taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
03CDBE749318B507CE72FD15FEEAF8F4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071279/files/figure.png	https://doi.org/10.5281/zenodo.8071279	FIG. 10. — Dorippinae H. Milne Edwards, 1837 n. stat.: habitus: A, B, Dorippe quadridens (Fabricius,1793):A, ♂ 36.3 × 38.1 mm, China Sea near Singapore, Hee Huat, ZRC 1984. 6308; B, ♀ 29.5 × 30 mm, NW Madagascar, Ambaro Bay, MNHN-IU-2018-5193 (= MNHN-B18279). C, D, Dorippe sinica Chen, 1980, China, Guangdong, Nanao Island, ZRC 1999.0470: C, ♂ 36.2 × 39.5 mm; D, ♀ 34.8 × 38.9 mm, specimen brushed.	FIG. 10. — Dorippinae H. Milne Edwards, 1837 n. stat.: habitus: A, B, Dorippe quadridens (Fabricius,1793):A, ♂ 36.3 × 38.1 mm, China Sea near Singapore, Hee Huat, ZRC 1984. 6308; B, ♀ 29.5 × 30 mm, NW Madagascar, Ambaro Bay, MNHN-IU-2018-5193 (= MNHN-B18279). C, D, Dorippe sinica Chen, 1980, China, Guangdong, Nanao Island, ZRC 1999.0470: C, ♂ 36.2 × 39.5 mm; D, ♀ 34.8 × 38.9 mm, specimen brushed.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE749318B507CE72FD15FEEAF8F4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071285/files/figure.png	https://doi.org/10.5281/zenodo.8071285	FIG. 12. — Dorippinae H. Milne Edwards,1837 n. stat.. Dorippe quadridens (Fabricius,1793):A-D, ovigerous ♀ 38.1 × 33.7 mm, South China Sea,Hee Huat,ZRC 1984.6307:A, habitus; B, anterior region, ventral view; C, thoracic sternum and pleon; D, vulvae. E, ovigerous ♀ 28.0 × 29.0 mm, NW Madagascar, Ambaro Bay, MNHN-IU-2018-5193 (= MNHN-B18279): thoracic sternum and vulvae.	FIG. 12. — Dorippinae H. Milne Edwards,1837 n. stat.. Dorippe quadridens (Fabricius,1793):A-D, ovigerous ♀ 38.1 × 33.7 mm, South China Sea,Hee Huat,ZRC 1984.6307:A, habitus; B, anterior region, ventral view; C, thoracic sternum and pleon; D, vulvae. E, ovigerous ♀ 28.0 × 29.0 mm, NW Madagascar, Ambaro Bay, MNHN-IU-2018-5193 (= MNHN-B18279): thoracic sternum and vulvae.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE749318B507CE72FD15FEEAF8F4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071289/files/figure.png	https://doi.org/10.5281/zenodo.8071289	FIG. 14. — Dorippinae H. Milne Edwards,1837 n. stat.:A-G: Dorippe frascone (Herbst,1785),Philippines,Panglao Island,ZRC 2008.0076:A-D, ♂ 29.7 × 31.0 mm, A, habitus; B, ventral view; C, thoracic sternum with pleon; D, G1; E-G, prepubertal ♀ 23.3 × 24.4 mm: E, habitus;F, ventral view; G, vulvae; H, I, Dorippe irrorata Manning & Holthuis, 1986, holotype, ♂ 21.5 × 22.0 mm, Andaman Sea, south of Mergui Archipelago, USNM 172495: H, habitus; I, ventral surface.	FIG. 14. — Dorippinae H. Milne Edwards,1837 n. stat.:A-G: Dorippe frascone (Herbst,1785),Philippines,Panglao Island,ZRC 2008.0076:A-D, ♂ 29.7 × 31.0 mm, A, habitus; B, ventral view; C, thoracic sternum with pleon; D, G1; E-G, prepubertal ♀ 23.3 × 24.4 mm: E, habitus;F, ventral view; G, vulvae; H, I, Dorippe irrorata Manning & Holthuis, 1986, holotype, ♂ 21.5 × 22.0 mm, Andaman Sea, south of Mergui Archipelago, USNM 172495: H, habitus; I, ventral surface.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE749318B507CE72FD15FEEAF8F4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071291/files/figure.png	https://doi.org/10.5281/zenodo.8071291	FIG. 15. — Dorippinae H. Milne Edwards,1837 n. stat.: A-D, J, Dorippe trilobata Manning,1993,holotype, ♂ 20.5 × 21.1 mm, off Mitchell River,Admiralty Gulf,AM P27124: A, habitus; B, ventral surface; C, carapace; D, pleon; J, callosity. E-I: Dorippe glabra Manning, 1993, holotype, ovigerous ♀ 24.3 × 26.7 mm, Chambers Bay, Australia, AM P13363: E, habitus with detached right cheliped, right P2 and P3, left P3; F, carapace; G, ventral surface; H, pleon; I, callosity.	FIG. 15. — Dorippinae H. Milne Edwards,1837 n. stat.: A-D, J, Dorippe trilobata Manning,1993,holotype, ♂ 20.5 × 21.1 mm, off Mitchell River,Admiralty Gulf,AM P27124: A, habitus; B, ventral surface; C, carapace; D, pleon; J, callosity. E-I: Dorippe glabra Manning, 1993, holotype, ovigerous ♀ 24.3 × 26.7 mm, Chambers Bay, Australia, AM P13363: E, habitus with detached right cheliped, right P2 and P3, left P3; F, carapace; G, ventral surface; H, pleon; I, callosity.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE749318B507CE72FD15FEEAF8F4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071283/files/figure.png	https://doi.org/10.5281/zenodo.8071283	FIG. 11. — Dorippinae H. Milne Edwards, 1837 n. stat.. Dorippe quadridens (Fabricius, 1793): A-C, ♂ 38.1 × 36.3 mm, China Sea near Singapore, Hee Huat, ZRC 1984.6308. A, anterior region; B, thoracic sternum with pleon; C, thoracic sternum without pleon, G1. D, ♂ 16.6 × 17.0 mm, NW Madagascar, Ambaro Bay, MNHN-IU-2018-5193 (= MNHN-B18279): thoracic sternum without pleon, G1 and G2.	FIG. 11. — Dorippinae H. Milne Edwards, 1837 n. stat.. Dorippe quadridens (Fabricius, 1793): A-C, ♂ 38.1 × 36.3 mm, China Sea near Singapore, Hee Huat, ZRC 1984.6308. A, anterior region; B, thoracic sternum with pleon; C, thoracic sternum without pleon, G1. D, ♂ 16.6 × 17.0 mm, NW Madagascar, Ambaro Bay, MNHN-IU-2018-5193 (= MNHN-B18279): thoracic sternum without pleon, G1 and G2.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE749318B507CE72FD15FEEAF8F4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071273/files/figure.png	https://doi.org/10.5281/zenodo.8071273	FIG. 8. — Particular structures of some dorippids: A, B, rim and strip along posterior margin of carapace: A, Dorippoides facchino (Herbst, 1785), ovigerous ♀ 20.3 × 26.2 mm, South China Sea, ZRC 1984.5347; B, Paradorippe granulata (De Haan, 1841), ♂ 23.6 × 27.3 mm, NE Taiwan, ZRC 2001.0014; C, retention of female pleon by process of sternite 8 overhanging pleonal somite 2: Dorippe quadridens (Fabricius, 1793), ♀ 38.1 × 33.7 mm, South China Sea, Hee Huat, ZRC 1984.6307; D, retention of telson engaged between edges of sternite 5: Dorippoides facchino (Herbst, 1785), ovigerous ♀ 20.3 × 26.2 mm, same data as A. Abbreviations: cx3-cx5, coxae of P3-P5; p, process of sternite 8; p2, p3, pereiopod 2, 3; r, rim; s, strip; sp, spur-like process on P3 ischium; 1, pleonal somite 1; 7, 8, dorsally exposed thoracic sternites 7, 8.	FIG. 8. — Particular structures of some dorippids: A, B, rim and strip along posterior margin of carapace: A, Dorippoides facchino (Herbst, 1785), ovigerous ♀ 20.3 × 26.2 mm, South China Sea, ZRC 1984.5347; B, Paradorippe granulata (De Haan, 1841), ♂ 23.6 × 27.3 mm, NE Taiwan, ZRC 2001.0014; C, retention of female pleon by process of sternite 8 overhanging pleonal somite 2: Dorippe quadridens (Fabricius, 1793), ♀ 38.1 × 33.7 mm, South China Sea, Hee Huat, ZRC 1984.6307; D, retention of telson engaged between edges of sternite 5: Dorippoides facchino (Herbst, 1785), ovigerous ♀ 20.3 × 26.2 mm, same data as A. Abbreviations: cx3-cx5, coxae of P3-P5; p, process of sternite 8; p2, p3, pereiopod 2, 3; r, rim; s, strip; sp, spur-like process on P3 ischium; 1, pleonal somite 1; 7, 8, dorsally exposed thoracic sternites 7, 8.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE749318B507CE72FD15FEEAF8F4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071277/files/figure.png	https://doi.org/10.5281/zenodo.8071277	FIG. 9. — Particular structures of some dorippids: A, spur-like process on ischium of P3 and sometimes P2: Dorippoides facchino (Herbst, 1785), postpubertal ♀ 19.0 × 22.0 mm,Pondicherry Bay,Maindron coll.1882,MNHN-IU-2018-5199 (= MNHN-B19817).B, erect spine on sternite 8: Neodorippe callida (Fabricius,1795), ovigerous ♀ 13.5 × 14.3 mm, Amoy, MNHN-IU-2021-8738 (= MNHN-B11170). C, D, callosity at base of P3: C, Dorippe sinica Chen, 1980, ♀ 34.8 × 38.9 mm, China, Guangdong, Nanao Island, ZRC 1999.0470; D, Dorippe tenuipes Chen, 1980, ♂ 13.2 × 14.2 mm, South China Sea, ZRC 1999.0009 (the granules on P2 and P3 are not visible in the photograph). Abbreviations: c, callosity; cx2-cx5, coxae of P2-P5; e, erect spine; m, articulating membrane; p, process of sternite 8; pl6, exposed pleurite 6; p2-p5, pereiopods 2-5; r, rim; s, strip; sp, spur-like process; t, special texture area.	FIG. 9. — Particular structures of some dorippids: A, spur-like process on ischium of P3 and sometimes P2: Dorippoides facchino (Herbst, 1785), postpubertal ♀ 19.0 × 22.0 mm,Pondicherry Bay,Maindron coll.1882,MNHN-IU-2018-5199 (= MNHN-B19817).B, erect spine on sternite 8: Neodorippe callida (Fabricius,1795), ovigerous ♀ 13.5 × 14.3 mm, Amoy, MNHN-IU-2021-8738 (= MNHN-B11170). C, D, callosity at base of P3: C, Dorippe sinica Chen, 1980, ♀ 34.8 × 38.9 mm, China, Guangdong, Nanao Island, ZRC 1999.0470; D, Dorippe tenuipes Chen, 1980, ♂ 13.2 × 14.2 mm, South China Sea, ZRC 1999.0009 (the granules on P2 and P3 are not visible in the photograph). Abbreviations: c, callosity; cx2-cx5, coxae of P2-P5; e, erect spine; m, articulating membrane; p, process of sternite 8; pl6, exposed pleurite 6; p2-p5, pereiopods 2-5; r, rim; s, strip; sp, spur-like process; t, special texture area.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE749318B507CE72FD15FEEAF8F4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071287/files/figure.png	https://doi.org/10.5281/zenodo.8071287	FIG. 13. — Dorippinae H. Milne Edwards, 1837 n. stat.. Dorippe sinica Chen, 1980: A, B, ♂ 36.2 × 39.5 mm, China, Guangdong, Nanao Island, ZRC 1999.0470: A, ventral view, thoracic sternum and pleon; B, thoracic sternum,G1 and G2; C, D, ♀ 34.8 × 38.9 mm, same data as A, B; C, thoracic sternum and pleon; D, thoracic sternum and vulvae.	FIG. 13. — Dorippinae H. Milne Edwards, 1837 n. stat.. Dorippe sinica Chen, 1980: A, B, ♂ 36.2 × 39.5 mm, China, Guangdong, Nanao Island, ZRC 1999.0470: A, ventral view, thoracic sternum and pleon; B, thoracic sternum,G1 and G2; C, D, ♀ 34.8 × 38.9 mm, same data as A, B; C, thoracic sternum and pleon; D, thoracic sternum and vulvae.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE749318B507CE72FD15FEEAF8F4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071329/files/figure.png	https://doi.org/10.5281/zenodo.8071329	FIG. 31. — Diversity of G1s in the seven dorippid subfamilies (schematic drawings): A, Dorippinae n. stat. (Dorippe quadridens); B, Dorippoidinae n. subfam. (Dorippoides facchino); C, Medorippinae n. subfam.(Medorippe lanata); D-F, Heikeopsinae n.subfam.:D, Heikeopsis?japonica; E, Neodorippe callida; F, Nobilum histrio; G, Paradorippinae n. subfam. (Paradorippe cathayana); H, Philippidorippinae n. subfam. (Philippidorippe philippinensis); I, Phyllodorippinae n. subfam. (Phyllodorippe armata). (A-H, after Holthuis & Manning 1990; I, after Manning & Holthuis 1981). From Sin et al. (2009: fig. 4). Setae not always figured.	FIG. 31. — Diversity of G1s in the seven dorippid subfamilies (schematic drawings): A, Dorippinae n. stat. (Dorippe quadridens); B, Dorippoidinae n. subfam. (Dorippoides facchino); C, Medorippinae n. subfam.(Medorippe lanata); D-F, Heikeopsinae n.subfam.:D, Heikeopsis?japonica; E, Neodorippe callida; F, Nobilum histrio; G, Paradorippinae n. subfam. (Paradorippe cathayana); H, Philippidorippinae n. subfam. (Philippidorippe philippinensis); I, Phyllodorippinae n. subfam. (Phyllodorippe armata). (A-H, after Holthuis & Manning 1990; I, after Manning & Holthuis 1981). From Sin et al. (2009: fig. 4). Setae not always figured.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE749318B507CE72FD15FEEAF8F4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071331/files/figure.png	https://doi.org/10.5281/zenodo.8071331	FIG. 32. — Diversity of vulvae in the seven dorippid subfamilies (schematic drawings): A, Dorippinae n. stat. (Dorippe quadridens); B, Dorippoidinae n. subfam. (Dorippoides facchino); C, Medorippinae n.subfam.(Medorippe lanata); D-F, Heikeopsinae n. subfam.;D, Heikeopsis?japonica; E, Neodorippe callida; F, Nobilum histrio; G, Paradorippinae n. subfam. (Paradorippe cathayana); H, Philippidorippinae n. subfam. (Philippidorippe philippinensis); I, Phyllodorippinae n. subfam. (Phyllodorippe armata, ♀ 13.0 × 19.0 mm, Guinean Trawling Survey, MNHN-IU-2021-8731 [= MNHN-B24199]). Setae represented only on B. A-H, modified from Holthuis & Manning (1990).	FIG. 32. — Diversity of vulvae in the seven dorippid subfamilies (schematic drawings): A, Dorippinae n. stat. (Dorippe quadridens); B, Dorippoidinae n. subfam. (Dorippoides facchino); C, Medorippinae n.subfam.(Medorippe lanata); D-F, Heikeopsinae n. subfam.;D, Heikeopsis?japonica; E, Neodorippe callida; F, Nobilum histrio; G, Paradorippinae n. subfam. (Paradorippe cathayana); H, Philippidorippinae n. subfam. (Philippidorippe philippinensis); I, Phyllodorippinae n. subfam. (Phyllodorippe armata, ♀ 13.0 × 19.0 mm, Guinean Trawling Survey, MNHN-IU-2021-8731 [= MNHN-B24199]). Setae represented only on B. A-H, modified from Holthuis & Manning (1990).	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE749318B507CE72FD15FEEAF8F4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071337/files/figure.png	https://doi.org/10.5281/zenodo.8071337	FIG. 35. — Schematic illustrations of female reproductive systems in representatives of four dorippid subfamilies:A, B, Dorippinae n. stat.: A, Dorippe sinica Chen, 1980, as interpreted by Hayer et al. (2016a: fig. 2); B, Dorippe quadridens (Fabricius, 1793) and D. sinica, as interpreted by Vehof et al. (2017: fig. 2A); C, Medorippinae n. subfam.: Medorippe lanata (Linnaeus, 1767), as interpreted by Vehof et al. (2017: fig. 2B); D, Paradorippinae n. subfam.: Paradorippe granulata (De Haan, 1841), as interpreted by Vehof et al. (2018b: fig. 3). E, Heikeopsinae n. subfam.: Heikeopsis japonica (von Siebold, 1824), with the same pattern shared by Neodorippe callida (Fabricius, 1798) and Nobilum histrio (Nobili, 1903), as interpreted by Vehof (2020: fig. 4). Abbreviations: a, apodeme; amb, anteromedian bursa; bu, bursa;cu, cuticle;cv, cuticular valves;bu, bursa; ev, extension of vulva;ge, glandular epithelium; mu, musculature;oc, oocyte; od, oviduct; ov, ovary; plb, posterolateral bursa; sr, seminal receptacle; vg, vagina; v, vulva.	FIG. 35. — Schematic illustrations of female reproductive systems in representatives of four dorippid subfamilies:A, B, Dorippinae n. stat.: A, Dorippe sinica Chen, 1980, as interpreted by Hayer et al. (2016a: fig. 2); B, Dorippe quadridens (Fabricius, 1793) and D. sinica, as interpreted by Vehof et al. (2017: fig. 2A); C, Medorippinae n. subfam.: Medorippe lanata (Linnaeus, 1767), as interpreted by Vehof et al. (2017: fig. 2B); D, Paradorippinae n. subfam.: Paradorippe granulata (De Haan, 1841), as interpreted by Vehof et al. (2018b: fig. 3). E, Heikeopsinae n. subfam.: Heikeopsis japonica (von Siebold, 1824), with the same pattern shared by Neodorippe callida (Fabricius, 1798) and Nobilum histrio (Nobili, 1903), as interpreted by Vehof (2020: fig. 4). Abbreviations: a, apodeme; amb, anteromedian bursa; bu, bursa;cu, cuticle;cv, cuticular valves;bu, bursa; ev, extension of vulva;ge, glandular epithelium; mu, musculature;oc, oocyte; od, oviduct; ov, ovary; plb, posterolateral bursa; sr, seminal receptacle; vg, vagina; v, vulva.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE749318B507CE72FD15FEEAF8F4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071339/files/figure.png	https://doi.org/10.5281/zenodo.8071339	FIG. 36. — Schematic reconstructions of some female reproductive systems in Brachyura:A, in a podotreme crab,with spermatheca as cuticular invagination derived from a split between plates of intersegmental phragma 7/8, independent of the rest of reproductive system (gonopore on P3 coxa), and with external fertilisation; B, in Dorippe sinica (Dorippinae n. stat.), with glandular and cuticle chambers independent of seminal receptacle (not connected to ovary) and extension of vulva where fertilisation can take place (from Hayer et al. 2016a: fig. 8A). In Paradorippinae n. subfam. (not shown, but see Fig. 35D), without seminal receptacle, the fertilisation occurs externally in vulva opening (according to Vehof et al. 2018b: fig. 3; Vehof 2020); C, in Ethusa mascarone (Ethusidae), with glandular and cuticle chambers largely independent of the seminal receptacle (according to Hayer et al. 2016b: fig. 8); D, in a generalised eubrachyuran (one of several patterns), with seminal receptacle directly connected to ovary via oviduct, with vulvar opening on sternite 6 and with internal fertilisation. Abbreviations: as, spermathecal aperture; cc, cuticule chamber; cx3, coxa of P3 with coxal gonopore;gc, glandular chamber; m, musculature; od, oviduct; ov, ovary; P3, third pereiopod; sp, spermatheca; sr, seminal receptacle; vg, vagina; v, vulva.	FIG. 36. — Schematic reconstructions of some female reproductive systems in Brachyura:A, in a podotreme crab,with spermatheca as cuticular invagination derived from a split between plates of intersegmental phragma 7/8, independent of the rest of reproductive system (gonopore on P3 coxa), and with external fertilisation; B, in Dorippe sinica (Dorippinae n. stat.), with glandular and cuticle chambers independent of seminal receptacle (not connected to ovary) and extension of vulva where fertilisation can take place (from Hayer et al. 2016a: fig. 8A). In Paradorippinae n. subfam. (not shown, but see Fig. 35D), without seminal receptacle, the fertilisation occurs externally in vulva opening (according to Vehof et al. 2018b: fig. 3; Vehof 2020); C, in Ethusa mascarone (Ethusidae), with glandular and cuticle chambers largely independent of the seminal receptacle (according to Hayer et al. 2016b: fig. 8); D, in a generalised eubrachyuran (one of several patterns), with seminal receptacle directly connected to ovary via oviduct, with vulvar opening on sternite 6 and with internal fertilisation. Abbreviations: as, spermathecal aperture; cc, cuticule chamber; cx3, coxa of P3 with coxal gonopore;gc, glandular chamber; m, musculature; od, oviduct; ov, ovary; P3, third pereiopod; sp, spermatheca; sr, seminal receptacle; vg, vagina; v, vulva.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE749318B507CE72FD15FEEAF8F4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071341/files/figure.png	https://doi.org/10.5281/zenodo.8071341	FIG. 37. — Schematic representation of character mapping of sperm storage organs in the different dorippid subfamilies recognised here (except Phyllodorippinae n.subfam.not studied) and in a typical eubrachyuran:ovary with oocytes; oviduct; cuticular bursae (thin black line), of equal or unequal size; seminal receptacle either bilateral twin- or bilateral single,and either completely cuticular (thin black line) or partially glandular (thick black line). The twin system occurs in Dorippinae n. stat., Dorippoidinae n. subfam., Paradorippinae n. subfam. and Philippidorippinae n. subfam.; the single system occurs in Heikeopsinae n. subfam.,Medorippinae n. subfam.and in the typical Eubrachyura (most, see e.g. Ethusidae). Modified from Vehof 2020: figs 20-22).	FIG. 37. — Schematic representation of character mapping of sperm storage organs in the different dorippid subfamilies recognised here (except Phyllodorippinae n.subfam.not studied) and in a typical eubrachyuran:ovary with oocytes; oviduct; cuticular bursae (thin black line), of equal or unequal size; seminal receptacle either bilateral twin- or bilateral single,and either completely cuticular (thin black line) or partially glandular (thick black line). The twin system occurs in Dorippinae n. stat., Dorippoidinae n. subfam., Paradorippinae n. subfam. and Philippidorippinae n. subfam.; the single system occurs in Heikeopsinae n. subfam.,Medorippinae n. subfam.and in the typical Eubrachyura (most, see e.g. Ethusidae). Modified from Vehof 2020: figs 20-22).	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE749318B507CE72FD15FEEAF8F4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071333/files/figure.png	https://doi.org/10.5281/zenodo.8071333	FIG. 33. — Callosities in Dorippoidinae n. subfam. (Dorippoides) and Dorippinae n. stat. (Dorippe): A, callosity reduced to a thin calcified band: Dorippoides facchino (Herbst, 1785), ♂ 25.3 × 32.0 mm, Malaysia, Johore, Pontian, ZRC 1991.6672. B, no apparent callosity in male Dorippoides nudipes Manning & Holthuis, 1986, ♂ 16.2 × 19.4 mm, Iran, ZRC 2017.1227. C, callosity as a thick, elongated bulge: Dorippe quadridens (Fabricius, 1793), ♂ 39.0 × 41.4 mm, Thailand, Pattani Province, ZRC 2003.0126. D-F, callosity as a hemicircular structure: close-up views from different angles: D, E. Dorippe sinica Chen, 1980. D, ♀ 36.2 × 39.5 mm, China, Guangdong, Nanao Island, ZRC 1999.0470; E, ♂ 36.6 × 38.6 mm, Japan, Kochi, Shikoku, SMF 57855. F, D. frascone (Herbst, 1785), ♂ 29.7 × 31.0 mm, Philippines, Exp. Panglao 2004, ZRC 2008.0076. G, H, callosity as an arched double cup: D. tenuipes Chen, 1980, South China Sea, ZRC 1999.0009: G, ovigerous ♀ 17.3 × 19.6 mm; H, ♂ 13.2 × 14.2 mm. Abbreviations: b, branchiostegite; c, callosity; cx2-cx5, P2-P5 coxae; f, sulcated part of coxa; l, onepiece lateroventral part of coxa; m, membrane; p, pleon; pr, process of retention of female pleonal somite 2; pl6, exposed pleurite 6; P2-P5, pereiopods 2-5; r, rim of carapace posterior margin; s, strip along posterior rim; 1, 2, pleonal somites 1, 2; t, bottom of callosity with special texture; 7, 8, thoracic sternites 7, 8.	FIG. 33. — Callosities in Dorippoidinae n. subfam. (Dorippoides) and Dorippinae n. stat. (Dorippe): A, callosity reduced to a thin calcified band: Dorippoides facchino (Herbst, 1785), ♂ 25.3 × 32.0 mm, Malaysia, Johore, Pontian, ZRC 1991.6672. B, no apparent callosity in male Dorippoides nudipes Manning & Holthuis, 1986, ♂ 16.2 × 19.4 mm, Iran, ZRC 2017.1227. C, callosity as a thick, elongated bulge: Dorippe quadridens (Fabricius, 1793), ♂ 39.0 × 41.4 mm, Thailand, Pattani Province, ZRC 2003.0126. D-F, callosity as a hemicircular structure: close-up views from different angles: D, E. Dorippe sinica Chen, 1980. D, ♀ 36.2 × 39.5 mm, China, Guangdong, Nanao Island, ZRC 1999.0470; E, ♂ 36.6 × 38.6 mm, Japan, Kochi, Shikoku, SMF 57855. F, D. frascone (Herbst, 1785), ♂ 29.7 × 31.0 mm, Philippines, Exp. Panglao 2004, ZRC 2008.0076. G, H, callosity as an arched double cup: D. tenuipes Chen, 1980, South China Sea, ZRC 1999.0009: G, ovigerous ♀ 17.3 × 19.6 mm; H, ♂ 13.2 × 14.2 mm. Abbreviations: b, branchiostegite; c, callosity; cx2-cx5, P2-P5 coxae; f, sulcated part of coxa; l, onepiece lateroventral part of coxa; m, membrane; p, pleon; pr, process of retention of female pleonal somite 2; pl6, exposed pleurite 6; P2-P5, pereiopods 2-5; r, rim of carapace posterior margin; s, strip along posterior rim; 1, 2, pleonal somites 1, 2; t, bottom of callosity with special texture; 7, 8, thoracic sternites 7, 8.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE749327B507CD91F8F2FA11FAD5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071289/files/figure.png	https://doi.org/10.5281/zenodo.8071289	FIG. 14. — Dorippinae H. Milne Edwards,1837 n. stat.:A-G: Dorippe frascone (Herbst,1785),Philippines,Panglao Island,ZRC 2008.0076:A-D, ♂ 29.7 × 31.0 mm, A, habitus; B, ventral view; C, thoracic sternum with pleon; D, G1; E-G, prepubertal ♀ 23.3 × 24.4 mm: E, habitus;F, ventral view; G, vulvae; H, I, Dorippe irrorata Manning & Holthuis, 1986, holotype, ♂ 21.5 × 22.0 mm, Andaman Sea, south of Mergui Archipelago, USNM 172495: H, habitus; I, ventral surface.	FIG. 14. — Dorippinae H. Milne Edwards,1837 n. stat.:A-G: Dorippe frascone (Herbst,1785),Philippines,Panglao Island,ZRC 2008.0076:A-D, ♂ 29.7 × 31.0 mm, A, habitus; B, ventral view; C, thoracic sternum with pleon; D, G1; E-G, prepubertal ♀ 23.3 × 24.4 mm: E, habitus;F, ventral view; G, vulvae; H, I, Dorippe irrorata Manning & Holthuis, 1986, holotype, ♂ 21.5 × 22.0 mm, Andaman Sea, south of Mergui Archipelago, USNM 172495: H, habitus; I, ventral surface.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE749327B507CD91F8F2FA11FAD5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071279/files/figure.png	https://doi.org/10.5281/zenodo.8071279	FIG. 10. — Dorippinae H. Milne Edwards, 1837 n. stat.: habitus: A, B, Dorippe quadridens (Fabricius,1793):A, ♂ 36.3 × 38.1 mm, China Sea near Singapore, Hee Huat, ZRC 1984. 6308; B, ♀ 29.5 × 30 mm, NW Madagascar, Ambaro Bay, MNHN-IU-2018-5193 (= MNHN-B18279). C, D, Dorippe sinica Chen, 1980, China, Guangdong, Nanao Island, ZRC 1999.0470: C, ♂ 36.2 × 39.5 mm; D, ♀ 34.8 × 38.9 mm, specimen brushed.	FIG. 10. — Dorippinae H. Milne Edwards, 1837 n. stat.: habitus: A, B, Dorippe quadridens (Fabricius,1793):A, ♂ 36.3 × 38.1 mm, China Sea near Singapore, Hee Huat, ZRC 1984. 6308; B, ♀ 29.5 × 30 mm, NW Madagascar, Ambaro Bay, MNHN-IU-2018-5193 (= MNHN-B18279). C, D, Dorippe sinica Chen, 1980, China, Guangdong, Nanao Island, ZRC 1999.0470: C, ♂ 36.2 × 39.5 mm; D, ♀ 34.8 × 38.9 mm, specimen brushed.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE749327B507CD91F8F2FA11FAD5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071283/files/figure.png	https://doi.org/10.5281/zenodo.8071283	FIG. 11. — Dorippinae H. Milne Edwards, 1837 n. stat.. Dorippe quadridens (Fabricius, 1793): A-C, ♂ 38.1 × 36.3 mm, China Sea near Singapore, Hee Huat, ZRC 1984.6308. A, anterior region; B, thoracic sternum with pleon; C, thoracic sternum without pleon, G1. D, ♂ 16.6 × 17.0 mm, NW Madagascar, Ambaro Bay, MNHN-IU-2018-5193 (= MNHN-B18279): thoracic sternum without pleon, G1 and G2.	FIG. 11. — Dorippinae H. Milne Edwards, 1837 n. stat.. Dorippe quadridens (Fabricius, 1793): A-C, ♂ 38.1 × 36.3 mm, China Sea near Singapore, Hee Huat, ZRC 1984.6308. A, anterior region; B, thoracic sternum with pleon; C, thoracic sternum without pleon, G1. D, ♂ 16.6 × 17.0 mm, NW Madagascar, Ambaro Bay, MNHN-IU-2018-5193 (= MNHN-B18279): thoracic sternum without pleon, G1 and G2.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE749327B507CD91F8F2FA11FAD5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071277/files/figure.png	https://doi.org/10.5281/zenodo.8071277	FIG. 9. — Particular structures of some dorippids: A, spur-like process on ischium of P3 and sometimes P2: Dorippoides facchino (Herbst, 1785), postpubertal ♀ 19.0 × 22.0 mm,Pondicherry Bay,Maindron coll.1882,MNHN-IU-2018-5199 (= MNHN-B19817).B, erect spine on sternite 8: Neodorippe callida (Fabricius,1795), ovigerous ♀ 13.5 × 14.3 mm, Amoy, MNHN-IU-2021-8738 (= MNHN-B11170). C, D, callosity at base of P3: C, Dorippe sinica Chen, 1980, ♀ 34.8 × 38.9 mm, China, Guangdong, Nanao Island, ZRC 1999.0470; D, Dorippe tenuipes Chen, 1980, ♂ 13.2 × 14.2 mm, South China Sea, ZRC 1999.0009 (the granules on P2 and P3 are not visible in the photograph). Abbreviations: c, callosity; cx2-cx5, coxae of P2-P5; e, erect spine; m, articulating membrane; p, process of sternite 8; pl6, exposed pleurite 6; p2-p5, pereiopods 2-5; r, rim; s, strip; sp, spur-like process; t, special texture area.	FIG. 9. — Particular structures of some dorippids: A, spur-like process on ischium of P3 and sometimes P2: Dorippoides facchino (Herbst, 1785), postpubertal ♀ 19.0 × 22.0 mm,Pondicherry Bay,Maindron coll.1882,MNHN-IU-2018-5199 (= MNHN-B19817).B, erect spine on sternite 8: Neodorippe callida (Fabricius,1795), ovigerous ♀ 13.5 × 14.3 mm, Amoy, MNHN-IU-2021-8738 (= MNHN-B11170). C, D, callosity at base of P3: C, Dorippe sinica Chen, 1980, ♀ 34.8 × 38.9 mm, China, Guangdong, Nanao Island, ZRC 1999.0470; D, Dorippe tenuipes Chen, 1980, ♂ 13.2 × 14.2 mm, South China Sea, ZRC 1999.0009 (the granules on P2 and P3 are not visible in the photograph). Abbreviations: c, callosity; cx2-cx5, coxae of P2-P5; e, erect spine; m, articulating membrane; p, process of sternite 8; pl6, exposed pleurite 6; p2-p5, pereiopods 2-5; r, rim; s, strip; sp, spur-like process; t, special texture area.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE749327B507CD91F8F2FA11FAD5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071333/files/figure.png	https://doi.org/10.5281/zenodo.8071333	FIG. 33. — Callosities in Dorippoidinae n. subfam. (Dorippoides) and Dorippinae n. stat. (Dorippe): A, callosity reduced to a thin calcified band: Dorippoides facchino (Herbst, 1785), ♂ 25.3 × 32.0 mm, Malaysia, Johore, Pontian, ZRC 1991.6672. B, no apparent callosity in male Dorippoides nudipes Manning & Holthuis, 1986, ♂ 16.2 × 19.4 mm, Iran, ZRC 2017.1227. C, callosity as a thick, elongated bulge: Dorippe quadridens (Fabricius, 1793), ♂ 39.0 × 41.4 mm, Thailand, Pattani Province, ZRC 2003.0126. D-F, callosity as a hemicircular structure: close-up views from different angles: D, E. Dorippe sinica Chen, 1980. D, ♀ 36.2 × 39.5 mm, China, Guangdong, Nanao Island, ZRC 1999.0470; E, ♂ 36.6 × 38.6 mm, Japan, Kochi, Shikoku, SMF 57855. F, D. frascone (Herbst, 1785), ♂ 29.7 × 31.0 mm, Philippines, Exp. Panglao 2004, ZRC 2008.0076. G, H, callosity as an arched double cup: D. tenuipes Chen, 1980, South China Sea, ZRC 1999.0009: G, ovigerous ♀ 17.3 × 19.6 mm; H, ♂ 13.2 × 14.2 mm. Abbreviations: b, branchiostegite; c, callosity; cx2-cx5, P2-P5 coxae; f, sulcated part of coxa; l, onepiece lateroventral part of coxa; m, membrane; p, pleon; pr, process of retention of female pleonal somite 2; pl6, exposed pleurite 6; P2-P5, pereiopods 2-5; r, rim of carapace posterior margin; s, strip along posterior rim; 1, 2, pleonal somites 1, 2; t, bottom of callosity with special texture; 7, 8, thoracic sternites 7, 8.	FIG. 33. — Callosities in Dorippoidinae n. subfam. (Dorippoides) and Dorippinae n. stat. (Dorippe): A, callosity reduced to a thin calcified band: Dorippoides facchino (Herbst, 1785), ♂ 25.3 × 32.0 mm, Malaysia, Johore, Pontian, ZRC 1991.6672. B, no apparent callosity in male Dorippoides nudipes Manning & Holthuis, 1986, ♂ 16.2 × 19.4 mm, Iran, ZRC 2017.1227. C, callosity as a thick, elongated bulge: Dorippe quadridens (Fabricius, 1793), ♂ 39.0 × 41.4 mm, Thailand, Pattani Province, ZRC 2003.0126. D-F, callosity as a hemicircular structure: close-up views from different angles: D, E. Dorippe sinica Chen, 1980. D, ♀ 36.2 × 39.5 mm, China, Guangdong, Nanao Island, ZRC 1999.0470; E, ♂ 36.6 × 38.6 mm, Japan, Kochi, Shikoku, SMF 57855. F, D. frascone (Herbst, 1785), ♂ 29.7 × 31.0 mm, Philippines, Exp. Panglao 2004, ZRC 2008.0076. G, H, callosity as an arched double cup: D. tenuipes Chen, 1980, South China Sea, ZRC 1999.0009: G, ovigerous ♀ 17.3 × 19.6 mm; H, ♂ 13.2 × 14.2 mm. Abbreviations: b, branchiostegite; c, callosity; cx2-cx5, P2-P5 coxae; f, sulcated part of coxa; l, onepiece lateroventral part of coxa; m, membrane; p, pleon; pr, process of retention of female pleonal somite 2; pl6, exposed pleurite 6; P2-P5, pereiopods 2-5; r, rim of carapace posterior margin; s, strip along posterior rim; 1, 2, pleonal somites 1, 2; t, bottom of callosity with special texture; 7, 8, thoracic sternites 7, 8.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE749327B502CE38FA94FEE9F813.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071291/files/figure.png	https://doi.org/10.5281/zenodo.8071291	FIG. 15. — Dorippinae H. Milne Edwards,1837 n. stat.: A-D, J, Dorippe trilobata Manning,1993,holotype, ♂ 20.5 × 21.1 mm, off Mitchell River,Admiralty Gulf,AM P27124: A, habitus; B, ventral surface; C, carapace; D, pleon; J, callosity. E-I: Dorippe glabra Manning, 1993, holotype, ovigerous ♀ 24.3 × 26.7 mm, Chambers Bay, Australia, AM P13363: E, habitus with detached right cheliped, right P2 and P3, left P3; F, carapace; G, ventral surface; H, pleon; I, callosity.	FIG. 15. — Dorippinae H. Milne Edwards,1837 n. stat.: A-D, J, Dorippe trilobata Manning,1993,holotype, ♂ 20.5 × 21.1 mm, off Mitchell River,Admiralty Gulf,AM P27124: A, habitus; B, ventral surface; C, carapace; D, pleon; J, callosity. E-I: Dorippe glabra Manning, 1993, holotype, ovigerous ♀ 24.3 × 26.7 mm, Chambers Bay, Australia, AM P13363: E, habitus with detached right cheliped, right P2 and P3, left P3; F, carapace; G, ventral surface; H, pleon; I, callosity.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE749327B502CE38FA94FEE9F813.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071279/files/figure.png	https://doi.org/10.5281/zenodo.8071279	FIG. 10. — Dorippinae H. Milne Edwards, 1837 n. stat.: habitus: A, B, Dorippe quadridens (Fabricius,1793):A, ♂ 36.3 × 38.1 mm, China Sea near Singapore, Hee Huat, ZRC 1984. 6308; B, ♀ 29.5 × 30 mm, NW Madagascar, Ambaro Bay, MNHN-IU-2018-5193 (= MNHN-B18279). C, D, Dorippe sinica Chen, 1980, China, Guangdong, Nanao Island, ZRC 1999.0470: C, ♂ 36.2 × 39.5 mm; D, ♀ 34.8 × 38.9 mm, specimen brushed.	FIG. 10. — Dorippinae H. Milne Edwards, 1837 n. stat.: habitus: A, B, Dorippe quadridens (Fabricius,1793):A, ♂ 36.3 × 38.1 mm, China Sea near Singapore, Hee Huat, ZRC 1984. 6308; B, ♀ 29.5 × 30 mm, NW Madagascar, Ambaro Bay, MNHN-IU-2018-5193 (= MNHN-B18279). C, D, Dorippe sinica Chen, 1980, China, Guangdong, Nanao Island, ZRC 1999.0470: C, ♂ 36.2 × 39.5 mm; D, ♀ 34.8 × 38.9 mm, specimen brushed.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE749327B502CE38FA94FEE9F813.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071277/files/figure.png	https://doi.org/10.5281/zenodo.8071277	FIG. 9. — Particular structures of some dorippids: A, spur-like process on ischium of P3 and sometimes P2: Dorippoides facchino (Herbst, 1785), postpubertal ♀ 19.0 × 22.0 mm,Pondicherry Bay,Maindron coll.1882,MNHN-IU-2018-5199 (= MNHN-B19817).B, erect spine on sternite 8: Neodorippe callida (Fabricius,1795), ovigerous ♀ 13.5 × 14.3 mm, Amoy, MNHN-IU-2021-8738 (= MNHN-B11170). C, D, callosity at base of P3: C, Dorippe sinica Chen, 1980, ♀ 34.8 × 38.9 mm, China, Guangdong, Nanao Island, ZRC 1999.0470; D, Dorippe tenuipes Chen, 1980, ♂ 13.2 × 14.2 mm, South China Sea, ZRC 1999.0009 (the granules on P2 and P3 are not visible in the photograph). Abbreviations: c, callosity; cx2-cx5, coxae of P2-P5; e, erect spine; m, articulating membrane; p, process of sternite 8; pl6, exposed pleurite 6; p2-p5, pereiopods 2-5; r, rim; s, strip; sp, spur-like process; t, special texture area.	FIG. 9. — Particular structures of some dorippids: A, spur-like process on ischium of P3 and sometimes P2: Dorippoides facchino (Herbst, 1785), postpubertal ♀ 19.0 × 22.0 mm,Pondicherry Bay,Maindron coll.1882,MNHN-IU-2018-5199 (= MNHN-B19817).B, erect spine on sternite 8: Neodorippe callida (Fabricius,1795), ovigerous ♀ 13.5 × 14.3 mm, Amoy, MNHN-IU-2021-8738 (= MNHN-B11170). C, D, callosity at base of P3: C, Dorippe sinica Chen, 1980, ♀ 34.8 × 38.9 mm, China, Guangdong, Nanao Island, ZRC 1999.0470; D, Dorippe tenuipes Chen, 1980, ♂ 13.2 × 14.2 mm, South China Sea, ZRC 1999.0009 (the granules on P2 and P3 are not visible in the photograph). Abbreviations: c, callosity; cx2-cx5, coxae of P2-P5; e, erect spine; m, articulating membrane; p, process of sternite 8; pl6, exposed pleurite 6; p2-p5, pereiopods 2-5; r, rim; s, strip; sp, spur-like process; t, special texture area.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE749327B502CE38FA94FEE9F813.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071333/files/figure.png	https://doi.org/10.5281/zenodo.8071333	FIG. 33. — Callosities in Dorippoidinae n. subfam. (Dorippoides) and Dorippinae n. stat. (Dorippe): A, callosity reduced to a thin calcified band: Dorippoides facchino (Herbst, 1785), ♂ 25.3 × 32.0 mm, Malaysia, Johore, Pontian, ZRC 1991.6672. B, no apparent callosity in male Dorippoides nudipes Manning & Holthuis, 1986, ♂ 16.2 × 19.4 mm, Iran, ZRC 2017.1227. C, callosity as a thick, elongated bulge: Dorippe quadridens (Fabricius, 1793), ♂ 39.0 × 41.4 mm, Thailand, Pattani Province, ZRC 2003.0126. D-F, callosity as a hemicircular structure: close-up views from different angles: D, E. Dorippe sinica Chen, 1980. D, ♀ 36.2 × 39.5 mm, China, Guangdong, Nanao Island, ZRC 1999.0470; E, ♂ 36.6 × 38.6 mm, Japan, Kochi, Shikoku, SMF 57855. F, D. frascone (Herbst, 1785), ♂ 29.7 × 31.0 mm, Philippines, Exp. Panglao 2004, ZRC 2008.0076. G, H, callosity as an arched double cup: D. tenuipes Chen, 1980, South China Sea, ZRC 1999.0009: G, ovigerous ♀ 17.3 × 19.6 mm; H, ♂ 13.2 × 14.2 mm. Abbreviations: b, branchiostegite; c, callosity; cx2-cx5, P2-P5 coxae; f, sulcated part of coxa; l, onepiece lateroventral part of coxa; m, membrane; p, pleon; pr, process of retention of female pleonal somite 2; pl6, exposed pleurite 6; P2-P5, pereiopods 2-5; r, rim of carapace posterior margin; s, strip along posterior rim; 1, 2, pleonal somites 1, 2; t, bottom of callosity with special texture; 7, 8, thoracic sternites 7, 8.	FIG. 33. — Callosities in Dorippoidinae n. subfam. (Dorippoides) and Dorippinae n. stat. (Dorippe): A, callosity reduced to a thin calcified band: Dorippoides facchino (Herbst, 1785), ♂ 25.3 × 32.0 mm, Malaysia, Johore, Pontian, ZRC 1991.6672. B, no apparent callosity in male Dorippoides nudipes Manning & Holthuis, 1986, ♂ 16.2 × 19.4 mm, Iran, ZRC 2017.1227. C, callosity as a thick, elongated bulge: Dorippe quadridens (Fabricius, 1793), ♂ 39.0 × 41.4 mm, Thailand, Pattani Province, ZRC 2003.0126. D-F, callosity as a hemicircular structure: close-up views from different angles: D, E. Dorippe sinica Chen, 1980. D, ♀ 36.2 × 39.5 mm, China, Guangdong, Nanao Island, ZRC 1999.0470; E, ♂ 36.6 × 38.6 mm, Japan, Kochi, Shikoku, SMF 57855. F, D. frascone (Herbst, 1785), ♂ 29.7 × 31.0 mm, Philippines, Exp. Panglao 2004, ZRC 2008.0076. G, H, callosity as an arched double cup: D. tenuipes Chen, 1980, South China Sea, ZRC 1999.0009: G, ovigerous ♀ 17.3 × 19.6 mm; H, ♂ 13.2 × 14.2 mm. Abbreviations: b, branchiostegite; c, callosity; cx2-cx5, P2-P5 coxae; f, sulcated part of coxa; l, onepiece lateroventral part of coxa; m, membrane; p, pleon; pr, process of retention of female pleonal somite 2; pl6, exposed pleurite 6; P2-P5, pereiopods 2-5; r, rim of carapace posterior margin; s, strip along posterior rim; 1, 2, pleonal somites 1, 2; t, bottom of callosity with special texture; 7, 8, thoracic sternites 7, 8.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE749327B502CE38FA94FEE9F813.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071285/files/figure.png	https://doi.org/10.5281/zenodo.8071285	FIG. 12. — Dorippinae H. Milne Edwards,1837 n. stat.. Dorippe quadridens (Fabricius,1793):A-D, ovigerous ♀ 38.1 × 33.7 mm, South China Sea,Hee Huat,ZRC 1984.6307:A, habitus; B, anterior region, ventral view; C, thoracic sternum and pleon; D, vulvae. E, ovigerous ♀ 28.0 × 29.0 mm, NW Madagascar, Ambaro Bay, MNHN-IU-2018-5193 (= MNHN-B18279): thoracic sternum and vulvae.	FIG. 12. — Dorippinae H. Milne Edwards,1837 n. stat.. Dorippe quadridens (Fabricius,1793):A-D, ovigerous ♀ 38.1 × 33.7 mm, South China Sea,Hee Huat,ZRC 1984.6307:A, habitus; B, anterior region, ventral view; C, thoracic sternum and pleon; D, vulvae. E, ovigerous ♀ 28.0 × 29.0 mm, NW Madagascar, Ambaro Bay, MNHN-IU-2018-5193 (= MNHN-B18279): thoracic sternum and vulvae.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE749327B502CE38FA94FEE9F813.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071289/files/figure.png	https://doi.org/10.5281/zenodo.8071289	FIG. 14. — Dorippinae H. Milne Edwards,1837 n. stat.:A-G: Dorippe frascone (Herbst,1785),Philippines,Panglao Island,ZRC 2008.0076:A-D, ♂ 29.7 × 31.0 mm, A, habitus; B, ventral view; C, thoracic sternum with pleon; D, G1; E-G, prepubertal ♀ 23.3 × 24.4 mm: E, habitus;F, ventral view; G, vulvae; H, I, Dorippe irrorata Manning & Holthuis, 1986, holotype, ♂ 21.5 × 22.0 mm, Andaman Sea, south of Mergui Archipelago, USNM 172495: H, habitus; I, ventral surface.	FIG. 14. — Dorippinae H. Milne Edwards,1837 n. stat.:A-G: Dorippe frascone (Herbst,1785),Philippines,Panglao Island,ZRC 2008.0076:A-D, ♂ 29.7 × 31.0 mm, A, habitus; B, ventral view; C, thoracic sternum with pleon; D, G1; E-G, prepubertal ♀ 23.3 × 24.4 mm: E, habitus;F, ventral view; G, vulvae; H, I, Dorippe irrorata Manning & Holthuis, 1986, holotype, ♂ 21.5 × 22.0 mm, Andaman Sea, south of Mergui Archipelago, USNM 172495: H, habitus; I, ventral surface.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE749327B502CE38FA94FEE9F813.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071283/files/figure.png	https://doi.org/10.5281/zenodo.8071283	FIG. 11. — Dorippinae H. Milne Edwards, 1837 n. stat.. Dorippe quadridens (Fabricius, 1793): A-C, ♂ 38.1 × 36.3 mm, China Sea near Singapore, Hee Huat, ZRC 1984.6308. A, anterior region; B, thoracic sternum with pleon; C, thoracic sternum without pleon, G1. D, ♂ 16.6 × 17.0 mm, NW Madagascar, Ambaro Bay, MNHN-IU-2018-5193 (= MNHN-B18279): thoracic sternum without pleon, G1 and G2.	FIG. 11. — Dorippinae H. Milne Edwards, 1837 n. stat.. Dorippe quadridens (Fabricius, 1793): A-C, ♂ 38.1 × 36.3 mm, China Sea near Singapore, Hee Huat, ZRC 1984.6308. A, anterior region; B, thoracic sternum with pleon; C, thoracic sternum without pleon, G1. D, ♂ 16.6 × 17.0 mm, NW Madagascar, Ambaro Bay, MNHN-IU-2018-5193 (= MNHN-B18279): thoracic sternum without pleon, G1 and G2.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE749327B502CE38FA94FEE9F813.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071287/files/figure.png	https://doi.org/10.5281/zenodo.8071287	FIG. 13. — Dorippinae H. Milne Edwards, 1837 n. stat.. Dorippe sinica Chen, 1980: A, B, ♂ 36.2 × 39.5 mm, China, Guangdong, Nanao Island, ZRC 1999.0470: A, ventral view, thoracic sternum and pleon; B, thoracic sternum,G1 and G2; C, D, ♀ 34.8 × 38.9 mm, same data as A, B; C, thoracic sternum and pleon; D, thoracic sternum and vulvae.	FIG. 13. — Dorippinae H. Milne Edwards, 1837 n. stat.. Dorippe sinica Chen, 1980: A, B, ♂ 36.2 × 39.5 mm, China, Guangdong, Nanao Island, ZRC 1999.0470: A, ventral view, thoracic sternum and pleon; B, thoracic sternum,G1 and G2; C, D, ♀ 34.8 × 38.9 mm, same data as A, B; C, thoracic sternum and pleon; D, thoracic sternum and vulvae.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE749322B50FCEB4FF17FDCAF813.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071293/files/figure.png	https://doi.org/10.5281/zenodo.8071293	FIG. 16. — Dorippoidinae n. subfam.: A, B, Dorippoides facchino (Herbst, 1785): A, ♂ 25.3 × 32 mm, Malaysia, Johore, Pontian, ZRC 1991.66-72; B, ovigerous ♀ 20.3 × 26.2 mm, South China Sea, Hee Huat, ZRC 1984.5347; C, D, Dorippoides nudipes Manning & Holthuis,1986: C, ♂ 16.2 × 19.4 mm, Iran,ZRC 2017.1227; D, ♀ 21.6 × 29.5 mm, South Africa, vicinity of Tugela River mouth, ZRC 2009.0885.	FIG. 16. — Dorippoidinae n. subfam.: A, B, Dorippoides facchino (Herbst, 1785): A, ♂ 25.3 × 32 mm, Malaysia, Johore, Pontian, ZRC 1991.66-72; B, ovigerous ♀ 20.3 × 26.2 mm, South China Sea, Hee Huat, ZRC 1984.5347; C, D, Dorippoides nudipes Manning & Holthuis,1986: C, ♂ 16.2 × 19.4 mm, Iran,ZRC 2017.1227; D, ♀ 21.6 × 29.5 mm, South Africa, vicinity of Tugela River mouth, ZRC 2009.0885.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE749322B50FCEB4FF17FDCAF813.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071297/files/figure.png	https://doi.org/10.5281/zenodo.8071297	FIG. 17. — Dorippoidinae n. subfam. Dorippoides facchino (Herbst, 1785): A-C, ♂ 25.3 × 32 mm, Malaysia, Johore, Pontian, ZRC 1991.66-72: A, anterior ventral view; B, thoracic sternum with pleon; C, without pleon. D, ovigerous ♀ 21 × 25 mm, South China Sea, MNHN-IU-2018-5195 (= MNHN-B18802): vulvae.	FIG. 17. — Dorippoidinae n. subfam. Dorippoides facchino (Herbst, 1785): A-C, ♂ 25.3 × 32 mm, Malaysia, Johore, Pontian, ZRC 1991.66-72: A, anterior ventral view; B, thoracic sternum with pleon; C, without pleon. D, ovigerous ♀ 21 × 25 mm, South China Sea, MNHN-IU-2018-5195 (= MNHN-B18802): vulvae.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE749322B50FCEB4FF17FDCAF813.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071299/files/figure.png	https://doi.org/10.5281/zenodo.8071299	FIG. 18. — Dorippoidinae n. subfam.: A-C, Dorippoides facchino (Herbst, 1785), ovigerous ♀ 20.3 × 26.2 mm, South China Sea, Hee Huat, ZRC 1984.5347: A, thoracic sternum with pleon; B, without pleon, vulvae. C, ovigerous ♀ 21 × 25 mm, South China Sea, trawl, 37 m, Chen leg. 1987, MNHN-IU-2018-5195 (= MNHN-B18802).D-F, Dorippoides nudipes Manning & Holthuis 1986: D, ♂ 16.2 × 19.4 mm, Iran, ZRC 2017.1227; thoracic sternum with pleon; E, F, ♀ 21.6 × 29.5 mm, South Africa, vicinity of Tugela River mouth, ZRC 2009.0885: E, thoracic sternum with pleon; F, thoracic sternum without pleon; vulvae. See the spurlike process on P3 ischium of females on A, B.	FIG. 18. — Dorippoidinae n. subfam.: A-C, Dorippoides facchino (Herbst, 1785), ovigerous ♀ 20.3 × 26.2 mm, South China Sea, Hee Huat, ZRC 1984.5347: A, thoracic sternum with pleon; B, without pleon, vulvae. C, ovigerous ♀ 21 × 25 mm, South China Sea, trawl, 37 m, Chen leg. 1987, MNHN-IU-2018-5195 (= MNHN-B18802).D-F, Dorippoides nudipes Manning & Holthuis 1986: D, ♂ 16.2 × 19.4 mm, Iran, ZRC 2017.1227; thoracic sternum with pleon; E, F, ♀ 21.6 × 29.5 mm, South Africa, vicinity of Tugela River mouth, ZRC 2009.0885: E, thoracic sternum with pleon; F, thoracic sternum without pleon; vulvae. See the spurlike process on P3 ischium of females on A, B.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE749322B50FCEB4FF17FDCAF813.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071277/files/figure.png	https://doi.org/10.5281/zenodo.8071277	FIG. 9. — Particular structures of some dorippids: A, spur-like process on ischium of P3 and sometimes P2: Dorippoides facchino (Herbst, 1785), postpubertal ♀ 19.0 × 22.0 mm,Pondicherry Bay,Maindron coll.1882,MNHN-IU-2018-5199 (= MNHN-B19817).B, erect spine on sternite 8: Neodorippe callida (Fabricius,1795), ovigerous ♀ 13.5 × 14.3 mm, Amoy, MNHN-IU-2021-8738 (= MNHN-B11170). C, D, callosity at base of P3: C, Dorippe sinica Chen, 1980, ♀ 34.8 × 38.9 mm, China, Guangdong, Nanao Island, ZRC 1999.0470; D, Dorippe tenuipes Chen, 1980, ♂ 13.2 × 14.2 mm, South China Sea, ZRC 1999.0009 (the granules on P2 and P3 are not visible in the photograph). Abbreviations: c, callosity; cx2-cx5, coxae of P2-P5; e, erect spine; m, articulating membrane; p, process of sternite 8; pl6, exposed pleurite 6; p2-p5, pereiopods 2-5; r, rim; s, strip; sp, spur-like process; t, special texture area.	FIG. 9. — Particular structures of some dorippids: A, spur-like process on ischium of P3 and sometimes P2: Dorippoides facchino (Herbst, 1785), postpubertal ♀ 19.0 × 22.0 mm,Pondicherry Bay,Maindron coll.1882,MNHN-IU-2018-5199 (= MNHN-B19817).B, erect spine on sternite 8: Neodorippe callida (Fabricius,1795), ovigerous ♀ 13.5 × 14.3 mm, Amoy, MNHN-IU-2021-8738 (= MNHN-B11170). C, D, callosity at base of P3: C, Dorippe sinica Chen, 1980, ♀ 34.8 × 38.9 mm, China, Guangdong, Nanao Island, ZRC 1999.0470; D, Dorippe tenuipes Chen, 1980, ♂ 13.2 × 14.2 mm, South China Sea, ZRC 1999.0009 (the granules on P2 and P3 are not visible in the photograph). Abbreviations: c, callosity; cx2-cx5, coxae of P2-P5; e, erect spine; m, articulating membrane; p, process of sternite 8; pl6, exposed pleurite 6; p2-p5, pereiopods 2-5; r, rim; s, strip; sp, spur-like process; t, special texture area.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE749322B50FCEB4FF17FDCAF813.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071329/files/figure.png	https://doi.org/10.5281/zenodo.8071329	FIG. 31. — Diversity of G1s in the seven dorippid subfamilies (schematic drawings): A, Dorippinae n. stat. (Dorippe quadridens); B, Dorippoidinae n. subfam. (Dorippoides facchino); C, Medorippinae n. subfam.(Medorippe lanata); D-F, Heikeopsinae n.subfam.:D, Heikeopsis?japonica; E, Neodorippe callida; F, Nobilum histrio; G, Paradorippinae n. subfam. (Paradorippe cathayana); H, Philippidorippinae n. subfam. (Philippidorippe philippinensis); I, Phyllodorippinae n. subfam. (Phyllodorippe armata). (A-H, after Holthuis & Manning 1990; I, after Manning & Holthuis 1981). From Sin et al. (2009: fig. 4). Setae not always figured.	FIG. 31. — Diversity of G1s in the seven dorippid subfamilies (schematic drawings): A, Dorippinae n. stat. (Dorippe quadridens); B, Dorippoidinae n. subfam. (Dorippoides facchino); C, Medorippinae n. subfam.(Medorippe lanata); D-F, Heikeopsinae n.subfam.:D, Heikeopsis?japonica; E, Neodorippe callida; F, Nobilum histrio; G, Paradorippinae n. subfam. (Paradorippe cathayana); H, Philippidorippinae n. subfam. (Philippidorippe philippinensis); I, Phyllodorippinae n. subfam. (Phyllodorippe armata). (A-H, after Holthuis & Manning 1990; I, after Manning & Holthuis 1981). From Sin et al. (2009: fig. 4). Setae not always figured.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE749322B50FCEB4FF17FDCAF813.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071331/files/figure.png	https://doi.org/10.5281/zenodo.8071331	FIG. 32. — Diversity of vulvae in the seven dorippid subfamilies (schematic drawings): A, Dorippinae n. stat. (Dorippe quadridens); B, Dorippoidinae n. subfam. (Dorippoides facchino); C, Medorippinae n.subfam.(Medorippe lanata); D-F, Heikeopsinae n. subfam.;D, Heikeopsis?japonica; E, Neodorippe callida; F, Nobilum histrio; G, Paradorippinae n. subfam. (Paradorippe cathayana); H, Philippidorippinae n. subfam. (Philippidorippe philippinensis); I, Phyllodorippinae n. subfam. (Phyllodorippe armata, ♀ 13.0 × 19.0 mm, Guinean Trawling Survey, MNHN-IU-2021-8731 [= MNHN-B24199]). Setae represented only on B. A-H, modified from Holthuis & Manning (1990).	FIG. 32. — Diversity of vulvae in the seven dorippid subfamilies (schematic drawings): A, Dorippinae n. stat. (Dorippe quadridens); B, Dorippoidinae n. subfam. (Dorippoides facchino); C, Medorippinae n.subfam.(Medorippe lanata); D-F, Heikeopsinae n. subfam.;D, Heikeopsis?japonica; E, Neodorippe callida; F, Nobilum histrio; G, Paradorippinae n. subfam. (Paradorippe cathayana); H, Philippidorippinae n. subfam. (Philippidorippe philippinensis); I, Phyllodorippinae n. subfam. (Phyllodorippe armata, ♀ 13.0 × 19.0 mm, Guinean Trawling Survey, MNHN-IU-2021-8731 [= MNHN-B24199]). Setae represented only on B. A-H, modified from Holthuis & Manning (1990).	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE749322B50FCEB4FF17FDCAF813.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071341/files/figure.png	https://doi.org/10.5281/zenodo.8071341	FIG. 37. — Schematic representation of character mapping of sperm storage organs in the different dorippid subfamilies recognised here (except Phyllodorippinae n.subfam.not studied) and in a typical eubrachyuran:ovary with oocytes; oviduct; cuticular bursae (thin black line), of equal or unequal size; seminal receptacle either bilateral twin- or bilateral single,and either completely cuticular (thin black line) or partially glandular (thick black line). The twin system occurs in Dorippinae n. stat., Dorippoidinae n. subfam., Paradorippinae n. subfam. and Philippidorippinae n. subfam.; the single system occurs in Heikeopsinae n. subfam.,Medorippinae n. subfam.and in the typical Eubrachyura (most, see e.g. Ethusidae). Modified from Vehof 2020: figs 20-22).	FIG. 37. — Schematic representation of character mapping of sperm storage organs in the different dorippid subfamilies recognised here (except Phyllodorippinae n.subfam.not studied) and in a typical eubrachyuran:ovary with oocytes; oviduct; cuticular bursae (thin black line), of equal or unequal size; seminal receptacle either bilateral twin- or bilateral single,and either completely cuticular (thin black line) or partially glandular (thick black line). The twin system occurs in Dorippinae n. stat., Dorippoidinae n. subfam., Paradorippinae n. subfam. and Philippidorippinae n. subfam.; the single system occurs in Heikeopsinae n. subfam.,Medorippinae n. subfam.and in the typical Eubrachyura (most, see e.g. Ethusidae). Modified from Vehof 2020: figs 20-22).	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE749322B50FCEB4FF17FDCAF813.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071333/files/figure.png	https://doi.org/10.5281/zenodo.8071333	FIG. 33. — Callosities in Dorippoidinae n. subfam. (Dorippoides) and Dorippinae n. stat. (Dorippe): A, callosity reduced to a thin calcified band: Dorippoides facchino (Herbst, 1785), ♂ 25.3 × 32.0 mm, Malaysia, Johore, Pontian, ZRC 1991.6672. B, no apparent callosity in male Dorippoides nudipes Manning & Holthuis, 1986, ♂ 16.2 × 19.4 mm, Iran, ZRC 2017.1227. C, callosity as a thick, elongated bulge: Dorippe quadridens (Fabricius, 1793), ♂ 39.0 × 41.4 mm, Thailand, Pattani Province, ZRC 2003.0126. D-F, callosity as a hemicircular structure: close-up views from different angles: D, E. Dorippe sinica Chen, 1980. D, ♀ 36.2 × 39.5 mm, China, Guangdong, Nanao Island, ZRC 1999.0470; E, ♂ 36.6 × 38.6 mm, Japan, Kochi, Shikoku, SMF 57855. F, D. frascone (Herbst, 1785), ♂ 29.7 × 31.0 mm, Philippines, Exp. Panglao 2004, ZRC 2008.0076. G, H, callosity as an arched double cup: D. tenuipes Chen, 1980, South China Sea, ZRC 1999.0009: G, ovigerous ♀ 17.3 × 19.6 mm; H, ♂ 13.2 × 14.2 mm. Abbreviations: b, branchiostegite; c, callosity; cx2-cx5, P2-P5 coxae; f, sulcated part of coxa; l, onepiece lateroventral part of coxa; m, membrane; p, pleon; pr, process of retention of female pleonal somite 2; pl6, exposed pleurite 6; P2-P5, pereiopods 2-5; r, rim of carapace posterior margin; s, strip along posterior rim; 1, 2, pleonal somites 1, 2; t, bottom of callosity with special texture; 7, 8, thoracic sternites 7, 8.	FIG. 33. — Callosities in Dorippoidinae n. subfam. (Dorippoides) and Dorippinae n. stat. (Dorippe): A, callosity reduced to a thin calcified band: Dorippoides facchino (Herbst, 1785), ♂ 25.3 × 32.0 mm, Malaysia, Johore, Pontian, ZRC 1991.6672. B, no apparent callosity in male Dorippoides nudipes Manning & Holthuis, 1986, ♂ 16.2 × 19.4 mm, Iran, ZRC 2017.1227. C, callosity as a thick, elongated bulge: Dorippe quadridens (Fabricius, 1793), ♂ 39.0 × 41.4 mm, Thailand, Pattani Province, ZRC 2003.0126. D-F, callosity as a hemicircular structure: close-up views from different angles: D, E. Dorippe sinica Chen, 1980. D, ♀ 36.2 × 39.5 mm, China, Guangdong, Nanao Island, ZRC 1999.0470; E, ♂ 36.6 × 38.6 mm, Japan, Kochi, Shikoku, SMF 57855. F, D. frascone (Herbst, 1785), ♂ 29.7 × 31.0 mm, Philippines, Exp. Panglao 2004, ZRC 2008.0076. G, H, callosity as an arched double cup: D. tenuipes Chen, 1980, South China Sea, ZRC 1999.0009: G, ovigerous ♀ 17.3 × 19.6 mm; H, ♂ 13.2 × 14.2 mm. Abbreviations: b, branchiostegite; c, callosity; cx2-cx5, P2-P5 coxae; f, sulcated part of coxa; l, onepiece lateroventral part of coxa; m, membrane; p, pleon; pr, process of retention of female pleonal somite 2; pl6, exposed pleurite 6; P2-P5, pereiopods 2-5; r, rim of carapace posterior margin; s, strip along posterior rim; 1, 2, pleonal somites 1, 2; t, bottom of callosity with special texture; 7, 8, thoracic sternites 7, 8.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE74932FB516CEBCFF17FD0EFD88.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071255/files/figure.png	https://doi.org/10.5281/zenodo.8071255	FIG. 1. — A typical dorippid crab, Heikeopsis japonica (von Siebold, 1824), the iconic ‘samurai crab’ known in Japan as ’heike-gani’, whose carapace looks like a human face: A, the species represented by De Haan (1839: pl. 31, fig. 1, as Dorippe japonica) in Fauna Japonica: top, a female; below, chelae of a male; B, lectotype selected by Yamaguchi & Baba (1993: 300, fig. 90-A.a-2, as Heikea japonica) from type series material labelled “Types, Japan, 1823, Leg. P. H. von Siebold”, RMNH CRUS D 822 (see Fransen et al. 1997: 83); C, ♂ 25.0 × 27.3 mm, ♀ 23.5 × 26.7 mm, Japan, Chaffanjon, 174-96, E.-L. Bouvier det. 1899 Dorippe japonica, dry condition, MNHN-IU-2000-4091 (= MNHN-B4091).	FIG. 1. — A typical dorippid crab, Heikeopsis japonica (von Siebold, 1824), the iconic ‘samurai crab’ known in Japan as ’heike-gani’, whose carapace looks like a human face: A, the species represented by De Haan (1839: pl. 31, fig. 1, as Dorippe japonica) in Fauna Japonica: top, a female; below, chelae of a male; B, lectotype selected by Yamaguchi & Baba (1993: 300, fig. 90-A.a-2, as Heikea japonica) from type series material labelled “Types, Japan, 1823, Leg. P. H. von Siebold”, RMNH CRUS D 822 (see Fransen et al. 1997: 83); C, ♂ 25.0 × 27.3 mm, ♀ 23.5 × 26.7 mm, Japan, Chaffanjon, 174-96, E.-L. Bouvier det. 1899 Dorippe japonica, dry condition, MNHN-IU-2000-4091 (= MNHN-B4091).	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE74932FB516CEBCFF17FD0EFD88.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071265/files/figure.png	https://doi.org/10.5281/zenodo.8071265	FIG. 5. — Terminology of main grooves on dorsal carapace of Dorippidae: A, diagrammatic figure of Holthuis & Manning (1990: fig. 1, based on a Dorippe species) without any mention of ‘precervical groove’. B-D, terminology used in the present paper and only partially used by Holthuis & Manning (1990). B, Dorippe tenuipes Chen, 1980: after Holthuis & Manning (1990: fig. 18a); C, Heikeopsis japonica (von Siebold, 1824): after Holthuis & Manning (1990: fig. 30a, as Heikea japonica); D, Neodorippe callida (Fabricius, 1795): after Holthuis & Manning (1990: fig. 39). The strip, which runs posteriorly along the rim, was only figured in C. Abbreviations: b.c., branchiocardiac groove; c.g., cervical groove; l.b., branchial lobe; p.g., precervical groove; r, rim; s, strip.	FIG. 5. — Terminology of main grooves on dorsal carapace of Dorippidae: A, diagrammatic figure of Holthuis & Manning (1990: fig. 1, based on a Dorippe species) without any mention of ‘precervical groove’. B-D, terminology used in the present paper and only partially used by Holthuis & Manning (1990). B, Dorippe tenuipes Chen, 1980: after Holthuis & Manning (1990: fig. 18a); C, Heikeopsis japonica (von Siebold, 1824): after Holthuis & Manning (1990: fig. 30a, as Heikea japonica); D, Neodorippe callida (Fabricius, 1795): after Holthuis & Manning (1990: fig. 39). The strip, which runs posteriorly along the rim, was only figured in C. Abbreviations: b.c., branchiocardiac groove; c.g., cervical groove; l.b., branchial lobe; p.g., precervical groove; r, rim; s, strip.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE74932FB516CEBCFF17FD0EFD88.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071302/files/figure.png	https://doi.org/10.5281/zenodo.8071302	FIG. 19. — Heikeopsinae n. subfam. A-D, Heikeopsis japonica (von Siebold, 1824): A, ♂ 22.5 × 24.3 mm, Japan, Tokushima, SMF 57856: habitus; B, ♀ 23.5 × 26.7 mm, Japan, Chaffanjon, MNHN-IU-2000-4091 (= MNHN-B4091): habitus; C, ♂ 25.0 × 27.3 mm, same data as B: ventral view; D, ♂ 24.2 × 25.4 mm, same data as B and C: thoracic sternum and pleon. E, F,?Heikeopsis aff. japonica, China, seas off Qingdao, ZRC 2002.0491: habitus: E, ♂ 21.9 × 24 mm; F, ♀ 23.4 × 26.4 mm.	FIG. 19. — Heikeopsinae n. subfam. A-D, Heikeopsis japonica (von Siebold, 1824): A, ♂ 22.5 × 24.3 mm, Japan, Tokushima, SMF 57856: habitus; B, ♀ 23.5 × 26.7 mm, Japan, Chaffanjon, MNHN-IU-2000-4091 (= MNHN-B4091): habitus; C, ♂ 25.0 × 27.3 mm, same data as B: ventral view; D, ♂ 24.2 × 25.4 mm, same data as B and C: thoracic sternum and pleon. E, F,?Heikeopsis aff. japonica, China, seas off Qingdao, ZRC 2002.0491: habitus: E, ♂ 21.9 × 24 mm; F, ♀ 23.4 × 26.4 mm.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE74932FB516CEBCFF17FD0EFD88.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071305/files/figure.png	https://doi.org/10.5281/zenodo.8071305	FIG. 20. — Heikeopsinae n. subfam. Neodorippe callida (Fabricius, 1798): A, young ♂ 10.1 × 10.3 mm, Singapore, Changi Point Beach, ZRC 2018.0445: habitus of young male; B, C, Andaman Sea, Phuket, Thailand, MNHN-IU-2016-10754 (ex ZRC 1998.1116): B, ♀ 14.0 × 15.0 mm, habitus; C, ♂ 13.5 × 14 mm: ventral view. D, E, ♂ 10.1 × 10.3 mm, same data as A, ZRC 2018.0445:D, thoracic sternum with pleon; E, without pleon, G1, press-buttons. F, ♀ 11.2 × 11.5 mm, same data as A, D, E, ZRC 2018.0445: anterior view. G, ♀ 11.2 × 11.5 mm, sama data as B, C, MNHN-IU-2016-10754: vulvae, press-buttons. H, ♀ 11.2 × 11.5 mm, same data as A, D, E, F, ZRC 2018.0445: vulvae.	FIG. 20. — Heikeopsinae n. subfam. Neodorippe callida (Fabricius, 1798): A, young ♂ 10.1 × 10.3 mm, Singapore, Changi Point Beach, ZRC 2018.0445: habitus of young male; B, C, Andaman Sea, Phuket, Thailand, MNHN-IU-2016-10754 (ex ZRC 1998.1116): B, ♀ 14.0 × 15.0 mm, habitus; C, ♂ 13.5 × 14 mm: ventral view. D, E, ♂ 10.1 × 10.3 mm, same data as A, ZRC 2018.0445:D, thoracic sternum with pleon; E, without pleon, G1, press-buttons. F, ♀ 11.2 × 11.5 mm, same data as A, D, E, ZRC 2018.0445: anterior view. G, ♀ 11.2 × 11.5 mm, sama data as B, C, MNHN-IU-2016-10754: vulvae, press-buttons. H, ♀ 11.2 × 11.5 mm, same data as A, D, E, F, ZRC 2018.0445: vulvae.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE74932FB516CEBCFF17FD0EFD88.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071307/files/figure.png	https://doi.org/10.5281/zenodo.8071307	FIG. 21. — Heikeopsinae n. subfam.Nobilum histrio (Nobili,1903), Malaysia, Johore,Pontian, ZRC 2002.0491:A-D, ♂ 21.0 × 22.4 mm:habitus; B, anterior ventral view; C, thoracic sternum with pleon; D, thoracic sternum without pleon, penis and G1. E, F, ovigerous ♀ 22.1 × 24.7 mm: thoracic sternum, pleon and vulvae.	FIG. 21. — Heikeopsinae n. subfam.Nobilum histrio (Nobili,1903), Malaysia, Johore,Pontian, ZRC 2002.0491:A-D, ♂ 21.0 × 22.4 mm:habitus; B, anterior ventral view; C, thoracic sternum with pleon; D, thoracic sternum without pleon, penis and G1. E, F, ovigerous ♀ 22.1 × 24.7 mm: thoracic sternum, pleon and vulvae.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE74932FB516CEBCFF17FD0EFD88.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071329/files/figure.png	https://doi.org/10.5281/zenodo.8071329	FIG. 31. — Diversity of G1s in the seven dorippid subfamilies (schematic drawings): A, Dorippinae n. stat. (Dorippe quadridens); B, Dorippoidinae n. subfam. (Dorippoides facchino); C, Medorippinae n. subfam.(Medorippe lanata); D-F, Heikeopsinae n.subfam.:D, Heikeopsis?japonica; E, Neodorippe callida; F, Nobilum histrio; G, Paradorippinae n. subfam. (Paradorippe cathayana); H, Philippidorippinae n. subfam. (Philippidorippe philippinensis); I, Phyllodorippinae n. subfam. (Phyllodorippe armata). (A-H, after Holthuis & Manning 1990; I, after Manning & Holthuis 1981). From Sin et al. (2009: fig. 4). Setae not always figured.	FIG. 31. — Diversity of G1s in the seven dorippid subfamilies (schematic drawings): A, Dorippinae n. stat. (Dorippe quadridens); B, Dorippoidinae n. subfam. (Dorippoides facchino); C, Medorippinae n. subfam.(Medorippe lanata); D-F, Heikeopsinae n.subfam.:D, Heikeopsis?japonica; E, Neodorippe callida; F, Nobilum histrio; G, Paradorippinae n. subfam. (Paradorippe cathayana); H, Philippidorippinae n. subfam. (Philippidorippe philippinensis); I, Phyllodorippinae n. subfam. (Phyllodorippe armata). (A-H, after Holthuis & Manning 1990; I, after Manning & Holthuis 1981). From Sin et al. (2009: fig. 4). Setae not always figured.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE74932FB516CEBCFF17FD0EFD88.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071337/files/figure.png	https://doi.org/10.5281/zenodo.8071337	FIG. 35. — Schematic illustrations of female reproductive systems in representatives of four dorippid subfamilies:A, B, Dorippinae n. stat.: A, Dorippe sinica Chen, 1980, as interpreted by Hayer et al. (2016a: fig. 2); B, Dorippe quadridens (Fabricius, 1793) and D. sinica, as interpreted by Vehof et al. (2017: fig. 2A); C, Medorippinae n. subfam.: Medorippe lanata (Linnaeus, 1767), as interpreted by Vehof et al. (2017: fig. 2B); D, Paradorippinae n. subfam.: Paradorippe granulata (De Haan, 1841), as interpreted by Vehof et al. (2018b: fig. 3). E, Heikeopsinae n. subfam.: Heikeopsis japonica (von Siebold, 1824), with the same pattern shared by Neodorippe callida (Fabricius, 1798) and Nobilum histrio (Nobili, 1903), as interpreted by Vehof (2020: fig. 4). Abbreviations: a, apodeme; amb, anteromedian bursa; bu, bursa;cu, cuticle;cv, cuticular valves;bu, bursa; ev, extension of vulva;ge, glandular epithelium; mu, musculature;oc, oocyte; od, oviduct; ov, ovary; plb, posterolateral bursa; sr, seminal receptacle; vg, vagina; v, vulva.	FIG. 35. — Schematic illustrations of female reproductive systems in representatives of four dorippid subfamilies:A, B, Dorippinae n. stat.: A, Dorippe sinica Chen, 1980, as interpreted by Hayer et al. (2016a: fig. 2); B, Dorippe quadridens (Fabricius, 1793) and D. sinica, as interpreted by Vehof et al. (2017: fig. 2A); C, Medorippinae n. subfam.: Medorippe lanata (Linnaeus, 1767), as interpreted by Vehof et al. (2017: fig. 2B); D, Paradorippinae n. subfam.: Paradorippe granulata (De Haan, 1841), as interpreted by Vehof et al. (2018b: fig. 3). E, Heikeopsinae n. subfam.: Heikeopsis japonica (von Siebold, 1824), with the same pattern shared by Neodorippe callida (Fabricius, 1798) and Nobilum histrio (Nobili, 1903), as interpreted by Vehof (2020: fig. 4). Abbreviations: a, apodeme; amb, anteromedian bursa; bu, bursa;cu, cuticle;cv, cuticular valves;bu, bursa; ev, extension of vulva;ge, glandular epithelium; mu, musculature;oc, oocyte; od, oviduct; ov, ovary; plb, posterolateral bursa; sr, seminal receptacle; vg, vagina; v, vulva.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE74932FB516CEBCFF17FD0EFD88.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071341/files/figure.png	https://doi.org/10.5281/zenodo.8071341	FIG. 37. — Schematic representation of character mapping of sperm storage organs in the different dorippid subfamilies recognised here (except Phyllodorippinae n.subfam.not studied) and in a typical eubrachyuran:ovary with oocytes; oviduct; cuticular bursae (thin black line), of equal or unequal size; seminal receptacle either bilateral twin- or bilateral single,and either completely cuticular (thin black line) or partially glandular (thick black line). The twin system occurs in Dorippinae n. stat., Dorippoidinae n. subfam., Paradorippinae n. subfam. and Philippidorippinae n. subfam.; the single system occurs in Heikeopsinae n. subfam.,Medorippinae n. subfam.and in the typical Eubrachyura (most, see e.g. Ethusidae). Modified from Vehof 2020: figs 20-22).	FIG. 37. — Schematic representation of character mapping of sperm storage organs in the different dorippid subfamilies recognised here (except Phyllodorippinae n.subfam.not studied) and in a typical eubrachyuran:ovary with oocytes; oviduct; cuticular bursae (thin black line), of equal or unequal size; seminal receptacle either bilateral twin- or bilateral single,and either completely cuticular (thin black line) or partially glandular (thick black line). The twin system occurs in Dorippinae n. stat., Dorippoidinae n. subfam., Paradorippinae n. subfam. and Philippidorippinae n. subfam.; the single system occurs in Heikeopsinae n. subfam.,Medorippinae n. subfam.and in the typical Eubrachyura (most, see e.g. Ethusidae). Modified from Vehof 2020: figs 20-22).	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE74932FB516CEBCFF17FD0EFD88.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071257/files/figure.png	https://doi.org/10.5281/zenodo.8071257	FIG. 2. — Colour print by Utagawa Kuniyoshi in 1851: The ghost of Taira Tomomori along with the anchor he drowned with, and heikegani with faces of fallen soldiers; A, Heikeopsis japonica (von Siebold, 1824), the ‘heike-gani’ or ‘samurai crab’, during the naval battle at Dan-no-ura (1185) in Japan’s Inland Sea; B, closeup view of the crabs, easily recognisable by their carapaces bearing like a human face, their last two legs, reduced and carried dorsally, and the dorsally visible pleon (See also Fig. 39).	FIG. 2. — Colour print by Utagawa Kuniyoshi in 1851: The ghost of Taira Tomomori along with the anchor he drowned with, and heikegani with faces of fallen soldiers; A, Heikeopsis japonica (von Siebold, 1824), the ‘heike-gani’ or ‘samurai crab’, during the naval battle at Dan-no-ura (1185) in Japan’s Inland Sea; B, closeup view of the crabs, easily recognisable by their carapaces bearing like a human face, their last two legs, reduced and carried dorsally, and the dorsally visible pleon (See also Fig. 39).	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE74932FB516CEBCFF17FD0EFD88.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071345/files/figure.png	https://doi.org/10.5281/zenodo.8071345	FIG. 39. — The ghost of Taira Tomomori along with the anchor he drowned with, and heikegani with faces of fallen soldiers, colour print by Utagawa Kuniyoshi illustrating the legend of the iconic ‘samurai crab’ known in Japan as ’heike-gani’,a dorippid whose current scientific name is Heikeopsis japonica (von Siebold, 1824).It depicts the naval battle of Dan-no-ura in the Japanese Inland Sea in 1185 between the two clans Heike and Gengi. Members of the Heike clan and generals tie themselves to the anchor of the ship to sink to the bottom of the sea and to rejoin the drowned. The souls of deceased Heike warriors were passed on to the crabs, and their faces were etched forever on the carapaces of the crabs. In Japan, Heike-gani crabs are considered to be the reincarnations of the samurai who died at Dan-no-ura.	FIG. 39. — The ghost of Taira Tomomori along with the anchor he drowned with, and heikegani with faces of fallen soldiers, colour print by Utagawa Kuniyoshi illustrating the legend of the iconic ‘samurai crab’ known in Japan as ’heike-gani’,a dorippid whose current scientific name is Heikeopsis japonica (von Siebold, 1824).It depicts the naval battle of Dan-no-ura in the Japanese Inland Sea in 1185 between the two clans Heike and Gengi. Members of the Heike clan and generals tie themselves to the anchor of the ship to sink to the bottom of the sea and to rejoin the drowned. The souls of deceased Heike warriors were passed on to the crabs, and their faces were etched forever on the carapaces of the crabs. In Japan, Heike-gani crabs are considered to be the reincarnations of the samurai who died at Dan-no-ura.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE749336B517CD91FD56FC90FBD6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071255/files/figure.png	https://doi.org/10.5281/zenodo.8071255	FIG. 1. — A typical dorippid crab, Heikeopsis japonica (von Siebold, 1824), the iconic ‘samurai crab’ known in Japan as ’heike-gani’, whose carapace looks like a human face: A, the species represented by De Haan (1839: pl. 31, fig. 1, as Dorippe japonica) in Fauna Japonica: top, a female; below, chelae of a male; B, lectotype selected by Yamaguchi & Baba (1993: 300, fig. 90-A.a-2, as Heikea japonica) from type series material labelled “Types, Japan, 1823, Leg. P. H. von Siebold”, RMNH CRUS D 822 (see Fransen et al. 1997: 83); C, ♂ 25.0 × 27.3 mm, ♀ 23.5 × 26.7 mm, Japan, Chaffanjon, 174-96, E.-L. Bouvier det. 1899 Dorippe japonica, dry condition, MNHN-IU-2000-4091 (= MNHN-B4091).	FIG. 1. — A typical dorippid crab, Heikeopsis japonica (von Siebold, 1824), the iconic ‘samurai crab’ known in Japan as ’heike-gani’, whose carapace looks like a human face: A, the species represented by De Haan (1839: pl. 31, fig. 1, as Dorippe japonica) in Fauna Japonica: top, a female; below, chelae of a male; B, lectotype selected by Yamaguchi & Baba (1993: 300, fig. 90-A.a-2, as Heikea japonica) from type series material labelled “Types, Japan, 1823, Leg. P. H. von Siebold”, RMNH CRUS D 822 (see Fransen et al. 1997: 83); C, ♂ 25.0 × 27.3 mm, ♀ 23.5 × 26.7 mm, Japan, Chaffanjon, 174-96, E.-L. Bouvier det. 1899 Dorippe japonica, dry condition, MNHN-IU-2000-4091 (= MNHN-B4091).	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE749337B512CDE8FBB4FC36FAD5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071311/files/figure.png	https://doi.org/10.5281/zenodo.8071311	FIG. 23. — Medorippinae n. subfam.: A, B,?Medorippe crosnieri Chen, 1988. Mozambique Channel, MAINBAZA Expedition: A, ♂ 21.1 × 25.9 mm, CP 3132, MNHN-IU-2009-1998, with dimorphic chelipeds; B, young ♂ 12.0 × 15.0 mm, CP 3130, MNHN-IU-2009-1999, with symmetrical chelipeds. Photos courtesy of T.-Y.Chan.C, Medorippe lanata (Linnaeus,1767),♂ 23.2 × 28.9 mm,no data,ZRC 2009.0412 (ex MNHN):P2 merus with spinulated dorsal margin;D,?Medorippe crosnieri Chen, 1988, ♂ 21.1 × 25.9 mm, same data as in A (MNHN-IU-2009-1998): P2 merus with smooth dorsal margin.Photographs made by S. Soubzmaigne.	FIG. 23. — Medorippinae n. subfam.: A, B,?Medorippe crosnieri Chen, 1988. Mozambique Channel, MAINBAZA Expedition: A, ♂ 21.1 × 25.9 mm, CP 3132, MNHN-IU-2009-1998, with dimorphic chelipeds; B, young ♂ 12.0 × 15.0 mm, CP 3130, MNHN-IU-2009-1999, with symmetrical chelipeds. Photos courtesy of T.-Y.Chan.C, Medorippe lanata (Linnaeus,1767),♂ 23.2 × 28.9 mm,no data,ZRC 2009.0412 (ex MNHN):P2 merus with spinulated dorsal margin;D,?Medorippe crosnieri Chen, 1988, ♂ 21.1 × 25.9 mm, same data as in A (MNHN-IU-2009-1998): P2 merus with smooth dorsal margin.Photographs made by S. Soubzmaigne.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE749337B512CDE8FBB4FC36FAD5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071309/files/figure.png	https://doi.org/10.5281/zenodo.8071309	FIG. 22. — Medorippinae n. subfam. Medorippe lanata (Linnaeus, 1767): A, ♂ 18.5 × 23.2 mm, Dahomey Coasts, ZRC 2009.0411: habitus; B, ovigerous ♀ 21.2 × 27.7 mm, Israel, ZRC 1999.0632: habitus; C-E, ♂ 18.5 × 23.2 mm, same data as A: C, anterior ventral view; D, thoracic sternum, press-button and G1; E, G1 and G2. F, ♂ 23.2 × 28.9 mm, no data, ZRC 2009.0412 (ex MNHN): thoracic sternum with pleon.	FIG. 22. — Medorippinae n. subfam. Medorippe lanata (Linnaeus, 1767): A, ♂ 18.5 × 23.2 mm, Dahomey Coasts, ZRC 2009.0411: habitus; B, ovigerous ♀ 21.2 × 27.7 mm, Israel, ZRC 1999.0632: habitus; C-E, ♂ 18.5 × 23.2 mm, same data as A: C, anterior ventral view; D, thoracic sternum, press-button and G1; E, G1 and G2. F, ♂ 23.2 × 28.9 mm, no data, ZRC 2009.0412 (ex MNHN): thoracic sternum with pleon.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE749337B512CDE8FBB4FC36FAD5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071263/files/figure.png	https://doi.org/10.5281/zenodo.8071263	FIG. 4. — Dorippid thoracic sternum exemplified here by Medorippe lanata (Linnaeus,1767),Mediterranean Sea,dehydrated specimens for skeletal preparation by Sylvie Secretan (MNHN): A, ♂; B, ♀. Abbreviations: b.p., press-button located in curved sternal suture 5/6; cx4, P4 coxa: e, episternite; G1, G2, first and second gonopods; j, sternum/pterygostome junction; m, membrane; m.o., Milne Edwards opening as pterygostomial slit; p, perforation at the end of sternal suture 3/4; r, sternal ridge; s.p., sternal prominence; v, vulva; 1-8, thoracic sternites 1-8; 2/3-7/8, sternal sutures 2/3-7/8. Photographs by Barry van Bakel.	FIG. 4. — Dorippid thoracic sternum exemplified here by Medorippe lanata (Linnaeus,1767),Mediterranean Sea,dehydrated specimens for skeletal preparation by Sylvie Secretan (MNHN): A, ♂; B, ♀. Abbreviations: b.p., press-button located in curved sternal suture 5/6; cx4, P4 coxa: e, episternite; G1, G2, first and second gonopods; j, sternum/pterygostome junction; m, membrane; m.o., Milne Edwards opening as pterygostomial slit; p, perforation at the end of sternal suture 3/4; r, sternal ridge; s.p., sternal prominence; v, vulva; 1-8, thoracic sternites 1-8; 2/3-7/8, sternal sutures 2/3-7/8. Photographs by Barry van Bakel.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE749337B512CDE8FBB4FC36FAD5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071329/files/figure.png	https://doi.org/10.5281/zenodo.8071329	FIG. 31. — Diversity of G1s in the seven dorippid subfamilies (schematic drawings): A, Dorippinae n. stat. (Dorippe quadridens); B, Dorippoidinae n. subfam. (Dorippoides facchino); C, Medorippinae n. subfam.(Medorippe lanata); D-F, Heikeopsinae n.subfam.:D, Heikeopsis?japonica; E, Neodorippe callida; F, Nobilum histrio; G, Paradorippinae n. subfam. (Paradorippe cathayana); H, Philippidorippinae n. subfam. (Philippidorippe philippinensis); I, Phyllodorippinae n. subfam. (Phyllodorippe armata). (A-H, after Holthuis & Manning 1990; I, after Manning & Holthuis 1981). From Sin et al. (2009: fig. 4). Setae not always figured.	FIG. 31. — Diversity of G1s in the seven dorippid subfamilies (schematic drawings): A, Dorippinae n. stat. (Dorippe quadridens); B, Dorippoidinae n. subfam. (Dorippoides facchino); C, Medorippinae n. subfam.(Medorippe lanata); D-F, Heikeopsinae n.subfam.:D, Heikeopsis?japonica; E, Neodorippe callida; F, Nobilum histrio; G, Paradorippinae n. subfam. (Paradorippe cathayana); H, Philippidorippinae n. subfam. (Philippidorippe philippinensis); I, Phyllodorippinae n. subfam. (Phyllodorippe armata). (A-H, after Holthuis & Manning 1990; I, after Manning & Holthuis 1981). From Sin et al. (2009: fig. 4). Setae not always figured.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE749337B512CDE8FBB4FC36FAD5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071331/files/figure.png	https://doi.org/10.5281/zenodo.8071331	FIG. 32. — Diversity of vulvae in the seven dorippid subfamilies (schematic drawings): A, Dorippinae n. stat. (Dorippe quadridens); B, Dorippoidinae n. subfam. (Dorippoides facchino); C, Medorippinae n.subfam.(Medorippe lanata); D-F, Heikeopsinae n. subfam.;D, Heikeopsis?japonica; E, Neodorippe callida; F, Nobilum histrio; G, Paradorippinae n. subfam. (Paradorippe cathayana); H, Philippidorippinae n. subfam. (Philippidorippe philippinensis); I, Phyllodorippinae n. subfam. (Phyllodorippe armata, ♀ 13.0 × 19.0 mm, Guinean Trawling Survey, MNHN-IU-2021-8731 [= MNHN-B24199]). Setae represented only on B. A-H, modified from Holthuis & Manning (1990).	FIG. 32. — Diversity of vulvae in the seven dorippid subfamilies (schematic drawings): A, Dorippinae n. stat. (Dorippe quadridens); B, Dorippoidinae n. subfam. (Dorippoides facchino); C, Medorippinae n.subfam.(Medorippe lanata); D-F, Heikeopsinae n. subfam.;D, Heikeopsis?japonica; E, Neodorippe callida; F, Nobilum histrio; G, Paradorippinae n. subfam. (Paradorippe cathayana); H, Philippidorippinae n. subfam. (Philippidorippe philippinensis); I, Phyllodorippinae n. subfam. (Phyllodorippe armata, ♀ 13.0 × 19.0 mm, Guinean Trawling Survey, MNHN-IU-2021-8731 [= MNHN-B24199]). Setae represented only on B. A-H, modified from Holthuis & Manning (1990).	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE749337B512CDE8FBB4FC36FAD5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071337/files/figure.png	https://doi.org/10.5281/zenodo.8071337	FIG. 35. — Schematic illustrations of female reproductive systems in representatives of four dorippid subfamilies:A, B, Dorippinae n. stat.: A, Dorippe sinica Chen, 1980, as interpreted by Hayer et al. (2016a: fig. 2); B, Dorippe quadridens (Fabricius, 1793) and D. sinica, as interpreted by Vehof et al. (2017: fig. 2A); C, Medorippinae n. subfam.: Medorippe lanata (Linnaeus, 1767), as interpreted by Vehof et al. (2017: fig. 2B); D, Paradorippinae n. subfam.: Paradorippe granulata (De Haan, 1841), as interpreted by Vehof et al. (2018b: fig. 3). E, Heikeopsinae n. subfam.: Heikeopsis japonica (von Siebold, 1824), with the same pattern shared by Neodorippe callida (Fabricius, 1798) and Nobilum histrio (Nobili, 1903), as interpreted by Vehof (2020: fig. 4). Abbreviations: a, apodeme; amb, anteromedian bursa; bu, bursa;cu, cuticle;cv, cuticular valves;bu, bursa; ev, extension of vulva;ge, glandular epithelium; mu, musculature;oc, oocyte; od, oviduct; ov, ovary; plb, posterolateral bursa; sr, seminal receptacle; vg, vagina; v, vulva.	FIG. 35. — Schematic illustrations of female reproductive systems in representatives of four dorippid subfamilies:A, B, Dorippinae n. stat.: A, Dorippe sinica Chen, 1980, as interpreted by Hayer et al. (2016a: fig. 2); B, Dorippe quadridens (Fabricius, 1793) and D. sinica, as interpreted by Vehof et al. (2017: fig. 2A); C, Medorippinae n. subfam.: Medorippe lanata (Linnaeus, 1767), as interpreted by Vehof et al. (2017: fig. 2B); D, Paradorippinae n. subfam.: Paradorippe granulata (De Haan, 1841), as interpreted by Vehof et al. (2018b: fig. 3). E, Heikeopsinae n. subfam.: Heikeopsis japonica (von Siebold, 1824), with the same pattern shared by Neodorippe callida (Fabricius, 1798) and Nobilum histrio (Nobili, 1903), as interpreted by Vehof (2020: fig. 4). Abbreviations: a, apodeme; amb, anteromedian bursa; bu, bursa;cu, cuticle;cv, cuticular valves;bu, bursa; ev, extension of vulva;ge, glandular epithelium; mu, musculature;oc, oocyte; od, oviduct; ov, ovary; plb, posterolateral bursa; sr, seminal receptacle; vg, vagina; v, vulva.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE749337B512CDE8FBB4FC36FAD5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071341/files/figure.png	https://doi.org/10.5281/zenodo.8071341	FIG. 37. — Schematic representation of character mapping of sperm storage organs in the different dorippid subfamilies recognised here (except Phyllodorippinae n.subfam.not studied) and in a typical eubrachyuran:ovary with oocytes; oviduct; cuticular bursae (thin black line), of equal or unequal size; seminal receptacle either bilateral twin- or bilateral single,and either completely cuticular (thin black line) or partially glandular (thick black line). The twin system occurs in Dorippinae n. stat., Dorippoidinae n. subfam., Paradorippinae n. subfam. and Philippidorippinae n. subfam.; the single system occurs in Heikeopsinae n. subfam.,Medorippinae n. subfam.and in the typical Eubrachyura (most, see e.g. Ethusidae). Modified from Vehof 2020: figs 20-22).	FIG. 37. — Schematic representation of character mapping of sperm storage organs in the different dorippid subfamilies recognised here (except Phyllodorippinae n.subfam.not studied) and in a typical eubrachyuran:ovary with oocytes; oviduct; cuticular bursae (thin black line), of equal or unequal size; seminal receptacle either bilateral twin- or bilateral single,and either completely cuticular (thin black line) or partially glandular (thick black line). The twin system occurs in Dorippinae n. stat., Dorippoidinae n. subfam., Paradorippinae n. subfam. and Philippidorippinae n. subfam.; the single system occurs in Heikeopsinae n. subfam.,Medorippinae n. subfam.and in the typical Eubrachyura (most, see e.g. Ethusidae). Modified from Vehof 2020: figs 20-22).	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE749332B51CCE38FA94FEC4FE89.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071311/files/figure.png	https://doi.org/10.5281/zenodo.8071311	FIG. 23. — Medorippinae n. subfam.: A, B,?Medorippe crosnieri Chen, 1988. Mozambique Channel, MAINBAZA Expedition: A, ♂ 21.1 × 25.9 mm, CP 3132, MNHN-IU-2009-1998, with dimorphic chelipeds; B, young ♂ 12.0 × 15.0 mm, CP 3130, MNHN-IU-2009-1999, with symmetrical chelipeds. Photos courtesy of T.-Y.Chan.C, Medorippe lanata (Linnaeus,1767),♂ 23.2 × 28.9 mm,no data,ZRC 2009.0412 (ex MNHN):P2 merus with spinulated dorsal margin;D,?Medorippe crosnieri Chen, 1988, ♂ 21.1 × 25.9 mm, same data as in A (MNHN-IU-2009-1998): P2 merus with smooth dorsal margin.Photographs made by S. Soubzmaigne.	FIG. 23. — Medorippinae n. subfam.: A, B,?Medorippe crosnieri Chen, 1988. Mozambique Channel, MAINBAZA Expedition: A, ♂ 21.1 × 25.9 mm, CP 3132, MNHN-IU-2009-1998, with dimorphic chelipeds; B, young ♂ 12.0 × 15.0 mm, CP 3130, MNHN-IU-2009-1999, with symmetrical chelipeds. Photos courtesy of T.-Y.Chan.C, Medorippe lanata (Linnaeus,1767),♂ 23.2 × 28.9 mm,no data,ZRC 2009.0412 (ex MNHN):P2 merus with spinulated dorsal margin;D,?Medorippe crosnieri Chen, 1988, ♂ 21.1 × 25.9 mm, same data as in A (MNHN-IU-2009-1998): P2 merus with smooth dorsal margin.Photographs made by S. Soubzmaigne.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE749332B51CCE38FA94FEC4FE89.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071309/files/figure.png	https://doi.org/10.5281/zenodo.8071309	FIG. 22. — Medorippinae n. subfam. Medorippe lanata (Linnaeus, 1767): A, ♂ 18.5 × 23.2 mm, Dahomey Coasts, ZRC 2009.0411: habitus; B, ovigerous ♀ 21.2 × 27.7 mm, Israel, ZRC 1999.0632: habitus; C-E, ♂ 18.5 × 23.2 mm, same data as A: C, anterior ventral view; D, thoracic sternum, press-button and G1; E, G1 and G2. F, ♂ 23.2 × 28.9 mm, no data, ZRC 2009.0412 (ex MNHN): thoracic sternum with pleon.	FIG. 22. — Medorippinae n. subfam. Medorippe lanata (Linnaeus, 1767): A, ♂ 18.5 × 23.2 mm, Dahomey Coasts, ZRC 2009.0411: habitus; B, ovigerous ♀ 21.2 × 27.7 mm, Israel, ZRC 1999.0632: habitus; C-E, ♂ 18.5 × 23.2 mm, same data as A: C, anterior ventral view; D, thoracic sternum, press-button and G1; E, G1 and G2. F, ♂ 23.2 × 28.9 mm, no data, ZRC 2009.0412 (ex MNHN): thoracic sternum with pleon.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE749332B51CCE38FA94FEC4FE89.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071293/files/figure.png	https://doi.org/10.5281/zenodo.8071293	FIG. 16. — Dorippoidinae n. subfam.: A, B, Dorippoides facchino (Herbst, 1785): A, ♂ 25.3 × 32 mm, Malaysia, Johore, Pontian, ZRC 1991.66-72; B, ovigerous ♀ 20.3 × 26.2 mm, South China Sea, Hee Huat, ZRC 1984.5347; C, D, Dorippoides nudipes Manning & Holthuis,1986: C, ♂ 16.2 × 19.4 mm, Iran,ZRC 2017.1227; D, ♀ 21.6 × 29.5 mm, South Africa, vicinity of Tugela River mouth, ZRC 2009.0885.	FIG. 16. — Dorippoidinae n. subfam.: A, B, Dorippoides facchino (Herbst, 1785): A, ♂ 25.3 × 32 mm, Malaysia, Johore, Pontian, ZRC 1991.66-72; B, ovigerous ♀ 20.3 × 26.2 mm, South China Sea, Hee Huat, ZRC 1984.5347; C, D, Dorippoides nudipes Manning & Holthuis,1986: C, ♂ 16.2 × 19.4 mm, Iran,ZRC 2017.1227; D, ♀ 21.6 × 29.5 mm, South Africa, vicinity of Tugela River mouth, ZRC 2009.0885.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE749332B51CCE38FA94FEC4FE89.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071325/files/figure.png	https://doi.org/10.5281/zenodo.8071325	FIG. 29. — Phyllodorippinae n. subfam. Phyllodorippe armata (Miers, 1881), Guinean Trawling Survey,ZRC 2009.0413 (ex MNHN-B24202). A, ♂ 12.3 × 16.2 mm, habitus; B-D, ovigerous ♀ 11.9 × 16.6 mm. B, habitus; C, thoracic sternum and pleon; D, thoracic sternum, vulvae and spine on sternite 8.	FIG. 29. — Phyllodorippinae n. subfam. Phyllodorippe armata (Miers, 1881), Guinean Trawling Survey,ZRC 2009.0413 (ex MNHN-B24202). A, ♂ 12.3 × 16.2 mm, habitus; B-D, ovigerous ♀ 11.9 × 16.6 mm. B, habitus; C, thoracic sternum and pleon; D, thoracic sternum, vulvae and spine on sternite 8.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE749332B51CCE38FA94FEC4FE89.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071327/files/figure.png	https://doi.org/10.5281/zenodo.8071327	FIG. 30. — Phyllodorippinae n. subfam. Phyllodorippe armata (Miers, 1881), ♂ 12.3 × 16.2 mm, Guinean Trawling Survey, ZRC 2009.0413 (ex MNHN-B24202): A, both chelae; B, thoracic sternum and pleon; C, thoracic sternum without pleon; D, G1 and press-buttons.	FIG. 30. — Phyllodorippinae n. subfam. Phyllodorippe armata (Miers, 1881), ♂ 12.3 × 16.2 mm, Guinean Trawling Survey, ZRC 2009.0413 (ex MNHN-B24202): A, both chelae; B, thoracic sternum and pleon; C, thoracic sternum without pleon; D, G1 and press-buttons.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE74933CB519CDE1FE76FAAAF813.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071279/files/figure.png	https://doi.org/10.5281/zenodo.8071279	FIG. 10. — Dorippinae H. Milne Edwards, 1837 n. stat.: habitus: A, B, Dorippe quadridens (Fabricius,1793):A, ♂ 36.3 × 38.1 mm, China Sea near Singapore, Hee Huat, ZRC 1984. 6308; B, ♀ 29.5 × 30 mm, NW Madagascar, Ambaro Bay, MNHN-IU-2018-5193 (= MNHN-B18279). C, D, Dorippe sinica Chen, 1980, China, Guangdong, Nanao Island, ZRC 1999.0470: C, ♂ 36.2 × 39.5 mm; D, ♀ 34.8 × 38.9 mm, specimen brushed.	FIG. 10. — Dorippinae H. Milne Edwards, 1837 n. stat.: habitus: A, B, Dorippe quadridens (Fabricius,1793):A, ♂ 36.3 × 38.1 mm, China Sea near Singapore, Hee Huat, ZRC 1984. 6308; B, ♀ 29.5 × 30 mm, NW Madagascar, Ambaro Bay, MNHN-IU-2018-5193 (= MNHN-B18279). C, D, Dorippe sinica Chen, 1980, China, Guangdong, Nanao Island, ZRC 1999.0470: C, ♂ 36.2 × 39.5 mm; D, ♀ 34.8 × 38.9 mm, specimen brushed.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE74933CB519CDE1FE76FAAAF813.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071313/files/figure.png	https://doi.org/10.5281/zenodo.8071313	FIG. 24. — Paradorippinae n. subfam. Paradorippe granulata (De Haan, 1841): habitus: A, ♂ 23.6 × 27.3 mm, NE Taiwan,I-Lan county, ZRC 2001.0014; B, ovigerous ♀ 23. 2 × 25.7 mm, Japan, off Hota, ZRC 1999.0082.	FIG. 24. — Paradorippinae n. subfam. Paradorippe granulata (De Haan, 1841): habitus: A, ♂ 23.6 × 27.3 mm, NE Taiwan,I-Lan county, ZRC 2001.0014; B, ovigerous ♀ 23. 2 × 25.7 mm, Japan, off Hota, ZRC 1999.0082.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE74933CB519CDE1FE76FAAAF813.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071315/files/figure.png	https://doi.org/10.5281/zenodo.8071315	FIG. 25. — Paradorippinae n. subfam. Paradorippe granulata (De Haan, 1841): A, C, E, ♂ 24.0 × 26.0 mm, China, Tuandao, off Quingdao, MNHN-IU-2016-10753. B, D, ♂ 23.6 × 27.3 mm, NE Taiwan, I-Lan county, ZRC 2001.0014; A, chelae; B, thoracic sternum and pleon; C, D, thoracic sternum without pleon, with G1, G2 and press-buttons; E, G1 and G2 in situ.	FIG. 25. — Paradorippinae n. subfam. Paradorippe granulata (De Haan, 1841): A, C, E, ♂ 24.0 × 26.0 mm, China, Tuandao, off Quingdao, MNHN-IU-2016-10753. B, D, ♂ 23.6 × 27.3 mm, NE Taiwan, I-Lan county, ZRC 2001.0014; A, chelae; B, thoracic sternum and pleon; C, D, thoracic sternum without pleon, with G1, G2 and press-buttons; E, G1 and G2 in situ.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE74933CB519CDE1FE76FAAAF813.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071319/files/figure.png	https://doi.org/10.5281/zenodo.8071319	FIG. 26. — Paradorippinae n. subfam. Paradorippe granulata (De Haan, 1841): A-C, ovigerous ♀ 23.2 × 25.7 mm, Japan, off Hota, ZRC 1999.0082. D, ♀ 22.7 × 24, 7 mm, China, Tuandao, off Quingdao, MNHN-IU-2016-10753. A, anterior ventral view; B, C, thoracic sternum and pleon; D, vulvae.	FIG. 26. — Paradorippinae n. subfam. Paradorippe granulata (De Haan, 1841): A-C, ovigerous ♀ 23.2 × 25.7 mm, Japan, off Hota, ZRC 1999.0082. D, ♀ 22.7 × 24, 7 mm, China, Tuandao, off Quingdao, MNHN-IU-2016-10753. A, anterior ventral view; B, C, thoracic sternum and pleon; D, vulvae.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE74933CB519CDE1FE76FAAAF813.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071337/files/figure.png	https://doi.org/10.5281/zenodo.8071337	FIG. 35. — Schematic illustrations of female reproductive systems in representatives of four dorippid subfamilies:A, B, Dorippinae n. stat.: A, Dorippe sinica Chen, 1980, as interpreted by Hayer et al. (2016a: fig. 2); B, Dorippe quadridens (Fabricius, 1793) and D. sinica, as interpreted by Vehof et al. (2017: fig. 2A); C, Medorippinae n. subfam.: Medorippe lanata (Linnaeus, 1767), as interpreted by Vehof et al. (2017: fig. 2B); D, Paradorippinae n. subfam.: Paradorippe granulata (De Haan, 1841), as interpreted by Vehof et al. (2018b: fig. 3). E, Heikeopsinae n. subfam.: Heikeopsis japonica (von Siebold, 1824), with the same pattern shared by Neodorippe callida (Fabricius, 1798) and Nobilum histrio (Nobili, 1903), as interpreted by Vehof (2020: fig. 4). Abbreviations: a, apodeme; amb, anteromedian bursa; bu, bursa;cu, cuticle;cv, cuticular valves;bu, bursa; ev, extension of vulva;ge, glandular epithelium; mu, musculature;oc, oocyte; od, oviduct; ov, ovary; plb, posterolateral bursa; sr, seminal receptacle; vg, vagina; v, vulva.	FIG. 35. — Schematic illustrations of female reproductive systems in representatives of four dorippid subfamilies:A, B, Dorippinae n. stat.: A, Dorippe sinica Chen, 1980, as interpreted by Hayer et al. (2016a: fig. 2); B, Dorippe quadridens (Fabricius, 1793) and D. sinica, as interpreted by Vehof et al. (2017: fig. 2A); C, Medorippinae n. subfam.: Medorippe lanata (Linnaeus, 1767), as interpreted by Vehof et al. (2017: fig. 2B); D, Paradorippinae n. subfam.: Paradorippe granulata (De Haan, 1841), as interpreted by Vehof et al. (2018b: fig. 3). E, Heikeopsinae n. subfam.: Heikeopsis japonica (von Siebold, 1824), with the same pattern shared by Neodorippe callida (Fabricius, 1798) and Nobilum histrio (Nobili, 1903), as interpreted by Vehof (2020: fig. 4). Abbreviations: a, apodeme; amb, anteromedian bursa; bu, bursa;cu, cuticle;cv, cuticular valves;bu, bursa; ev, extension of vulva;ge, glandular epithelium; mu, musculature;oc, oocyte; od, oviduct; ov, ovary; plb, posterolateral bursa; sr, seminal receptacle; vg, vagina; v, vulva.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE74933CB519CDE1FE76FAAAF813.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071341/files/figure.png	https://doi.org/10.5281/zenodo.8071341	FIG. 37. — Schematic representation of character mapping of sperm storage organs in the different dorippid subfamilies recognised here (except Phyllodorippinae n.subfam.not studied) and in a typical eubrachyuran:ovary with oocytes; oviduct; cuticular bursae (thin black line), of equal or unequal size; seminal receptacle either bilateral twin- or bilateral single,and either completely cuticular (thin black line) or partially glandular (thick black line). The twin system occurs in Dorippinae n. stat., Dorippoidinae n. subfam., Paradorippinae n. subfam. and Philippidorippinae n. subfam.; the single system occurs in Heikeopsinae n. subfam.,Medorippinae n. subfam.and in the typical Eubrachyura (most, see e.g. Ethusidae). Modified from Vehof 2020: figs 20-22).	FIG. 37. — Schematic representation of character mapping of sperm storage organs in the different dorippid subfamilies recognised here (except Phyllodorippinae n.subfam.not studied) and in a typical eubrachyuran:ovary with oocytes; oviduct; cuticular bursae (thin black line), of equal or unequal size; seminal receptacle either bilateral twin- or bilateral single,and either completely cuticular (thin black line) or partially glandular (thick black line). The twin system occurs in Dorippinae n. stat., Dorippoidinae n. subfam., Paradorippinae n. subfam. and Philippidorippinae n. subfam.; the single system occurs in Heikeopsinae n. subfam.,Medorippinae n. subfam.and in the typical Eubrachyura (most, see e.g. Ethusidae). Modified from Vehof 2020: figs 20-22).	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE74933AB565CDF1FF17FDAEF833.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071321/files/figure.png	https://doi.org/10.5281/zenodo.8071321	FIG. 27. — Philippidorippinae n. subfam. Philippidorippe philippinensis Chen, 1986, Philippines, Bohol Province, ZRC 2016.0240: A, ♂ 23.0 × 29.3 mm: habitus; B-D, ovigerous ♀ 25.2 × 33.2 mm; B, habitus; C, thoracic sternum with pleon, press-button; D, vulvae.	FIG. 27. — Philippidorippinae n. subfam. Philippidorippe philippinensis Chen, 1986, Philippines, Bohol Province, ZRC 2016.0240: A, ♂ 23.0 × 29.3 mm: habitus; B-D, ovigerous ♀ 25.2 × 33.2 mm; B, habitus; C, thoracic sternum with pleon, press-button; D, vulvae.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE74933AB565CDF1FF17FDAEF833.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071323/files/figure.png	https://doi.org/10.5281/zenodo.8071323	FIG. 28. — Philippidorippinae n. subfam.Philippidorippe philippinensis Chen,1986:A-D, ♂ 23.0 × 29.3 mm, Philippines,Bohol Province, ZRC 2016.0240:A, ventral view; B, anterior region; C, thoracic sternum with pleon; D, thoracic sternum, G1. E, ♂ 25.0 × 30.0 mm, Philippines, MUSORSTOM 3, MNHN-IU-2018-5201 (= MNHN-B18913): thoracic sternum, G1 and G2.	FIG. 28. — Philippidorippinae n. subfam.Philippidorippe philippinensis Chen,1986:A-D, ♂ 23.0 × 29.3 mm, Philippines,Bohol Province, ZRC 2016.0240:A, ventral view; B, anterior region; C, thoracic sternum with pleon; D, thoracic sternum, G1. E, ♂ 25.0 × 30.0 mm, Philippines, MUSORSTOM 3, MNHN-IU-2018-5201 (= MNHN-B18913): thoracic sternum, G1 and G2.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE74933AB565CDF1FF17FDAEF833.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071329/files/figure.png	https://doi.org/10.5281/zenodo.8071329	FIG. 31. — Diversity of G1s in the seven dorippid subfamilies (schematic drawings): A, Dorippinae n. stat. (Dorippe quadridens); B, Dorippoidinae n. subfam. (Dorippoides facchino); C, Medorippinae n. subfam.(Medorippe lanata); D-F, Heikeopsinae n.subfam.:D, Heikeopsis?japonica; E, Neodorippe callida; F, Nobilum histrio; G, Paradorippinae n. subfam. (Paradorippe cathayana); H, Philippidorippinae n. subfam. (Philippidorippe philippinensis); I, Phyllodorippinae n. subfam. (Phyllodorippe armata). (A-H, after Holthuis & Manning 1990; I, after Manning & Holthuis 1981). From Sin et al. (2009: fig. 4). Setae not always figured.	FIG. 31. — Diversity of G1s in the seven dorippid subfamilies (schematic drawings): A, Dorippinae n. stat. (Dorippe quadridens); B, Dorippoidinae n. subfam. (Dorippoides facchino); C, Medorippinae n. subfam.(Medorippe lanata); D-F, Heikeopsinae n.subfam.:D, Heikeopsis?japonica; E, Neodorippe callida; F, Nobilum histrio; G, Paradorippinae n. subfam. (Paradorippe cathayana); H, Philippidorippinae n. subfam. (Philippidorippe philippinensis); I, Phyllodorippinae n. subfam. (Phyllodorippe armata). (A-H, after Holthuis & Manning 1990; I, after Manning & Holthuis 1981). From Sin et al. (2009: fig. 4). Setae not always figured.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE74933AB565CDF1FF17FDAEF833.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071331/files/figure.png	https://doi.org/10.5281/zenodo.8071331	FIG. 32. — Diversity of vulvae in the seven dorippid subfamilies (schematic drawings): A, Dorippinae n. stat. (Dorippe quadridens); B, Dorippoidinae n. subfam. (Dorippoides facchino); C, Medorippinae n.subfam.(Medorippe lanata); D-F, Heikeopsinae n. subfam.;D, Heikeopsis?japonica; E, Neodorippe callida; F, Nobilum histrio; G, Paradorippinae n. subfam. (Paradorippe cathayana); H, Philippidorippinae n. subfam. (Philippidorippe philippinensis); I, Phyllodorippinae n. subfam. (Phyllodorippe armata, ♀ 13.0 × 19.0 mm, Guinean Trawling Survey, MNHN-IU-2021-8731 [= MNHN-B24199]). Setae represented only on B. A-H, modified from Holthuis & Manning (1990).	FIG. 32. — Diversity of vulvae in the seven dorippid subfamilies (schematic drawings): A, Dorippinae n. stat. (Dorippe quadridens); B, Dorippoidinae n. subfam. (Dorippoides facchino); C, Medorippinae n.subfam.(Medorippe lanata); D-F, Heikeopsinae n. subfam.;D, Heikeopsis?japonica; E, Neodorippe callida; F, Nobilum histrio; G, Paradorippinae n. subfam. (Paradorippe cathayana); H, Philippidorippinae n. subfam. (Philippidorippe philippinensis); I, Phyllodorippinae n. subfam. (Phyllodorippe armata, ♀ 13.0 × 19.0 mm, Guinean Trawling Survey, MNHN-IU-2021-8731 [= MNHN-B24199]). Setae represented only on B. A-H, modified from Holthuis & Manning (1990).	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE74933AB565CDF1FF17FDAEF833.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071341/files/figure.png	https://doi.org/10.5281/zenodo.8071341	FIG. 37. — Schematic representation of character mapping of sperm storage organs in the different dorippid subfamilies recognised here (except Phyllodorippinae n.subfam.not studied) and in a typical eubrachyuran:ovary with oocytes; oviduct; cuticular bursae (thin black line), of equal or unequal size; seminal receptacle either bilateral twin- or bilateral single,and either completely cuticular (thin black line) or partially glandular (thick black line). The twin system occurs in Dorippinae n. stat., Dorippoidinae n. subfam., Paradorippinae n. subfam. and Philippidorippinae n. subfam.; the single system occurs in Heikeopsinae n. subfam.,Medorippinae n. subfam.and in the typical Eubrachyura (most, see e.g. Ethusidae). Modified from Vehof 2020: figs 20-22).	FIG. 37. — Schematic representation of character mapping of sperm storage organs in the different dorippid subfamilies recognised here (except Phyllodorippinae n.subfam.not studied) and in a typical eubrachyuran:ovary with oocytes; oviduct; cuticular bursae (thin black line), of equal or unequal size; seminal receptacle either bilateral twin- or bilateral single,and either completely cuticular (thin black line) or partially glandular (thick black line). The twin system occurs in Dorippinae n. stat., Dorippoidinae n. subfam., Paradorippinae n. subfam. and Philippidorippinae n. subfam.; the single system occurs in Heikeopsinae n. subfam.,Medorippinae n. subfam.and in the typical Eubrachyura (most, see e.g. Ethusidae). Modified from Vehof 2020: figs 20-22).	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE749345B560CE9BFF17FBB3FE89.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071325/files/figure.png	https://doi.org/10.5281/zenodo.8071325	FIG. 29. — Phyllodorippinae n. subfam. Phyllodorippe armata (Miers, 1881), Guinean Trawling Survey,ZRC 2009.0413 (ex MNHN-B24202). A, ♂ 12.3 × 16.2 mm, habitus; B-D, ovigerous ♀ 11.9 × 16.6 mm. B, habitus; C, thoracic sternum and pleon; D, thoracic sternum, vulvae and spine on sternite 8.	FIG. 29. — Phyllodorippinae n. subfam. Phyllodorippe armata (Miers, 1881), Guinean Trawling Survey,ZRC 2009.0413 (ex MNHN-B24202). A, ♂ 12.3 × 16.2 mm, habitus; B-D, ovigerous ♀ 11.9 × 16.6 mm. B, habitus; C, thoracic sternum and pleon; D, thoracic sternum, vulvae and spine on sternite 8.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE749345B560CE9BFF17FBB3FE89.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071327/files/figure.png	https://doi.org/10.5281/zenodo.8071327	FIG. 30. — Phyllodorippinae n. subfam. Phyllodorippe armata (Miers, 1881), ♂ 12.3 × 16.2 mm, Guinean Trawling Survey, ZRC 2009.0413 (ex MNHN-B24202): A, both chelae; B, thoracic sternum and pleon; C, thoracic sternum without pleon; D, G1 and press-buttons.	FIG. 30. — Phyllodorippinae n. subfam. Phyllodorippe armata (Miers, 1881), ♂ 12.3 × 16.2 mm, Guinean Trawling Survey, ZRC 2009.0413 (ex MNHN-B24202): A, both chelae; B, thoracic sternum and pleon; C, thoracic sternum without pleon; D, G1 and press-buttons.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE749345B560CE9BFF17FBB3FE89.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071329/files/figure.png	https://doi.org/10.5281/zenodo.8071329	FIG. 31. — Diversity of G1s in the seven dorippid subfamilies (schematic drawings): A, Dorippinae n. stat. (Dorippe quadridens); B, Dorippoidinae n. subfam. (Dorippoides facchino); C, Medorippinae n. subfam.(Medorippe lanata); D-F, Heikeopsinae n.subfam.:D, Heikeopsis?japonica; E, Neodorippe callida; F, Nobilum histrio; G, Paradorippinae n. subfam. (Paradorippe cathayana); H, Philippidorippinae n. subfam. (Philippidorippe philippinensis); I, Phyllodorippinae n. subfam. (Phyllodorippe armata). (A-H, after Holthuis & Manning 1990; I, after Manning & Holthuis 1981). From Sin et al. (2009: fig. 4). Setae not always figured.	FIG. 31. — Diversity of G1s in the seven dorippid subfamilies (schematic drawings): A, Dorippinae n. stat. (Dorippe quadridens); B, Dorippoidinae n. subfam. (Dorippoides facchino); C, Medorippinae n. subfam.(Medorippe lanata); D-F, Heikeopsinae n.subfam.:D, Heikeopsis?japonica; E, Neodorippe callida; F, Nobilum histrio; G, Paradorippinae n. subfam. (Paradorippe cathayana); H, Philippidorippinae n. subfam. (Philippidorippe philippinensis); I, Phyllodorippinae n. subfam. (Phyllodorippe armata). (A-H, after Holthuis & Manning 1990; I, after Manning & Holthuis 1981). From Sin et al. (2009: fig. 4). Setae not always figured.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE749345B560CE9BFF17FBB3FE89.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071331/files/figure.png	https://doi.org/10.5281/zenodo.8071331	FIG. 32. — Diversity of vulvae in the seven dorippid subfamilies (schematic drawings): A, Dorippinae n. stat. (Dorippe quadridens); B, Dorippoidinae n. subfam. (Dorippoides facchino); C, Medorippinae n.subfam.(Medorippe lanata); D-F, Heikeopsinae n. subfam.;D, Heikeopsis?japonica; E, Neodorippe callida; F, Nobilum histrio; G, Paradorippinae n. subfam. (Paradorippe cathayana); H, Philippidorippinae n. subfam. (Philippidorippe philippinensis); I, Phyllodorippinae n. subfam. (Phyllodorippe armata, ♀ 13.0 × 19.0 mm, Guinean Trawling Survey, MNHN-IU-2021-8731 [= MNHN-B24199]). Setae represented only on B. A-H, modified from Holthuis & Manning (1990).	FIG. 32. — Diversity of vulvae in the seven dorippid subfamilies (schematic drawings): A, Dorippinae n. stat. (Dorippe quadridens); B, Dorippoidinae n. subfam. (Dorippoides facchino); C, Medorippinae n.subfam.(Medorippe lanata); D-F, Heikeopsinae n. subfam.;D, Heikeopsis?japonica; E, Neodorippe callida; F, Nobilum histrio; G, Paradorippinae n. subfam. (Paradorippe cathayana); H, Philippidorippinae n. subfam. (Philippidorippe philippinensis); I, Phyllodorippinae n. subfam. (Phyllodorippe armata, ♀ 13.0 × 19.0 mm, Guinean Trawling Survey, MNHN-IU-2021-8731 [= MNHN-B24199]). Setae represented only on B. A-H, modified from Holthuis & Manning (1990).	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE749383B5A3CDC6FA72FB8BFE29.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071259/files/figure.png	https://doi.org/10.5281/zenodo.8071259	FIG. 3. — Carapace diversity in the nine dorippid genera: A, Dorippe Weber, 1795 (D. quadridens); B, Dorippoides Serène & Romimohtarto, 1969 (D. facchino); C, Medorippe Manning & Holthuis, 1981 (?M. crosnieri); D, Heikeopsis Ng, Guinot & Davie, 2008 (H. aff. japonica); E, Neodorippe Serène & Romimohtarto, 1969 (N. callida); F, Nobilum Serène & Romimohtarto, 1969 (N. histrio); G, Paradorippe Serène & Romimohtarto, 1969 (P. cathayana); H, Philippidorippe Chen, 1986 (P. philippinensis); I, Phyllodorippe Manning & Holthuis, 1981 (P. armata). A-G, I: modified from Sin et al. (2009: fig. 3); H, from Chen (1986: pl. 1, fig. 3). The strip, which is shown here only in Dorippoides facchino (Herbst, 1785) and Medorippe?crosnieri Chen, 1988, is in fact present in all species of dorippids.	FIG. 3. — Carapace diversity in the nine dorippid genera: A, Dorippe Weber, 1795 (D. quadridens); B, Dorippoides Serène & Romimohtarto, 1969 (D. facchino); C, Medorippe Manning & Holthuis, 1981 (?M. crosnieri); D, Heikeopsis Ng, Guinot & Davie, 2008 (H. aff. japonica); E, Neodorippe Serène & Romimohtarto, 1969 (N. callida); F, Nobilum Serène & Romimohtarto, 1969 (N. histrio); G, Paradorippe Serène & Romimohtarto, 1969 (P. cathayana); H, Philippidorippe Chen, 1986 (P. philippinensis); I, Phyllodorippe Manning & Holthuis, 1981 (P. armata). A-G, I: modified from Sin et al. (2009: fig. 3); H, from Chen (1986: pl. 1, fig. 3). The strip, which is shown here only in Dorippoides facchino (Herbst, 1785) and Medorippe?crosnieri Chen, 1988, is in fact present in all species of dorippids.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE749383B5A3CEA6FDD6FB89FC97.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071293/files/figure.png	https://doi.org/10.5281/zenodo.8071293	FIG. 16. — Dorippoidinae n. subfam.: A, B, Dorippoides facchino (Herbst, 1785): A, ♂ 25.3 × 32 mm, Malaysia, Johore, Pontian, ZRC 1991.66-72; B, ovigerous ♀ 20.3 × 26.2 mm, South China Sea, Hee Huat, ZRC 1984.5347; C, D, Dorippoides nudipes Manning & Holthuis,1986: C, ♂ 16.2 × 19.4 mm, Iran,ZRC 2017.1227; D, ♀ 21.6 × 29.5 mm, South Africa, vicinity of Tugela River mouth, ZRC 2009.0885.	FIG. 16. — Dorippoidinae n. subfam.: A, B, Dorippoides facchino (Herbst, 1785): A, ♂ 25.3 × 32 mm, Malaysia, Johore, Pontian, ZRC 1991.66-72; B, ovigerous ♀ 20.3 × 26.2 mm, South China Sea, Hee Huat, ZRC 1984.5347; C, D, Dorippoides nudipes Manning & Holthuis,1986: C, ♂ 16.2 × 19.4 mm, Iran,ZRC 2017.1227; D, ♀ 21.6 × 29.5 mm, South Africa, vicinity of Tugela River mouth, ZRC 2009.0885.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE74938CB5ACCC18FE56FDEAFB77.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071259/files/figure.png	https://doi.org/10.5281/zenodo.8071259	FIG. 3. — Carapace diversity in the nine dorippid genera: A, Dorippe Weber, 1795 (D. quadridens); B, Dorippoides Serène & Romimohtarto, 1969 (D. facchino); C, Medorippe Manning & Holthuis, 1981 (?M. crosnieri); D, Heikeopsis Ng, Guinot & Davie, 2008 (H. aff. japonica); E, Neodorippe Serène & Romimohtarto, 1969 (N. callida); F, Nobilum Serène & Romimohtarto, 1969 (N. histrio); G, Paradorippe Serène & Romimohtarto, 1969 (P. cathayana); H, Philippidorippe Chen, 1986 (P. philippinensis); I, Phyllodorippe Manning & Holthuis, 1981 (P. armata). A-G, I: modified from Sin et al. (2009: fig. 3); H, from Chen (1986: pl. 1, fig. 3). The strip, which is shown here only in Dorippoides facchino (Herbst, 1785) and Medorippe?crosnieri Chen, 1988, is in fact present in all species of dorippids.	FIG. 3. — Carapace diversity in the nine dorippid genera: A, Dorippe Weber, 1795 (D. quadridens); B, Dorippoides Serène & Romimohtarto, 1969 (D. facchino); C, Medorippe Manning & Holthuis, 1981 (?M. crosnieri); D, Heikeopsis Ng, Guinot & Davie, 2008 (H. aff. japonica); E, Neodorippe Serène & Romimohtarto, 1969 (N. callida); F, Nobilum Serène & Romimohtarto, 1969 (N. histrio); G, Paradorippe Serène & Romimohtarto, 1969 (P. cathayana); H, Philippidorippe Chen, 1986 (P. philippinensis); I, Phyllodorippe Manning & Holthuis, 1981 (P. armata). A-G, I: modified from Sin et al. (2009: fig. 3); H, from Chen (1986: pl. 1, fig. 3). The strip, which is shown here only in Dorippoides facchino (Herbst, 1785) and Medorippe?crosnieri Chen, 1988, is in fact present in all species of dorippids.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE74938CB5ACCC3EFB14FD40F813.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071347/files/figure.png	https://doi.org/10.5281/zenodo.8071347	FIG. 40. — Fossil crabs: A, the poorly known true dorippid, Dorippe judicis Gripp, 1964 (pl. 17, fig. 7a), Lower Miocene of northern Germany, generic status to be reappraised: carapace and pleon partially dorsally exposed (photocopy from Gripp 1964: pl. 17, fig. 7a); B-F, Goniochele angulata Bell, 1858; B-D, F, Eocene London Clay, Ypresian, Isle of Sheppey, Kent; E, Ypresian, Forest, Brussels, Belgium: B, carapace with preserved lateral spines, except long epibranchial spine, broken, concealed with matrix (Griffiths’ collection); C, carapace of paralectotype (lateral spines lost) with well-preserved protruding concave rim of posterior margin and narrow male pleon, with first three somites dorsally exposed (Wetherell’s collection, NHM PL OR 59085); D, E, two views of male thoracic sternum, with triangular first sternites and press-buttons on suture 5/6 (D, Jeff Saward collection);E, original from Collins & Smith 1993: pl. 2, fig. 3, as Silvacarcinus laurae Collins & Smith, 1993; F, ventral view of a female, with narrow elongated sternites 1-4 and wider last sternites (Griffiths’ collection, NHM 36652). Abbreviation: v, enormous vulvae. All, courtesy of Barry van Bakel.	FIG. 40. — Fossil crabs: A, the poorly known true dorippid, Dorippe judicis Gripp, 1964 (pl. 17, fig. 7a), Lower Miocene of northern Germany, generic status to be reappraised: carapace and pleon partially dorsally exposed (photocopy from Gripp 1964: pl. 17, fig. 7a); B-F, Goniochele angulata Bell, 1858; B-D, F, Eocene London Clay, Ypresian, Isle of Sheppey, Kent; E, Ypresian, Forest, Brussels, Belgium: B, carapace with preserved lateral spines, except long epibranchial spine, broken, concealed with matrix (Griffiths’ collection); C, carapace of paralectotype (lateral spines lost) with well-preserved protruding concave rim of posterior margin and narrow male pleon, with first three somites dorsally exposed (Wetherell’s collection, NHM PL OR 59085); D, E, two views of male thoracic sternum, with triangular first sternites and press-buttons on suture 5/6 (D, Jeff Saward collection);E, original from Collins & Smith 1993: pl. 2, fig. 3, as Silvacarcinus laurae Collins & Smith, 1993; F, ventral view of a female, with narrow elongated sternites 1-4 and wider last sternites (Griffiths’ collection, NHM 36652). Abbreviation: v, enormous vulvae. All, courtesy of Barry van Bakel.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE74938CB5ACCC3EFB14FD40F813.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071259/files/figure.png	https://doi.org/10.5281/zenodo.8071259	FIG. 3. — Carapace diversity in the nine dorippid genera: A, Dorippe Weber, 1795 (D. quadridens); B, Dorippoides Serène & Romimohtarto, 1969 (D. facchino); C, Medorippe Manning & Holthuis, 1981 (?M. crosnieri); D, Heikeopsis Ng, Guinot & Davie, 2008 (H. aff. japonica); E, Neodorippe Serène & Romimohtarto, 1969 (N. callida); F, Nobilum Serène & Romimohtarto, 1969 (N. histrio); G, Paradorippe Serène & Romimohtarto, 1969 (P. cathayana); H, Philippidorippe Chen, 1986 (P. philippinensis); I, Phyllodorippe Manning & Holthuis, 1981 (P. armata). A-G, I: modified from Sin et al. (2009: fig. 3); H, from Chen (1986: pl. 1, fig. 3). The strip, which is shown here only in Dorippoides facchino (Herbst, 1785) and Medorippe?crosnieri Chen, 1988, is in fact present in all species of dorippids.	FIG. 3. — Carapace diversity in the nine dorippid genera: A, Dorippe Weber, 1795 (D. quadridens); B, Dorippoides Serène & Romimohtarto, 1969 (D. facchino); C, Medorippe Manning & Holthuis, 1981 (?M. crosnieri); D, Heikeopsis Ng, Guinot & Davie, 2008 (H. aff. japonica); E, Neodorippe Serène & Romimohtarto, 1969 (N. callida); F, Nobilum Serène & Romimohtarto, 1969 (N. histrio); G, Paradorippe Serène & Romimohtarto, 1969 (P. cathayana); H, Philippidorippe Chen, 1986 (P. philippinensis); I, Phyllodorippe Manning & Holthuis, 1981 (P. armata). A-G, I: modified from Sin et al. (2009: fig. 3); H, from Chen (1986: pl. 1, fig. 3). The strip, which is shown here only in Dorippoides facchino (Herbst, 1785) and Medorippe?crosnieri Chen, 1988, is in fact present in all species of dorippids.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE74938CB5ACCEA6FAD3FAE6F814.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071279/files/figure.png	https://doi.org/10.5281/zenodo.8071279	FIG. 10. — Dorippinae H. Milne Edwards, 1837 n. stat.: habitus: A, B, Dorippe quadridens (Fabricius,1793):A, ♂ 36.3 × 38.1 mm, China Sea near Singapore, Hee Huat, ZRC 1984. 6308; B, ♀ 29.5 × 30 mm, NW Madagascar, Ambaro Bay, MNHN-IU-2018-5193 (= MNHN-B18279). C, D, Dorippe sinica Chen, 1980, China, Guangdong, Nanao Island, ZRC 1999.0470: C, ♂ 36.2 × 39.5 mm; D, ♀ 34.8 × 38.9 mm, specimen brushed.	FIG. 10. — Dorippinae H. Milne Edwards, 1837 n. stat.: habitus: A, B, Dorippe quadridens (Fabricius,1793):A, ♂ 36.3 × 38.1 mm, China Sea near Singapore, Hee Huat, ZRC 1984. 6308; B, ♀ 29.5 × 30 mm, NW Madagascar, Ambaro Bay, MNHN-IU-2018-5193 (= MNHN-B18279). C, D, Dorippe sinica Chen, 1980, China, Guangdong, Nanao Island, ZRC 1999.0470: C, ♂ 36.2 × 39.5 mm; D, ♀ 34.8 × 38.9 mm, specimen brushed.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE74938DB5ADCC27FF17FC9DFD28.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071279/files/figure.png	https://doi.org/10.5281/zenodo.8071279	FIG. 10. — Dorippinae H. Milne Edwards, 1837 n. stat.: habitus: A, B, Dorippe quadridens (Fabricius,1793):A, ♂ 36.3 × 38.1 mm, China Sea near Singapore, Hee Huat, ZRC 1984. 6308; B, ♀ 29.5 × 30 mm, NW Madagascar, Ambaro Bay, MNHN-IU-2018-5193 (= MNHN-B18279). C, D, Dorippe sinica Chen, 1980, China, Guangdong, Nanao Island, ZRC 1999.0470: C, ♂ 36.2 × 39.5 mm; D, ♀ 34.8 × 38.9 mm, specimen brushed.	FIG. 10. — Dorippinae H. Milne Edwards, 1837 n. stat.: habitus: A, B, Dorippe quadridens (Fabricius,1793):A, ♂ 36.3 × 38.1 mm, China Sea near Singapore, Hee Huat, ZRC 1984. 6308; B, ♀ 29.5 × 30 mm, NW Madagascar, Ambaro Bay, MNHN-IU-2018-5193 (= MNHN-B18279). C, D, Dorippe sinica Chen, 1980, China, Guangdong, Nanao Island, ZRC 1999.0470: C, ♂ 36.2 × 39.5 mm; D, ♀ 34.8 × 38.9 mm, specimen brushed.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE74938DB5ADCDF9FCD5FE49FA75.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071255/files/figure.png	https://doi.org/10.5281/zenodo.8071255	FIG. 1. — A typical dorippid crab, Heikeopsis japonica (von Siebold, 1824), the iconic ‘samurai crab’ known in Japan as ’heike-gani’, whose carapace looks like a human face: A, the species represented by De Haan (1839: pl. 31, fig. 1, as Dorippe japonica) in Fauna Japonica: top, a female; below, chelae of a male; B, lectotype selected by Yamaguchi & Baba (1993: 300, fig. 90-A.a-2, as Heikea japonica) from type series material labelled “Types, Japan, 1823, Leg. P. H. von Siebold”, RMNH CRUS D 822 (see Fransen et al. 1997: 83); C, ♂ 25.0 × 27.3 mm, ♀ 23.5 × 26.7 mm, Japan, Chaffanjon, 174-96, E.-L. Bouvier det. 1899 Dorippe japonica, dry condition, MNHN-IU-2000-4091 (= MNHN-B4091).	FIG. 1. — A typical dorippid crab, Heikeopsis japonica (von Siebold, 1824), the iconic ‘samurai crab’ known in Japan as ’heike-gani’, whose carapace looks like a human face: A, the species represented by De Haan (1839: pl. 31, fig. 1, as Dorippe japonica) in Fauna Japonica: top, a female; below, chelae of a male; B, lectotype selected by Yamaguchi & Baba (1993: 300, fig. 90-A.a-2, as Heikea japonica) from type series material labelled “Types, Japan, 1823, Leg. P. H. von Siebold”, RMNH CRUS D 822 (see Fransen et al. 1997: 83); C, ♂ 25.0 × 27.3 mm, ♀ 23.5 × 26.7 mm, Japan, Chaffanjon, 174-96, E.-L. Bouvier det. 1899 Dorippe japonica, dry condition, MNHN-IU-2000-4091 (= MNHN-B4091).	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE74938DB5ADCDF9FCD5FE49FA75.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071302/files/figure.png	https://doi.org/10.5281/zenodo.8071302	FIG. 19. — Heikeopsinae n. subfam. A-D, Heikeopsis japonica (von Siebold, 1824): A, ♂ 22.5 × 24.3 mm, Japan, Tokushima, SMF 57856: habitus; B, ♀ 23.5 × 26.7 mm, Japan, Chaffanjon, MNHN-IU-2000-4091 (= MNHN-B4091): habitus; C, ♂ 25.0 × 27.3 mm, same data as B: ventral view; D, ♂ 24.2 × 25.4 mm, same data as B and C: thoracic sternum and pleon. E, F,?Heikeopsis aff. japonica, China, seas off Qingdao, ZRC 2002.0491: habitus: E, ♂ 21.9 × 24 mm; F, ♀ 23.4 × 26.4 mm.	FIG. 19. — Heikeopsinae n. subfam. A-D, Heikeopsis japonica (von Siebold, 1824): A, ♂ 22.5 × 24.3 mm, Japan, Tokushima, SMF 57856: habitus; B, ♀ 23.5 × 26.7 mm, Japan, Chaffanjon, MNHN-IU-2000-4091 (= MNHN-B4091): habitus; C, ♂ 25.0 × 27.3 mm, same data as B: ventral view; D, ♂ 24.2 × 25.4 mm, same data as B and C: thoracic sternum and pleon. E, F,?Heikeopsis aff. japonica, China, seas off Qingdao, ZRC 2002.0491: habitus: E, ♂ 21.9 × 24 mm; F, ♀ 23.4 × 26.4 mm.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE74938DB5ADCDABFA12FB2CFAF5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071255/files/figure.png	https://doi.org/10.5281/zenodo.8071255	FIG. 1. — A typical dorippid crab, Heikeopsis japonica (von Siebold, 1824), the iconic ‘samurai crab’ known in Japan as ’heike-gani’, whose carapace looks like a human face: A, the species represented by De Haan (1839: pl. 31, fig. 1, as Dorippe japonica) in Fauna Japonica: top, a female; below, chelae of a male; B, lectotype selected by Yamaguchi & Baba (1993: 300, fig. 90-A.a-2, as Heikea japonica) from type series material labelled “Types, Japan, 1823, Leg. P. H. von Siebold”, RMNH CRUS D 822 (see Fransen et al. 1997: 83); C, ♂ 25.0 × 27.3 mm, ♀ 23.5 × 26.7 mm, Japan, Chaffanjon, 174-96, E.-L. Bouvier det. 1899 Dorippe japonica, dry condition, MNHN-IU-2000-4091 (= MNHN-B4091).	FIG. 1. — A typical dorippid crab, Heikeopsis japonica (von Siebold, 1824), the iconic ‘samurai crab’ known in Japan as ’heike-gani’, whose carapace looks like a human face: A, the species represented by De Haan (1839: pl. 31, fig. 1, as Dorippe japonica) in Fauna Japonica: top, a female; below, chelae of a male; B, lectotype selected by Yamaguchi & Baba (1993: 300, fig. 90-A.a-2, as Heikea japonica) from type series material labelled “Types, Japan, 1823, Leg. P. H. von Siebold”, RMNH CRUS D 822 (see Fransen et al. 1997: 83); C, ♂ 25.0 × 27.3 mm, ♀ 23.5 × 26.7 mm, Japan, Chaffanjon, 174-96, E.-L. Bouvier det. 1899 Dorippe japonica, dry condition, MNHN-IU-2000-4091 (= MNHN-B4091).	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE74938DB5ADCDABFA12FB2CFAF5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071302/files/figure.png	https://doi.org/10.5281/zenodo.8071302	FIG. 19. — Heikeopsinae n. subfam. A-D, Heikeopsis japonica (von Siebold, 1824): A, ♂ 22.5 × 24.3 mm, Japan, Tokushima, SMF 57856: habitus; B, ♀ 23.5 × 26.7 mm, Japan, Chaffanjon, MNHN-IU-2000-4091 (= MNHN-B4091): habitus; C, ♂ 25.0 × 27.3 mm, same data as B: ventral view; D, ♂ 24.2 × 25.4 mm, same data as B and C: thoracic sternum and pleon. E, F,?Heikeopsis aff. japonica, China, seas off Qingdao, ZRC 2002.0491: habitus: E, ♂ 21.9 × 24 mm; F, ♀ 23.4 × 26.4 mm.	FIG. 19. — Heikeopsinae n. subfam. A-D, Heikeopsis japonica (von Siebold, 1824): A, ♂ 22.5 × 24.3 mm, Japan, Tokushima, SMF 57856: habitus; B, ♀ 23.5 × 26.7 mm, Japan, Chaffanjon, MNHN-IU-2000-4091 (= MNHN-B4091): habitus; C, ♂ 25.0 × 27.3 mm, same data as B: ventral view; D, ♂ 24.2 × 25.4 mm, same data as B and C: thoracic sternum and pleon. E, F,?Heikeopsis aff. japonica, China, seas off Qingdao, ZRC 2002.0491: habitus: E, ♂ 21.9 × 24 mm; F, ♀ 23.4 × 26.4 mm.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE74938DB5AEC91BF8D1FE05FBD6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071259/files/figure.png	https://doi.org/10.5281/zenodo.8071259	FIG. 3. — Carapace diversity in the nine dorippid genera: A, Dorippe Weber, 1795 (D. quadridens); B, Dorippoides Serène & Romimohtarto, 1969 (D. facchino); C, Medorippe Manning & Holthuis, 1981 (?M. crosnieri); D, Heikeopsis Ng, Guinot & Davie, 2008 (H. aff. japonica); E, Neodorippe Serène & Romimohtarto, 1969 (N. callida); F, Nobilum Serène & Romimohtarto, 1969 (N. histrio); G, Paradorippe Serène & Romimohtarto, 1969 (P. cathayana); H, Philippidorippe Chen, 1986 (P. philippinensis); I, Phyllodorippe Manning & Holthuis, 1981 (P. armata). A-G, I: modified from Sin et al. (2009: fig. 3); H, from Chen (1986: pl. 1, fig. 3). The strip, which is shown here only in Dorippoides facchino (Herbst, 1785) and Medorippe?crosnieri Chen, 1988, is in fact present in all species of dorippids.	FIG. 3. — Carapace diversity in the nine dorippid genera: A, Dorippe Weber, 1795 (D. quadridens); B, Dorippoides Serène & Romimohtarto, 1969 (D. facchino); C, Medorippe Manning & Holthuis, 1981 (?M. crosnieri); D, Heikeopsis Ng, Guinot & Davie, 2008 (H. aff. japonica); E, Neodorippe Serène & Romimohtarto, 1969 (N. callida); F, Nobilum Serène & Romimohtarto, 1969 (N. histrio); G, Paradorippe Serène & Romimohtarto, 1969 (P. cathayana); H, Philippidorippe Chen, 1986 (P. philippinensis); I, Phyllodorippe Manning & Holthuis, 1981 (P. armata). A-G, I: modified from Sin et al. (2009: fig. 3); H, from Chen (1986: pl. 1, fig. 3). The strip, which is shown here only in Dorippoides facchino (Herbst, 1785) and Medorippe?crosnieri Chen, 1988, is in fact present in all species of dorippids.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE74938DB5AEC91BF8D1FE05FBD6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071309/files/figure.png	https://doi.org/10.5281/zenodo.8071309	FIG. 22. — Medorippinae n. subfam. Medorippe lanata (Linnaeus, 1767): A, ♂ 18.5 × 23.2 mm, Dahomey Coasts, ZRC 2009.0411: habitus; B, ovigerous ♀ 21.2 × 27.7 mm, Israel, ZRC 1999.0632: habitus; C-E, ♂ 18.5 × 23.2 mm, same data as A: C, anterior ventral view; D, thoracic sternum, press-button and G1; E, G1 and G2. F, ♂ 23.2 × 28.9 mm, no data, ZRC 2009.0412 (ex MNHN): thoracic sternum with pleon.	FIG. 22. — Medorippinae n. subfam. Medorippe lanata (Linnaeus, 1767): A, ♂ 18.5 × 23.2 mm, Dahomey Coasts, ZRC 2009.0411: habitus; B, ovigerous ♀ 21.2 × 27.7 mm, Israel, ZRC 1999.0632: habitus; C-E, ♂ 18.5 × 23.2 mm, same data as A: C, anterior ventral view; D, thoracic sternum, press-button and G1; E, G1 and G2. F, ♂ 23.2 × 28.9 mm, no data, ZRC 2009.0412 (ex MNHN): thoracic sternum with pleon.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE74938DB5AEC91BF8D1FE05FBD6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071325/files/figure.png	https://doi.org/10.5281/zenodo.8071325	FIG. 29. — Phyllodorippinae n. subfam. Phyllodorippe armata (Miers, 1881), Guinean Trawling Survey,ZRC 2009.0413 (ex MNHN-B24202). A, ♂ 12.3 × 16.2 mm, habitus; B-D, ovigerous ♀ 11.9 × 16.6 mm. B, habitus; C, thoracic sternum and pleon; D, thoracic sternum, vulvae and spine on sternite 8.	FIG. 29. — Phyllodorippinae n. subfam. Phyllodorippe armata (Miers, 1881), Guinean Trawling Survey,ZRC 2009.0413 (ex MNHN-B24202). A, ♂ 12.3 × 16.2 mm, habitus; B-D, ovigerous ♀ 11.9 × 16.6 mm. B, habitus; C, thoracic sternum and pleon; D, thoracic sternum, vulvae and spine on sternite 8.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE74938EB5AECC16FBB3FEEFF813.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071259/files/figure.png	https://doi.org/10.5281/zenodo.8071259	FIG. 3. — Carapace diversity in the nine dorippid genera: A, Dorippe Weber, 1795 (D. quadridens); B, Dorippoides Serène & Romimohtarto, 1969 (D. facchino); C, Medorippe Manning & Holthuis, 1981 (?M. crosnieri); D, Heikeopsis Ng, Guinot & Davie, 2008 (H. aff. japonica); E, Neodorippe Serène & Romimohtarto, 1969 (N. callida); F, Nobilum Serène & Romimohtarto, 1969 (N. histrio); G, Paradorippe Serène & Romimohtarto, 1969 (P. cathayana); H, Philippidorippe Chen, 1986 (P. philippinensis); I, Phyllodorippe Manning & Holthuis, 1981 (P. armata). A-G, I: modified from Sin et al. (2009: fig. 3); H, from Chen (1986: pl. 1, fig. 3). The strip, which is shown here only in Dorippoides facchino (Herbst, 1785) and Medorippe?crosnieri Chen, 1988, is in fact present in all species of dorippids.	FIG. 3. — Carapace diversity in the nine dorippid genera: A, Dorippe Weber, 1795 (D. quadridens); B, Dorippoides Serène & Romimohtarto, 1969 (D. facchino); C, Medorippe Manning & Holthuis, 1981 (?M. crosnieri); D, Heikeopsis Ng, Guinot & Davie, 2008 (H. aff. japonica); E, Neodorippe Serène & Romimohtarto, 1969 (N. callida); F, Nobilum Serène & Romimohtarto, 1969 (N. histrio); G, Paradorippe Serène & Romimohtarto, 1969 (P. cathayana); H, Philippidorippe Chen, 1986 (P. philippinensis); I, Phyllodorippe Manning & Holthuis, 1981 (P. armata). A-G, I: modified from Sin et al. (2009: fig. 3); H, from Chen (1986: pl. 1, fig. 3). The strip, which is shown here only in Dorippoides facchino (Herbst, 1785) and Medorippe?crosnieri Chen, 1988, is in fact present in all species of dorippids.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE74938EB5AECC16FBB3FEEFF813.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071309/files/figure.png	https://doi.org/10.5281/zenodo.8071309	FIG. 22. — Medorippinae n. subfam. Medorippe lanata (Linnaeus, 1767): A, ♂ 18.5 × 23.2 mm, Dahomey Coasts, ZRC 2009.0411: habitus; B, ovigerous ♀ 21.2 × 27.7 mm, Israel, ZRC 1999.0632: habitus; C-E, ♂ 18.5 × 23.2 mm, same data as A: C, anterior ventral view; D, thoracic sternum, press-button and G1; E, G1 and G2. F, ♂ 23.2 × 28.9 mm, no data, ZRC 2009.0412 (ex MNHN): thoracic sternum with pleon.	FIG. 22. — Medorippinae n. subfam. Medorippe lanata (Linnaeus, 1767): A, ♂ 18.5 × 23.2 mm, Dahomey Coasts, ZRC 2009.0411: habitus; B, ovigerous ♀ 21.2 × 27.7 mm, Israel, ZRC 1999.0632: habitus; C-E, ♂ 18.5 × 23.2 mm, same data as A: C, anterior ventral view; D, thoracic sternum, press-button and G1; E, G1 and G2. F, ♂ 23.2 × 28.9 mm, no data, ZRC 2009.0412 (ex MNHN): thoracic sternum with pleon.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE74938EB5AECEE4FF17FBB8F9B5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071309/files/figure.png	https://doi.org/10.5281/zenodo.8071309	FIG. 22. — Medorippinae n. subfam. Medorippe lanata (Linnaeus, 1767): A, ♂ 18.5 × 23.2 mm, Dahomey Coasts, ZRC 2009.0411: habitus; B, ovigerous ♀ 21.2 × 27.7 mm, Israel, ZRC 1999.0632: habitus; C-E, ♂ 18.5 × 23.2 mm, same data as A: C, anterior ventral view; D, thoracic sternum, press-button and G1; E, G1 and G2. F, ♂ 23.2 × 28.9 mm, no data, ZRC 2009.0412 (ex MNHN): thoracic sternum with pleon.	FIG. 22. — Medorippinae n. subfam. Medorippe lanata (Linnaeus, 1767): A, ♂ 18.5 × 23.2 mm, Dahomey Coasts, ZRC 2009.0411: habitus; B, ovigerous ♀ 21.2 × 27.7 mm, Israel, ZRC 1999.0632: habitus; C-E, ♂ 18.5 × 23.2 mm, same data as A: C, anterior ventral view; D, thoracic sternum, press-button and G1; E, G1 and G2. F, ♂ 23.2 × 28.9 mm, no data, ZRC 2009.0412 (ex MNHN): thoracic sternum with pleon.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE74938EB5AECEE4FF17FBB8F9B5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071325/files/figure.png	https://doi.org/10.5281/zenodo.8071325	FIG. 29. — Phyllodorippinae n. subfam. Phyllodorippe armata (Miers, 1881), Guinean Trawling Survey,ZRC 2009.0413 (ex MNHN-B24202). A, ♂ 12.3 × 16.2 mm, habitus; B-D, ovigerous ♀ 11.9 × 16.6 mm. B, habitus; C, thoracic sternum and pleon; D, thoracic sternum, vulvae and spine on sternite 8.	FIG. 29. — Phyllodorippinae n. subfam. Phyllodorippe armata (Miers, 1881), Guinean Trawling Survey,ZRC 2009.0413 (ex MNHN-B24202). A, ♂ 12.3 × 16.2 mm, habitus; B-D, ovigerous ♀ 11.9 × 16.6 mm. B, habitus; C, thoracic sternum and pleon; D, thoracic sternum, vulvae and spine on sternite 8.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE74938EB5AECEE4FF17FBB8F9B5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071311/files/figure.png	https://doi.org/10.5281/zenodo.8071311	FIG. 23. — Medorippinae n. subfam.: A, B,?Medorippe crosnieri Chen, 1988. Mozambique Channel, MAINBAZA Expedition: A, ♂ 21.1 × 25.9 mm, CP 3132, MNHN-IU-2009-1998, with dimorphic chelipeds; B, young ♂ 12.0 × 15.0 mm, CP 3130, MNHN-IU-2009-1999, with symmetrical chelipeds. Photos courtesy of T.-Y.Chan.C, Medorippe lanata (Linnaeus,1767),♂ 23.2 × 28.9 mm,no data,ZRC 2009.0412 (ex MNHN):P2 merus with spinulated dorsal margin;D,?Medorippe crosnieri Chen, 1988, ♂ 21.1 × 25.9 mm, same data as in A (MNHN-IU-2009-1998): P2 merus with smooth dorsal margin.Photographs made by S. Soubzmaigne.	FIG. 23. — Medorippinae n. subfam.: A, B,?Medorippe crosnieri Chen, 1988. Mozambique Channel, MAINBAZA Expedition: A, ♂ 21.1 × 25.9 mm, CP 3132, MNHN-IU-2009-1998, with dimorphic chelipeds; B, young ♂ 12.0 × 15.0 mm, CP 3130, MNHN-IU-2009-1999, with symmetrical chelipeds. Photos courtesy of T.-Y.Chan.C, Medorippe lanata (Linnaeus,1767),♂ 23.2 × 28.9 mm,no data,ZRC 2009.0412 (ex MNHN):P2 merus with spinulated dorsal margin;D,?Medorippe crosnieri Chen, 1988, ♂ 21.1 × 25.9 mm, same data as in A (MNHN-IU-2009-1998): P2 merus with smooth dorsal margin.Photographs made by S. Soubzmaigne.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE74938FB5AFCC1CFF17FDAAFD17.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071259/files/figure.png	https://doi.org/10.5281/zenodo.8071259	FIG. 3. — Carapace diversity in the nine dorippid genera: A, Dorippe Weber, 1795 (D. quadridens); B, Dorippoides Serène & Romimohtarto, 1969 (D. facchino); C, Medorippe Manning & Holthuis, 1981 (?M. crosnieri); D, Heikeopsis Ng, Guinot & Davie, 2008 (H. aff. japonica); E, Neodorippe Serène & Romimohtarto, 1969 (N. callida); F, Nobilum Serène & Romimohtarto, 1969 (N. histrio); G, Paradorippe Serène & Romimohtarto, 1969 (P. cathayana); H, Philippidorippe Chen, 1986 (P. philippinensis); I, Phyllodorippe Manning & Holthuis, 1981 (P. armata). A-G, I: modified from Sin et al. (2009: fig. 3); H, from Chen (1986: pl. 1, fig. 3). The strip, which is shown here only in Dorippoides facchino (Herbst, 1785) and Medorippe?crosnieri Chen, 1988, is in fact present in all species of dorippids.	FIG. 3. — Carapace diversity in the nine dorippid genera: A, Dorippe Weber, 1795 (D. quadridens); B, Dorippoides Serène & Romimohtarto, 1969 (D. facchino); C, Medorippe Manning & Holthuis, 1981 (?M. crosnieri); D, Heikeopsis Ng, Guinot & Davie, 2008 (H. aff. japonica); E, Neodorippe Serène & Romimohtarto, 1969 (N. callida); F, Nobilum Serène & Romimohtarto, 1969 (N. histrio); G, Paradorippe Serène & Romimohtarto, 1969 (P. cathayana); H, Philippidorippe Chen, 1986 (P. philippinensis); I, Phyllodorippe Manning & Holthuis, 1981 (P. armata). A-G, I: modified from Sin et al. (2009: fig. 3); H, from Chen (1986: pl. 1, fig. 3). The strip, which is shown here only in Dorippoides facchino (Herbst, 1785) and Medorippe?crosnieri Chen, 1988, is in fact present in all species of dorippids.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE74938FB5AFCC1CFF17FDAAFD17.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071307/files/figure.png	https://doi.org/10.5281/zenodo.8071307	FIG. 21. — Heikeopsinae n. subfam.Nobilum histrio (Nobili,1903), Malaysia, Johore,Pontian, ZRC 2002.0491:A-D, ♂ 21.0 × 22.4 mm:habitus; B, anterior ventral view; C, thoracic sternum with pleon; D, thoracic sternum without pleon, penis and G1. E, F, ovigerous ♀ 22.1 × 24.7 mm: thoracic sternum, pleon and vulvae.	FIG. 21. — Heikeopsinae n. subfam.Nobilum histrio (Nobili,1903), Malaysia, Johore,Pontian, ZRC 2002.0491:A-D, ♂ 21.0 × 22.4 mm:habitus; B, anterior ventral view; C, thoracic sternum with pleon; D, thoracic sternum without pleon, penis and G1. E, F, ovigerous ♀ 22.1 × 24.7 mm: thoracic sternum, pleon and vulvae.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
03CDBE74938FB5AFCC1CFF17FDAAFD17.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/8071305/files/figure.png	https://doi.org/10.5281/zenodo.8071305	FIG. 20. — Heikeopsinae n. subfam. Neodorippe callida (Fabricius, 1798): A, young ♂ 10.1 × 10.3 mm, Singapore, Changi Point Beach, ZRC 2018.0445: habitus of young male; B, C, Andaman Sea, Phuket, Thailand, MNHN-IU-2016-10754 (ex ZRC 1998.1116): B, ♀ 14.0 × 15.0 mm, habitus; C, ♂ 13.5 × 14 mm: ventral view. D, E, ♂ 10.1 × 10.3 mm, same data as A, ZRC 2018.0445:D, thoracic sternum with pleon; E, without pleon, G1, press-buttons. F, ♀ 11.2 × 11.5 mm, same data as A, D, E, ZRC 2018.0445: anterior view. G, ♀ 11.2 × 11.5 mm, sama data as B, C, MNHN-IU-2016-10754: vulvae, press-buttons. H, ♀ 11.2 × 11.5 mm, same data as A, D, E, F, ZRC 2018.0445: vulvae.	FIG. 20. — Heikeopsinae n. subfam. Neodorippe callida (Fabricius, 1798): A, young ♂ 10.1 × 10.3 mm, Singapore, Changi Point Beach, ZRC 2018.0445: habitus of young male; B, C, Andaman Sea, Phuket, Thailand, MNHN-IU-2016-10754 (ex ZRC 1998.1116): B, ♀ 14.0 × 15.0 mm, habitus; C, ♂ 13.5 × 14 mm: ventral view. D, E, ♂ 10.1 × 10.3 mm, same data as A, ZRC 2018.0445:D, thoracic sternum with pleon; E, without pleon, G1, press-buttons. F, ♀ 11.2 × 11.5 mm, same data as A, D, E, ZRC 2018.0445: anterior view. G, ♀ 11.2 × 11.5 mm, sama data as B, C, MNHN-IU-2016-10754: vulvae, press-buttons. H, ♀ 11.2 × 11.5 mm, same data as A, D, E, F, ZRC 2018.0445: vulvae.	2023-06-05	Guinot, Danièle		Zenodo	biologists	Guinot, Danièle			
