identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03C987A6FF93FFE83AB6FD441947DE5D.text	03C987A6FF93FFE83AB6FD441947DE5D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Scolytodes Ferrari 1867	<div><p>Scolytodes Ferrari</p><p>urn:lsid:zoobank.org:act: 0745C155-3489-4576-8A9B-8CA2026EFEE2</p><p>Scolytodes is readily recognized by the sharply carinate lateral margins of the pronotum and by the separated procoxae. The protibiae have two large lateral teeth near its distal end (socketed teeth embedded in cuticle, see fig. 8 in Jordal, 1998b). The eyes are entire, weakly sinuate, or emarginate, and the antennal funicle is either 5- or 6- segmented. In most species the frons of the female is distinctly ornamented, with various patterns of setae and impressions (Jordal 2013), while the male has a nearly glabrous and convex frons.</p></div>	https://treatment.plazi.org/id/03C987A6FF93FFE83AB6FD441947DE5D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Jordal, Bjarte H.	Jordal, Bjarte H. (2018): Hidden gems in museum cabinets: new species and new distributional records of Scolytodes (Coleoptera: Scolytinae). Zootaxa 4504 (1): 76-104, DOI: 10.11646/zootaxa.4504.1.4
03C987A6FF93FFEB3AB6FEA41DA9DC78.text	03C987A6FF93FFEB3AB6FEA41DA9DC78.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Scolytodes cavus Jordal 2018	<div><p>Scolytodes cavus Jordal, sp. nov.</p><p>urn:lsid:zoobank.org:act: 03593C14-850C-47C5-8055-832C751AEC3B</p><p>(Figs 1, 4, 7)</p><p>Type material. Holotype, female: Peru, Huanuco Dept, N side Cerro Carpish, 2400m, Chinchap site 670, 9¯ 15 Jan. 1983, cloud forest, A. Newton &amp; M. Thayer . Allotype: same data as holotype. HT and AT deposited in FMNH .</p><p>Diagnosis. Interstriae 10 sharply elevated to the level of metacoxa, interstriae 9 sharply elevated from level of</p><p>metacoxa to near apex; protibiae without additional mesal tooth. Female frons deeply excavated between eyes to epistoma, a weak median carina from epistoma to centre of impressed area, dorsal vestiture of long subplumose setae reaching level of antennal insertion. Similar to S. excavatus, but female frons with much longer dorsal vestiture.</p><p>Description female. Length 2.4 mm, 2.2 × longer than wide; colour dark brown. Head. Eyes weakly sinuate, separated above by 3.2 × their width. Frons deeply, sharply, concave from edge of eyes between vertex and epistoma; surface finely pubescent, with a weak carina running from epistoma towards centre of frons; vestiture consisting of few long and golden setae reaching level of antennal insertion. Antennal club densely pubescent, sutures not clearly marked. Funiculus likely 6-segmented. Pronotum weakly reticulate, faint punctures spaced by 2¯4 × their diameter. Vestiture consisting of 8 erect setae (4–2–2). Elytra smooth, striae not impressed, punctures small, shallow, densely placed in irregular rows; interstriae approximately 4 × wider than striae, punctures slightly smaller than those in striae, entirely confused. Vestiture consisting of about 3¯5 erect setae on every odd interstriae.</p><p>Legs. Procoxae separated by 0.5 × and mesocoxae 0.8 × the width of one procoxa. Protibiae straight and narrow, distal tooth 1 twice as long as 2, with 2¯3 additional tiny granules along the edge towards base; protibial mucro very short and curved posteriorly. Meso- and metatibiae with 5–6 lateral socketed teeth on distal third. Ventral vestiture. Setae on metasternum and metanepisternum mainly simple, some bifid.</p><p>Male. Similar to female except smaller (2.0 mm long), frons convex, slightly impressed just above epistoma, surface weakly reticulated, with few shallow punctures; vestiture consisting of densely placed tiny setae on lower third; eyes separated by 2.1 × their width.</p><p>Key (Wood 2007). The first couplet fails due to the combination of short interstria 10 and no mesal tooth on protibiae.</p><p>Etymology. The Latin name cavus, a masculine nominative adjective, meaning concave, refers to the excavated female frons.</p><p>Biology and distribution. This species is only known from the type locality in Peru, in high altitude cloud forest.</p></div>	https://treatment.plazi.org/id/03C987A6FF93FFEB3AB6FEA41DA9DC78	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Jordal, Bjarte H.	Jordal, Bjarte H. (2018): Hidden gems in museum cabinets: new species and new distributional records of Scolytodes (Coleoptera: Scolytinae). Zootaxa 4504 (1): 76-104, DOI: 10.11646/zootaxa.4504.1.4
03C987A6FF90FFEA3AB6FC3A1DA9D90D.text	03C987A6FF90FFEA3AB6FC3A1DA9D90D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Scolytodes excavatus Jordal 2018	<div><p>Scolytodes excavatus Jordal, sp. nov.</p><p>urn:lsid:zoobank.org:act: 26C36B3E-E231-4662-9EAC-7D3FB9D76EB0</p><p>(Figs 2, 5, 8)</p><p>Type material. Holotype, female: Peru, Huanuco Dept, Cordillera Azul, 37km NE Tingo Maria, 1600m, site673, 11¯ 14jan. 1983, montane rain forest, A. Newton &amp; M. Thayer . Allotype: same data as holotype. Paratype: Peru, Huanuco Dept, Cordillera Azul, 39km NE Tingo Maria, 1700m, site672, 11¯ 14jan. 1983, montane rain forest, A. Newton &amp; M. Thayer. HT, AT and PT deposited in FMNH .</p><p>Diagnosis. Interstriae 10 sharply elevated to the level of metacoxa, interstriae 9 sharply elevated from level of metacoxa to near apex; protibiae without additional mesal tooth. Female frons deeply excavated between eyes to epistoma, a short obscure median carina just above epistoma, vestiture in excavated area minute. Similar to S. cavus, but the female frons lacks long dorsal vestiture.</p><p>Description female. Length 1.9 mm, 2.2 × longer than wide; colour dark brown. Head. Eyes weakly sinuate, separated above by 2.9 × their width. Frons weakly concave from edge of eyes between vertex and epistoma; surface finely pubescent, with an obscure short longitudinal carina just above epistoma; vestiture consisting of minute pubescence in impressed area, with longer setae close to and on epistoma. Antennal club densely pubescent, sutures not clearly marked. Funiculus likely 6-segmented. Pronotum weakly reticulate, faint punctures spaced by 2¯4 × their diameter. Vestiture consisting of 8 erect setae (4–2–2). Elytra smooth, striae not impressed, punctures small, shallow, densely placed in irregular rows; interstriae approximately 4 × wider than striae, punctures slightly smaller than those in striae, entirely confused. Vestiture consisting of about 3¯5 erect setae on every odd interstriae.</p><p>Legs. Procoxae separated by 0.4 × and mesocoxae 0.8 × the width of one procoxa. Protibiae straight and narrow, distal tooth 1 two times longer than tooth 2, with 2¯3 additional tiny granules along the edge towards base; protibial mucro very short and curved posteriorly. Meso- and metatibiae with 5–6 lateral socketed teeth on distal third. Ventral vestiture. Setae on metasternum and metanepisternum mainly simple.</p><p>Male. Similar to female except smaller, 1.7 mm long, lower frons flat and impressed just above epistoma, surface shiny, with few shallow punctures, glabrous; eyes separated by 1.9¯2.0 × their width.</p><p>Key (Wood 2007). The first couplet fails due to the combination of short interstria 10 and no mesal tooth on protibiae.</p><p>Etymology. The Latin name excavatus, a masculine nominative participle, meaning excavated, referring to the concave female frons.</p><p>Biology and distribution. This species is only known from the type locality in Peru, in medium altitude rain forest.</p></div>	https://treatment.plazi.org/id/03C987A6FF90FFEA3AB6FC3A1DA9D90D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Jordal, Bjarte H.	Jordal, Bjarte H. (2018): Hidden gems in museum cabinets: new species and new distributional records of Scolytodes (Coleoptera: Scolytinae). Zootaxa 4504 (1): 76-104, DOI: 10.11646/zootaxa.4504.1.4
03C987A6FF91FFEA3AB6F9C918EEDE2A.text	03C987A6FF91FFEA3AB6F9C918EEDE2A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Scolytodes speculofrons Jordal 2018	<div><p>Scolytodes speculofrons Jordal, sp. nov.</p><p>urn:lsid:zoobank.org:act: 20F90265-A1D8-45F2-B2AA-B940D709BB7A</p><p>(Figs 3, 6, 9)</p><p>Type material. Holotype, female: Peru, Huanuco Dept, Cordillera Azul, 39km NE Tingo Maria, 1700m, site672, 11¯ 14jan. 1983, montane rain forest, A. Newton &amp; M. Thayer . Allotype and Paratypes (4): same data as holotype. All types deposited in FMNH .</p><p>Diagnosis. Interstriae 10 sharply elevated to near apex; protibiae without additional mesal tooth. Female frons flat with a longitudinal and slightly elevated shiny area surrounded by setae forming a bulb-shaped wreath. Closely related to S. cenchros Jordal, but the female frons has longer setae, the vertex is shinier and obscurely punctured, and pronotum has erect setae. The female frons and protibiae are very similar to S. tucumani Wood, but the body of both sexes is much stouter, the pronotum more narrowly rounded, and elytral punctures are less impressed.</p><p>Description female. Length 2.1¯ 2.2 mm, 2.1¯2.2 × longer than wide; colour dark brown. Head. Eyes entire, separated above by 2.4¯2.6 × their width. Frons flat from upper level of eyes to epistoma, median fourth on lower half slightly elevated, shiny and impunctate; vestiture consisting of fine long hair-like setae outside shiny area, almost reaching epistoma. Antennal club densely pubescent, setae semi-erect, sutures weakly marked. Funiculus 6- segmented. Pronotum reticulate, faint irregular punctures spaced by 2¯5 × their diameter. Vestiture consisting of 6 erect setae (4–0–2). Elytra smooth, striae not impressed, punctures tiny, shallow, separated in regular rows by 2¯3 × their diameter; interstriae approximately 5¯8 × wider than striae, punctures slightly smaller than those in striae, mainly in rows. Glabrous. Legs. Procoxae separated by 0.5 × and mesocoxae 1 × the width of one procoxa. Protibiae slightly broader towards apex, broadest at level of tooth 2, distal teeth 1 and 2 of equal length, with 4¯5 tiny teeth and granules along the outer edge towards base; protibial mucro very short and straight. Meso- and metatibiae with 5–6 lateral, socketed, thin teeth on distal half. Ventral vestiture. Setae on metasternum and metanepisternum simple.</p><p>Male. Similar to female except smaller, 1.9¯2.0 mm long, lower frons flat and impressed just above epistoma, surface reticulate, shallowly punctured, 4¯6 longer setae on epistoma and near antennal insertion; eyes separated by 2.1 × their width.</p><p>Key (Wood 2007). There are several ambiguous couplets in the key, including couplet 2 (protibial teeth) and 35 (pronotum reticulate). One may reach couplet 39, but S. pseudopiceus Wood has much longer frontal setae in the female, and other differences.</p><p>Etymology. The name speculofrons, a dative singular form of the Latin noun speculo, meaning glass, and frons (front of the head), referring to the impunctate shiny frons and epistoma.</p><p>Biology and distribution. This species is only known from the type locality in Peru, in medium altitude rain forest.</p></div>	https://treatment.plazi.org/id/03C987A6FF91FFEA3AB6F9C918EEDE2A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Jordal, Bjarte H.	Jordal, Bjarte H. (2018): Hidden gems in museum cabinets: new species and new distributional records of Scolytodes (Coleoptera: Scolytinae). Zootaxa 4504 (1): 76-104, DOI: 10.11646/zootaxa.4504.1.4
03C987A6FF91FFED3AB6FEE4180ADB5F.text	03C987A6FF91FFED3AB6FEE4180ADB5F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Scolytodes potens Jordal 2018	<div><p>Scolytodes potens Jordal, sp. nov.</p><p>urn:lsid:zoobank.org:act: 9EF26835-0B73-4B6A-9149-A4C1844D4599</p><p>(Figs 10, 13, 16)</p><p>Type material. Holotype, male(?): Panama, Chiriqui Prov, La Culebra Trail, 1750m, 3km N. Boquete, 9 Aug. 1995, AR Gillogly . Holotype deposited in TAMU .</p><p>Diagnosis. Interstriae 10 elevated to level of ventrite 2, interstriae 9 sharply elevated to apex; protibiae without additional mesal tooth. Male mandibles greatly enlarged. Closely related to S. montanus Jordal, but is much larger, nearly glabrous, punctures on pronotum much smaller, interstriae 9 is sharply raised for its entire length, and the posterior face of protibiae is lacking an additional mesal tooth.</p><p>Description male. Length 3.1 mm, 2.0 × longer than wide; colour reddish brown. Head. Eyes entire, separated above by 4.2 × their width. Frons convex, slightly impressed above epistoma, shiny, with scant, simple, hair-like setae. Antennal club with three clearly marked segments, the first two corneous. Funiculus 6-segmented. Pronotum smooth, shiny, densely punctured. Vestiture consisting of 8 erect setae (4–2–2). Elytra smooth, striae strongly impressed, punctures tiny, shallow, separated in regular rows by 2¯3 × their diameter, interspersed by smaller punctures of the same size as interstrial punctures; interstriae approximately 4 × wider than striae, punctures two-thirds the size of those in striae, mainly in rows, some intermixed with striae. Appears glabrous, but tiny strial and interstrial setae on declivity, some longer erect setae at apex and on interstriae 9. Legs. Procoxae separated by 1.3 × and mesocoxae 1.4 × the width of one procoxa. Protibiae broader towards apex, broadest at level of tooth 2, distal teeth 1 and 2 of equal length, tooth 2 with socketed denticle exposed, with 5¯6 tiny additional spines or granules along the edge towards base; protibial mucro very short and straight. Meso- and metatibiae with 7–8 lateral, socketed, thin teeth on distal half. Ventral vestiture. Setae on metasternum and metanepisternum simple.</p><p>Female. Not known.</p><p>Key (Wood 1982). Belongs to a unique group of Scolytodes not included in Wood’s (1982) key. With some uncertainty one may navigate to couplet 9, but this species is not close to S. plumeriae Wood.</p><p>Etymology. The name potens, a Latin noun (third declension) meaning strong or powerful, referring to the large size, stout body shape, and huge mandibles.</p><p>Biology and distribution. This species is only known from the type locality at medium altitude in Panama.</p></div>	https://treatment.plazi.org/id/03C987A6FF91FFED3AB6FEE4180ADB5F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Jordal, Bjarte H.	Jordal, Bjarte H. (2018): Hidden gems in museum cabinets: new species and new distributional records of Scolytodes (Coleoptera: Scolytinae). Zootaxa 4504 (1): 76-104, DOI: 10.11646/zootaxa.4504.1.4
03C987A6FF96FFED3AB6FBEF183CDF22.text	03C987A6FF96FFED3AB6FBEF183CDF22.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Scolytodes minimus Jordal 2018	<div><p>Scolytodes minimus Jordal, sp. nov.</p><p>urn:lsid:zoobank.org:pub: ABE697DE-EB0F-424D-BC5C-FA8CF36F27DA</p><p>(Figs 11, 14, 17)</p><p>Type material. Holotype, female: Panama, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-78.916664&amp;materialsCitation.latitude=9.316667" title="Search Plazi for locations around (long -78.916664/lat 9.316667)">San Blas Prov</a>, Nusagandi, 320m, 9°19’ N, 78°55’W, 27¯ 28 May 1995, FIT, Gillogly &amp; Stockwell. Type deposited in TAMU.</p><p>Diagnosis. Interstriae 10 elevated to level of ventrite 1; protibiae with an additional mesal tooth. Superficially similar to S. parvulus Wood and other species in the acares group, but differs strongly by having distinctly impressed striae and coarse granules at the base of interstrial setae. Also superficially similar to S. montanus Jordal and S. potens Jordal, but much smaller, with scattered, long, erect setae on elytra.</p><p>Description female. Length 1.4 mm, 2.2 × longer than wide; colour brown. Head. Eyes entire, separated above by 2.4 × their width. Frons weakly concave on a circular area occupying three-quarter the width frons on lower half; impressed area densely punctate with fine setae, impunctate and glabrous above. Antennal club setose, without sutures. Funiculus 5-segmented. Pronotum smooth, shiny, punctures very faint. Vestiture consisting of 6 erect setae (4–2–0). Elytra smooth, shiny, striae strongly impressed, punctures small, deep, separated in regular rows by 1 × their diameter; interstriae approximately 4¯5 × wider than striae, punctures tiny, obscure. Vestiture consisting of 5¯7 erect, long setae on each odd interstriae, with a coarse granule at base of setae. Legs. Procoxae separated by 0.7 × and mesocoxae 1.1 × the width of one procoxa. Protibiae narrow, distal teeth 1 and 2 of equal length, 2¯3 tiny granules along the edge towards base; protibial mucro obtuse. Mesotibiae with 6 lateral, socketed teeth on distal half, metatibiae with 5 lateral, socketed teeth on distal third. Ventral vestiture. Setae on metasternum and metanepisternum simple.</p><p>Male. Not known.</p><p>Key (Wood 1982). With much hesitation the species may key to couplet 22 and 23, without matching S. c lusiae Wood or similar species.</p><p>Etymology. The Latin name minimus is a masculine nominative adjective, meaning smallest, and refers to the much smaller size compared to S. montanus Jordal and S. potens Jordal</p><p>Biology and distribution. This species is only known from the type locality at low altitude in Panama.</p></div>	https://treatment.plazi.org/id/03C987A6FF96FFED3AB6FBEF183CDF22	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Jordal, Bjarte H.	Jordal, Bjarte H. (2018): Hidden gems in museum cabinets: new species and new distributional records of Scolytodes (Coleoptera: Scolytinae). Zootaxa 4504 (1): 76-104, DOI: 10.11646/zootaxa.4504.1.4
03C987A6FF95FFEE3AB6F8F41EC6DD63.text	03C987A6FF95FFEE3AB6F8F41EC6DD63.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Scolytodes setosicauda Jordal 2018	<div><p>Scolytodes setosicauda Jordal, sp. nov.</p><p>urn:lsid:zoobank.org:pub: ABE697DE-EB0F-424D-BC5C-FA8CF36F27DA</p><p>(Figs 12, 15, 18)</p><p>Type material. Holotype, female: Ecuador, Napo Prov, Quito-Baeza road, 4100m, March 1983, Leg. L. Huggert. Type deposited in ZMLU.</p><p>Diagnosis. Interstriae 10 very short, not sharply elevated; protibiae with tooth 2 socketed and exposed, with a large additional mesal tooth. Unique by its sulcate declivity with densely placed, short, stiff bristles.</p><p>Description female. Length 2.7 mm, 2.2 × longer than wide; colour reddish brown. Head. Eyes entire, separated above by 3.6 × their width. Frons convex, reticulate, densely and deeply punctured, except shiny and impunctate on lower third. Antennal club apically and laterally setose, first segment only partially indicated by few scale-like setae, the second suture asymmetrically procurved, marked by coarse setae, posterior two segments with scant fine setae. Funiculus 6-segmented. Pronotum densely punctured on posterior fourth, gradually replaced anteriorly by low granules and transverse asperities; anterior margin forming an elevated rim of coarse asperities. Vestiture consisting of fine recumbent setae, without longer erect setae (0–0–0). Elytra on disc shiny, striae 1 impressed, others not, punctures large, deep, separated in regular rows by 0.5¯1 × their diameter; interstriae approximately 1.5¯2 × wider than striae, punctures half the size of those in striae. On declivity, striae 1¯3 impressed, sulcate. Vestiture on disc and lateral parts of declivity consisting of long, fine interstrial setae, and slightly shorter strial setae, on declivity replaced by short, semi-erect, thick, bristle-like setae with a coarse granule at base of most setae; setae towards the elytral apex and along the sutural interstriae confused, densely placed.</p><p>Legs. Procoxae separated by approximately 0.5 × the width of one procoxa (difficult to observe). Protibiae slightly broader towards apex, distal teeth 1 and 2 large and of equal length, tooth 2 with socketed denticle exposed, 4 additional small teeth along the edge towards base; protibial mucro large and curved laterally. Meso- and metatibiae with 6¯7 lateral socketed teeth on distal half. Ventral vestiture. Setae on metasternum and metanepisternum generally simple, a few bifid setae close to elytra, some plumose, short scales on ventral side.</p><p>Male. Not known.</p><p>Key (Wood 2007). Couplet 102 is inconclusive and the alternative possible couplets, 103 and 117, do not match the specimen. None of the species included in this part of the key match.</p><p>Etymology. The name is composed of the Latin masculine adjective setosi, meaning bristly, and the feminine noun cauda, meaning tail, referring to the peculiar pattern of dense setae on the elytral declivity.</p><p>Biology and distribution. This species is only known from the type locality in high altitude elfin forest in Ecuador. This is one of the highest altitude records for any bark beetle.</p></div>	https://treatment.plazi.org/id/03C987A6FF95FFEE3AB6F8F41EC6DD63	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Jordal, Bjarte H.	Jordal, Bjarte H. (2018): Hidden gems in museum cabinets: new species and new distributional records of Scolytodes (Coleoptera: Scolytinae). Zootaxa 4504 (1): 76-104, DOI: 10.11646/zootaxa.4504.1.4
03C987A6FF95FFE13AB6FD3D1D8DD99D.text	03C987A6FF95FFE13AB6FD3D1D8DD99D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Scolytodes monticola Jordal 2018	<div><p>Scolytodes monticola Jordal, sp. nov.</p><p>urn:lsid:zoobank.org:act: 724AE2DC-50BA-4E43-A7C1-425F37963923</p><p>(Figs 19, 22, 25)</p><p>Type material. Holotype, male: Ecuador, Pich./ Napo, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-78.19806&amp;materialsCitation.latitude=-0.31" title="Search Plazi for locations around (long -78.19806/lat -0.31)">Quito-Baeza Pass</a>, 4000m, 00°18’36’’S, 78°11’53’’W, 6.XI.1999 -232b, R. Anderson, elfin forest. Note: HT was broken and glued by the author. HT deposited in FSCA.</p><p>Diagnosis. Interstriae 10 short; protibiae with tooth 2 socketed and exposed, a large additional mesal tooth present. Closely related to S. setosicauda except smaller in size, declivity not sulcate, and all setae hair-like.</p><p>Description male. Length 2.1 mm, 2.4 × longer than wide; colour brown. Head. Eyes entire, separated above by 3.4 × their width. Frons convex, reticulate, densely and deeply punctured, particularly close to epistoma. Antennal club apically and laterally setose, suture 1 and 2 strongly procurved, each marked by few long setae, segment 1 and 2 corneous. Funiculus 6-segmented. Pronotum densely punctured on posterior third, replaced anteriorly by low transverse asperities; anterior margin forming a weak elevated rim of asperities. Vestiture consisting of fine recumbent setae, without longer erect setae (0–0–0). Elytra shiny, slightly rugose, striae not impressed, punctures deep, separated in irregular rows by 1¯2 × their diameter; interstriae approximately 1.5¯2 × wider than striae, punctures slightly smaller and more widely spaced than of those in striae. Vestiture consisting of rows of long, erect, fine interstrial setae and slightly shorter semi-recumbent interstrial setae, which on sutural interstriae on declivity are more densely placed and confused. Legs. Procoxae separated by 0.4 × and mesocoxae 0.7 × the width of one procoxa. Protibiae slightly broader towards apex, distal teeth 1 and 2 large, of equal length, tooth 2 with socketed denticle exposed, 4¯5 additional small teeth along the edge towards base; protibial mucro long and curved laterally. Meso- and metatibiae with 7 lateral socketed teeth on distal half. Ventral vestiture. Setae on metasternum and metanepisternum simple.</p><p>Female. Not known.</p><p>Key (Wood 2007). Same as for S. setosicauda .</p><p>Etymology. The name monticola is a Latin noun (masculine and feminine), meaning mountaineer or mountain dwelling, referring to the high altitude elfin forest habitat around 4000 m a.s.l.</p><p>Biology and distribution. This species is only known from the type locality in high altitude elfin forest in Ecuador.</p></div>	https://treatment.plazi.org/id/03C987A6FF95FFE13AB6FD3D1D8DD99D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Jordal, Bjarte H.	Jordal, Bjarte H. (2018): Hidden gems in museum cabinets: new species and new distributional records of Scolytodes (Coleoptera: Scolytinae). Zootaxa 4504 (1): 76-104, DOI: 10.11646/zootaxa.4504.1.4
03C987A6FF9AFFE13AB6FA241DA9DE43.text	03C987A6FF9AFFE13AB6FA241DA9DE43.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Scolytodes aureifrons Jordal 2018	<div><p>Scolytodes aureifrons Jordal, sp. nov.</p><p>urn:lsid:zoobank.org:act: A8B5A3CB-D011-4EF9-B5AB-B9074AF738EC</p><p>(Figs 20, 23, 26)</p><p>Type material. Holotype, female: Panama, Canal Zone, Albrook forest site, Fort Clayton. Lot No. 177, March 7/ 8, 1968, R. Hutton, black light trap. HT deposited in USNM.</p><p>Diagnosis. Interstriae 10 elevated to level of metacoxae; protibiae with an additional mesal tooth. Pronotum with a weak summit on anterior third. Distinguished from S. ingavorus Wood by the short strial and interstrial setae, and narrow pronotal asperities.</p><p>Description female. Length 1.2 mm, 2.3 × longer than wide; colour light brown. Head. Eyes weakly sinuate, separated above by 2.3 × their width. Frons flattened from upper level of eyes to epistoma; vestiture consisting of coarse plumose setae in two-thirds of the flattened area, setae reaching epistoma, with a small median shiny area on lower median third which is barely visible. Antennal club anteriorly densely pubescent, first two segments corneous, suture 1 partially marked laterally by coarse setae and a partial septum, suture 2 strongly procurved and marked by dense setae. Funiculus 5(4?)-segmented. Pronotum strongly reticulate, shallow punctures in posterolateral areas spaced by their diameter. Vestiture consisting of 4 anterior erect setae (4–0–0). Elytra smooth, striae not impressed, punctures shallow, in regular rows, separated by 1¯1.5 × their diameter; interstriae as wide as striae, punctures slightly smaller than those in striae, half the size of strial punctures. Vestiture consisting of very short coarse strial and interstrial setae. Legs. Procoxae separated by 0.3 × and mesocoxae 0.8 × the width of one procoxa. Protibiae narrow, distal teeth 1 and 2 of equal length, with 2¯3 tiny granules along the edge towards base; protibial mucro obtuse. Meso- and metatibiae with 5 lateral, socketed, thin teeth on distal third. Ventral vestiture. Setae on metanepisternum bifid, on metasternum simple.</p><p>Male. Unknown.</p><p>Key (Wood 1982). Keys to couplet 49, S. ingavorus, but differs as noted in the diagnosis.</p><p>Etymology. The name aureus is a Latin masculine nominative adjective meaning golden (in a beautiful sense), referring to the golden setae in the female frons, thereby aureifrons .</p><p>Biology and distribution. This species is only known from the type locality in Panama, in low altitude rain forest.</p></div>	https://treatment.plazi.org/id/03C987A6FF9AFFE13AB6FA241DA9DE43	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Jordal, Bjarte H.	Jordal, Bjarte H. (2018): Hidden gems in museum cabinets: new species and new distributional records of Scolytodes (Coleoptera: Scolytinae). Zootaxa 4504 (1): 76-104, DOI: 10.11646/zootaxa.4504.1.4
03C987A6FF9AFFE03AB6FE1D1C66DBE8.text	03C987A6FF9AFFE03AB6FE1D1C66DBE8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Scolytodes piliscapus Jordal 2018	<div><p>Scolytodes piliscapus Jordal, sp. nov.</p><p>urn:lsid:zoobank.org:act: 0AB74528-BA08-4336-9FD6-9DD586F1C277</p><p>(Figs 21, 24, 27)</p><p>Type material. Holotype, female: Panama, Chiriqui Pr., Rava la Fortua Est. Cientifia, el. 1150m, 23-VII-1994, UV light, Stockwell &amp; Gillogly. HT deposited in TAMU.</p><p>Diagnosis. Interstriae 10 sharply elevated to near apex; protibiae with a very minute additional mesal tooth near tarsal insertion. Female scapus with long setae on its anterior face. Remotely related to S. perplexus Schedl, but differs by the long setae on the scapus, distinct asperities on the pronotum, and the lack of strial setae. In Central America it groups with species near S. genialis Wood in which all have confused interstrial punctures.</p><p>Description female. Length 2.3 mm, 2.2 × longer than wide; colour brown. Head. Eyes entire, separated above by 1.9 × their width. Frons weakly concave from upper level of eyes to epistoma, vestiture consisting of fine setae in entire impressed area except lower median quarter smooth and glabrous, setae along upper margin longer than below and longer than setae on scapus. Antennal club with two obliquely procurved sutures marked by setae, first segment corneous, suture 1 marked by darker colour, but not true septum. Funiculus 6-segmented, scapus with long setae on anterior side. Pronotum weakly reticulate, shallow punctures spaced by their diameter, small and obscure at base of asperities on anterior half. Vestiture consisting of 8 erect setae (4–2–2), and scant, short, fine, interspersed setae. Elytra generally smooth and shiny, striae not impressed, punctures distinct, in regular rows, separated by 1¯1.5 × their diameter; interstriae 3 × wider than striae, punctures half the size of those in striae; additional tiny punctures associated with larger strial and interstrial punctures. Vestiture consisting of erect interstrial setae, which are shorter and more densely placed on declivity; a tiny granule at base of many setae. Legs. Procoxae separated by 0.4 × and mesocoxae 0.8 × the width of one procoxa. Protibiae rather narrow, distal teeth 1 and 2 of equal length, with 2¯3 tiny teeth or granules along the edge towards base; protibial mucro long, thin and curved. Meso- and metatibiae with 7 lateral, socketed, thin teeth on distal half. Ventral vestiture. Setae on metasternum and metanepisternum simple.</p><p>Male. Unknown.</p><p>Key (Wood 1982). Keys to couplet 7 with no match because interstrial punctures are in rows.</p><p>Etymology. The name piliscapus is composed by the Latin noun pilus, meaning hair-like appendage, and the anatomical part of the antenna, scapus, meaning shank, referring to the setose extended part of the scapus.</p><p>Biology and distribution. This species is only known from the type locality in Panama, in medium altitude rain forest.</p></div>	https://treatment.plazi.org/id/03C987A6FF9AFFE03AB6FE1D1C66DBE8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Jordal, Bjarte H.	Jordal, Bjarte H. (2018): Hidden gems in museum cabinets: new species and new distributional records of Scolytodes (Coleoptera: Scolytinae). Zootaxa 4504 (1): 76-104, DOI: 10.11646/zootaxa.4504.1.4
03C987A6FF9BFFE53AB6FBB71C53D8BD.text	03C987A6FF9BFFE53AB6FBB71C53D8BD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Scolytodes curvicostatus Jordal 2018	<div><p>Scolytodes curvicostatus Jordal, sp. nov.</p><p>urn:lsid:zoobank.org:act: BC1B067C-6418-466E-BE89-9A3DB3534D94</p><p>(Figs 28, 31, 34)</p><p>Type material. Holotype, female: Costa Rica, San Jose / Cartago, km 69, Int. Amer. Hwy. nr. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-83.85833&amp;materialsCitation.latitude=9.658333" title="Search Plazi for locations around (long -83.85833/lat 9.658333)">Tres de Junio</a>, 2600m 09°39’30’’N, 83°51’30’’W, 7.vi.1997, R. Anderson, wet cloud forest litter, 97-004A. HT deposited in FSCA.</p><p>Diagnosis. Interstriae 10 sharply elevated to level of ventrite 1; protibiae with a distinct additional mesal tooth near tarsal insertion. Lateral carina on pronotum strongly curved dorso-ventrally; pronotum from above appearing constricted on posterior fourth. Distinguished from a group of distantly related species near S. semipunctatus Wood and S. clusiacolens Wood by erect interstrial setae in rows combined with the presence of very short strial setae on the elytra.</p><p>Description female. Length 2.7 mm, 2.45 × longer than wide; colour reddish brown to black. Head. Eyes entire, separated above by 2.8 × their width. Frons convex on upper two-thirds, weakly impressed below; surface reticulate with few scattered punctures; vestiture consisting of a brush of short golden setae in impressed area, denser on epistoma. Antennal club with two obliquely procurved sutures marked by fine setae, first two segments corneous. Funiculus 6-segmented. Pronotum strongly reticulate, small punctures spaced by 3¯5 × their diameter. Vestiture consisting of 6 erect setae (4–2–0), and scant, short, fine, interspersed setae. Elytra generally smooth and shiny, striae weakly impressed, punctures distinct, in regular rows, separated by 1¯2 × their diameter; interstriae 3 × wider than striae, punctures minute. Vestiture consisting of erect interstrial setae of variable length, and much shorter recumbent strial setae. Legs. Procoxae separated by 0.8 × and mesocoxae 1.2 × the width of one procoxa. Protibiae narrow, distal teeth 1 and 2 of equal length, with 4¯5 tiny additional granules along the edge towards base; protibial mucro obtuse. Meso- and metatibiae with 6 lateral, socketed teeth on distal third. Ventral vestiture. Setae on metasternum and metanepisternum simple.</p><p>Male. Unknown. The sex of the type (female) was determined based on the single sclerotized terminal tergite (seven in total).</p><p>Key (Wood 1982). Keys to couplet 25, S. clusiacolens, with no further match.</p><p>Etymology. The name is composed by the Latin masculine genitive adjective curvi (from curvus) and the masculine nominative adjective costatus, referring to the strongly dorsoventrally curved lateral costa of the pronotum.</p><p>Biology and distribution. This species is only known from the type locality in Costa Rica, in high altitude cloud forest.</p></div>	https://treatment.plazi.org/id/03C987A6FF9BFFE53AB6FBB71C53D8BD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Jordal, Bjarte H.	Jordal, Bjarte H. (2018): Hidden gems in museum cabinets: new species and new distributional records of Scolytodes (Coleoptera: Scolytinae). Zootaxa 4504 (1): 76-104, DOI: 10.11646/zootaxa.4504.1.4
03C987A6FF9EFFE53AB6F9441C53DD47.text	03C987A6FF9EFFE53AB6F9441C53DD47.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Scolytodes vellus Jordal 2018	<div><p>Scolytodes vellus Jordal, sp. nov.</p><p>urn:lsid:zoobank.org:act: BDA91FB3-12CB-448C-8C9C-B0F5AA92700C</p><p>(Figs 29, 32, 35)</p><p>Type material. Holotype, female: Costa Rica, San Jose, Zurqui de Moravia, 1600m, XI-1-9-1998, FIT, C.W. &amp; L.B.O’Brien. HT deposited in CAS.</p><p>Diagnosis. Interstriae 10 sharply elevated to the level of metacoxa; protibiae with a small additional mesal tooth near tarsal insertion. Pronotum on anterior half with dense asperities. Elytral declivity unique by its dense ground vestiture consisting of short scale-like setae, and interstriae 1, 3 and 5 carinate.</p><p>Description female. Length 1.7 mm, 2.5 × longer than wide; colour reddish brown. Head. Eyes weakly sinuate, separated above by 2.8 × their width. Frons weakly concave, surface weakly reticulate, without punctures; vestiture consisting of scattered long setae from vertex and lateral edge, reaching level of antennal insertion. Antennae not clearly visible, club possibly with two obliquely procurved sutures marked by dense setae. Pronotum strongly reticulate, obscure and transversely elongated punctures only on posterior fourth, anterior twothirds with recurved, sub-contiguous asperities. Vestiture consisting of 8 erect setae (4–2–2). Elytra smooth on disc, rough on declivity, striae impressed, more so on declivity than disc, punctures on disc large, in regular rows, separated by 0.5 × their diameter; interstriae 1.5 × wider than striae, without punctures, on declivity interstriae 1, 3 and 5 strongly elevated. Vestiture consisting of erect interstrial setae of variable length, particularly on the declivity and sides, and minute strial setae; additional, densely placed, scale-like setae on declivity. Legs. Procoxae separated by 0.8 × and mesocoxae 1.3 × the width of one procoxa. Protibiae narrow, distal teeth 1 and 2 small, of equal length, with 3¯4 tiny additional granules along the edge towards base, each with a coarse setae at its base; protibial mucro straight. Mesotibiae with 5 lateral, socketed teeth on distal half, metatibiae with 4 teeth on distal fourth. Ventral vestiture. Setae on metasternum simple, on metanepisternum trifid and plumose.</p><p>Male. Unknown.</p><p>Key (Wood 1982). Goes with much hesitation to couplet 53, with no further match.</p><p>Etymology. The name vellus is a Latin noun, meaning pelt or wool of an animal, referring to the declivity which is densely clothed by short, yellow, scale-like setae.</p><p>Biology and distribution. This species is only known from the type locality in Costa Rica, in medium altitude cloud forest.</p></div>	https://treatment.plazi.org/id/03C987A6FF9EFFE53AB6F9441C53DD47	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Jordal, Bjarte H.	Jordal, Bjarte H. (2018): Hidden gems in museum cabinets: new species and new distributional records of Scolytodes (Coleoptera: Scolytinae). Zootaxa 4504 (1): 76-104, DOI: 10.11646/zootaxa.4504.1.4
03C987A6FF9EFFE43AB6FDE11818D99D.text	03C987A6FF9EFFE43AB6FDE11818D99D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Scolytodes granulatus Jordal 2018	<div><p>Scolytodes granulatus Jordal, sp. nov.</p><p>urn:lsid:zoobank.org:act: EDE969FB-E9C4-41A6-A2AB-A64D2BAE456E</p><p>(Figs 30, 33, 36)</p><p>Type material. Holotype: Panama, Veraguas Prov, 8 km W. Santa Fe, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-81.11361&amp;materialsCitation.latitude=8.507222" title="Search Plazi for locations around (long -81.11361/lat 8.507222)">Cerro Tute</a>, el 3000 ft, 08°30’26’’N, 81°06’49’’W, 24-vii-1999, J.B. Woolley 99/054. HT deposited in TAMU.</p><p>Diagnosis. Interstriae 10 sharply elevated to just before level of metacoxa; protibiae with a small additional mesal tooth near tarsal insertion. A unique species with sharp granules associated with erect setae on odd interstriae on posterior half of the elytra, and the combination of an extremely reticulated pronotum and shiny elytra.</p><p>Description (female?). Length 1.3 mm, 2.3 × longer than wide; colour dark brown. Head. Eyes entire, separated above by 2.9 × their width. Frons convex, surface strongly reticulate and weakly punctured close to vertex, smooth and impunctate below; vestiture consisting of scattered short setae on lower half. Antennal club with two transverse sutures marked by dense setae. Funicular segments not visible. Pronotum strongly reticulate, shallow punctures on posterior two-thirds, replaced anteriorly by low asperities or granules. Vestiture consisting of 6 erect setae (4–2–0). Elytra smooth and shiny, striae slightly impressed, punctures large, separated in regular rows by 0.2¯0.5 × their diameter; interstriae as wide as striae, impunctate, sharp granules associated with setae on oddnumbered interstriae, larger and sharper on posterolateral area. Vestiture consisting of erect interstrial setae on odd- numbered interstriae, and minute to obscure strial and interstrial setae near apex. Legs. Procoxae separated by 0.7 × and mesocoxae 1.2 × the width of one procoxa. Protibiae narrow, distal teeth 1 and 2 of equal length, with 3¯4 tiny additional granules along the edge towards base; protibial mucro straight, short. Meso- and metatibiae with 5 lateral, socketed teeth on distal half, and third, respectively. Ventral vestiture. Setae on metasternum and metanepisternum simple, regularly placed setae on venter pointing posteriorly.</p><p>Key (Wood 1982). Keys to couplet 53, with no further match (if using the key in Wood 2007 one may key to couplet 98).</p><p>Etymology. The name granulatus is a Latin adjective (nominative, masculine) meaning granular (formed of small grains), referring to the distinct and sharp granules on odd interstriae on the posterior half of the elytra.</p><p>Biology and distribution. This species is only known from the type locality in Panama, at 900 m altitude.</p></div>	https://treatment.plazi.org/id/03C987A6FF9EFFE43AB6FDE11818D99D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Jordal, Bjarte H.	Jordal, Bjarte H. (2018): Hidden gems in museum cabinets: new species and new distributional records of Scolytodes (Coleoptera: Scolytinae). Zootaxa 4504 (1): 76-104, DOI: 10.11646/zootaxa.4504.1.4
03C987A6FF9FFFE43AB6FA241F70DE1B.text	03C987A6FF9FFFE43AB6FA241F70DE1B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Scolytodes bipilosus Jordal 2018	<div><p>Scolytodes bipilosus Jordal, sp. nov.</p><p>urn:lsid:zoobank.org:act: 35A942A9-DBEE-4DB5-AB6A-A61DB97CE6B3</p><p>(Figs 37, 40, 43)</p><p>Type material. Holotype: Panama, Panama Prov, 8 km NE Cerro Jéfe, 27 Mar. 1976, 700 m, H.P. Stockwell. HT deposited in TAMU.</p><p>Diagnosis. One of very few species with the combination of a long interstriae 10 which is sharply elevated to the level of ventrite 4, and protibiae having a small additional mesal tooth near tarsal insertion. Elytra with interstrial spatulate setae on posterior half, intermixed with fine strial and interstrial setae.</p><p>Description (female?). Length 1.6 mm, 2.3 × longer than wide; colour brown. Head. Eyes entire, separated above by 1.4 × their width. Frons convex, surface finely reticulate on upper half, smooth and impunctate below on median half; vestiture consisting of about 30 scattered short setae around margin of impunctate area and on epistoma. Antennal club small, with three slightly procurved sutures on anterior face, two sutures on posterior face. Funiculus 5-segmented. Pronotum weakly reticulated, with shallow, small punctures almost reaching anterior margin. Vestiture consisting of 10 erect setae (4–4–2) and short fine semi-recumbent setae arising from punctures. Elytra smooth and shiny, striae not impressed, punctures irregular, small and shallow, separated in regular rows by 2 × their diameter; interstriae 2¯4 × wider than striae, punctures minute, in rows on disc, confused on declivity. Vestiture consisting of recumbent fine strial and interstrial setae in rows, increasing in length and progressively confused posteriorly, with additional curved spatulate setae on each interstriae on the posterior half of elytra. Legs. Procoxae and mesocoxae separated by 0.8 × the width of one procoxa. Protibiae narrow, distal teeth 1 and 2 of equal length, with 5 tiny additional short teeth or granules along the edge towards base; protibial mucro short, curved posteriorly. Meso- and metatibiae with 7 and 6 lateral, socketed teeth on distal half and third, respectively. Ventral vestiture. Setae on anterior part of metasternum and all setae on metanepisternum trifid, elsewhere simple.</p><p>Key (Wood 1982). Keys to couplet 6, with no further match (to couplet 1 in Wood 2007), due to the mismatch between a long interstriae 10 and a mesal tooth present on protibia.</p><p>Etymology. Composed by the Latin nominative, masculine adjective pilosus, meaning hairy or shaggy, and the prefix bi -, referring to the two types of elytral setae on interstriae and striae, respectively.</p><p>Biology and distribution. This species is only known from the type locality in Panama, at approximately 600¯ 800 m altitude (Cerro Jefe).</p></div>	https://treatment.plazi.org/id/03C987A6FF9FFFE43AB6FA241F70DE1B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Jordal, Bjarte H.	Jordal, Bjarte H. (2018): Hidden gems in museum cabinets: new species and new distributional records of Scolytodes (Coleoptera: Scolytinae). Zootaxa 4504 (1): 76-104, DOI: 10.11646/zootaxa.4504.1.4
03C987A6FF9FFFE73AB6FEA51F91DC30.text	03C987A6FF9FFFE73AB6FEA51F91DC30.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Scolytodes asperatus Jordal 2018	<div><p>Scolytodes asperatus Jordal, sp. nov.</p><p>urn:lsid:zoobank.org:act: AE010FAA-66EB-490C-ADE1-C41BD0C78994</p><p>(Figs 38, 41, 44)</p><p>Type material. Holotype, female: Mexico, Oaxaca, Mpio San Miguel del Puerto, Arriba de Sta. Ma. Xadani, Cerro Campaa, 05/29/2006. Host trunk Ficus sp, TH Atkinson, THA-793 . Allotype and paratypes (4): same data as HT. HT and AT deposited in CNIN, 2 PTs in UTIC, 2 PT in USNM.</p><p>Diagnosis. Interstriae 10 sharply elevated to the level of metacoxa; protibiae with a small additional mesal tooth near tarsal insertion. Distinguished from other species with dense asperities on the anterior part of pronotum, in particular S. pubescens Wood, by the smooth and glabrous central area of the female frons, and by the regular rows of erect setae on all elytral interstriae.</p><p>Description female. Length 2.0¯ 2.1 mm, 2.2¯2.3 × longer than wide; bicoloured, dark brown on anterior half of pronotum, along elytral suture and elytral margins, light brown elsewhere. Head. Eyes entire, separated above by 2.7¯2.9 × their width. Frons flattened on lateral margins, weakly concave on lower one-fourth, a central spadelike area smooth and impunctate, slightly protuberant with lateral edges sub-carinate. Vestiture consisting of a wreath of golden setae around the spade-like structure. Antennal club small, slightly asymmetrical, with two obliquely procurved sutures. Funiculus 6-segmented. Pronotum shiny, deep punctures on posterior fourth separated by less than their diameter, replaced on anterior three-fourth by coarse asperities, forming on anterior margin a low serrated ridge. Vestiture consisting of 10 erect setae (4–4–2) and shorter semi-recumbent setae in area with asperities. Elytra generally smooth and shiny except for minute granules at base of setae, striae not impressed, punctures small and shallow, separated in regular rows by 1¯2 × their diameter; interstriae 3¯5 × wider than striae, punctures obscure and confused, particularly so on declivity. Vestiture consisting of recumbent fine strial and interstrial setae, with additional longer bristle-like setae on each interstriae. Legs. Procoxae separated by 0.2¯0.3 × and mesocoxae 0.7¯0.8 × the width of one procoxa. Protibiae broad, distal teeth 1 and 2 of subequal length, with 4 additional rugae or granules along the sharply formed edge towards base; protibial mucro short, curved posteriorly. Meso- and metatibiae with 7 lateral, socketed teeth on distal half and third, respectively. Ventral vestiture. Setae on metasternum simple, on metanepisternum bifid.</p><p>Male. Similar to females except slightly smaller (length 1.8¯2.0 mm), the frons is convex, strongly reticulated and granulated, with fine short setae evenly distributed.</p><p>Key (Wood 1982). Keys to couplet 49, near S, pubescens Wood, but is distinguished (in the key) by the glabrous central area of the female frons.</p><p>Etymology. The Latin name asperatus (masculine nominative participle), meaning rough, refer to the asperate pronotum and regular rows of stiff elytral setae.</p><p>Biology and distribution. This species is only known from the type locality in Oaxaca, Mexico. The type series was collected under bark of a Ficus tree trunk.</p></div>	https://treatment.plazi.org/id/03C987A6FF9FFFE73AB6FEA51F91DC30	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Jordal, Bjarte H.	Jordal, Bjarte H. (2018): Hidden gems in museum cabinets: new species and new distributional records of Scolytodes (Coleoptera: Scolytinae). Zootaxa 4504 (1): 76-104, DOI: 10.11646/zootaxa.4504.1.4
03C987A6FF9CFFF93AB6FCC21C62D928.text	03C987A6FF9CFFF93AB6FCC21C62D928.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Scolytodes flavifrons Jordal 2018	<div><p>Scolytodes flavifrons Jordal, sp. nov.</p><p>urn:lsid:zoobank.org:act: 83B82BF9-B502-485E-BB44-C265C934F4F1</p><p>(Figs 39, 42, 45)</p><p>Type material. Holotype, female: Panama, Canal Zone, Albrook Forest Site , Fort Clayton, Lot no. 147, November 9/10, 1967, Hutton and Llaurado, black light trap . Allotype, male: same data as HT, except lot no. 155, Dec. 6/7, R. Hutton; Paratype, female: lot no. 159, Dec. 21/22. HT and AT deposited in USNM, 1 PT in UTIC.</p><p>Diagnosis. Interstriae 10 sharply elevated to level of ventrite 3; protibiae without an additional mesal tooth. Female frons unique by its deep glabrous concavity appearing orange and velvet-like, and by the truncated anterior margin of the pronotum.</p><p>Description female. Length 2.1¯ 2.2 mm, 1.9¯2.0 × longer than wide; colour dark brown. Head. Eyes entire, separated above by 2.1¯2.3 × their width. Frons deeply concave from epistoma to vertex, lateral margins sharp, touching upper edge of eyes, impressed area smooth and impunctate, appearing orange and velvet-like in texture; glabrous. Antennal club setose, two sutures obscurely indicated, a faint septum present at suture 1. Funiculus 6- segmented. Pronotum weakly reticulated, subshining, shallow fine punctures separated by 2¯3 × their diameter, reaching anterior margin; anterior half with low asperities. Vestiture consisting of 6 erect setae (4–0–2). Elytra smooth and shiny, striae not impressed, punctures moderately large, shallow, in irregular rows, separated by 1¯2 × their diameter; interstriae 4¯6 × wider than striae, punctures smaller and confused, particularly so on declivity. Glabrous. Legs. Procoxae separated by 0.5¯0.6 × and mesocoxae 1.0 × the width of one procoxa. Protibiae narrow, distal teeth 1 and 2 large, of equal length, with 3¯4 additional rugae or sharp granules along towards base; protibial mucro obtuse. Meso- and metatibiae with 7 and 6 lateral, socketed teeth on distal half and third, respectively. Ventral vestiture. Setae on metasternum and metanepisternum simple.</p><p>Male. Similar to females except frons convex, weakly reticulated, with faint punctures; eyes separated above by 2.5 × their width.</p><p>Key (Wood 1982). Keys to couplet 10, with no further match.</p><p>Etymology. Composed by the Latin nominative, masculine adjective flavus, meaning yellow, and frons, meaning front of head, referring to the large yellow and velvet like concavity in the female frons.</p><p>Biology and distribution. This species is only known from the lowland type locality in Panama, attracted to black light.</p></div>	https://treatment.plazi.org/id/03C987A6FF9CFFF93AB6FCC21C62D928	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Jordal, Bjarte H.	Jordal, Bjarte H. (2018): Hidden gems in museum cabinets: new species and new distributional records of Scolytodes (Coleoptera: Scolytinae). Zootaxa 4504 (1): 76-104, DOI: 10.11646/zootaxa.4504.1.4
03C987A6FF82FFF93AB6F9F71C62DE6F.text	03C987A6FF82FFF93AB6F9F71C62DE6F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Scolytodes capillus Jordal 2018	<div><p>Scolytodes capillus Jordal, sp. nov.</p><p>urn:lsid:zoobank.org:act: D86D9BBB-D43C-4C25-A8A4-F37FD5FB7B88</p><p>(Figs 46, 49, 52)</p><p>Type material. Holotype, female: Panama, Canal Zone, Albrook Forest Site, Fort Clayton, May 2¯3 1968, blacklight trap, Hutton leg. Allotype, male: same data as HT, except lot no. 165, Jan 11/12 1968. Paratypes (3 females): same data as HT (1), except March 7/8 (1), May 25/26 1967, Hutton &amp; Llaurado (1). HT, AT and one PT deposited in USNM, 2 PTs in UTIC .</p><p>Diagnosis. Interstriae 10 sharply elevated to level of ventrite 3; protibiae without an additional mesal tooth. Distinguished from the most similar species, S. phoebeae Wood and S. amoenus Wood, by the much larger size and the broadly rounded, sub-truncated pronotum and elytral apex, and further from S. phoebeae by distinct asperities on pronotum, and from S. amoenus by the much longer and abundant setae in the female frons.</p><p>Description female. Length 3.1¯ 3.3 mm, 2.1¯2.2 × longer than wide; colour dark brown. Head. Eyes entire, separated above by 2.7¯2.8 × their width. Frons presumably flat from epistoma to vertex, entirely covered by a dense long brush of golden setae from vertex, along anterior eye margin, reaching epistoma. Antennal club setose with two sutures, first suture slightly procurved in the middle, marked by a dark septum, second suture procurved, obscurely marked. Funiculus 6-segmented. Pronotum weakly reticulated, subshining, shallow fine punctures separated by 1¯2 × their diameter, reaching anterior margin but more obscure on anterior half between low asperities. Vestiture consisting of 2¯4 erect setae (2/4–0–0). Elytra smooth and shiny, striae not impressed, punctures small, shallow, in irregular rows, separated by 1¯3 × their diameter; interstriae 4¯6 × wider than striae, punctures slightly smaller than strial punctures, confused, on declivity inseparable from and confused with strial punctures. Glabrous. Legs. Procoxae separated by 0.2¯0.3 × and mesocoxae 0.7¯0.8 × the width of one procoxa. Protibiae narrow, distal tooth 1 larger than 2, with 4-5 additional low transverse rugae or irregular granules along towards base; protibial mucro obtuse. Meso- and metatibiae with 8¯9 and 6¯7 lateral, socketed teeth, on distal half and third, respectively. Ventral vestiture. Setae on metasternum and metanepisternum simple.</p><p>Male. Similar to females except smaller (length 2.9 mm), frons convex, eyes separated above by 3.2 × their width, procoxae separated by 0.4 × their width.</p><p>Key (Wood 1982). Keys to couplet 10, with no further match.</p><p>Etymology. The name capillus is a Latin nominative noun (singular), meaning head of hair. The noun is a composite of the nouns caput, meaning head, and pilus, meaning hair.</p><p>Biology and distribution. This species is only known from the lowland type locality in Panama, attracted to black light.</p></div>	https://treatment.plazi.org/id/03C987A6FF82FFF93AB6F9F71C62DE6F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Jordal, Bjarte H.	Jordal, Bjarte H. (2018): Hidden gems in museum cabinets: new species and new distributional records of Scolytodes (Coleoptera: Scolytinae). Zootaxa 4504 (1): 76-104, DOI: 10.11646/zootaxa.4504.1.4
03C987A6FF82FFF83AB6FE391FEEDB50.text	03C987A6FF82FFF83AB6FE391FEEDB50.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Scolytodes rufus Jordal 2018	<div><p>Scolytodes rufus Jordal, sp. nov.</p><p>urn:lsid:zoobank.org:act: BD49DB5D-F5D8-49A6-88CA-ACFECE4E6B5C</p><p>(Figs 47, 50, 53)</p><p>Type material. Holotype, female: Brazil, Bahia Camacan, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-39.5635&amp;materialsCitation.latitude=-15.390483" title="Search Plazi for locations around (long -39.5635/lat -15.390483)">Serra Bonita Reserve</a>, 800m, GIS 15°23.429S, 39°33.810W, 10 May 2013, Cognato, Smith and Flechtmann, leg. Allotype and 8 paratypes: same data as HT. HT, AT and 2 PTs deposited in MZUSP, 2 PTs in UNESP, 4 PTs in ZMBN .</p><p>Diagnosis. Interstriae 10 sharply elevated to the level of metacoxae; protibiae with a small additional mesal tooth near tarsal insertion. Pronotum with coarse asperities on anterior two-thirds; female frons with long protuberant setae around a longitudinally shiny and impunctate callus.</p><p>Description female. Length 1.4¯ 1.6 mm, 2.4¯2.5 × longer than wide; colour light to dark reddish brown.</p><p>Head. Eyes entire, separated above by 2.5¯2.7 × their width. Frons flat from vertex to epistoma, strongly punctured in entire area except a longitudinal, narrow, impunctate callus on lower half; vestiture consisting of long golden setae pointing forward, forming a complete circle. Antennal club with two oblique sutures marked by fine setae. Funiculus 5-segmented. Pronotum weakly reticulated, subshining, shallow fine punctures separated by 2¯4 × their diameter, replaced on anterior two-thirds by coarse asperities. Vestiture consisting of 4 erect setae (4–0–0). Elytra smooth and shiny, striae not impressed, punctures moderately large, shallow, separated in irregular rows by 1¯2 × their diameter; interstriae 3 × wider than striae, punctures slightly smaller, in rows. Vestiture consisting of less than 20 erect setae on interstriae 3, 7 and 9. Legs. Procoxae separated by 0.6 × and mesocoxae 0.9 × the width of one procoxa. Protibiae narrow, distal tooth 1 slightly larger than 2, with 3¯4 additional granules or small spines towards base; protibial mucro obtuse. Meso- and metatibiae with 6¯7 lateral, socketed teeth, on distal half and third, respectively. Ventral vestiture. Setae on upper metasternum and metanepisternum trifid, occasionally bifid, simple below. Setae on ventrites 3¯5 coarse, directed caudally.</p><p>Male. Similar to females except smaller (length 1.3¯ 1.4 mm, 2.3 × longer than wide), frons convex with about 10 erect pointing setae on lower half.</p><p>Key (Wood 2007). Keys to couplet 59, with no further match.</p><p>Etymology. The name rufus is a Latin nominative masculine adjective, meaning reddish or ruddy, referring to the distinctly reddish and dark yellow colour of apparently fully matured adult beetles.</p><p>Biology and distribution. This species is only known from the type locality in the Atlantic rainforest of Brazil, collected from an unknown host plant.</p></div>	https://treatment.plazi.org/id/03C987A6FF82FFF83AB6FE391FEEDB50	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Jordal, Bjarte H.	Jordal, Bjarte H. (2018): Hidden gems in museum cabinets: new species and new distributional records of Scolytodes (Coleoptera: Scolytinae). Zootaxa 4504 (1): 76-104, DOI: 10.11646/zootaxa.4504.1.4
03C987A6FF83FFFA3AB6FBE21C54D9C4.text	03C987A6FF83FFFA3AB6FBE21C54D9C4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Scolytodes johnsoni Jordal 2018	<div><p>Scolytodes johnsoni Jordal, sp. nov.</p><p>urn:lsid:zoobank.org:act: 46FBAEEF-D79E-4A62-8011-74066380862C</p><p>(Figs 48, 51, 54)</p><p>Type material. Holotype, female: French Guiana, Kaw Mts, ex Cecropia petioles, 4.6.2015, A. Johnson . Allotype and 7 paratypes: same data as holotype. AT, HT and 2 PTs deposited in FSCA, 4 PTs in ZMBN .</p><p>Diagnosis. Interstriae 10 sharply elevated to the level of ventrite 4; protibiae without an additional mesal tooth, lateral tooth 2 longer than tooth 1. Female frons only slightly impressed, with dense, golden, short, semi-erect setae in an area from half way between upper eyes and antennal insertion, to epistoma. Distinguished from the three most similar species S. suturalis Wood, S. nitidissimus (Eggers) and S. ingae (Backman) by the smoother male frons not having a transverse elevation on epistoma, and by the less impressed female frons. It is further distinguished from S. suturalis Wood by the smaller size.</p><p>Description female. Length 1.9¯ 2.2 mm, 2.0¯2.1 × longer than wide; colour black. Head. Eyes entire, separated above by 2.6¯2.7 × their width. Frons weakly impressed from half way between upper level of eyes and antennal insertion to epistoma, scattered small punctures above, impressed area more densely punctured below and covered by short semi-erect golden setae. Antennal club with two obliquely procurved sutures marked by dense setae, segment one and two partly corneous and shiny. Funiculus 6-segmented. Pronotum weakly reticulated, subshining, punctures small, deep, separated by 2¯3 × their diameter, reduced in size anteriorly and mixed with faint asperities on anterior fourth. Vestiture consisting of 8 erect setae (4–2–2). Elytra shiny, striae 1 impressed on disc, 1¯3 impressed on declivity, punctures small, separated in irregular rows by 2¯3 × their diameter, additional microscopic punctures present on the anterior edge of each main puncture; interstriae about 5 × wider than striae, punctures smaller, widely separated, in rows, many associated with supporting granules for setae. Vestiture consisting of interstrial rows of erect, coarse, almost spatulate setae; microscopic setae associated with strial micropunctures. Legs. Procoxae and mesocoxae separated by 1.1¯1.3 × the width of one procoxa. Protibiae broad, distal tooth 1 slightly shorter than 2, with 3¯4 additional granules or small spines decreasing in size towards base, a fine semi-transparent extension of cuticle connecting teeth 2¯4; protibial mucro large, curved posteriorly. Mesotibiae with 6¯9 lateral, socketed teeth on distal two-thirds, metatibiae with 6¯7 teeth on distal half. Ventral vestiture. Setae on metasternum and metanepisternum long simple; occasional trifid setae along the anterior margin of upper metasternum. Setae on ventrites 3¯5 coarse, directed caudally.</p><p>Male. Similar to females except slightly smaller (length 1.6¯ 2.2 mm, 1.9¯2.0 × longer than wide), frons convex, generally smooth, slightly transversely wrinkled on lower third, with fine punctures from vertex to epistoma; glabrous, except scant setae on epistoma. Eyes separated by 2.4¯2.6 × their width.</p><p>Key (Wood 2007). Keys to couplet 9, S. imitans (= S. nitidissimus). However, the description of S. imitans in Wood (2007), including the key, does not match the holotype from Guadeloupe. It is possible that specimens in Wood’s collection identified as S. imitans from Venezuela (Wood 2007) is conspecific to S. johnsoni .</p><p>Etymology. Named after the collector, Andrew Johnson, a recently graduated student of bark beetle systematics.</p><p>Biology and distribution. This species is only known from the type locality in French Guyana, collected from Cecropia petioles. This species belongs to a complex of many very similar species, all breeding in the leaf petioles of Cecropia .</p></div>	https://treatment.plazi.org/id/03C987A6FF83FFFA3AB6FBE21C54D9C4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Jordal, Bjarte H.	Jordal, Bjarte H. (2018): Hidden gems in museum cabinets: new species and new distributional records of Scolytodes (Coleoptera: Scolytinae). Zootaxa 4504 (1): 76-104, DOI: 10.11646/zootaxa.4504.1.4
03C987A6FF81FFFA3AB6F99C1831DE1B.text	03C987A6FF81FFFA3AB6F99C1831DE1B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Scolytodes longipilus Jordal 2018	<div><p>Scolytodes longipilus Jordal, sp. nov.</p><p>urn:lsid:zoobank.org:act: E284AE39-7EA0-4477-B1E5-F30CC3717C9D</p><p>(Figs 55, 58, 61)</p><p>Type material. Holotype: Colombia, Cundinamarca, P. &amp; B. Wygodzinky. Road Bogota to Choachi, 3250¯3000 meters, July 19, 1967 . Holotype deposited in AMNH .</p><p>Diagnosis. Interstriae 10 obscurely elevated to the level of metacoxa; protibiae with a sharp additional mesal tooth just inside the lateral margin between tooth 1 and 2; tooth 2 socketed. Strial punctures on elytra confused, volcano shaped; interstriae not clearly marked, cuticle wrinkled. Distinguished from all species with similarly granulated pronotum by the much longer interstrial setae (except Scolytodes opacus Wood, which has impressed striae 1 and tuberculate interstriae 1¯3 on posterior half of elytra), and by the much larger size (except Scolytodes major (Eggers), which has short elytral setae and smooth cuticle).</p><p>Description, female? Length 4.1 mm, 2.1 × longer than wide; colour black. Head. Eyes entire, slightly bulging, separated above by 3.1 × their width. Frons flattened, smooth with small punctures, strongly reticulated, a low carina run from each lateral edge of the epistoma towards the anterior margin of the eyes near antennal insertion; vestiture consisting of scattered fine setae. Antennal club with two triangularly procurved sutures marked by dense short setae. Funiculus 6-segmented. Pronotum strongly reticulated, dull; punctures small, volcano shaped, reaching anterior margin, puncture on anterior two-thirds associated with low, shiny, crescent shaped asperities. Vestiture consisting of long, fine setae in no particular pattern (0–0–0). Elytra sub-shining, surface wrinkled, strial and interstrial punctures of variable size, confused, volcano shaped. Vestiture consisting of interstrial rows of erect, long, fine setae, and intermixed strial and interstrial setae about one-third as long. Legs. Procoxae separated by 0.5 × and mesocoxae 1.2 × the width of one procoxa. Protibiae broad on distal half, distal teeth 1 and 2 of equal length, tooth 2 with socketed denticle exposed, 4¯5 additional small spines towards base; protibial mucro large, curved posteriorly. Mesotibiae with 10 lateral, socketed teeth on distal two-thirds, metatibiae with 10¯11 teeth on distal half. Ventral vestiture. Setae on ventrites, metasternum and metanepisternum long, simple.</p><p>Key (Wood 2007). Keys to couplet 123, S. opacus, but stria 1 is not impressed on declivity, interstriae are without tubercles, and size is much larger.</p><p>Etymology. The species name is composed by longi, a Latin masculine adjective in its genitive form, meaning extended or prolonged, and pilus, a Latin masculine noun, meaning hair. The name refer to the exceedingly long interstrial setae on the elytra, with shorter setae mixed in between</p><p>Biology and distribution. This species is only known from the high altitude type locality in Colombia</p></div>	https://treatment.plazi.org/id/03C987A6FF81FFFA3AB6F99C1831DE1B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Jordal, Bjarte H.	Jordal, Bjarte H. (2018): Hidden gems in museum cabinets: new species and new distributional records of Scolytodes (Coleoptera: Scolytinae). Zootaxa 4504 (1): 76-104, DOI: 10.11646/zootaxa.4504.1.4
03C987A6FF81FFFC3AB6FEA51FAADB84.text	03C987A6FF81FFFC3AB6FEA51FAADB84.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Scolytodes prolatus , Jordal 2018	<div><p>Scolytodes prolatus Jordal, sp. nov.</p><p>urn:lsid:zoobank.org:act: F7E8C491-2320-4FCB-BF2A-50D050505E99</p><p>(Figs 56, 59, 62)</p><p>Type material. Holotype, male: Honduras, Franc. Mor., P.N. La Tigra, 23.2 km N. Tegucigalpa, 1950 m, oak/ cloud forest, 29.06.1994, R. Anderson . Paratype, male: Olancho, La Muralla, 14 km N, La Union, 1510-1550 m, transition cloud for. Litt. Berl., 16– 17.8.1994, R. Anderson. Types deposited in FSCA .</p><p>Diagnosis. Interstriae 10 sharply elevated to the level of metacoxa, a faint trace continue to level of ventrite 2; protibiae with one additional mesal tooth just inside the lateral margin between tooth 1 and 2; lateral tooth 2 socketed. Closely related to S. clusiacolens, but the new species has elytral striae more deeply impressed, the male pronotum is strongly dilated, and very fine setae are present only on odd interstriae.</p><p>Description, male. Length 2.7–2.9 mm, 2.5 × longer than wide; colour black. Head. Eyes sinuate, separated above by 4.0–4.2 × their width. Frons convex, finely reticulated, dull, smooth with many small, shallow punctures; epistomal area strongly impressed; vestiture consisting of scattered fine setae (in punctures). Antennal club with two obliquely procurved sutures marked by dense short setae. Funiculus 6-segmented. Pronotum very finely reticulated, shiny; punctures small, separated on average by twice their diameter, punctures replaced by fine asperities on anterior one-sixth; anterior row of asperities forming a finely serrated margin. Vestiture consisting of very fine, long, setae (4–2–2, abraded on PT). Elytra smooth, sub-shining, striae narrowly, deeply impressed, punctures slightly elongated, near contiguous on disc, smaller and more widely separated on declivity; interstriae 4–5 × wider than striae, punctures much smaller and widely separated. Vestiture consisting of rows of widely spaced, erect, long, fine setae on odd interstriae, intermixed with minute interstrial and strial setae on declivity.</p><p>Legs. Procoxae separated by 0.6¯0.7 × and mesocoxae 0.9–1.0 × the width of one procoxa. Protibiae narrow, distal teeth 1 and 2 of equal length, 4¯5 additional small granules along lateral edge towards base; protibial mucro not present (obtuse). Meso- and metatibiae with 6–7 lateral, socketed teeth on distal half and thirds, respectively. Ventral vestiture. Scattered setae on ventrites, metasternum and metanepisternum fine, simple.</p><p>Key (Wood 1982). Keys to couplet 25b, S. clusiacolens Wood, but differ as noted in the diagnosis.</p><p>Etymology. The name prolatus is a Latin masculine nominative participle of the verb profero, meaning extending, stretch out or extend, referring to the dilated anterior half of the pronotum.</p><p>Biology and distribution. This species is only known from two medium altitude localities in Honduras, in cloud forests dominated by oak. One specimen was collected by extraction from a litter sample in a Berlese funnel. The close relative, S. clusiacolens, breeds in Clusia .</p></div>	https://treatment.plazi.org/id/03C987A6FF81FFFC3AB6FEA51FAADB84	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Jordal, Bjarte H.	Jordal, Bjarte H. (2018): Hidden gems in museum cabinets: new species and new distributional records of Scolytodes (Coleoptera: Scolytinae). Zootaxa 4504 (1): 76-104, DOI: 10.11646/zootaxa.4504.1.4
03C987A6FF87FFFF3AB6FC5C1E8AD90D.text	03C987A6FF87FFFF3AB6FC5C1E8AD90D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Scolytodes bicarinatus Jordal 2018	<div><p>Scolytodes bicarinatus Jordal, sp. nov.</p><p>urn:lsid:zoobank.org:act: 6EB59A17-F5AA-4DAF-9116-73132CE8C7E8</p><p>(Figs 57, 60, 63)</p><p>Type material. Holotype, female: Panama, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-79.377&amp;materialsCitation.latitude=9.211" title="Search Plazi for locations around (long -79.377/lat 9.211)">Cerro Azul</a>, Cerro Jefe, [GIS] 9.211, -79.377, 827m, 18.Aug.2008, SM Smith, AD Smith, AR Gillogly, leg, ex. very wet Cecropia petioles . Allotype, male: same data as holotype. Types deposited in MIUP .</p><p>Diagnosis. Interstriae 10 sharply elevated to apex; protibiae without additional mesal tooth. Female frons with a pair of broad, subcontiguous carinae which affiliates this species with those in the cecropiavorus species group (Jordal 1998b). Distinguished from all species in this group by the male frons having no impression just above the epistoma, and by the the female frons which has shorter vestiture and broader carinae.</p><p>Description, female. Length 1.9 mm, 2.2 × longer than wide; colour dark brown to black. Head. Eyes entire, separated above by 2.0 × their width. Frons weakly concave, with a pair of broad, round, parallel carinae on lower two-thirds; surface finely punctured above and between carinae, vestiture consisting of scattered fine setae in each puncture, and longer whitish sub-plumose setae surrounding the impressed area, none of which reaching the epistomal margin. Antennal club setose, no sutures marked. Funiculus 6-segmented. Pronotum very finely reticulated, shiny; punctures small, shallow, reaching anterior margin, separated on average by 2¯3 × their diameter. Glabrous, except six erect setae (4–0–2). Elytra smooth, shining, striae not impressed, punctures in rows, small, shallow, separated by 2 × their diameter; interstriae 4–5 × wider than striae, punctures in rows, almost as large as those in striae, more widely separated. Glabrous. Legs. Procoxae separated by 0.6 × and mesocoxae 0.8 × the width of one procoxa. Protibiae broaden distally, distal teeth 1 and 2 of subequal length and curved posteriorly, edge between these teeth slightly extended, with 3¯4 triangular rugae along lateral edge decreasing in size towards base; protibial mucro small, curved posteriorly. Meso- and metatibiae with 7–8 lateral, socketed teeth on distal half. Ventral vestiture. Scattered, fine, simple setae on ventrites, metasternum and metanepisternum.</p><p>Male. Similar to female except slightly smaller (length 1.7 mm), frons convex, smooth, flattened on epistoma, with fine punctures; vestiture consisting of scant minute setae, slightly longer and denser near epistomal lobe.</p><p>Key (Wood 1982). Keys to couplet 13 or 16, S. cecropiavorus Wood, or S. cecropii (Scedl) but differ in the female and male frons as noted in the diagnosis.</p><p>Etymology. The name bicarinatus is a masculine Latin nominative adjective, meaning bicarinate, referring to the two keels running in parallel in the female frons.</p><p>Biology and distribution. This species is only known from the type locality in Panama, in wet secondary forest at medium altitude dominated by Cecropia trees typical for forest gaps.</p></div>	https://treatment.plazi.org/id/03C987A6FF87FFFF3AB6FC5C1E8AD90D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Jordal, Bjarte H.	Jordal, Bjarte H. (2018): Hidden gems in museum cabinets: new species and new distributional records of Scolytodes (Coleoptera: Scolytinae). Zootaxa 4504 (1): 76-104, DOI: 10.11646/zootaxa.4504.1.4
03C987A6FF84FFFF3AB6FAED19B6DE05.text	03C987A6FF84FFFF3AB6FAED19B6DE05.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Scolytodes venustulus Wood 1967	<div><p>Scolytodes venustulus Wood</p><p>(Figs 64, 66, 68)</p><p>Scolytodes venustulus Wood, 1967: 124 .</p><p>Material examined. Holotype, male: Panama, Volcan Chiriqui, 5500ft, I-14-1964, SL Wood, ex unknown sapling, [USNM]. New record: Costa Rica, San Jose, Zurqui de Moravia, 1600m, viii-25-31-1998, FIT, C.W. &amp; L.B.O’Brien [FSCA].</p><p>Diagnosis. Interstriae 10 sharply elevated to level of ventrite 1; protibiae with a distinct additional mesal tooth near tarsal insertion. Elytral striae with punctures in longitudinal pairs, spatulate setae on uneven declivital interstriae only.</p><p>Description female. Length 1.6 mm, 2.4 × longer than wide; colour brown. Head. Eyes entire, separated above by 2.1 × their width. Frons convex on upper two-thirds, slightly impressed on median third of lower onethird, forming a triangular impression; surface reticulate on vertex, shiny below, with few scattered punctures; vestiture consisting of few short golden setae in impressed area, setae longer closer to epistoma. Antennal club with two obliquely procurved sutures marked by long white setae, club generally setose. Funiculus 6-segmented. Pronotum strongly reticulate, punctures spaced by 1¯2 × their diameter, very faint asperities near anterior margin. Vestiture consisting of 4 anterior erect setae (4–0–0). Elytra generally smooth and shiny, striae not impressed, punctures small, in longitudinal pairs, each pair separated in irregular rows by 1¯2 × the diameter of one puncture; interstriae 3 × wider than striae, punctures of same size as in striae, separated by 3¯4 × their diameter. Vestiture consisting of erect, slightly spatulate setae on mainly declivital interstriae 1, 3, 5 and 7, and much shorter, fine strial and interstrial setae. Legs. Procoxae separated by 0.5 × and mesocoxae 0.9 × the width of one procoxa. Protibiae relatively short, distal teeth 1 and 2 of equal length, with 3 tiny additional spines or granules along the edge towards base; protibial mucro thin, short and curved posteriorly. Meso- and metatibiae with 6 lateral, rather coarse, socketed teeth on distal half. Ventral vestiture. Setae on metasternum and metanepisternum simple.</p><p>Male. Frons less impressed, glabrous. The male holotype has slightly larger elytral punctures.</p><p>Biology and distribution. This species is known from the type locality at medium altitude in Panama (Chiriquí, 1800m), and in Costa Rica (Zurquí de Moravia, 1600m), presumably in cloud forest.</p></div>	https://treatment.plazi.org/id/03C987A6FF84FFFF3AB6FAED19B6DE05	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Jordal, Bjarte H.	Jordal, Bjarte H. (2018): Hidden gems in museum cabinets: new species and new distributional records of Scolytodes (Coleoptera: Scolytinae). Zootaxa 4504 (1): 76-104, DOI: 10.11646/zootaxa.4504.1.4
03C987A6FF84FFF13AB6FEDC1DBEDA7F.text	03C987A6FF84FFF13AB6FEDC1DBEDA7F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Scolytodes pelicipennis (Schedl 1952)	<div><p>Scolytodes pelicipennis (Schedl)</p><p>(Figs 65, 67, 69)</p><p>Hexacolus pelicipennis Schedl, 1952: 356 .</p><p>Material examined. Holotype, male: Costa Rica, Jimenez, Halbins. [Peninsula de] Osa, E. Reimoser leg [NHMW]. New record: Mexico, Oaxaca, Finca Montecarlo, Sta. Ma. Xadani, [GIS] 15.993, -96.104, 916m, 24-VI- 2009, ex Ficus sp., T.H. Atkinson THA-879. Host: trunk of Ficus sp, TH Atkinson, THA-793 [UTIC].</p><p>Diagnosis. Interstriae 10 sharply elevated to the level of metacoxa, protibiae with a small additional mesal tooth near tarsal insertion. Elytral striae increasingly impressed posteriorly, punctures increasingly larger. Distinguished from S. impressus Wood (Costa Rica) and S. striatulus Wood (Caribbean) by having only a few erect setae on odd-numbered interstriae only, and by the much larger strial punctures, and further from S. impressus by the more abundant pronotal asperities, and long abundant setae in the female frons.</p><p>Description female. Length 1.4¯ 1.6 mm, 2.2¯2.3 × longer than wide; colour dark reddish brown. Head. Eyes entire, separated above by 2.6¯2.8 × their width. Frons flattened, with a central impunctate area surrounded by long golden setae, progressively shorter towards the impunctate area. Antennal club with two oblique sutures marked by setae. Funiculus 6-segmented. Pronotum shiny, large punctures on posterior fourth separated by less than half their diameter, replaced on anterior three-fourth by coarse asperities, forming a low serrated ridge on anterior margin. Vestiture consisting of 8 erect setae (4–2–2) and shorter semi-recumbent setae in area with asperities. Elytra shiny, striae strongly and progressively impressed posteriorly, punctures large and deep, subconfluent; interstriae as wide as striae, punctures small, in rows. Vestiture consisting of scattered long spatulate setae on odd-numbered interstriae, with short fine setae on each declivital interstriae. Legs. Procoxae separated by 0.3¯0.4 × and mesocoxae 0.7¯0.8 × the width of one procoxa. Protibiae with distal tooth 1 slightly larger than 2, with 4 additional granules along the lateral edge towards base; protibial mucro short, curved posteriorly. Meso- and metatibiae with 6 and 5 lateral, socketed teeth on distal half. Ventral vestiture. Setae on metasternum and metanepisternum mainly simple, bifid close to anterior margin.</p><p>Male. Similar to females except slightly smaller (1.3¯ 1.5 mm), and the frons is convex, strongly reticulate and punctured, with fine short setae near epistoma.</p><p>Biology and distribution. Previously known only from a single male specimen from the west coast of Costa Rica. This is the first record from Mexico and the first of a female. Specimens were collected under bark of a Ficus tree.</p></div>	https://treatment.plazi.org/id/03C987A6FF84FFF13AB6FEDC1DBEDA7F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Jordal, Bjarte H.	Jordal, Bjarte H. (2018): Hidden gems in museum cabinets: new species and new distributional records of Scolytodes (Coleoptera: Scolytinae). Zootaxa 4504 (1): 76-104, DOI: 10.11646/zootaxa.4504.1.4
03C987A6FF8AFFF13AB6FCA11907DDC5.text	03C987A6FF8AFFF13AB6FCA11907DDC5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Scolytodes bolivianus Eggers 1928	<div><p>Scolytodes bolivianus Eggers, 1928</p><p>Scolytodes aequipunctatus Eggers, 1943 . syn. nov.</p><p>The holotype of S. aequipunctatus at MNHN was compared to the holotype at USNM (photo: http:// www.barkbeetles.info) and one of Eggers paratypes (co-types) (NHMW) of bolivianus and these were identical except that S. bolivianus types are younger individuals which are more lightly coloured.</p></div>	https://treatment.plazi.org/id/03C987A6FF8AFFF13AB6FCA11907DDC5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Jordal, Bjarte H.	Jordal, Bjarte H. (2018): Hidden gems in museum cabinets: new species and new distributional records of Scolytodes (Coleoptera: Scolytinae). Zootaxa 4504 (1): 76-104, DOI: 10.11646/zootaxa.4504.1.4
03C987A6FF8AFFF13AB6FD9C1924DF5A.text	03C987A6FF8AFFF13AB6FD9C1924DF5A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Scolytodes canalis Wood 1974	<div><p>Scolytodes canalis Wood, 1974</p><p>Scolytodes amabilis Wood, 1975 . syn. nov.</p><p>Wood (1975) described S. amabilis with no comparison to S. canalis, a species which he described the year before. Female holotypes of the two species were collected in the same locality and are nearly identical; they may differ in the female frons by the length of the long median setae and by the less pronounced rows of elevated granules on the lower part, but these differences are very small. A new collection from Guatemala (Table 1) indicates slight variation in both traits. The two high altitude locations are only separated by 300 km.</p></div>	https://treatment.plazi.org/id/03C987A6FF8AFFF13AB6FD9C1924DF5A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Jordal, Bjarte H.	Jordal, Bjarte H. (2018): Hidden gems in museum cabinets: new species and new distributional records of Scolytodes (Coleoptera: Scolytinae). Zootaxa 4504 (1): 76-104, DOI: 10.11646/zootaxa.4504.1.4
03C987A6FF8AFFF13AB6FAD11E63DC07.text	03C987A6FF8AFFF13AB6FAD11E63DC07.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Scolytodes ingae (Blackman 1943) Jordal 2018	<div><p>Scolytodes ingae (Blackman 1943), stat. nov.</p><p>Prinosceles ingae Blackman, 1943, resurrected from synonymy in S. maurus Blandford.</p><p>Scolytodes trigonus Jordal, 2013 . syn. nov.</p><p>The holotype of Prinosceles ingae, deposited in USNM (photo: http://www.barkbeetles.info), was compared to the recently described S. trigonus and found to be identical. Scolytodes ingae was placed in synonymy with S. maurus by Wood (2007). However, S. maurus has a largely smooth pronotum, and a smooth male frons. The male holotypes of S. ingae and S. trigonus have a triangular field in the frons that becomes rough and irregularly elevated near the epistoma. They also have a tiny puncture associated with each main strial puncture on the elytra, and the interstrial granules are much larger and consistent compared to S. maurus . Another synonym of S. maurus in South America, S. medius (Eggers), looks like S. maurus, but cannot be evaluated properly due to the frons is glued in both the type (USNM) and paratypes (NHMW).</p></div>	https://treatment.plazi.org/id/03C987A6FF8AFFF13AB6FAD11E63DC07	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Jordal, Bjarte H.	Jordal, Bjarte H. (2018): Hidden gems in museum cabinets: new species and new distributional records of Scolytodes (Coleoptera: Scolytinae). Zootaxa 4504 (1): 76-104, DOI: 10.11646/zootaxa.4504.1.4
03C987A6FF8AFFF23AB6FFE61C83DBB3.text	03C987A6FF8AFFF23AB6FFE61C83DBB3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Scolytodes major (Eggers 1928)	<div><p>Scolytodes major (Eggers, 1928)</p><p>Prionosceles major Eggers, 1928: 86 .</p><p>Scolytodes argentinensis (Eggers, 1928: 87) . syn. nov.</p><p>Scolytodes similis (Eggers, 1928: 87) . syn. nov.</p><p>Scolytodes boliviensis (Eggers, 1928: 88), synonymy of S. similis (see Bright 2014). Types of the four species in USNM (photo: http://www.barkbeetles.info) were compared to paratypes at NHMW and have only slight variation in size, ranging from 2.5 mm ( similis) to 3.9 mm ( major). The shape and placement of setae and punctures are identical and the minor variation observed in the frons and pronotum are typical sexual dimorphism. Type localities in the four species are closely situated locations in Bolivia and Argentina, but the full range of this species is much larger with several new country records reported here (see below). Everything else being equal, the name S. major (Eggers, 1928) is given priority as the first mentioned name of the four taxa (page priority).</p></div>	https://treatment.plazi.org/id/03C987A6FF8AFFF23AB6FFE61C83DBB3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Jordal, Bjarte H.	Jordal, Bjarte H. (2018): Hidden gems in museum cabinets: new species and new distributional records of Scolytodes (Coleoptera: Scolytinae). Zootaxa 4504 (1): 76-104, DOI: 10.11646/zootaxa.4504.1.4
