencar, Isabel D. C. C., Limeira-De-Oliveira, Francisco, Azevedo, Celso O. (2018): New species and records of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Cerrado, Caatinga and relicts of the Atlantic Forest from northeastern Brazil. Zootaxa 4462 (1): 1-40, DOI: 10.11646/zootaxa.4462.1.1
