taxonID	type	description	language	source
03CC1E69A337FFD7B924F93DB77EFED7.taxon	discussion	Remarks In our previous publications (Páll-Gergely et al. 2016, 2017, 2020, 2021; Páll-Gergely & Asami 2017; Páll-Gergely & Auffenberg 2019; Páll-Gergely & Hunyadi 2022; Páll-Gergely 2023), we treated this group as a family of its own right. However, the recent molecular phylogeny (Páll-Gergely et al. 2024) suggested treating Alycaeinae as a family would result in numerous paraphyletic groups, and therefore, Cyclophoridae should be maintained at the family rank, and Alycaeinae is to be treated as one of its subfamilies.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A335FFD7B92BFC7CB4C2FBFB.taxon	discussion	Remarks Chamalycaeus mizoramensis sp. nov. is classified in its own species group, as this species is not similar to any other species of Alycaeinae in the region.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A335FFD7B915FE75B74CFCCF.taxon	discussion	Remarks Chamalycaeus is characterised by a smooth (or pitted) protoconch and a spirally striated R 1. Most species classified in Chamalycaeus here are transferred from the genus Dicharax (see Páll-Gergely et al. 2020). Although no spiral striation was found on the protoconch of Chamalycaeus oglei (Godwin-Austen, 1914) (see Páll-Gergely et al. 2020), it is moved to the genus Metalycaeus Pilsbry, 1900 due to its similarity with M. brahma (Godwin-Austen, 1886) and other Himalayan species of Metalycaeus. Furthermore, Chamalycaeus perplexus (Godwin-Austen, 1914) is moved to Metalycaeus due to its similarity with M. polygonoma.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A335FFD9BA94FB88B43FFDF7.taxon	description	Fig. 4	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A335FFD9BA94FB88B43FFDF7.taxon	diagnosis	Diagnosis A relatively large species of Chamalycaeus with a ca half whorl-long R 2 + R 3, and R 3 with a low central swelling and a fringed aperture.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A335FFD9BA94FB88B43FFDF7.taxon	etymology	Etymology This species is named after the state of Mizoram, where the Blue Mountain is situated.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A335FFD9BA94FB88B43FFDF7.taxon	materials_examined	Type material examined Holotype INDIA – Mizoram • empty shell (D: 4.8 mm, H: 3 mm); Blue Mountain, Lawngtlai district; 22.727 ° N, 93.135 ° E; 1400 m a. s. l.; 27 Jan. 2019; N. A. Aravind leg.; AR 15; NZSI LM 1715.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A335FFD9BA94FB88B43FFDF7.taxon	description	Description Shell off-white / ivory in colour, not translucent; shell outline slightly oval in dorsal view; spire slightly elevated, low conical; body whorl rounded; protoconch low, finely granular, spiral striae lacking, consisting of nearly 2 whorls; R 1 of 1.75 whorls, with rather irregular, dense, low ribs (last half whorl of R 1 with ca 55 – 60 ribs) and weaker, denser spiral striation; boundary between R 1 and R 2 not conspicuous because R 2 ribs are only slightly higher and sharper than R 1 ribs, but R 2 area more inflated than R 1; sutural tube conspicuously shorter than R 2, probably due to teratological condition; R 2 with ca 50 – 52 straight, sharp but low ribs; R 2 + R 3 ca half whorl, R 3 shorter than R 2, boundary between R 2 and R 3 clearly visible due to a slight constriction; R 3 with long and low central swelling; R 3 ribs stand approximately as densely as R 1 ribs, with the exception of the beginning of R 3 with more widely-spaced ribbing; aperture strongly oblique to shell axis, rounded with 5 small bays on the palatal side, resulting in a fringed peristome; boundary between inner and outer peristomes is not conspicuous but clearly visible, inner peristome thickened, slightly protruding and expanded; outer peristome thin, sharp, slightly expanded, especially towards umbilicus; umbilicus relatively narrow, ca one fourth of shell width, rounded. MEASUREMENTS. D: 4.8 mm, H: 3 mm.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A335FFD9BA94FB88B43FFDF7.taxon	distribution	Distribution Known only from the Blue Mountain region.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A335FFD9BA94FB88B43FFDF7.taxon	diagnosis	Differential diagnosis No other species of Chamalycaeus resembles Chamalycaeus mizoramensis sp. nov. The relatively large size, the fringed aperture, the dense, fine ribbing on R 1 and R 3, and the long R 2 + R 3 (ca half whorl) are characteristic of this species.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A33BFFD9B916FD94B565FCA0.taxon	discussion	Remarks Three tiny species from Myanmar belong here, which are characterized by widely-spaced R 1 ribs, a short R 2, a low and blunt R 3 swelling, and a circular aperture.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A33BFFD9B91CFCA1B7B0FAF1.taxon	description	Fig. 5 A – D	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A33BFFD9B91CFCA1B7B0FAF1.taxon	diagnosis	Diagnosis See under C. humilis (W. T. Blanford, 1862) and C. irmatallus Páll-Gergely, 2021.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A33BFFD9B91CFCA1B7B0FAF1.taxon	materials_examined	Type locality “ ad Thyet-Mio ”.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A33BFFDBB937FA92B2AEFDD3.taxon	description	Fig. 5 E – I	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A33BFFDBB937FA92B2AEFDD3.taxon	materials_examined	Type material examined MYANMAR • holotype (single shell mentioned in the original description: Fig. 5 E – I); River Bank, Myanoung, Pegu; NHMUK 1906.4. 4.69. Type locality “ ad Akouktoung, ad ripas fluminis Irawaddi, in provincia Burmana Pegu ”.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A33BFFDBB937FA92B2AEFDD3.taxon	diagnosis	Differential diagnosis This species has a comparatively smaller aperture than C. armillatus. Chamalycaeus irmatallus has denser and finer R 1 ribs, more expanded outer peristome and a less inflated R 3 swelling.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A33BFFDBB937FA92B2AEFDD3.taxon	discussion	Remarks Only a single, corroded shell is known, therefore the important traits (presence or absence of spiral striation on the teleoconch) could not be examined without doubt. We classified this species in the genus Dicharax in our previous catalogue, but during the revision of Himalayan Dicharax, we had to reconsider our decision. Since the shell shape of A. humilis is most similar to the geographically closely occurring Chamalycaeus armillatus and C. irmatallus, we move this species to Chamalycaeus.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A339FFDBB916FA27B347F8A6.taxon	discussion	Remarks The species belonging here are characterized by a low conical, colourless shell, a strongly ribbed R 1 (rib density relatively low), and an upper incision in the aperture. The aperture shape (fringed to circular) and the shape of R 3 swelling are highly variable. Some species of this species group have been classified in the genus Dicharax (see Páll-Gergely et al. 2020), because spiral striation could not be observed on them, partly because the examined shells were corroded, and partly because the spiral striation on the shells are naturally weak and difficult to observe.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A339FFDBB930FD77B78DFA23.taxon	description	Fig. 5 J – N	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A339FFDBB930FD77B78DFA23.taxon	materials_examined	Type material examined THAILAND • holotype; Surat Thani Province, Limestone Mt, 6 km S of Na San; 8 ° 39.583 ′ N, 99 ° 23.883 ′ E; 100 m a. s. l.; 3 Jun. 1987; F. G. Thompson leg.; UF 279508 • 11 paratypes; same data as for preceding; UF 551216. Additional material examined MYANMAR • 6 shells (see Páll-Gergely et al. 2021: fig. 10); Thayet-myo, Pegu; Blanford coll.; NHMUK 1906.4. 4.71. Type locality “ Thailand, Surat Thani Province, limestone mtn., 6 km S of Na San, 8 ° 39.583 ′ N, 99 ° 23.883 ′ E; 100 m a. s. l. ”.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A339FFDBB930FD77B78DFA23.taxon	diagnosis	Differential diagnosis Chamalycaeus armillatus has a more rounded body whorl, longer R 2 and R 3, and a thicker peristome with greater distance between the edges of the inner and outer peristomes (Páll-Gergely et al. 2021). See also under C. humilis.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A339FFDDBAD2F8DBB4A1FA62.taxon	description	Fig. 6	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A339FFDDBAD2F8DBB4A1FA62.taxon	diagnosis	Diagnosis Differs from the congeners by the long R 3 swelling having straight anterior part.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A339FFDDBAD2F8DBB4A1FA62.taxon	materials_examined	Type material examined INDIA • 2 syntypes of A. asaluensis (Fig. 6 A – E); Dihung, N. Cachar; Godwin-Austen coll.; NHMUK 1903.7. 1.2636 • holotype of A. barowliensis (single shell mentioned in the original description: Fig. 6 F – J); Barowli R. Durrang, Assam; Godwin-Austen coll.; NHMUK 1903.7. 1.2723. Type localities “ Barowli River, Akha Hills, Durrang, Assam ” (A. barowliensis); “ Neuglo ”, “ Phulong ” and “ Dihung River, N. Cachar, north of Asalu ” (A. asaluensis).	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A339FFDDBAD2F8DBB4A1FA62.taxon	diagnosis	Differential diagnosis Dicharax aspidentatus Gittenberger, Choki Gyeltshen & Sherub Sherub, 2024 has a higher spire and larger distance between the inner and outer peristomes. Chamalycaeus crispatus (Godwin-Austen, 1871) has a constriction anterior to the R 3 swelling, is glossier and less densely ribbed. Dicharax rugosus (Godwin-Austen, 1914) has a higher spire, a shorter R 3 swelling and more strongly fringed peristome.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A339FFDDBAD2F8DBB4A1FA62.taxon	discussion	Remarks Only the corroded holotype of A. barowliensis is known, which is very similar to C. asaluensis, but has a slightly more globose shell (and narrower umbilicus) and more widely-spaced ribs on R 1. Nevertheless, given their geographic proximity and that all other traits are identical, these differences are best explained by intraspecific variability. This species was included in the genus Dicharax due to the absence of spiral striation (Páll-Gergely et al. 2020). However, the shell shape and other characters agree with this species group, and therefore, it is reclassified in the genus Chamalycaeus.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A33FFFDFBACAF9E6B518F9F3.taxon	description	Fig. 7	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A33FFFDFBACAF9E6B518F9F3.taxon	diagnosis	Diagnosis The narrow and high R 3 swelling distinguishes this species from other congeners of the same species group.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A33FFFDFBACAF9E6B518F9F3.taxon	materials_examined	Type material examined INDIA • 5 syntypes of A. crispatus (Fig. 7 A – E); Shibak, Habiang Garo Hills; NHMUK 1903.7. 1.2635 • 11 syntypes of A. crispatus; same container as previous (probably same locality); NHMUK 1903.7. 1.2759 • holotype of A. crispatus var. minimus (single shell mentioned in the original description: Fig. 7 K – O); Habiang Garo Hills, W. Khasi; Godwin-Austen leg.; NHMUK 1906.4. 4.176 • 2 syntypes of A. crispatus var. rywukensis (Fig. 7 F – J); Rywuk, Garo Hills, South base of; Godwin-Austen coll.; NHMUK 1903.7. 1.2637. Type localities “ Khasia, Jaintia and N. Cachar Hills ” (A. crispatus); “ Habiang Garo Hills, West Khasi ” (A. crispatus var. minimus); “ Rywuk Valley of the Garo Hills ” (A. crispatus var. rywukensis).	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A33FFFDFBACAF9E6B518F9F3.taxon	diagnosis	Differential diagnosis Dicharax rugosus has a higher spire, and a more strongly and densely ribbed R 1. See also under C. asaluensis.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A33FFFDFBACAF9E6B518F9F3.taxon	discussion	Remarks Godwin-Austen (1914: 372) stated in his previous paper (Godwin-Austen 1871: 93) that the Alycaeus crispatus variety from north Cachar was renamed A. asaluensis. The originally figured sample (Godwin-Austen 1871: pl. 4 fig. 1) is from Shibak, Gabir Valley (Godwin-Austen 1914: 372). Alycaeus crispatus and A. crispatus var. minimus do not differ in any important shell characters, and therefore, the latter is a junior synonym of the former. Alycaeus crispatus var. rywukensis is smaller than the previous two and has a ribbed instead of glossy R 3. Nevertheless, we interpret this as intraspecific variability. Gittenberger et al. (2024) reported this species from Bhutan. We agree with their identification based on the photos they provided. This species was included in the genus Dicharax due to the absence of spiral striation (Páll-Gergely et al. 2020). However, the shell shape and other characters agree with this species group, and therefore, it is reclassified in the genus Chamalycaeus. Furthermore, re-examination of the syntypes of A. crispatus var. makarsae revealed spiral striation on the teleoconch.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A33DFFC1BAD7F997B556FBFC.taxon	description	Fig. 8	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A33DFFC1BAD7F997B556FBFC.taxon	diagnosis	Diagnosis The low R 3 without conspicuous swelling distinguishes this species from all congeners of this species group.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A33DFFC1BAD7F997B556FBFC.taxon	materials_examined	Type material examined INDIA • 1 syntype of A. logtakensis (Fig. 8 A – E); on a low hill, Logtak Lake, Munipur; Godwin-Austen coll.; “ figured ”; NHMUK 1903.7. 1.2639 • 6 syntypes of A. tanghali; Munipur; figured by Godwin-Austen; NHMUK 1903.7. 1.2671 (Fig. 8 F – J). Type localities “ Logtak Lake, Munipur (No. 2639 B. M. Coll.), on a low hill near the northern shore ” (A. logtakensis); “ Munipur. Exact locality not recorded; somewhere on the northern side of the valley ” (A. tanghali).	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A33DFFC1BAD7F997B556FBFC.taxon	diagnosis	Differential diagnosis The most similar species is C. sculpturus, but that species has a less fringed aperture and a more rounded body whorl.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A33DFFC1BAD7F997B556FBFC.taxon	discussion	Remarks There are no conchological differences between A. logtakensis and A. tanghali, and they are both from central Manipur State. Their descriptions were published in the same work but A. tanghali appeared a few pages later, therefore A. logtakensis is chosen to be the valid name. Godwin-Austen (1914) compared A. tanghali only with A. sculptilis. Alycaeus logtakensis was included in the genus Dicharax because no spiral striation was observed on the corroded syntype (Páll-Gergely et al. 2020). However, re-examination of the syntype of A. logtakensis revealed spiral striation on the teleoconch. Chamalycaeus tanghali was already classified in Chamalycaeus due to spiral striation on the teleoconch.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A323FFC5BAF4FB8DB591FCA9.taxon	description	Figs 9 – 10, 12 A	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A323FFC5BAF4FB8DB591FCA9.taxon	diagnosis	Diagnosis A small species of Chamalycaeus with a ca quarter whorl-long R 2 + R 3, an R 3 with a low central swelling (situated slightly closer to the peristome), a strongly expanded outer peristome, and a rounded inner peristome.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A323FFC5BAF4FB8DB591FCA9.taxon	etymology	Etymology The specific epithet refers to the reflected outer peristome, which is characteristic of this species.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A323FFC5BAF4FB8DB591FCA9.taxon	materials_examined	Type material examined Holotype INDIA – Mizoram • 1 empty shell (D: 1.9 mm, H: 1.4 mm, Figs 9, 12 A); Blue Mountain, Lawngtlai district; 22.727 ° N, 93.135 ° E; 1400 m a. s. l.; 27 Jan. 2019; N. A. Aravind leg.; Ar 55; NZSI LM 1723. Paratype INDIA – Mizoram • 1 empty shell; same data as for holotype; NZSI LM 1724. Additional material examined INDIA – Mizoram • 2 empty shells; same data as for holotype; Ar 56; NZSI LM 1725 • 1 empty shell (Fig. 10); same data as for holotype; AR 44; NZSI LM 1913. – Tripura • 1 empty shell (Fig. 11); South Tripura, Purba Patichhari, Towards Garji from Santirbazar; 23 ° 24.227 ′ N, 91 ° 30.294 ′ E; 31 Jan. 2021; Nipu Kumar Das leg.; NZSI LM 1906.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A323FFC5BAF4FB8DB591FCA9.taxon	description	Description Shell small, white, fresh shell possibly translucent; shell outline rather rounded in dorsal view; spire slightly elevated, low conical; body whorl rounded; protoconch low, finely granular, spiral striae lacking, consisting of ca 1.5 whorls; R 1 of slightly under 1.5 whorls, with strong, sharp, but relatively low radial ribs that become more widely-spaced towards R 2; last half whorl of R 1 with ca 18 ribs, space between last two R 1 ribs twice as large as distance between two ribs half whorl behind the end of R 2; spiral striation absent; boundary between R 1 and R 2 conspicuous due to the sudden increase in rib density; R 2 with ca 16 straight, sharp but low ribs; R 2 + R 3 ca quarter whorl or slightly more; R 3 and R 2 of comparable length, boundary between R 2 and R 3 clearly visible due to a slight constriction; R 3 with moderately long swelling that is situated closer to R 3; R 3 nearly smooth, with ca 3 weak ribs between swelling and outer peristome; aperture oblique to shell axis, rounded; boundary between inner and outer peristomes conspicuous; inner peristome strongly thickened, protruding and expanded; outer peristome thin, sharp, strongly reflected; umbilicus narrow, ca one fifth of shell width, rounded. MEASUREMENTS. D: 1.9 – 2.5 mm, H: 1.4 – 1.9 mm.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A323FFC5BAF4FB8DB591FCA9.taxon	distribution	Distribution Known only from the Blue Mountain region.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A323FFC5BAF4FB8DB591FCA9.taxon	diagnosis	Differential diagnosis We have not found any similar species in the historical collection other than C. sculpturus, with which this species is sympatric. Chamalycaeus sculpturus is similar in shell shape and sculpture, and the ratios of regions. However, its peristome is dominated by the inner peristome, which is fringed and thickened, while the outer peristome is not expanded. In contrast, C. reflectus sp. nov. has a smooth (not fringed) inner, and a strongly expanded outer peristome. Furthermore, the upper apertural incision on C. crispatus is stronger and the R 1 ribs (most clearly visible in the area just preceding R 2) are denser. The peristome of C. sculptilis (Fig. 13 A – D) is similar to that of C. reflectus, but that species is larger and has a strongly keeled body whorl.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A323FFC5BAF4FB8DB591FCA9.taxon	discussion	Remarks The single shell Ar 44 (Fig. 10) is larger than the two type specimens and has a longer R 2, while the sculpture and the morphology of the inner and outer peristomes of the type specimens and the Ar 44 specimen is identical. We need more extensive material to understand whether the observed differences represent an intraspecific diversity or differences between species. The two Ar 56 specimens are identical with the Ar 44 specimen but are strongly corroded. The single shell from Tripura (Fig. 11) is similar to the holotype, but has a less expanded outer peristome, and denser R 1 and R 2 ribs. Nevertheless, for the time being, we consider it conspecific with C. reflectus sp. nov.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A326FFC4B914FEECB791FA91.taxon	description	Fig. 13 A – D	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A326FFC4B914FEECB791FA91.taxon	diagnosis	Diagnosis Differs from all other species of this genus by the keeled body whorl and the finely beaded peristome. See also under Dicharax bhucarinatus Gittenberger & Sherub Sherub, 2024.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A326FFC4B914FEECB791FA91.taxon	materials_examined	Type material examined MYANMAR • neotype (designated by Preece et al. 2022: fig. 13 a – d); Thyet Myo; Bens. coll.; UMZC I. 102845. A • 2 shells; same data as for preceding; UMZC I. 102845. B, I. 102845. C Additional material examined MYANMAR • 1 shell; Pegu, Thayet-myo; “ typical ”, “ aperture figured ”; NHMUK 1906.4. 4.70. Type locality “ raro ad Thyet-Mio prope fluvium Irawadi, non procul a finibus provinciae Burmanicae Britannicae ” [rarely at Thyet-Mio near the river Irawadi, not far from the borders of British Burma] (translation by Preece et al. 2022).	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A326FFCBBAD3FAB2B4C4FD1F.taxon	description	Figs 14 – 16	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A326FFCBBAD3FAB2B4C4FD1F.taxon	materials_examined	Type material examined INDIA • 1 syntype of A. sculpturus (Fig. 14 A – E); Sikhami, NE Munipur; NHMUK 1903.7. 1.2666 • 4 syntypes of A. sculpturus (Fig. 14 F – J); Mungching, Munipur; NHMUK 1903.7. 1.2667 • 8 syntypes of A. crispatus var. makarsae (Fig. 14 K – O); Makarsa, Khasi; Godwin-Austen leg.; NHMUK 1903.7. 1.2638. Additional material examined INDIA – Mizoram • 1 empty shell (Fig. 15); Blue Mountain, Lawngtlai district; 22.727 ° N, 93.135 ° E; 1400 m a. s. l.; 27 Jan. 2019; N. A. Aravind leg.; Ar 68; NZSI LM 1720 • 1 empty shell; same data as for preceding; Ar 61; NZSI LM 1721 • 2 complete shells (Fig. 16) + 2 broken shells; same data as for preceding; Ar 74; NZSI LM 1722. Type localities “ on the hill ranges from near Tellizo Peak to the eastward, and on Mungching Hill in Munipur ” (A. sculpturus); “ Makarsa, N. Khasi Hills (or more correct, Maokarsa; the common Khasi prefix “ Mao ” meaning a stone) ” (A. crispatus var. makarsae).	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A326FFCBBAD3FAB2B4C4FD1F.taxon	discussion	Remarks The type specimens of A. sculpturus and A. crispatus var. makarsae show no notable differences; therefore, the latter is a junior synonym of the former. The specimens collected in the Blue Mountain (Figs 15 – 16) are slightly smaller than typical C. sculpturus and A. crispatus var. makarsae shells and have multiple tiny knobs along the peristome but otherwise are very similar to those species. These knobs are probably homologous with the tips of triangle-shaped structures (termed chevrons in Gittenberger et al. 2024) in the aperture of typical C. sculpturus shells and many other alycaeine species (i. e., two tiny knobs of one shell are homologous with the two tips of a single triangle in the other). A similar case was observed in a Chinese species; typical Dicharax fimbriatus (Bavay & Dautzenberg, 1912) shells had triangular thickenings along the peristome, while its synonym, Chamalycaeus plicilabris multidentatus Yen, 1939, had multiple tiny knobs along the peristome (Páll-Gergely et al. 2017). This species was included in the genus Dicharax due to the absence of spiral striation (Páll-Gergely et al. 2020). However, re-examination of the syntypes of A. sculpturus and Alycaeus crispatus var. makarsae revealed some spiral striation on the teleoconch.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A329FFCBB93EFD2CB4F3F800.taxon	description	Fig. 17	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A329FFCBB93EFD2CB4F3F800.taxon	diagnosis	Diagnosis This species can be distinguished from other species of this species group by the long and inflated R 3 swelling. Chamalycaeus sylheticus is the most similar species; for comparisons, see under that species.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A329FFCBB93EFD2CB4F3F800.taxon	materials_examined	Type material examined INDIA • 1 syntype of A. kezamaensis (Fig. 17 F – J); Kezama, Naga Hills; Godwin-Austen coll.; NHMUK 1903.7. 1.2556 • 4 syntypes of A. theobaldi var. solidus (Fig. 17 A – E); Garo Hills; NHMUK 1903.7. 1.2560. Type localities “ Kezama, Anghami-Naga Hills ” (A. kezamaensis); “ Garo Hills ” (A. theobaldi solidus).	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A329FFCBB93EFD2CB4F3F800.taxon	discussion	Remarks Alycaeus kezamaensis is very slightly smaller than the syntypes of A. theobaldi solidus, but all other traits (shell and aperture shape, morphology and ratios of shell regions, sculpture) are identical. Therefore, they represent the same species. Alycaeus kezamaensis was introduced later in the same publication, therefore the valid name is Chamalycaeus solidus. Chamalycaeus solidus is a species of its own right, because it differs in important traits from D. theobaldi, namely the latter is more depressed, its R 2 is longer, and the R 3 swelling is lower and shorter (and they belong to different genera). This species was included in the genus Dicharax due to the absence of spiral striation on the corroded syntypes of Alycaeus theobaldi var. solidus and Alycaeus kezamaensis (Páll-Gergely et al. 2020). Nevertheless, the other conchological characters agree with species of this species group, and therefore, this species is reclassified in Chamalycaeus.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A32FFFCDBACFFEECB7FBFBC1.taxon	description	Fig. 13 F – I	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A32FFFCDBACFFEECB7FBFBC1.taxon	materials_examined	Type material examined BANGLADESH • holotype (single shell mentioned in the original description: Fig. 13 F – I); S Sylhet Hills; W. Channel leg.; NHMUK 1903.7. 1.55. Type locality “ South Sylhet Hills ”.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A32FFFCDBACFFEECB7FBFBC1.taxon	diagnosis	Differential diagnosis The most similar species is C. solidus in terms of shell shape and sculpture, but that species is slightly larger, has a more elevated spire, and a more bulging R 3 swelling.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A32FFFCDBACFFEECB7FBFBC1.taxon	discussion	Remarks This species was included in the genus Dicharax in spite of the presence of spiral striation, because the R 2 ribs were typical of that genus (Páll-Gergely et al. 2020). However, the critical revision of Himalayan species of Chamalycaeus and Dicharax revealed that C. sylheticus belongs to this species group of Chamalycaeus due to other conchological traits (colourless shells, strong and widely-spaced R 1 ribs, upper apertural notch).	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A32FFFCDB998F8ACB30AF86C.taxon	discussion	Remarks The species belonging here usually have a conical or elongated conical shell and a thin peristome.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A32FFFCDB908F9B1B597F89E.taxon	type_taxon	Type species Cyclostoma constrictum Benson, 1851, by original designation.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A32EFFCEB969F95CB7AAFD25.taxon	description	Fig. 19 A – H	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A32EFFCEB969F95CB7AAFD25.taxon	diagnosis	Diagnosis This species can be distinguished from others due to the practically smooth, conical shell, and the thin peristome.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A32EFFCEB969F95CB7AAFD25.taxon	materials_examined	Type material examined INDIA • 1 syntype (Fig. 19 A – D); UMZC I. 102730 • 6 possible syntypes (one of them separated in a different vial and marked with a “ T ”); Darjiling; NHMUK 1906.4. 4.44 (Fig. 19 E – H). The latter lot was not considered type by Preece et al. (2022). Type locality “ in valle Rungun ”.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A32CFFCEB97DFD25B4B3F9E5.taxon	description	Fig. 20 A – D	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A32CFFCEB97DFD25B4B3F9E5.taxon	diagnosis	Diagnosis This species can be recognized based on the high conical shell shape, widely-spaced ribs, and thin peristome. R 2 with several (ca 10) low ribs.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A32CFFCEB97DFD25B4B3F9E5.taxon	materials_examined	Type material examined INDIA • holotype (see Fig. 20 A – D and Preece et al. 2022); UMZC I. 103745. Additional material examined INDIA • 5 shells; Rungun Valley, Darjiling; NHMUK 1906.4.4.41. Type locality “ ad Darjiling Himalayae Sikkimensis ”.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A32CFFF0B91DF966B284FE01.taxon	description	Fig. 20 E – I	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A32CFFF0B91DF966B284FE01.taxon	materials_examined	Type material examined INDIA • 18 syntypes (Fig. 20 E – I); Toruputu Peak, Dafla Hills, Dikrang Valley, Dafla; Godwin-Austen leg.; NHMUK 1903.7. 1.2596. Type locality “ Dikrang Valley, Dafla Hills ”.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A32CFFF0B91DF966B284FE01.taxon	diagnosis	Differential diagnosis In size, shape and R 1 sculpture it is similar to C. constrictus, but has only 4 – 5 R 2 ribs.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A312FFF0B91EFE02B7E3F9A1.taxon	description	Fig. 21 A – E	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A312FFF0B91EFE02B7E3F9A1.taxon	diagnosis	Diagnosis This species is characterized by a conical shell and dense ribbing on the entire surface. The peristome is thin, but the umbilicus is slightly covered by a reflected outer peristome.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A312FFF0B91EFE02B7E3F9A1.taxon	materials_examined	Type material examined INDIA • 1 syntype of C. elegans (Fig. 21 A – E); Shengorh Pk, Dafla Hills; Godwin-Austen coll.; NHMUK 1903.7. 1.2594 • 1 syntype of A. khunhoensis (Fig. 21 F – J); Khunho Peak, Naga Hills; Godwin-Austen leg.; NHMUK 1903.7. 1.2520. Type localities “ Shengorh Peak, Dafla Hills ” (C. elegans); “ Khunho Peak, Naga Hills ” (C. khunhoensis).	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A312FFF0B91EFE02B7E3F9A1.taxon	discussion	Remarks This species is similar to C. constrictus, but is characterized by a thinner peristome and finer ribbing. Cycloryx granum has similar sculpture, but it is more globular in shape. Cycloryx khunhoensis has slightly less dense ribbing than C. elegans, which we interpret as part of the intraspecific variability.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A312FFF2BAFAF9A1B267FDBB.taxon	description	Fig. 19 I – Q	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A312FFF2BAFAF9A1B267FDBB.taxon	diagnosis	Diagnosis This species can be recognized based on the short R 3.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A312FFF2BAFAF9A1B267FDBB.taxon	materials_examined	Type material examined Photos of the holotype (Fig. 19 I – L) examined. Additional material examined INDIA • 2 shells (Fig. 19 M – Q); North Sikkim, 2 km N of Lachen, towards Gurudongmar Lake; 27.74487 ° N, 88.54439 ° E; 2749 m a. s. l.; 27 Nov. 2019; N. A. Aravind leg.; ATREE / 2019 / LS 6001, ATREE / 2019 / LS 6002. Type locality “ Gasa Dzongkhag, 9 km S of Gasa, 2400 m a. s. l., 27 ° 50 ′ N 89 ° 43 ′ E ” (Bhutan).	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A310FFF2B991F9C3B370F92C.taxon	discussion	Remarks The species belonging here usually have a globular shell and a strongly expanded outer peristome.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A310FFF2B91CFDCFB531F9BF.taxon	description	Fig. 20 J – N	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A310FFF2B91CFDCFB531F9BF.taxon	diagnosis	Diagnosis A species with somewhat questionable status (see Remarks). The only known shell is characterized by a somewhat conical-ovoid shell, smooth initial whorls, widely-spaced ribbing in the lower whorls and thin peristome.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A310FFF2B91CFDCFB531F9BF.taxon	materials_examined	Type material examined MYANMAR • holotype (single shell mentioned in the original description: Fig. 20 J – N); Shan Hills, in the same slide as N. 22; Fedden leg.; NHMUK 1906.5. 5.87. Type locality “ Shan States ”.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A310FFF2B91CFDCFB531F9BF.taxon	discussion	Remarks It is larger than any known species in this group. It is most similar to C. constrictus but has weaker ribbing (upper whorls smooth, although the holotype is strongly corroded). The more obvious separation between the inner and outer peristomes of this species compared to the figured syntype of C. constrictus is not a stable character, because in the other available lot of C. constrictus (NHMUK 1906.4.4.41) there are shells with similar peristome to that of the holotype of C. tenellus. A single shell is known of this species and very few of C. constrictus. If more material becomes available, it may happen that C. tenellus will be moved to the synonymy of C. constrictus.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A310FFF4B935F95CB5C7FB5D.taxon	description	Fig. 22 A – E	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A310FFF4B935F95CB5C7FB5D.taxon	materials_examined	Type material examined Lectotype (hereby selected: Fig. 22 A – E) INDIA • Japvo Peak, Naga Hills; elev. 9890 f; Godwin-Austen leg.; acc. no. 1830, NHMUK 1903.7.1.2591.1. Paralectotypes INDIA • 5 specs; same data as for lectotype; NHMUK 1903.7. 1.2591.2 to 1903.7.1.2591.6 • several specs in two vials, one of the vials with a single shell and a pink wool; Munipur side of the Burrail; NHMUK 1903.7.1.2606. Type localities “ Japvo Peak, Naga Hills, 9890 ft ”; “ Munipur side of the Burrail ”; Angaoluo Peak.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A310FFF4B935F95CB5C7FB5D.taxon	diagnosis	Differential diagnosis Densely ribbed, similar to C. granum, but has wider spaces between R 3 ribs.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A310FFF4B935F95CB5C7FB5D.taxon	discussion	Remarks In the type collection in the same box we found the three vials (1: Japvo peak, labelled as “ var. minor ”, 2591; 2: several shells with label “ 2606 ”; 3: single shell, also with label “ 2606 ” and a pink wool [which usually indicates that it is the “ type ”]). Besides the size, the difference between shells from the two localities is that R 3 is much more densely ribbed in “ var. minor “ (from Japvo Peak). In the original description, the type locality was given as “ Japvo Peak, Naga Hills, 9890 ft ”, and it is later added that “ obtained also on the Munipur side of the Burrail, No. 2606 B. M.; and Angaoluo Peak, No. 2749 B. M. ”. One understanding of Godwin-Austen’s concept is to accept only the lot from Japvo Peak as syntypes, and none of the shells from the other two sites. However, in that case specimens from Burrail would be excluded from the type series, which would be strange for a species named “ burrailensis ”. Moreover, although not mentioned in the original description, the shells from Japvo Peak were labelled as “ var. minor, 2591 ”, suggesting, that the ones from Burrail should be considered “ typical ”. Lastly, the description states that there is a “ distant fine costulation between sutural tube and aperture ”, which unambiguously refers to the larger form from Burrail (NHMUK 1903.7.1.2606). Therefore, we accept shells belonging to all three lots as syntypes. The single shell from Angaoluo Peak was also found in the general collection, and it is similar to the ones from Burrail in size and formation of the aperture, but the R 1 ribbing is approximately half as dense. Since the syntypes were collected at three different localities, and are somewhat different in shell traits, we hereby select a lectotype to avoid further nomenclatural confusion.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A316FFF6B936FAEDB70EFE26.taxon	description	Fig. 23 A – E	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A316FFF6B936FAEDB70EFE26.taxon	diagnosis	Diagnosis This species can be recognized by its large size, the reflected outer peristome in the direction of the umbilicus, and the thick peristome (protruding inner peristome). See also under Cycloryx rohiniae sp. nov.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A316FFF6B936FAEDB70EFE26.taxon	materials_examined	Type material examined INDIA • 2 syntypes (Fig. 23 A – E); Sadia, Assam; NHMUK 1903.7. 1.2516. Type locality “ Sadia, Assam, 350 ft ”.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A316FFF6B936FAEDB70EFE26.taxon	discussion	Remarks Similar to C. major, but it is larger and has a thicker peristome with more protruding inner peristome. Similar in size to A. otiphorus, but has a more globular shell, stronger ribbing on the apical whorls, and ribs on R 3.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A314FFF6BA84FE5AB43BFCDA.taxon	description	Fig. 24 A – D	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A314FFF6BA84FE5AB43BFCDA.taxon	diagnosis	Diagnosis This species can be recognized based on the very widely-spaced R 1 ribs and the smooth R 2 with numerous slender microtunnels.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A314FFF6BA84FE5AB43BFCDA.taxon	materials_examined	Type locality “ Samdrup Jongkhar Dzongkhag, 9 km N of Samdrup Jongkhar, 1008 m a. s. l., warm broadleaf forest, 26 ° 52 ′ N 91 ° 29 ′ E ” (Bhutan).	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A314FFF9B91DFC6FB599FE05.taxon	description	Fig. 22 F – O	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A314FFF9B91DFC6FB599FE05.taxon	diagnosis	Diagnosis Shell globular, ribs dense on the entire shell.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A314FFF9B91DFC6FB599FE05.taxon	materials_examined	Type material examined INDIA • 1 syntype of A. granum (Fig. 22 F – J); Margarita, near Sadia, Assam; Godwin-Austen coll.; NHMUK 1903.7. 1.2511 • 1 syntype of A. multicostulatus (Fig. 22 K – O); Munipur, head of Lanier Rr; Godwin-Austen leg.; NHMUK 1903.7. 1.2557. Type localities “ Margarita, foot of Eastern Naga Hills ” (A. granum); “ Head of the Lanier River, Lahupa Naga Hills, N. E. Manipur ” (A. multicostulatus).	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A314FFF9B91DFC6FB599FE05.taxon	discussion	Remarks Types of C. granum are much smaller than C. multicostulatus, but there is no difference between those shells other than shell size. Specimens from the Japvo Peak, Naga Hills (labelled as “ A. burrailensis var. minor ”, see also A. burrailensis) are intermediate in size between C. granum and C. multicostulatus. Therefore, we treat the latter as a junior synonym of the former.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A31BFFF8BAA1FE06B5F4FE7F.taxon	description	Fig. 24 E – H	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A31BFFF8BAA1FE06B5F4FE7F.taxon	diagnosis	Diagnosis This species can be recognized by the widely-spaced R 1 ribs and the smooth R 2 with numerous slender microtunnels.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A31BFFF8BAA1FE06B5F4FE7F.taxon	materials_examined	Type locality “ Haa Dzongkhag, Gakiling, Rangtse Nye, 27 ° 05 ′ N 89 ° 09 ′ E, 890 m a. s. l. ” (Bhutan).	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A31BFFF8BAA1FE06B5F4FE7F.taxon	diagnosis	Differential diagnosis Differs from C. sajumbiclausus by the presence of spiral striation.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A31AFFF8B923FE0CB510F901.taxon	description	Fig. 25	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A31AFFF8B923FE0CB510F901.taxon	diagnosis	Diagnosis This species can be recognized based on the dense R 1 ribs, the R 3 with widely-spaced ribs, the thick peristome (protruding inner peristome) and that the outer peristome is not reflected abruptly towards umbilicus (i. e., it does not form a sharp angle below the umbilicus).	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A31AFFF8B923FE0CB510F901.taxon	materials_examined	Type material examined MYANMAR • 4 syntypes of A. graphicus (Fig. 25 A – E); Moditoung, Tongoop Pass, Arakan; NHMUK 1906.4. 4.42 • 2 syntypes of A. difficilis (Fig. 25 F – J); Shan States; Fedden leg.; NHMUK 1906.5. 5.22. Type localities “ in montibus Arakanensibus provinciam Burmanam Pegu ad Arakan secernentibus ” (A. graphicus); “ Shan Hills ” (A. difficilis).	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A31AFFF8B923FE0CB510F901.taxon	discussion	Remarks Alycaeus graphicus and A. difficilis are identical in all shell traits (except minor difference in rib density), therefore they are treated as synonyms. The ribs of this species are more widely-spaced than those of C. major, although no other differences were found.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A31AFFFCB910F902B4C5FD2E.taxon	description	Figs 26, 27 F – J	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A31AFFFCB910F902B4C5FD2E.taxon	diagnosis	Diagnosis This species can be recognized by the dense R 1 ribbing, the regular ribbing on R 3, and the angle below the umbilicus, formed by the reflected outer peristome.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A31AFFFCB910F902B4C5FD2E.taxon	materials_examined	Type material examined INDIA • 8 syntypes of A. major (Fig. 26 A – E); Mangat Valley, Jiantia Hills; Godwin-Austen leg.; NHMUK 1903.7. 1.2518 • 17 syntypes of A. graphicus var. variabilis (Fig. 26 F – J); Lhota Naga; NHMUK 1903.7. 1.2607 • 1 syntype of A. thompsoni (Fig. 26 K – O); Munipur; Godwin-Austen coll.; NHMUK 1903.7. 1.2550. Additional material examined INDIA – Mizoram • 1 empty shell (Fig. 27 F – J); Blue Mountain, Lawngtlai district; 22.727 ° N, 93.135 ° E; 1400 m a. s. l.; 27 Jan. 2019; N. A. Aravind leg.; Ar 53; NZSI LM 1716. Type localities “ Mangut Valley, Jaintia Hills ” (A. major); “ Lhota Naga ” (A. graphicus var. variabilis); “ Munipur ” (A. thompsoni).	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A31AFFFCB910F902B4C5FD2E.taxon	diagnosis	Differential diagnosis Cycloryx graphicus has more widely-spaced ribs than C. major, and the outer peristome does not form a sharp angle below the umbilicus in that species.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A31AFFFCB910F902B4C5FD2E.taxon	discussion	Remarks The specimen from Blue Mountain is practically identical to the type specimens of P. major in all shell characters (size, shape, rib density). Specimens labelled as “ A. graphicus var. ” from the Burroi Gorge, Dafla Hills (NHMUK 1903.07.1.2598) are also identical, although most specimens are larger. Although slightly larger, A. thompsoni and A. graphicus var. variabilis also have similar traits. Therefore, they are treated as synonyms of Cycloryx major.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A31EFFFCB907FD52B335F836.taxon	description	Fig. 28 Alycaeus margarita Theobald in Hanley & Theobald, 1874: 39, pl. 95 fig. 10.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A31EFFFCB907FD52B335F836.taxon	diagnosis	Diagnosis This species is characterized by a globular shell, and widely-spaced, strong ribs on the entire shell surface.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A31EFFFCB907FD52B335F836.taxon	materials_examined	Type material examined MYANMAR • 1 syntype of A. margarita (Fig. 28 A – D); Upper Salwin; NHMUK 1888.12. 4.892 • 1 syntype of A. margarita; Upper Salwin; NHMUK 1888.12. 4.893 • 3 syntypes of A. graphiarius (Fig. 28 E – I, mentioned as 1 syntype in Páll-Gergely et al. 2020); Shan States; Fedden leg.; Blanford coll.; NHMUK 1906.4. 4.21 (two of them are A. margarita, the third shell belongs to another species, see remarks). Type locality “ Shan Provinces ” (A. margarita); not specified for A. graphiarius.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A31EFFFCB907FD52B335F836.taxon	discussion	Remarks Two syntypes of A. graphiarius agreed with the original description (Godwin-Austen, 1914) by having widely-spaced ribs, but the third shell had much denser ribbing, and therefore, it clearly belongs to a different species. The two sparsely ribbed syntypes of A. graphiarius are identical to those of A. margarita in all shell characters (size, shape, density of ribs), therefore the former is a junior synonym of the latter.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A31DFFE1B975F9C5B2A5FD95.taxon	description	Fig. 29	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A31DFFE1B975F9C5B2A5FD95.taxon	diagnosis	Diagnosis This species can be recognized based on the large, conical shell, and the weak sculpture. See also under Cycloryx rohiniae sp. nov.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A31DFFE1B975F9C5B2A5FD95.taxon	materials_examined	Type material examined INDIA • 1 syntype of A. otiphorus (see Preece et al. 2022: fig. 33 e); no locality data; UMZC I. 102555 • 12 syntypes of A. summus (Fig. 29 F – J); Rechila Pk, W. Bhutan; Godwin-Austen leg.; NHMUK 1903.7. 1.2573. Additional material examined INDIA • single shell from the R. McAndrew collection (Fig. 29 A – D); UMZC I. 102560 • 1 shell of A. otiphorus; Darjiling; “ compared with typical sp in Museum Cambridge ”; NHMUK 1903.7.1.2565. Type localities “ ad Pankabari (1000 ped. alt.) et in valle Rungun (4000 ped.) prope Darjiling Himalayanum ” (A. otiphorus); “ Rechila Peak, Western Bhutan ” (A. summus).	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A31DFFE1B975F9C5B2A5FD95.taxon	discussion	Remarks Alycaeus summus is identical to A. otiphorus, therefore the former is a junior synonym of the latter. Unlike other members of this species group, A. otiphorus has a rather conical shell (instead of conical-globular), but it is classified here due to the expanded and reflected outer peristome.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A303FFE1BAC3FDB5B26DFAA6.taxon	description	Fig. 23 F – J	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A303FFE1BAC3FDB5B26DFAA6.taxon	diagnosis	Diagnosis This species can be distinguished from all congeners based on the lamella-like ribs on R 3.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A303FFE1BAC3FDB5B26DFAA6.taxon	materials_examined	Type material examined INDIA • 2 syntypes (Fig. 23 F – J); Toruputu Pk, Dafla Hills; NHMUK 1903.7. 1.2595. Type locality “ Torúpútú Peak, Dafla Hills ”.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A303FFE1BAC3FDB5B26DFAA6.taxon	diagnosis	Differential diagnosis The most similar species is C. burrailensis, but that species has denser R 1 ribs.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A303FFE0BAD1FADBB348FB7A.taxon	description	Fig. 27 A – E	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A303FFE0BAD1FADBB348FB7A.taxon	diagnosis	Diagnosis A large species of Cycloryx with smooth R 3.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A303FFE0BAD1FADBB348FB7A.taxon	etymology	Etymology The new species is named after Ms Rohini Nilekani, author, environmentalist and acclaimed philanthropist.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A303FFE0BAD1FADBB348FB7A.taxon	materials_examined	Type material examined Holotype INDIA – Mizoram • empty shell (D: 3.7 mm, H: 4.1 mm, Fig. 27 A – E); Blue Mountain, Lawngtlai district; 22.727 ° N, 93.135 ° E; 1400 m a. s. l.; 27 Jan. 2019; N. A. Aravind leg.; Ar 37; NZSI LM 1717. Paratype INDIA – Mizoram • 1 empty shell; same data as for holotype; Ar 34; NZSI LM 1718 • 1 empty shell; same data as for holotype; Ar 35; NZSI LM 1719.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A303FFE0BAD1FADBB348FB7A.taxon	description	Description Shell large for the genus, all available shells slightly corroded, original colour may be greenish-yellowish, semitranslucent; shell conical or conical-globular, height and width approximately equivalent; body whorl rounded; protoconch smooth, glossy, consisting of 1.5 or slightly fewer whorls; R 1 of ca 2 – 2.25 whorls, regularly ribbed, rib density gradually decreases towards end of R 2 (last half whorl with ca 16 ribs); ribs thread-like, low; fine, dense spiral striation discernible between ribs; boundary between R 1 and R 2 not conspicuous, but clearly visible due to increase in rib density (R 2 ribs twice as dense as ribs at the end of R 1); R 2 practically smooth, its “ ribs ” (6 – 7 in number) are not elevated from the surface, represented as slender, light stripes; sutural tube short, cylindrical; R 2 + R 3 ca quarter whorl, R 3 ca 3 times as long as R 2; boundary between R 2 and R 3 clearly visible due to a shallow constriction; R 3 with a low central, long swelling, smooth, only fine growth lines visible but no ribs; aperture slightly oblique to shell axis, rounded; inner and outer peristomes clearly separated; inner peristome slightly protruding and expanded; outer peristome strongly expanded, sharp, and reflected in direction of umbilicus; umbilicus open, narrow, the reflected outer peristome curves into umbilicus, terminating at its centre. OPERCULUM. A single operculum of the Ar 35 shells was examined. The inner side is smooth, without central swelling, outer side multispiral, without elevated lamina. MEASUREMENTS. D: 3.7 mm, H: 3.5 – 4.1 mm.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A303FFE0BAD1FADBB348FB7A.taxon	distribution	Distribution Known only from the Blue Mountain region.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A303FFE0BAD1FADBB348FB7A.taxon	diagnosis	Differential diagnosis The only species with similar size and lack of R 3 ribs is C. otiphorus, which has a more pointed (more triangular) shell, and has denser ribs on the apical part. Cycloryx dihingensis, which is similar in shell shape, has strong R 3 ribs. All these species inhabit Assam, which geographically is distant from Mizoram.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A302FFE0BAA6FB0FB71DF92E.taxon	description	Fig. 24 I – L	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A302FFE0BAA6FB0FB71DF92E.taxon	diagnosis	Diagnosis This species can be recognized based on the widely-spaced R 1 ribs and the smooth R 2 with numerous slender microtunnels. It differs from C. haumbiclausus based on the absence of spiral striation.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A302FFE0BAA6FB0FB71DF92E.taxon	materials_examined	Type locality “ Samdrup Jongkhar Dzongkhag, 4 km NW of Samdrup Jongkhar, 260 m a. s. l., 26 ° 49 ′ N 91 ° 28 ′ E ” (Bhutan).	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A302FFE0B930F95CB20DF877.taxon	type_taxon	Type species	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A301FFE3BA71FD67B552FB2D.taxon	description	Fig. 30 A – D	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A301FFE3BA71FD67B552FB2D.taxon	materials_examined	Type locality “ between Duenmang Tsachu and Gongphu Zero Point, 24 km SE of Zhemgang, 335 m a. s. l., 27 ° 02 ’ N 90 ° 48 ’ E, scree in warm broadleaf forest ” (Bhutan).	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A301FFE3BA71FD67B552FB2D.taxon	diagnosis	Differential diagnosis Probably the most similar species is D. rugosus due to the similar shell and aperture shape as well as rib density. However, that species has a shorter distance between the inner and outer peristomes and has a wider umbilicus. For the differences with other Bhutanese species see Gittenberger et al. (2024). See also under Chamalycaeus asaluensis and D. umbiclausus.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A301FFE3BACBFB5DB5BBF886.taxon	description	Fig. 31 A – E	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A301FFE3BACBFB5DB5BBF886.taxon	materials_examined	Type material examined MYANMAR • holotype (Fig. 31 A – E); Shan State, Hopong centre 7.4 km towards Namsang, rd. # 4 N 5 km, near Parpant Cave; 20 ° 850.9630 ′ N, 97 ° 814.2670 ′ E; 1170 m a. s. l.; 6 Oct. 2018; A. Hunyadi, K. Okubo and J. U. Otani, leg.; HA (to be deposited in the HNHM). Type locality “ Myanmar, Shan State, Hopong centre 7.4 km towards Namsang, rd. # 4 N 5 km, near Parpant Cave, 20 ° 850.9630 ’ N, 97 ° 814.2670 ’ E, 1170 m a. s. l. ”	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A301FFE3BACBFB5DB5BBF886.taxon	diagnosis	Differential diagnosis This species differs from its congeners in this species group by the strongly expanded outer peristome and the smaller distance between the two peristomes. Some species of Chamalycaeus are also similar to Dicharax floridus, but their outer peristome is less expanded.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A301FFE3B9BFFEECB216FDE3.taxon	discussion	Remarks This species group is characterized by a usually strongly ribbed shell, a fringed aperture, and most species (exception: D. floridus) have a large space between the inner and outer peristomes. The R 3 swelling of D. obscurus is similar to that of the species belonging to the species group 5, but it is included in this group due to the fringed aperture and the large space between the two peristomes. See also remarks under Dicharax species group 2 regarding D. bicrenatus.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A301FFE7B92FF8BBB50DFB64.taxon	description	Fig. 32	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A301FFE7B92FF8BBB50DFB64.taxon	diagnosis	Diagnosis The elevated spire, rather smooth R 1 (with the exception of strong, widely-spaced ribs along the suture), the long R 3 swelling with straight anterior part, and the strongly fringed aperture distinguish this species from its congeners.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A301FFE7B92FF8BBB50DFB64.taxon	materials_examined	Type material examined INDIA • 14 syntypes of A. jaintiacus (Fig. 32 A – E); Nongjinghi, Jaintia Hills; Godwin-Austen leg.; NHMUK 1903.7. 1.2686 • 4 syntypes of A. jaintiacus var. crassus in 2 vials (Fig. 32 K – O); Nonjinghi, Jaintia; Godwin-Austen coll.; NHMUK 1903.7. 1.2752 • 1 syntype of A. serratus (Fig. 32 F – J); Laisen Valley, Jiantia Hills; NHMUK 1903.7. 1.2487. Type localities “ in Nongjinghi, Jaintia ” (A. jaintiacus); “ in Nongjinghi, Jaintia, 4563 feet ” (A. jaintiacus var. crassus); “ Laisen Trigl. station, Munipur Hills ” (A. serratus).	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A301FFE7B92FF8BBB50DFB64.taxon	diagnosis	Differential diagnosis The most similar is D. obscurus; for comparisons, see under that species. Species of the Dicharax species group 7 (D. chennelli and similar species) resemble D. jaintiacus, but the distance between their two peristomes is smaller, their R 3 swelling is more inflated and usually have R 3 depressions. In contrast, D. jaintiacus has a less inflated R 3 without depressions.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A301FFE7B92FF8BBB50DFB64.taxon	discussion	Remarks Dicharax jaintiacus var. crassus is practically identical to the nominotypical form, therefore it is treated here as a junior synonym. Alycaeus serratus is slightly smaller and has a shallower R 3 swelling, but all other shell characters are very similar, and they were described from the same geographic region; therefore, Alycaeus serratus is hereby synonymised with D. jaintiacus. Alycaeus serratus was not compared with A. jaintiacus in its original description.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A305FFE7B92EFAE5B20EF85B.taxon	description	Fig. 33 A – E	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A305FFE7B92EFAE5B20EF85B.taxon	materials_examined	Type material examined INDIA • 16 syntypes (Fig. 33 A – E); E. Naga; R. H. Beddome coll.; NHMUK 1912.4. 16.273. Type locality “ Eastern Naga ”.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A305FFE7B92EFAE5B20EF85B.taxon	diagnosis	Differential diagnosis Dicharax dohertyi (of which no specimens were available, see under D. dohertyi) and D. rugosus are similar in sculpture and aperture formation, but they have a higher spire. Dicharax trashidentatus is very similar to D. muspratti in shell and aperture shape and shell sculpture, but it also has a higher spire and denser R 1 ribs. Dicharax rugosus is also similar due to the strong R 1 ribs, the large distance between the inner and outer peristomes and the fringed aperture, but it has a higher spire.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A304FFE9B92AFEECB347FE05.taxon	description	Fig. 31 F – J	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A304FFE9B92AFEECB347FE05.taxon	materials_examined	Type material examined INDIA • 1 syntype (Fig. 31 F – J); Cherra Poonje; Godwin-Austen leg.; NHMUK 1913.3. 16.8. Type locality “ Cherra Poonje ”.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A304FFE9B92AFEECB347FE05.taxon	diagnosis	Differential diagnosis The most similar species in this species group is D. jaintiacus, which has a more globular shell, possesses more widely-spaced R 1 ribs, and a less expanded outer peristome. Species of the Dicharax species group 7 are similar to D. obscurus in the formation of the R 3 swelling, but the distance between their two peristomes is smaller and their R 3 swelling is more inflated and usually have depressions and / or grooves.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A30BFFE9B920FE06B772F836.taxon	description	Fig. 33 F – N	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A30BFFE9B920FE06B772F836.taxon	diagnosis	Diagnosis The combination of the strongly ribbed R 1, the rather centrally located R 3 swelling, the large distance between the inner and outer peristomes and the fringed aperture distinguishes this species from its congeners.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A30BFFE9B920FE06B772F836.taxon	materials_examined	Type material examined BHUTAN • holotype of Dicharax sajodentatus (Fig. 33 K – N) (photographs examined). INDIA • 3 syntypes of D. rugosus (figured specimen indicated by pink wool; Fig. 33 F – J); Burroi Gorge, Dafla; NHMUK 1903.7. 1.2641. Type localities “ Burroi Gorge, Dafla Hills ” (A. rugosus); “ Samdrup Jongkhar Dzongkhag: N of Bangtar, Phuntshothang, 26 ° 53 ’ N 91 ° 42 ’ E, 375 m a. s. l. ” (D. sajodentatus).	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A30BFFE9B920FE06B772F836.taxon	diagnosis	Differential diagnosis For differences with newly described Bhutanese species (D. zhemdentatus, D. trashidentatus, D. umbiclausus and D. aspidentatus) see Gittenberger et al. (2024) (under the name D. sajodentatus). See also under C. asaluensis, C. crispatus, C. dohertyi and D. muspratti.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A30BFFE9B920FE06B772F836.taxon	discussion	Remarks During the examination of A. rugosus, we found traces of a spiral striation on the teleoconch, based on which we classified that species in the genus Metalycaeus (see Páll-Gergely et al. 2020). However, the re-examination revealed that those spiral lines were probably part of lower shell layers of the corroded syntypes, and moreover, Gittenberger et al. (2024) described several similar species from Bhutan that can undoubtedly be assigned to Dicharax. The comparison of syntypes of D. rugosus and D. sajodentatus revealed that their shell shape, sculpture and formation of the peristome are identical; therefore, we treat the latter as a synonym of the former.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A30AFFE8B9BFF95CB484F85F.taxon	discussion	Remarks This species group is characterized by a strongly elevated, short R 3 swelling that is situated very close to the outer peristome, and a slightly to strongly crenulated peristome. Dicharax bicrenatus is classified in this species group due to the small distance between the R 3 swelling and the outer peristome. However, it also has a large distance between the two peristomes, which is characteristic of Dicharax species group 1.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A30AFFE8BA00FEECB79BFCBF.taxon	description	Fig. 29 E – H	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A30AFFE8BA00FEECB79BFCBF.taxon	materials_examined	Type locality “ Trashigang Dzongkhag: 3 km E of Trashigang, 825 m a. s. l., 27 ° 21 ’’ N 91 ° 35 ’ E ” (Bhutan).	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A30AFFE8BA00FEECB79BFCBF.taxon	diagnosis	Differential diagnosis In shell shape and sculpture this species is practically identical with D. zhemdentatus. The only difference is that the chevrons (for the terminology see Gittenberger et al. 2024) reach the peristome edge, and do not in D. zhemdentatus. Therefore, we maintain this as a valid species until intermediate forms become available. For the differences with other Bhutanese species see Gittenberger et al. (2024). See also under Dicharax muspratti.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A30AFFE8BA7AFCC1B244FB64.taxon	description	Fig. 29 M – P	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A30AFFE8BA7AFCC1B244FB64.taxon	materials_examined	Type locality “ Samdrup Jongkhar Dzongkhag: 20 km N of Samdrup Jongkhar, 1680 m a. s. l., 26 ° 59 ’ N 91 ° 31 ’ E ” (Bhutan).	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A30AFFE8BA7AFCC1B244FB64.taxon	diagnosis	Differential diagnosis The most similar species is D. aspidentatus, which has a slightly open umbilicus, more widely-spaced ribs, and a lower R 3 swelling. For other differences, see Gittenberger et al. 2024.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A30AFFE8BA9FFB1BB7A0F92A.taxon	description	Fig. 30 I – L	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A30AFFE8BA9FFB1BB7A0F92A.taxon	materials_examined	Type locality “ Zhemgang Dzongkhag: Ngangla - Trong (= Tong), 2 km N of Ngangla, c. 17 km NE of Panbang, 1085 m a. s. l., 26 ° 58 ’ N 91 ° 03 ’ E ” (Bhutan).	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A30AFFE8BA9FFB1BB7A0F92A.taxon	diagnosis	Differential diagnosis For diagnosis and differences with other Bhutanese species see Gittenberger et al. (2024). See also under D. trashidentatus.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A309FFEBB933FEECB50EFC18.taxon	description	Fig. 34 A – E	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A309FFEBB933FEECB50EFC18.taxon	diagnosis	Diagnosis The combination of the small shell size, the large distance between the inner and outer peristomes, the depressed shell, the fine ribbing, and the short R 3 swelling defines this species.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A309FFEBB933FEECB50EFC18.taxon	materials_examined	Type material examined INDIA • 7 syntypes in two vials (Fig. 34 A – E); Kopamedza, Naga Hills; NHMUK 1903.7. 1.2490. Type locality “ Kopamedza Peak, Naga Hill, 8 – 9,000 feet, in forest ”.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A309FFEBBA95FC29B7E5FA41.taxon	description	Fig. 35 A – D	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A309FFEBBA95FC29B7E5FA41.taxon	materials_examined	Type locality “ Thimphu Dzongkhag: road W of quarry near Bjimina (13 km SW of Thimphu), 2700 m a. s. l., 27 ° 24 ’ N 89 ° 34 ’ E ” (Bhutan).	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A309FFEBBA95FC29B7E5FA41.taxon	diagnosis	Differential diagnosis The combination of the densely ribbed R 1, the elevated, short R 3 swelling situated close to the outer peristome, the large distance between the inner and outer peristomes is true for this species and for D. lunabulbus. However, the latter species is much smaller, it has a more undulated inner peristome, a larger distance between the inner and outer peristomes, and more widely-spaced R 2 ribs. Dicharax vanderbijli has a smaller distance between the inner and outer peristomes, and has more lamella-like and widely-spaced ribs.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A309FFEDB924F9C2B508FC38.taxon	description	Fig. 34 F – J	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A309FFEDB924F9C2B508FC38.taxon	materials_examined	Type material examined INDIA • 2 syntypes (Fig. 34 F – J); Mokarsa, Khasi Hills; Godwin-Austen coll.; NHMUK 1903.7. 1.2642. Type locality “ On Burrail Range, N. Cachar, at ca 5000 feet ”.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A309FFEDB924F9C2B508FC38.taxon	diagnosis	Differential diagnosis Dicharax daflaensis has a smooth R 2, while D. crenatus possesses curved R 2 ribs. Moreover, the R 3 swelling of D. daflaensis is more conical, i. e., less abruptly elevating. Dicharax tangmaiensis has a more strongly undulated inner peristome, and a smooth R 2 surface. While in D. crenatus the R 2 and R 3 are of comparable length, in D. davisi and D. ochraceus the R 3 is much longer than the R 2.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A309FFEDB924F9C2B508FC38.taxon	discussion	Remarks In the original description, Godwin-Austen (1871) gave “ Burrail Range, N. Cachar ” as the type locality. Later (Godwin-Austen 1914: 389) explained that he found two shells in his collection (from Mokarsa in the NW Khasi Hills, no. 2642), based on which the drawings of A. crenatus were made in the original description (Godwin-Austen 1871), but their locality was not mentioned in the original description. Based on ICZN Art. 72.4.1.1, the two shells from Mokarsa are also part of the type series. Even if Godwin-Austen referred to one of the shells as “ the type shell ” (Godwin-Austen 1914: 389), this does not constitute a valid lectotype designation (Art 74.5), because the condition of “ unambiguousity ” has not been met (we do not know which of the two shells were selected to be “ the type ”). Godwin-Austen (1874) noted that “ A. crenatus was found as far east as Shiroifurar, also at Kezakenomih and Yémai ”. Those samples were not examined by us.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A30FFFEDB908FC49B77BF8D4.taxon	description	Fig. 36 A – J	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A30FFFEDB908FC49B77BF8D4.taxon	materials_examined	Type material examined MYANMAR • 2 syntypes of D. cucullatus; Shan States; NHMUK 1888.12.4.951 - 952 • 4 syntypes of A. davisi; Shan States; Woodthorpe leg.; NHMUK 1903.7. 1.1630. Type localities “ Shan States ” (A. cucullatus); “ Siam and Shan boundary ” (A. davisi).	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A30FFFEDB908FC49B77BF8D4.taxon	diagnosis	Differential diagnosis The most similar species is D. ochraceus, which is smaller, and has more widely-spaced R 2 ribs (Jirapatrasilp et al. 2021). See also under D. digitatus.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A30FFFEDB908FC49B77BF8D4.taxon	discussion	Remarks This is a variable species regarding the aperture shape and the height of R 3 swelling (see Jirapatrasilp et al. 2021).	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A30FFFECB91BF875B753FBDC.taxon	description	Fig. 34 K – O	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A30FFFECB91BF875B753FBDC.taxon	diagnosis	Diagnosis The high and short R 3 swelling, which is situated close to the outer peristome, the strongly fringed inner peristome, the large distance between the inner and outer peristomes, and the protruding finger-like projections on the inner peristome distinguish this species from all congeners.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A30FFFECB91BF875B753FBDC.taxon	materials_examined	Material examined INDIA • 1 shell (probably not syntype, but figured by Godwin-Austen 1914; Fig. 34 K – O); Rechila Pk, Sikkim; W. Robert leg.; NHMUK 1903.7. 1.1253. Type locality “ apud Darjeeling in vallo Rungno fluminis Himalayae Sikkimensis ”.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A30FFFECB91BF875B753FBDC.taxon	diagnosis	Differential diagnosis Other species of this species group may also have finger-like projections of the peristome (D. cucullatus, D. ochraceus, D. plectocheilus, D. tangmaiensis), but in those cases the outer peristome ‘ follows’ the projections, while in D. digitatus the outer peristome stops at a single, straight line. See also under D. bicrenatus.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A30EFFECBA72FB6CB2E8F990.taxon	description	Fig. 35 E – H	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A30EFFECBA72FB6CB2E8F990.taxon	materials_examined	Type locality “ Chhukha Dzongkhag: SE of Gedu, 26 ° 54 ’ N 89 ° 33 ’ E, 1988 m a. s. l. ” (Bhutan).	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A30EFFECBA72FB6CB2E8F990.taxon	diagnosis	Differential diagnosis Dicharax vanderbijli has more widely-spaced R 1 ribs, smaller distance between the inner and outer peristomes, and a less fringed aperture. See also under D. bicrenatus and D. bjiminaensis.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A30EFF91B932F9B0B5E4FEEC.taxon	description	Fig. 36 K – O	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A30EFF91B932F9B0B5E4FEEC.taxon	materials_examined	Type material examined MYANMAR • 2 syntypes (Fig. 36 K – O); Ruby mine Disr., Up. Burma; Doherty leg.; NHMUK 1903.7. 1.2684. Type locality “ Ruby Mines District, Upper Burmah ”.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A30EFF91B932F9B0B5E4FEEC.taxon	diagnosis	Differential diagnosis See under D. cucullatus (the most similar species) and D. digitatus.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A373FF91B90DFE9CB54CFB57.taxon	description	Fig. 37 A – D	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A373FF91B90DFE9CB54CFB57.taxon	description	figs 4, 4 a – c. Alycaeus (Dicharax) plectochilus [sic] – Kobelt 1902: 375. — Gude 1921: 264 – 265. Alycaeus plectocheilus, large var. – Godwin-Austen 1914: 342 – 343, pl. 133 figs 3, 3 a – c. Chamalycaeus (Dicharax) plectochilus – Ramakrishna et al. 2010: 65. Dicharax plectocheilus – Páll-Gergely et al. 2017: 54. — Preece et al. 2022: 80, fig. 38 e – f. — Gittenberger et al. 2024: 214, figs 39 – 40, 43. Dicharax (?) plectocheilus – Páll-Gergely et al. 2020: 101.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A373FF91B90DFE9CB54CFB57.taxon	materials_examined	Type material examined INDIA • 2 syntypes (Fig. 37 A – D); Darjiling, Rungun Valley; Blanford coll.; NHMUK 1906.4. 4.184. Type locality “ in valle Rungun ”.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A373FF91B90DFE9CB54CFB57.taxon	diagnosis	Differential diagnosis There are two other species in this species group with smooth R 2 surface: D. subdigitatus and D. tangmaiensis. The former has a larger shell, has a larger distance between the R 3 swelling and the peristome, and less protruding peristomal projections; the latter is much larger than D. plectocheilus and has a less elevated R 3 swelling. See also under D. digitatus.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A373FF93B939FAEBB36FFE26.taxon	description	Fig. 37 E – I	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A373FF93B939FAEBB36FFE26.taxon	materials_examined	Type material examined INDIA • 3 syntypes; Shengorh Peak, Dafla Hills; Godwin-Austen coll.; NHMUK 1903.7. 1.2498. Type locality “ Shengorh Peak ” and “ Tánir ridge at 4000 feet ”.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A373FF93B939FAEBB36FFE26.taxon	diagnosis	Differential diagnosis Dicharax tangmaiensis is larger, and has a shorter distance between the R 3 swelling and the peristome. See also under D. plectocheilus.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A373FF93B939FAEBB36FFE26.taxon	discussion	Remarks The shell regions and the sculpture are similar to D. digitatus, but the undulation of the peristome started anterior to the R 3 swelling. Moreover, the R 3 swelling is situated at a greater distance from the peristome edge. Therefore, D. subdigitatus is a species in its own right, instead of a subspecies of D. daflaensis.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A371FF93B9BFF9EDB5B5F8A5.taxon	discussion	Remarks This species group is characterized by the relatively large (around 5 mm), usually finely ribbed shell, two R 3 swellings, and a rounded (not fringed) aperture. One species is known from the Himalayan region (D. mutatus), while the other three are known from central Myanmar (D. bifrons, D. spatiosus, D. woodthorpei).	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A371FF93B939FE5AB296FB86.taxon	description	Fig. 37 J – N	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A371FF93B939FE5AB296FB86.taxon	diagnosis	Diagnosis A large species with smooth R 2. See under D. digitatus and D. plectocheilus and D. subdigitatus.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A371FF93B939FE5AB296FB86.taxon	materials_examined	Type material examined CHINA • holotype (Fig. 37 J – N); Tong-Mai Town, Bo-Mi County, Tibet Autonomous Region; 20 Jun. 1980; Chen De-Niu leg.; CASIZ TM 0010054 • 1 paratype; same data as for holotype; CASIZ TM 0010056. Type locality “ Tongmai Town, (30 ° 01 ’ N, 95 ° E), Bomi County, Tibet Autonomous Region, China ”.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A371FF93BA89FBBBB7A8FA5C.taxon	description	Fig. 35 I – L	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A371FF93BA89FBBBB7A8FA5C.taxon	materials_examined	Type locality “ Paro Dzongkhag: 12 km SW of Paro, 3680 m a. s. l., 27 ° 22 ’ N 89 ° 19 ’ E ” (Bhutan).	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A371FF93BA89FBBBB7A8FA5C.taxon	diagnosis	Differential diagnosis Most similar to D. lunabulbus and D. bjiminaensis. For comparisons with those species, see the respective species.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A371FF95B975F8A6B376FE7D.taxon	description	Fig. 38 A – E	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A371FF95B975F8A6B376FE7D.taxon	materials_examined	Type material examined MYANMAR • 3 syntypes (Fig. 38 A – E); Shan States; NHMUK 1888.12.4.956 - 958. Type locality “ Shan States ”.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A371FF95B975F8A6B376FE7D.taxon	diagnosis	Differential diagnosis The most similar species is D. mutatus, which has a longer R 2 and more pronounced R 3 swellings.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A377FF95B921FE0EB7A8FB68.taxon	description	Fig. 38 F – J	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A377FF95B921FE0EB7A8FB68.taxon	materials_examined	Type material examined INDIA • 9 syntypes (Fig. 38 F – J); Toruputu Peak, Dafla Hills; Godwin-Austen coll.; NHMUK 1903.7. 1.2495. Type locality “ On Torúpútú, Tánir, and Shengorh Peaks, at 6 – 7000 feet elevation, in the dead leaves and moss about the roots of the forest ”.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A377FF95B921FE0EB7A8FB68.taxon	diagnosis	Differential diagnosis Dicharax spatiosus is larger, has a longer R 2, a more elevated main R 3 swelling, and a clearer distinction between the inner and outer peristomes. For differences with the most similar species, D. bifrons, see under that species.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A377FF95BAD0FB19B426F83D.taxon	description	Fig. 39 A – E	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A377FF95BAD0FB19B426F83D.taxon	diagnosis	Diagnosis The large shell, long R 2, two elevated swellings on R 3, and the circular (not fringed) aperture distinguishes this species from all congeners. See also under D. mutatus and also the original description of D. spatiosus for further comparisons.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A377FF95BAD0FB19B426F83D.taxon	materials_examined	Type material examined MYANMAR • holotype (Fig. 39 A – E); Shan State, 7.5 km N of Pinlaung, Tar Kge, near “ Big Bang Cave ”; 20 ° 810.2730 ′ N, 96 ° 847.4420 ′ E; 1540 m a. s. l.; A. Hunyadi, K. Okubo and J. U. Otani leg.; coll. HA (to be deposited in the HNHM). Type locality “ Myanmar, Shan State, Pinlaung centre N 7.5 km, Tar Kge, near „ Big Bang Cave ”, 20810.2730 N, 96847.4420 E, 1540 m a. s. l. ”.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A376FF97B934FEECB787FC5E.taxon	description	Fig. 39 F – J	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A376FF97B934FEECB787FC5E.taxon	diagnosis	Diagnosis This species differs from the other three species of the same species group by the smooth R 2.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A376FF97B934FEECB787FC5E.taxon	materials_examined	Type material examined MYANMAR • 22 syntypes in two vials (Fig. 39 F – J); Fort Stedman, Burma; Woodthorpe leg.; NHMUK 1903.7. 1.3064. Additional material examined MYANMAR • 2 shells; Burma; E. R. Sykes coll.; acc. no. 1825; NHMUK 20150126. Type locality “ Fort Stedman, Burma ”.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A376FF97B934FEECB787FC5E.taxon	diagnosis	Differential diagnosis There are several species with smooth R 2 surface (mostly in Dicharax species group 8), but D. woodthorpei can be distinguished from them by the relatively large shell, the double R 3 swelling and the rounded aperture.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A375FF97B937FAA4B578F863.taxon	description	Fig. 40 A – E	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A375FF97B937FAA4B578F863.taxon	materials_examined	Type material examined INDIA • holotype (single shell mentioned in the original description: Fig. 40 A – E); Tsanspu Valley; Oakes leg.; NHMUK 1903.7. 1.3582. Type locality “ Tsanspu Valley ”.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A375FF97B937FAA4B578F863.taxon	diagnosis	Differential diagnosis The most similar species is D. magnificus, which has a stronger R 1 sculpture, a more strongly fringed aperture and a sharper keel on the umbilical side of R 2.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A375FF97B9BFFBE3B523FAA7.taxon	discussion	Remarks This species group is characterized by the relatively large (around 5 mm), finely ribbed R 1, smooth R 2, two R 3 swellings, and the fringed aperture. Species of the Dicharax species group 3 also have double R 3 swellings, but have rounded apertures. Both species are known from the Indian Himalaya.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A37BFF99B9BFFA97B283F939.taxon	discussion	Remarks This species group is characterized by shells with an elevated spire and relatively low R 3 swelling. Other traits are variable. It is likely that D. globosus and D. vestitus are relatives, because other than the shell height they share the strong ribs, which are peculiarly oblique. Dicharax burchi, and D. verrucosus are also similar in the R 3 swelling that is situated close to the aperture. Dicharax conicus may not be related to the other high-spired species, and can be recognized by its peculiar outer peristome.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A37BFF99B937FEECB46CFAF3.taxon	description	Fig. 40 F – P	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A37BFF99B937FEECB46CFAF3.taxon	materials_examined	Type material examined INDIA • 1 syntype (Fig. 40 F – J); Yamney Valley, Abor Hills; NHMUK 1903.7. 1.3115. Additional material examined INDIA • 1 shell (Fig. 40 K – P); Arunachal Pradesh, 2 km from Kalek towards Rottung, base of the dry bamboo clumps, in semi-evergreen forest; 28 ° 6.067 ′ N, 95 ° 10.378 ′ E; 784 m a. s. l.; 24 April 2022; Aravind N. A. and Surya Narayanan leg.; NZSI LM 1914 • 7 shells; same data as for preceding; ATREE. Type locality “ Yamne valley, Abor Hills ”.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A37BFF99B937FEECB46CFAF3.taxon	diagnosis	Differential diagnosis See under Dicharax duoculmen.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A37BFF99B937FEECB46CFAF3.taxon	discussion	Remarks Based on the spiral striae visible on the protoconch of the syntype, this species has been classified in Metalycaeus (see Páll-Gergely et al. 2020). Re-examination of that syntype revealed that those spiral striae are parts of lower shell layers; therefore, those are not homologous with those of species of Metalycaeus, which are raised striae. Furthermore, additional, fresh shells collected at Kalek-Rottung Road clearly lack spiral striation, showing that this is a species of Dicharax. Dicharax magnificus is the type species of Raptomphalus Godwin-Austen, 1914, and as a consequence, that taxon is a synonym of Dicharax (instead of Metalycaeus).	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A37BFF9BBAD4F949B2EDFDD2.taxon	description	Fig. 41 A – E	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A37BFF9BBAD4F949B2EDFDD2.taxon	materials_examined	Type material examined THAILAND • holotype; Khon Kaen Province, Tham Phaya Nakharat; 16 ° 48 ′ 32.8 ″ N, 101 ° 57 ′ 23.9 ″ E; 21 Jul. 2020; P. Tongkerd and A. Pholyotha leg.; CUMZ 7428. For other specimens, see Jirapatrasilp et al. (2021). Additional material examined MYANMAR • 1 shell (Fig. 41 A – E); Shan State, ca 6 km E of Hsihseng centre, right side of rd + 500 m on unpaved rd, limestone hill; 20 ° 8.002 ′ N, 97 ° 18.024 ′ E; 1000 m a. s. l.; 7 Oct. 2018; A. Hunyadi, K. Okubo and J. U. Otani leg.; coll. HA (locality code: 2018 / 41). Type locality “ Tham Phaya Nakharat, Khon Kaen Province, Thailand, 16 ° 48 ’ 32.8 ” N, 101 ° 57 ’ 23.9 ” E ”.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A37BFF9BBAD4F949B2EDFDD2.taxon	diagnosis	Differential diagnosis Differs from the most similar D. verrucosus by the more elevated spire and the longer R 3.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A379FF9BB922FD75B346F8FB.taxon	description	Fig. 42	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A379FF9BB922FD75B346F8FB.taxon	diagnosis	Diagnosis Besides the elevated spire, this species can be distinguished from its congeners based on the nearly smooth R 1, and the peculiar outer peristome, which is strongly expanded and turns anteriorly.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A379FF9BB922FD75B346F8FB.taxon	materials_examined	Type material examined INDIA • 12 syntypes of A. conicus (Fig. 42 A – E); Samiamri, E of the Kopili R.; Godwin-Austen leg.; NHMUK 1903.7. 1.2674 • 12 syntypes of A. conicus var. nanus (Fig. 42 F – J); Jatinga Valley, N. Cachar; NHMUK 1903.7. 1.2675. Additional material examined INDIA • 1 shell (included with type lot, but not mentioned in the original description and not considered as part of the type series); Khasi Hills; NHMUK 1903.7. 1.2565. Type localities “ Was abundant on the Limestone Hill east of Kopili river, North Cachar District, and was occasionally also found in other places, but rare ” (A. conicus); “ Jatinga Valley, North Cachar Hills ” (A. conicus var. nanus).	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A379FF9BB922FD75B346F8FB.taxon	discussion	Remarks Besides a slight difference in shell size, we found no important conchological traits that differ between A. conicus and A. conicus var. nanus; therefore, we see no reason to maintain the latter as a subspecies.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A379FF9DB924F88EB79FFC90.taxon	description	Fig. 43 A – E	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A379FF9DB924F88EB79FFC90.taxon	diagnosis	Diagnosis Differs from all congeners by the deep longitudinal grooves of the R 3.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A379FF9DB924F88EB79FFC90.taxon	materials_examined	Type material examined INDIA • 13 syntypes (Fig. 43 A – E); Phunggum, Lahupa Naga Hills, Munipur; NHMUK 1903.7. 1.2486. Type locality “ Phunggum, a Naga village at head of the Lanier valley, at 5,000 feet ”.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A379FF9DB924F88EB79FFC90.taxon	discussion	Remarks It appears that the outer peristome of this species is situated close to the R 3 swelling. Such extended space between the inner and outer peristomes is visible in other species as well, and most extreme in the cases of D. bicrenatus, D. digitatus, D. rugosus and D. umbiclausus. However, in D. globulus the relatively large area between the two peristomes bears deep longitudinal grooves between the aperture and the R 3 swelling.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A37FFF9DBADEFCB1B527FA33.taxon	description	Fig. 41 F – J	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A37FFF9DBADEFCB1B527FA33.taxon	materials_examined	Type material examined MYANMAR • holotype (Fig. 41 F – J); Shan State, Taunggyi, hill above Aye Say Tee Pagoda, Dragon Cave; 20 ° 47.489 ′ N, 97 ° 03.036 ′ E; 1380 m a. s. l.; A. Hunyadi, K. Okubo and J. U. Otani leg.; HNHM 104862. See other specimens in Páll-Gergely et al. (2021). Type locality “ Myanmar, Shan State, Taunggyi, hill above Aye Say Tee Pagoda, Dragon Cave, 20 ° 47.489 ’ N, 97 ° 03.036 ’ E, 1380 m a. s. l. ”	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A37FFF9DBADEFCB1B527FA33.taxon	diagnosis	Differential diagnosis See under D. burchi and in the original description.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A37FFF9FBAC7FA50B586FDC6.taxon	description	Fig. 43 F – J	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A37FFF9FBAC7FA50B586FDC6.taxon	materials_examined	Type material examined MYANMAR • holotype (single shell mentioned in the original description: Fig. 43 F – J); Moditoung; NHMUK 1906.4. 4.53. Additional material examined MYANMAR • 2 additional non-type specimens in the same lot from Alori Khyoung and Mamya Khyoung; NHMUK 1906.4.4.53. Type locality “ in montibus Arakanensibus ”.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A37FFF9FBAC7FA50B586FDC6.taxon	diagnosis	Differential diagnosis Probably the most closely related species is D. globulus due to the peculiar oblique R 1 ribs. That species, however, has longitudinal grooves on R 3, and a fringed aperture.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A37DFF9FB9BFFADBB774F9AE.taxon	discussion	Remarks This species group is characterized by large, depressed shells and rounded (not fringed) apertures. Species of this species group are distributed from Myanmar (Ayeyarwady and Yangon Regions) to the Naga Hills.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A37DFF9FBAE6FD7AB478FAA6.taxon	description	Fig. 43 K – O	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A37DFF9FBAE6FD7AB478FAA6.taxon	materials_examined	Type material examined MYANMAR • 2 syntypes (Fig. 43 K – O); Baumi, Akyab; NHMUK 1888.12.4.251 - 252. Type locality “ Baumi, Akyab ”.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A37DFF9FBAE6FD7AB478FAA6.taxon	discussion	Remarks Dicharax vestitus akyabensis is smaller and has a thinner peristome than the nominotypical subspecies, which probably indicates intraspecific variability. Nevertheless, we maintain the two subspecies until more material becomes available.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A37DFF85B90CF9D3B230FD0D.taxon	description	Figs 44 – 49	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A37DFF85B90CF9D3B230FD0D.taxon	materials_examined	Type material examined MYANMAR • possible syntypes of A. glaber (Fig. 44 A – E); Akyab; W. T. Blanford coll.; NHMUK 1906.4. 4.181 • 2 shells of (A. glaber); Akyab, Arakan; NHMUK 20191066 • 8 syntypes of A. edei in 2 vials (Fig. 44 F – J); Naraindhur, Cachar; F. Ede leg.; NHMUK 1903.7. 1.1665 • holotype of A. levis (single shell mentioned in the original description: Fig. 44 K – O); Munipur; Godwin-Austen coll.; NHMUK 1903.7. 1.2631. Additional material examined INDIA – Mizoram • 1 empty shell; Blue Mountain, Lawngtlai district; 22.727 ° N, 93.135 ° E; 1400 m a. s. l.; 27 Jan. 2019; N. A. Aravind leg.; Ar 29; NZSI LM 1730 • 1 empty shell; same data as for preceding; Ar 39; NZSI LM 1731 • 1 empty shell; same data as for preceding; Ar 52; NZSI LM 1732 • 1 empty shell; same data as for preceding; Ar 30; NZSI LM 1733 • 1 empty shell; same data as for preceding; Ar 64 (Fig. 48); NZSI LM 1734 • 2 empty shells; same data as for preceding; Ar 54 (Fig. 46); NZSI LM 1735 • 2 empty shells; same data as for preceding; Ar 20 (Fig. 47 B – D [operculum]); NZSI LM 1736 • 1 empty shell (Fig. 45); same data as for preceding; Ar 19; NZSI LM 1737 • 1 empty shell; same data as for preceding; Ar 65; NZSI LM 1738 • 1 empty shell; same data as for preceding; Ar 41 (Fig. 47 A); ZSI LM 1739 • 1 empty shell; same data as for preceding; Ar 28; ZSI LM 1905. Type localities “ Akyab, Arakan; the hills south of the harbour ” (A. glaber); “ Naraindhur, Cachar, No. 1665 B. M. ” [probably Narainpur Village, Rajabazar Tehsil, Cachar District: 24 ° 30.4 ′ N, 92 ° 52.2 ′ E] (A. edei); “ Gaziphimi, Lahupa Naga Hills, Munipur ” [probably Gaziphema, Naga Hills, Manipur: 25 ° 30 ′ N, 94 ° 23.3 ′ E] (A. levis).	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A37DFF85B90CF9D3B230FD0D.taxon	diagnosis	Differential diagnosis There are few species of Dicharax of this size. For comparisons with other species, see under D. magnus and D. ingrami. Additional information on the morphology A cross sectional view of R 2 was examined in a specimen (Fig. 48): anterior crust forms a significantly elevated rib, which has an anterior projection (i. e., the R 2 ribs are curved towards the aperture); the anterior crust folds over the much weaker posterior crust; microtunnel narrow compared to the ribs, its cross sectional view subcircular. A D. glaber shell from the Blue Mountain apparently has short chevrons on the inner surface of its aperture (Fig. 47 A). The tendency towards a fringed aperture appears in several species (see other examples in Páll-Gergely et al. 2017, 2021). Operculum (Fig. 47 B – D): colour chocolate brown, outer surface multispiral, without elevated lamina, inner side with a strongly elevated, knob-like nipple.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A37DFF85B90CF9D3B230FD0D.taxon	distribution	Distribution This species is known from northeastern Myanmar and the neighbouring Indian areas (Fig. 49).	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A37DFF85B90CF9D3B230FD0D.taxon	discussion	Remarks The type specimens of A. glaber, A. edei and A. levis are very similar, and the notable differences (i. e., the more rounded shape of A. glaber and the strongly reflected outer peristome of A. edei) are considered to be intraspecific variability. The shells from the Blue Mountain are identical to the type specimens of A. edei with the exception of the strongly reflected outer peristome (similar cases were reported for D. notus (Godwin-Austen, 1914), see Páll-Gergely et al. 2021: fig. 26).	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A365FF87B91AFEECB321F977.taxon	description	Fig. 50	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A365FF87B91AFEECB321F977.taxon	description	1 a – c; 1914: 421. Alycaeus nagaensis – Godwin-Austen 1884: pl. 51 figs 3, 7; 1886: 195, pl. 44 figs 3, 3 a – c; 1914: 396 – 397, pl. 143 figs 2, 2 a – b. Alycaeus (Chamalycaeus) ingrami – Kobelt 1902: 357. — Gude 1921: 228 – 229. Alycaeus (Chamalycaeus) nagaensis – Kobelt 1902: 359. — Gude 1921: 230. Alycaeus anonymus – Gude 1921: 205. Chamalycaeus (Chamalycaeus) nagaensis – Ramakrishna et al. 2010: 54. Dicharax anonymus – Páll-Gergely et al. 2020: 53. Dicharax (?) ingrami – Páll-Gergely et al. 2020: 90. Dicharax (?) nagaensis – Páll-Gergely et al. 2020: 98.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A365FF87B91AFEECB321F977.taxon	materials_examined	Type material examined MYANMAR • 7 shells, probable syntypes of A. ingrami (figured by Godwin-Austen 1886 and Fig. 50 A – E); Tongoop, Arakan; Blanford coll.; NHMUK 1906.4. 4.68 • 1 shell of A. ingrami; Arakan, “ authentic ”, “ type var. ”, 1944, No. 450; NHMUK • 2 syntypes of A. anonymus (Fig. 50 K – O); Akouktoung, Pegu; NHMUK 1906.4.4.67 a • 7 syntypes of A. nagaensis in 2 vials (Fig. 50 F – J); Asalu, N. Cachar; NHMUK 1903.7. 1.2615. Type localities “ prope Tongoop in Arakan ” (A. ingrami); “ Akouk-toung, Pegu: Type; also Thoudaung and Yenandoung, Pegu ” (A. anonymus); “ Neighbourhood of Asálú, rather local in its distribution, but abundant ” (A. nagaensis).	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A365FF87B91AFEECB321F977.taxon	diagnosis	Differential diagnosis Dicharax glaber has glossy initial whorls and R 3, while those are regularly ribbed in D. ingrami (although the R 3 is glossy in the type specimens of A. nagaensis). No other differences have been found between D. glaber and D. ingrami, and the difference regarding the R 3 sculpture is rather minor, therefore, they may be considered synonyms. However, since we know of no specimens showing intermediate character states, we maintain both species as valid. See also under D. magnus.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A365FF87B91AFEECB321F977.taxon	discussion	Remarks The type specimens of A. anonymus and A. ingrami are very similar, the only differences are that the former has a slightly keeled body whorl and a stronger sculpture. Nevertheless, these differences are considered intraspecific variability. Godwin-Austen (1914), when describing A. anonymus, did not compare it with any other species. The type specimens of A. nagaensis also do not differ from those of A. anonymus and A. ingrami except for their larger size, which is not sufficient for species level distinction. Therefore, both A. nagaensis and A. anonymus are junior synonyms of D. ingrami.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A365FF89B920F908B7C3FDC9.taxon	description	Fig. 51 A – E	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A365FF89B920F908B7C3FDC9.taxon	materials_examined	Type material examined INDIA • 1 syntype (Fig. 51 A – E); Naga Hills; Beddome fr.; NHMUK 1903.7. 1.1480. Type locality “ Naga Hills, 150 miles eastward of Kohima ”.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A365FF89B920F908B7C3FDC9.taxon	diagnosis	Differential diagnosis This species is much larger than D. glaber, has ribbed initial whorls, and the outer peristome partly covers the umbilicus. The same is true when comparing A. magnus with D. ingrami, but both species have ribbed initial whorls.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A36BFF89B9BFF8DDB52DF859.taxon	discussion	Remarks This species group is characterized by a strongly swollen R 3, which has its more elevated posterior end (“ tip ”) close to the constriction between R 2 and R 3.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A36BFF89B902FD7AB2AFF8AC.taxon	description	Fig. 51 F – N	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A36BFF89B902FD7AB2AFF8AC.taxon	diagnosis	Diagnosis This species differs from all other congeners of this species group by the wide umbilicus and less inflated R 2.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A36BFF89B902FD7AB2AFF8AC.taxon	materials_examined	Type material examined MYANMAR • holotype of A. sandowayensis (single shell mentioned in the original description: Fig. 51 F – J); Mai-i, Sandoway Dist., Arakan; Stoliczka leg.; NHMUK 1903.7. 1.2558 • 3 syntypes of A. umbonalis (Fig. 51 K – N); Burmah; UMZC I. 103005. Additional material examined MYANMAR • 5 shells of A. umbonalis; Akoutong, Burma; Blanford coll.; NHMUK. Type localities “ ad Akaouktong, prope ripas fluvii Irawadi ” (A. umbonalis); “ Mai-i, Sandoway District, Arakan ” (A. sandowayensis).	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A36BFF89B902FD7AB2AFF8AC.taxon	discussion	Remarks The holotype of A. sandowayensis is a smaller shell than typical A. umbonalis, but since all shell characters are identical, we treat it as a synonym of the latter species. In the original description of A. sandowayensis, Godwin-Austen (1914) did not compare it with any other species.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A36AFF88B928FEECB5A7FC62.taxon	description	Fig. 52 A – E	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A36AFF88B928FEECB5A7FC62.taxon	description	Alycaeus chennelli var. – Godwin-Austen 1914: 387.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A36AFF88B928FEECB5A7FC62.taxon	materials_examined	Type material examined INDIA • 8 syntypes (Fig. 52 A – E); Piknui, Naga Hills; A. Chennell leg.; NHMUK 1903.7. 1.2612 • 10 shells of “ chennelli var. ”; Lhota Naga Hills; A. Chennell leg.; NHMUK 1903.7. 1.2613. Type localities “ Piknúi, Naga Hills ”; “ Lhota Naga Hills ” (chennelli var.).	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A36AFF88B928FEECB5A7FC62.taxon	diagnosis	Differential diagnosis The only stable character which distinguishes D. chennelli from D. diagonius is the presence of a lower apertural incision in the former, whilst it is absent in the latter.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A36AFF88BAC5FBE6B7E3F8FD.taxon	description	Fig. 53 A – E	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A36AFF88BAC5FBE6B7E3F8FD.taxon	materials_examined	Type material examined BHUTAN • 12 syntypes (Fig. 53 A – E); Damsang, W. Bhutan; Robert leg.; NHMUK 1903.7. 1.2677. Type locality “ Damsang Peak, Western Bhutan Hills ”.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A36AFF88BAC5FBE6B7E3F8FD.taxon	diagnosis	Differential diagnosis The most similar species is D. notatus, which has a more inflated R 3 and denser R 1 ribs. Dicharax nowgongensis (Godwin-Austen, 1914) is also similar, but its basal lip is rounded.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A36AFF88BAC5FBE6B7E3F8FD.taxon	discussion	Remarks The specimens figured by Gittenberger et al. (2024) from Bhutan differ from the type specimens in the denser R 1 ribbing and the ribbed R 3 (vs smooth in D. damsangensis). Therefore, they likely represent an undescribed species.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A36AFF8DB92FF88DB739FCE2.taxon	description	Fig. 52 F – J	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A36AFF8DB92FF88DB739FCE2.taxon	materials_examined	Type material examined INDIA • 10 syntypes (Fig. 52 F – J); Diyung Valley, N of Asalu, N. Cachar; Godwin-Austen coll.; NHMUK 1903.7. 1.2678. Type locality “ The Diyung valley, north of Asálú, in Cachar District ”.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A36AFF8DB92FF88DB739FCE2.taxon	diagnosis	Differential diagnosis This species has a longer and more pointed R 3 swelling and more widely-spaced R 1 than Dicharax subculmen (Godwin-Austen, 1893). See also under D. chennelli and D. habiangensis (Godwin-Austen, 1914).	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A36FFF8DBAC0FC65B2B9F86A.taxon	description	Fig. 54	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A36FFF8DBAC0FC65B2B9F86A.taxon	materials_examined	Type material examined INDIA • holotype (single shell mentioned in the original description: Fig. 54 A – E) of A. habiangensis; Habiang Garo, W. Khasi; Godwin-Austen leg.; NHMUK 1903.7. 1.2649 • 2 syntypes of A. generosus (Fig. 54 F – J); Khasi Hills, Godwin-Austen coll.; NHMUK 1903.7. 1.2566. Type localities “ Habiang Garo, on the West Khasi border ” (A. habiangensis); “ Khasi Hills ” (A. generosus).	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A36FFF8DBAC0FC65B2B9F86A.taxon	diagnosis	Differential diagnosis Both D. subculmen and D. diagonius have a longer R 2, a ribbed R 1, and a less bulging R 3. See also under D. nitidus.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A36FFF8DBAC0FC65B2B9F86A.taxon	discussion	Remarks The only difference between the two syntypes of A. generosus and the single, subadult syntype of A. habiangensis is the very slightly shallower R 3 dent of the latter, suggesting that they represent the same species. The original description of A. habiangensis immediately followed that of A. generosus, and in the face of the nearly identical shells, they were not compared with each other; except for the umbilical view, they were not illustrated from the same views. We selected A. habiangensis to be the accepted name because it is known from a more precise locality than A. generosus.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A36DFF8FB960FEECB74DFB78.taxon	description	Fig. 53 F – J	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A36DFF8FB960FEECB74DFB78.taxon	description	1914: 374 – 375, pl. 145 figs 5, 5 a – b. Alycaeus (Dicharax) hebes – Kobelt 1902: 371. — Gude 1921: 255. Chamalycaeus (Dicharax) hebes – Ramakrishna et al. 2010: 61. Dicharax hebes – Páll-Gergely et al. 2017: 10, fig. 5; 2020: 64, fig. 13 a; 2024, fig. 12 d. — Preece et al. 2022: 80, fig. 31 c – d.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A36DFF8FB960FEECB74DFB78.taxon	diagnosis	Diagnosis The relatively large shell, the nearly smooth R 1, the formation of the R 3 swelling and the rounded aperture distinguishes this species from its congeners.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A36DFF8FB960FEECB74DFB78.taxon	materials_examined	Type material examined INDIA • neotype (designated by Preece et al. 2022: Fig. 53 F – J); Teria Ghat is in the Khasi Hills; Theobald leg.; NHMUK 1888.12. 4.909. Additional material examined INDIA • 3 shells; Khasi Berge; Möllendorff coll.; SMF 109244 • 1 shell; Nongphrian, Khasi Hills; Dr Alfred Oberwimmer coll.; NHMW (mixed sample with D. theobaldi) • 1 shell; same data as for preceding; NHMW 14715 (mixed sample with D. theobaldi). Type locality “ ad Teria Ghát ”.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A36DFFB1B913FB09B7AEFD12.taxon	description	Fig. 55	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A36DFFB1B913FB09B7AEFD12.taxon	description	Alycaeus nitidus – Sowerby 1877: pl. 3, species 25. — Godwin-Austen 1914: 421 – 422, pl. 151 figs 4, 4 a.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A36DFFB1B913FB09B7AEFD12.taxon	materials_examined	Type material examined MYANMAR • 3 possible syntypes of A. nitidus (Fig. 55 A – E); Manya Khyoung, Arakan; Blanford coll.; NHMUK 1906.4. 4.54 • 3 probable syntypes of A. kurzianus (labelled as “ kurtzianus ”; the locality does not match with the original description, but the specimens are identical to the figured one) (Fig. 55 F – J); Nr. Prome, Pegu; F. Stoliczka leg.; NHMUK 1903.7. 1.2700 • 1 syntype of A. kurzianus; Nattoung, Prome; W. Theobald coll.; NZSI M. 24974 • 6 syntypes of A. kurzianus; same data as for preceding; NZSI M. 25021. Type localities “ prope Tongoop in Arakan ” (A. nitidus); “ Nattoung in provincia Barmana, Prome dicta ” (A. kurzianus).	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A36DFFB1B913FB09B7AEFD12.taxon	diagnosis	Differential diagnosis The most similar species due to the tiny, glossy shell and the short R 2 is D. habiangensis, which has a larger R 3 swelling and a basally rounded aperture. See also under D. semivivus sp. nov.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A36DFFB1B913FB09B7AEFD12.taxon	discussion	Remarks Dicharax nitidus and D. kurzianus agree in all important traits (shell and aperture shape, shell size, length of shell regions, sculpture), and no notable difference was found between them. Therefore, D. kurzianus is a junior synonym of D. nitidus. The two species were not compared in the original description of D. kurzianus. Dicharax nitidus was described from Tongoop (Taungup Pass at 18 ° 40.1 ′ N, 94 ° 44.5 ′ E), whereas D. kurzianus from Nattoung, which could not be located on the map. According to the additional information about Nattung, it is in “ Mendon District, Pegu ”, and close to Prome (= Pyay). A mountain with the name “ Nattaung ” is located at 18 ° 48.6 ′ N, 97 ° 2.2 ′ E, ca 190 km east from Pyay, therefore it is probably not the same as Nattung. Nevertheless, the Taungup Pass and Pyay are situated geographically close to each other.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A353FFB1B91DFD37B4D2FA09.taxon	description	Fig. 53 K – O	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A353FFB1B91DFD37B4D2FA09.taxon	diagnosis	Diagnosis The elevated spire, relatively dense R 1 ribs and the strongly inflated R 3 swelling distinguishes this species from its congeners. See under D. damsangensis.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A353FFB1B91DFD37B4D2FA09.taxon	materials_examined	Type material examined INDIA • 4 syntypes (Fig. 53 K – O); Toruputu Peak, Dafla Hills; elev. 3000 ft; NHMUK 1903.7. 1.2672 • 2 syntypes; Dafla Hills; Godwin-Austen coll.; NHMUK 1903.7. 1.2544. Both samples are in the same box but in different vials. Type locality “ On the slopes of Torúpútú Peak at 3000 feet ”.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A353FFB0BAC5FA3AB5A7FAE5.taxon	description	Fig. 56	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A353FFB0BAC5FA3AB5A7FAE5.taxon	materials_examined	Type material examined INDIA • holotype (single shell mentioned in the original description: Fig. 56 A – G); Koliaghur nr Tezpur, Assam; NHMUK 1903.7. 1.2682. Type locality “ Koliaghur or Koliahur, Nowgoug District, Assam ”.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A353FFB0BAC5FA3AB5A7FAE5.taxon	diagnosis	Differential diagnosis Similar to D. chennelli, D. damsangensis, D. diagonius and D. vitreus, which have similar R 1 sculptures. Unfortunately, the holotype specimen is broken at its R 3, making it difficult to examine some key characters. Nevertheless, D. nowgongensis has a rounded aperture and a relatively low-spired shell, which differs from most congeners of the same species group.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A352FFB2BACFFA65B799F9D9.taxon	description	Figs 12 B – C, 57	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A352FFB2BACFFA65B799F9D9.taxon	diagnosis	Diagnosis A small species of Dicharax with glossy R 1 and R 3, a combined length of R 2 + R 3 reaching nearly half a whorl, and a strongly inflated R 3 bearing a single dent near the suture on the upper part of the R 3 swelling.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A352FFB2BACFFA65B799F9D9.taxon	etymology	Etymology The specific epithet ‘ praeda ’ (Latin for ‘ prey’) refers to the dent on R 3 resembling a bite trace. To be used as a noun in apposition.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A352FFB2BACFFA65B799F9D9.taxon	materials_examined	Type material examined Holotype INDIA – Mizoram • empty shell (D: 3 mm, H: 2.4 mm, Figs 12 B – C, 57); Blue Mountain, Lawngtlai district; 22.727 ° N, 93.135 ° E; 1400 m a. s. l.; 27 Jan. 2019; N. A. Aravind leg.; Ar 67; NZSI LM 1740. Paratype INDIA – Mizoram • 1 empty shell; Blue Mountain, Lawngtlai district; 22.727 ° N, 93.135 ° E; 1400 m a. s. l.; 27 Jan. 2019; N. A. Aravind leg.; Ar 67; NZSI LM 1741.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A352FFB2BACFFA65B799F9D9.taxon	description	Description Shell small, yellowish, glossy, semitranslucent; shell outline rather rounded in dorsal view except for the conspicuously inflated R 3; spire elevated, conical; body whorl rounded; protoconch low, glossy, consisting of ca 1.5 whorls; R 1 of 1.75 – 2 whorls, glossy, smooth, with some inconspicuous, widely-spaced, incomplete (i. e., present only near the suture) ribs on the last ca 0.75 whorl of R 1; spiral striation absent; boundary between R 1 and R 2 conspicuous due to the sudden change to a regularly ribbed surface; R 2 with ca 22 – 26 straight, sharp, lamella-like ribs; R 2 ribs slightly curved towards aperture, the spaces between the ribs are wider than the width of a single rib; R 2 + R 3 nearly half whorl; R 3 slightly longer than R 2; boundary between R 2 and R 3 clearly visible due to a deep constriction; entire R 3 strongly swollen, forming a blunt tip towards constriction; a shallow, inconspicuous, elongated dent is situated near the upper end of the swelling, nearly touching the suture; aperture strongly oblique to shell axis, rounded with a slight upper (parieto-palatal) incision; boundary between inner and outer peristomes conspicuous; inner peristome slightly thickened and protruding; outer peristome more conspicuous, moderately sharp, strongly expanded, especially on the upper palatal and basal side; umbilicus very narrow, slit-like. MEASUREMENTS. D: 2.9 – 3 mm, H: 2.4 mm.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A352FFB2BACFFA65B799F9D9.taxon	distribution	Distribution Known only from the Blue Mountain region.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A352FFB2BACFFA65B799F9D9.taxon	diagnosis	Differential diagnosis The most similar species is Dicharax diagonius, which has complete R 1 ribs on the entire region (absent on the first 1.5 whorl, and on the last ca 1 whorl present only near the suture in the new species), more curved R 2 ribs, and a more pointed tip of the R 3 swelling. Dicharax chennelli differs from the new species in the following characteristics: the ribs near the suture of R 1 are stronger in D. chennelli, and the R 2 ribs are more curved towards the aperture, forming an almost continuous flat area, while the R 2 ribs of the new species are rather straight, and therefore they do not form a flat surface. The dorsal side of the R 3 swelling of D. chennelli bears two dents, one situated on the upper part, and the lower at its middle, both being the same distance from the suture. In contrast, the upper dent nearly touches the suture in the new species, and the lower dent is not visible. Furthermore, the tip of the swelling (i. e., the part situated closest to the constriction) is sharper in D. chennelli, and blunter in the new species. Lastly, D. chennelli has a basal incision, while the aperture of the new species is perfectly rounded. Dicharax habiangensis has a shorter R 2 and R 3, the R 2 ribs are more curved, the dent on the R 3 is situated at some distance from the suture (touching the suture in the new species), and the entire R 3 is separated from the penultimate whorl by the deeper suture. Dicharax subculmen has a lower spire, denser ribbing on R 1, and a deeper dent on the R 3 swelling. Dicharax nowgongensis has stronger and more complete R 1 ribs, but the R 3 of the holotype of that species was broken.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A350FFB5B936F969B76DFDEA.taxon	description	Fig. 52 K – O	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A350FFB5B936F969B76DFDEA.taxon	diagnosis	Diagnosis The small shell, the rather weak R 1 ribs, and the relatively less swollen R 3 swelling distinguish this species from its congeners. See also under D. habiangensis and Dicharax praeda sp. nov.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A350FFB5B936F969B76DFDEA.taxon	materials_examined	Type material examined INDIA • 3 syntypes (Fig. 52 K – O); Naga Hills; Doherty leg.; NHMUK 1903.7. 1.2687. Type locality “ Naga Hills ”.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A357FFB5BAE9FD9FB5A7FBFA.taxon	description	Fig. 58	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A357FFB5BAE9FD9FB5A7FBFA.taxon	materials_examined	Type locality “ Dagana Dzongkhag: [302] Lhamoizingkha, Pubji near Quarry, ca 250 m a. s. l., 26 º 45 ’ N 89 º 48 ’ E ” (Bhutan).	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A357FFB5BAE9FD9FB5A7FBFA.taxon	diagnosis	Differential diagnosis Similar to D. chennelli and D. diagonius, but the R 2 ribs are seemingly more widely-spaced (although the photos in the original description of D. vitreus are of low quality). Moreover, D. vitreus lacks the basal incision in the aperture, which is present in D. chennelli.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A356FFB4B92CFE39B43EF80F.taxon	description	Fig. 59	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A356FFB4B92CFE39B43EF80F.taxon	diagnosis	Diagnosis This species is characterized by strong, widely-spaced R 1 ribs, a short R 2 + R 3, a smooth R 2, two low R 3 swellings and a slightly fringed peristome.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A356FFB4B92CFE39B43EF80F.taxon	materials_examined	Type material examined INDIA • holotype of A. akhaensis (single shell mentioned in the original description: Fig. 59 A – E); Akha Hills, Barowli River; Godwin-Austen coll.; NHMUK 1903.7. 1.2683 • 1 syntype of A. burroiensis (Fig. 59 F – J); Burroi Rr., Dafla; NHMUK 1903.7. 1.2653 • 3 corroded syntypes of A. gemma in two vials (Fig. 59 K – O); No. 7. camp, Dikrang Valley, Dafla; NHMUK 1903.7. 1.2601. Type localities “ Barowli Gorge, Durrang District, Assam, foot of the Akha Hills ” (A. akhaensis); “ Burroi Gorge, Dafla Hills ” (A. burroiensis); “ No. 7 Camp, Dikrang Valley, Dafla Hills ” (A. gemma).	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A356FFB4B92CFE39B43EF80F.taxon	diagnosis	Differential diagnosis Dicharax lahupaensis (Godwin-Austen, 1914) has a ribbed R 2, and its aperture is less fringed.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A356FFB4B92CFE39B43EF80F.taxon	discussion	Remarks Both A. akhaensis and A. burroiensis are represented in the NHM by single, corroded specimens. Acknowledging that the important characters (mostly the fine sculpture and the formation of R 3) could not be very clearly seen on the shells, we could not find any conchological differences that would justify their distinction. Therefore, we treat A. burroiensis as a junior synonym of Dicharax akhaensis. Alycaeus gemma also agrees in all important shell characters (ratios of shell regions, R 3 swellings, fringed aperture), but its shell has a rather oval outline in dorsal view. Nevertheless, this is also considered part of intraspecific variability, and therefore, A. gemma is also treated as a junior synonym of A. akhaensis. See also under D. khasiacus.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A356FFB4B9BFFEECB495FE08.taxon	diagnosis	Diagnosis This species group is characterized by mostly small (smaller than 3 mm), mostly glossy species. The aperture is not fringed or slightly fringed.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A354FFB9B92FFEECB474FA87.taxon	description	Fig. 60	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A354FFB9B92FFEECB474FA87.taxon	diagnosis	Diagnosis The characteristic, oblique main R 3 swelling, and the small secondary swelling distinguish this species from its congeners. See under Dicharax umashaankeri sp. nov.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A354FFB9B92FFEECB474FA87.taxon	materials_examined	Type material examined INDIA • 2 syntypes of A. birugosus (Fig. 60 A – E); Khasi Hills; Godwin-Austen leg.; NHMUK 1903.7.1.2628 • 1 syntype of Alycaeus birugosus var. minor (labelled as duorugosus var. minor); Garo Hills; NHMUK 1903.7.1.2755 • 1 syntype of A. canaliculus (Fig. 60 F – J); Teria Ghat, Khasi; Godwin-Austen leg.; NHMUK 1903.7.1.2764. Additional material examined INDIA • 7 shells (labelled as “ birugosus var. ”); Jawai, Jiantia Hills, 282 a; Godwin-Austen leg., NHMUK 1903.7.1.2571. Type localities “ Khasi Hills and Manipur ” (A. birugosus); “ Garo Hills ” (A. birugosus var. minor); “ Teria Ghat, foot of the Khasi Hills ” (A. canaliculus).	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A35BFFBDB92CFA84B4B4FE33.taxon	description	Fig. 61 A – E	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A35BFFBDB92CFA84B4B4FE33.taxon	materials_examined	Type material examined INDIA • lectotype (designated by Páll-Gergely et al. 2020) (Fig. 61 A – E); Toruputu Peak, Dafla Hills; Godwin-Austen coll.; NHMUK 1903.7. 1.2497.1 • 6 paralectotypes; same data as for preceding; NHMUK 1903.7.1.2497.2 - 7. Type locality “ Torúpútú Peak, 7000 feet ”.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A35BFFBDB92CFA84B4B4FE33.taxon	diagnosis	Differential diagnosis Within this species group, D. gemmula is the most similar due to the similarly shaped R 3 swelling, the smooth R 2 and the overall smooth shell surface. Nevertheless, the aperture of D. gemmula is not fringed. Dicharax lahupaensis is also similar, but it has elevated R 2 ribs (smooth in D. daflaensis), and its R 3 swelling is situated at a greater distance from the aperture. Dicharax pachitaensis (Godwin-Austen, 1886) has a less elevated R 3 swelling, stronger R 1 ribs, more rounded aperture, and a keeled umbilical part of R 2. See also under D. crenatus.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A35FFFBDB934FE57B739FB05.taxon	description	Fig. 62 A – E	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A35FFFBDB934FE57B739FB05.taxon	materials_examined	Type material examined INDIA • 7 syntypes (Fig. 62 A – E); Rechila Pk, Sikkim; W. Robert leg.; NHMUK 1903.7. 1.1251. Type locality “ Rechila Peak, Daling District, on Sikkim and Bhutan Boundary (10,300 ft.) ”.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A35FFFBDB934FE57B739FB05.taxon	diagnosis	Differential diagnosis The most similar species are D. lenticulus (Godwin-Austen, 1874), D. nongtungensis (Godwin-Austen, 1914) and D. politus (W. T. Blanford, 1865), due to the smooth shell surface and the long, inflated R 3 swelling. Dicharax lenticulus is larger and has a more depressed shell. Dicharax nongtungensis is smaller, more depressed, and has a more strongly expanded outer peristome. Lastly, D. politus is much more depressed, its R 2 ribs are separated (R 2 surface is smooth in D. dalingensis), and has a more inflated R 3 swelling. See also under D. ganeshaiahi sp. nov. and D. nattoungensis (Godwin-Austen, 1914).	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A35FFFA1BAEFFB05B4FFFA29.taxon	description	Figs 64, 65 A – C, 66 A – B	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A35FFFA1BAEFFB05B4FFFA29.taxon	diagnosis	Diagnosis A small species of Dicharax with a glossy shell, smooth, short R 2 (combined length of R 2 + R 3 ca quarter whorl, R 3 ca twice as long as R 2), an R 3 with a central, blunt swelling, a rounded aperture with shallow upper and lower incisions, and a strongly expanded outer peristome.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A35FFFA1BAEFFB05B4FFFA29.taxon	etymology	Etymology This species is named after Prof. K. N. Ganeshaiah, Founder, Trustee of Ashoka Trust for Research in Ecology and the Environment (ATREE), Bangalore, India. He has contributed immensely to the understanding of the ecology, evolution, and conservation of Indian flora and fauna. He is also a popular science communicator in the vernacular language and has published over 32 books in the Kannada language.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A35FFFA1BAEFFB05B4FFFA29.taxon	materials_examined	Type material examined Holotype INDIA – Mizoram • empty shell (D: 3.2 mm, H: 2.3 mm, Fig. 64); Blue Mountain, Lawngtlai district; 22.727 ° N, 93.135 ° E; 1400 m a. s. l.; 27 Jan. 2019; N. A. Aravind leg.; Ar 62; NZSI LM 1755. Paratypes INDIA – Mizoram • 8 empty shells; Blue Mountain, Lawngtlai district; 22.727 ° N, 93.135 ° E; 1400 m a. s. l.; 27 Jan. 2019; N. A. Aravind leg.; Ar 62; NZSI LM 1756 • 2 empty shells; same data as for preceding; Ar 47; NZSI LM 1757 • 1 empty shell (Fig. 65 A – C, 66 A – B); same data as for preceding; Ar 27; NZSI LM 1758 • 6 empty shells; same data as for preceding; Ar 72; NZSI LM 1759. Additional material examined INDIA – Mizoram • 1 corroded empty shell; same data as for holotype; Ar 47; NZSI LM 1760 • 6 corroded empty shells; same data as for holotype; Ar 72; NZSI LM 1761.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A35FFFA1BAEFFB05B4FFFA29.taxon	description	Description Shell small, yellowish or pinkish when fresh, semitranslucent; entire shell surface glossy; shell outline rounded in dorsal view, R 2 and end of R 3 conspicuously swollen, although not in all specimens; spire elevated, conical; body whorl rounded; protoconch low, smooth, consisting of 1.25 – 1.5 whorls; R 1 of ca 1.5 – 1.75 whorls, smooth, glossy, last ca quarter whorl of some specimens with weak, widely-spaced ribs, while in other specimens entire R 3 remains smooth; some weak ribs of the last ca quarter whorl visible inside umbilicus; boundary between R 1 and R 2 not conspicuous, in the case of specimens with ribbed end of R 1 the change between the two regions is more clearly visible due to the large increase in rib density; R 2 without elevated ribs, but very densely arranged lighter and darker alternating stripes (more than 40 in number); cross sectional view of R 2 was examined in a specimen (Fig. 66 C – D): anterior crust forms no rib, it is entirely flat; instead of folding over the posterior crust, the posterior crust covers the larger part of the microtunnels, and the anterior crust only covers the edge of the posterior crust; microtunnel wide compared to anterior and posterior crusts, its cross sectional view teardrop-shaped; R 2 + R 3 ca 90 degrees; R 3 ca twice as long as R 2; boundary between R 2 and R 3 clearly visible due to a shallow constriction; R 3 with a low, central, triangular swelling; aperture strongly oblique to shell axis, rounded with a shallow upper (parieto-palatal), and a shallow lower (basal) incision; inner and outer peristomes clearly separated (mostly at the parietal, palatal and columellar part and less so at the basal part); inner peristome slightly protruding; outer peristome strongly expanded and reflected; umbilicus ca one fourth of shell width, almost rounded, although its part facing R 2 is nearly straight, forming a slight keel on the umbilical side of R 2. OPERCULUM. Inner side smooth, without central nipple; outer side smooth, without any sculpture (Ar 27). MEASUREMENTS. D: 2.8 – 3.4 mm, H: 2.1 – 2.4 mm.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A35FFFA1BAEFFB05B4FFFA29.taxon	diagnosis	Differential diagnosis Dicharax dalingensis (Fig. 62 A – E) is similar in having a smooth R 2, but has longer R 2 and R 3, and a higher spire. Dicharax lenticulus (Fig. 63) also has a smooth R 2, but is much larger, and has a longer R 2 and R 3. Moreover, both D. dalingensis and D. lenticulus have an incision between the penultimate whorl and peristome, while the peristome is attached to the penultimate whorl in the new species. Dicharax pachitaensis (Fig. 61 K – O), which is probably the most similar species, also has a smooth R 2, but the combined length of R 2 and R 3 is usually (not always) longer than that of the new species. Moreover, D. pachitaensis has prominent ribs near the suture on the dorsal side, the R 3 swelling is situated a greater distance from the peristome, and its aperture has deeper upper and lower incisions.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A35FFFA1BAEFFB05B4FFFA29.taxon	discussion	Remarks The cross-sectional view of R 2 of two species of Alycaeinae with smooth R 2 is known: Dicharax moellendorffi (Kobelt & Möllendorff, 1886) (see Páll-Gergely et al. 2017) and Metalycaeus minatoi Páll-Gergely, 2017 (see Páll-Gergely & Asami 2017). Even though the R 2 surfaces are similar, the fine structure of the R 2 (i. e., how the anterior and posterior crusts fold over each other to make the microtunnels) is very different. In the case of D. moellendorffi, the anterior crust has a strong posterior projection that covers the anterior crust. In the case of Metalycaeus minatoi, the anterior and posterior crusts meet just above the microtunnel, and in the new species the posterior crust has an elongated anterior projection covering the microtunnel.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A343FFA3B97AFA59B4A3FC5A.taxon	description	Fig. 67	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A343FFA3B97AFA59B4A3FC5A.taxon	description	Alycaeus (Charax) subhumilis – Godwin-Austen 1914: 344 – 345, pl. 133 figs 2 – 2 c.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A343FFA3B97AFA59B4A3FC5A.taxon	diagnosis	Diagnosis This species is characterized by a tiny, glossy shell, smooth R 2 and a single, sharp R 3 swelling, R 2 + R 3 only slightly longer than a quarter whorl.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A343FFA3B97AFA59B4A3FC5A.taxon	materials_examined	Type material examined INDIA • neotype (designated by Preece et al. 2022: Fig. 67 A – E); Darjiling; Blanford coll.; NHMUK 1906.4. 4.55 • lectotype of A. subhumilis (designated by Zilch 1957: Fig. 67 F – I); Vorderindien: Darjiling; Möllendorff coll.; SMF 109224 • 1 paralectotype of A. subhumilis; same data as for preceding; SMF 109225. Type localities “ in valle Rungun ” (A. gemmula); “ in montibus Darjiling Indiae ” (A. subhumilis).	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A343FFA3B97AFA59B4A3FC5A.taxon	diagnosis	Differential diagnosis Dicharax lahupaensis is similar in general shell and aperture shape, but it has two R 3 swellings, and its R 2 has separate ribs (R 3 surface is smooth in D. gemmula). See also under D. daflaensis.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A343FFA3B97AFA59B4A3FC5A.taxon	discussion	Remarks No notable differences were found between the types of A. gemmula and A. subhumilis, therefore the latter is a junior synonym of the former.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A341FFA5B92CFBEFB5E1FDB9.taxon	description	Fig. 68	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A341FFA5B92CFBEFB5E1FDB9.taxon	diagnosis	Diagnosis Dicharax khasiacus is characterized by the rounded outline, the short, smooth R 2, and the two, anteriorly situated R 3 swellings.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A341FFA5B92CFBEFB5E1FDB9.taxon	materials_examined	Type material examined INDIA • 29 syntypes of A. khasiacus (Fig. 68 A – E); Lailangkote, Khasi Hills; NHMUK 1903.7. 1.2650 • 1 syntype of A. duorugosus (Fig. 68 F – J); Burrail; Godwin-Austen coll.; NHMUK 1903.7. 1.2771. Additional material examined INDIA • 4 shells; Jawai, Assam; Oberwimmer coll.; NHMW • 2 shells; same data as for preceding; NHMW 14711 • 4 shells; Jawai, India; Dr. Leo P. Rušnov coll., ex coll. Dr W. Blume ex coll. W. F. Webb; NHMW 15168. Type localities “ On the highest parts of the Khasi and Jiantia Hills ” (A. khasiacus); “ Burrail Range, Naga ” “ Also Angaoluo Trigonometrical Station, No. 2572; South Barak, No. 2629, and Munipur, No. 2654 B. M. ” (A. duorugosus).	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A341FFA5B92CFBEFB5E1FDB9.taxon	diagnosis	Differential diagnosis Dicharax lahupaensis has a ribbed R 2 (smooth in D. khasiacus) and a thicker peristome. Dicharax akhaensis is larger, has a shorter R 3 swelling, and a less fringed aperture.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A341FFA5B92CFBEFB5E1FDB9.taxon	discussion	Remarks The single examined syntype of A. duorugosus fits the morphological variability of the 29 syntypes of A. khasiacus; therefore the former is a junior synonym of the latter.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A347FFA5B938FDCAB48CFB16.taxon	description	Fig. 61 F – J	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A347FFA5B938FDCAB48CFB16.taxon	diagnosis	Diagnosis This species is characterized by a ribbed R 2, two low R 3 swellings, a fringed aperture and a strongly expanded outer peristome. See also under D. akhaensis, D. daflaensis, D. gemmula and D. khasiacus.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A347FFA5B938FDCAB48CFB16.taxon	materials_examined	Type material examined INDIA • 10 syntypes (Fig. 61 F – J); Gaziphimih, N. E. Munipur; NHMUK 1903.7. 1.2655. Type locality “ Gaziphimi, Lahupa Naga Hills, Munipur ”.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A347FFA4B92FFB2BB2B0FE05.taxon	description	Fig. 63	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A347FFA4B92FFB2BB2B0FE05.taxon	materials_examined	Type material examined INDIA • holotype (single shell mentioned in the original description: Fig 63 F – J); Darjeeling; Godwin-Austen coll.; NZSI M. 8075 • several shells (Fig. 63 A – E); Rissom Pk. (W. Bhutan); Godwin-Austen Coll.; NHMUK 1903.07. 01.2552. Type locality “ Darjeeling ”.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A347FFA4B92FFB2BB2B0FE05.taxon	diagnosis	Differential diagnosis Dicharax nongtungensis is smaller, has a thicker peristome and a narrower umbilicus. Dicharax politus has a more depressed shell and a more strongly inflated R 3 swelling. See also under D. dalingensis, D. ganeshaiahi sp. nov., and D. nattoungensis.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A347FFA4B92FFB2BB2B0FE05.taxon	discussion	Remarks This species was described based on a strongly corroded shell (Godwin-Austen 1874). Later, Godwin-Austen (1914) provided a more complete description and drawings based on multiple additional specimens. We received photos of the holotype (Fig. 63 F – J) from the Zoological Survey of India, and could confirm that the figured shell (Fig. 63 A – E) is conspecific with the holotype.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A346FFA4BAC0FE06B254FA88.taxon	description	Fig. 69 A – E	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A346FFA4BAC0FE06B254FA88.taxon	materials_examined	Type material examined INDIA • 2 syntypes (Fig. 69 A – E); head of Lanier River NE of Munipur; Godwin-Austen coll.; NHMUK 1903.7. 1.2485. Type locality “ Hills at head of the Lanier River, Naga Hills, ca. 5 – 6,000 feet ”.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A346FFA4BAC0FE06B254FA88.taxon	discussion	Remarks An easily recognisable species based on the three R 3 swellings.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A346FFA4BAC0FE06B254FA88.taxon	diagnosis	Differential diagnosis Due to the complicated R 3 swelling, this species is similar to some species of the Dicharax species group 7, but differs from most of those species by the more depressed shell, the rounded aperture, and the R 3 swelling separated into 3 parts. See also under D. umashaankeri sp. nov.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A346FFA6BAC5FAB8B58EFEF8.taxon	description	Fig. 70 A – E	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A346FFA6BAC5FAB8B58EFEF8.taxon	materials_examined	Type material examined MYANMAR • holotype (single shell mentioned in the original description: Fig. 70 A – E); Nattoung Hills, Mendon District, Pegu, Burma; Theobald coll.; NZSI M. 8036. Type locality “ Nattoung Hills ”.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A346FFA6BAC5FAB8B58EFEF8.taxon	diagnosis	Differential diagnosis Dicharax dalingensis, D. lenticulus and D. politus have a more inflated R 3 swelling preceded by a deeper constriction between R 2 and R 3. Dicharax nongtungensis has a more oval outline in dorsal view and a more expanded outer peristome (thicker peristome). Dicharax pusillus is similar in general shell shape, but is much smaller and has a more strongly expanded peristome.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A34BFFA9BAC9FEECB762FB51.taxon	description	Fig. 70 F – J	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A34BFFA9BAC9FEECB762FB51.taxon	materials_examined	Type material examined INDIA • 21 syntypes (Fig. 70 F – J); Nongtung; Godwin-Austen coll.; NHMUK 1903.7. 1.2692. Additional material examined INDIA • 2 shells; Nontung; Dr Leo P. Rušnov coll., ex coll. Dr. W. Blume, ex coll. W. F. Webb; NHMW 15144 • 4 shells; Nongtung, Burmah; NHMW 14890. Type locality “ Nongtung, Jaintia Hills ”.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A34BFFA9BAC9FEECB762FB51.taxon	diagnosis	Differential diagnosis Dicharax politus is more depressed, has a deeper constriction and a more inflated R 3 swelling. Dicharax pusillus is smaller, has a shorter R 2 and a less expanded outer peristome. See also under D. dalingensis, D. lenticulus and D. nattoungensis.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A34BFFA9BAC9FEECB762FB51.taxon	discussion	Remarks The specimen form Bhutan is indeed similar to the syntypes of D. nongtungensis, but has a less inflated R 3 swelling.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A34BFFA9B911FAF2B7ECF869.taxon	description	Fig. 71 A – E	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A34BFFA9B911FAF2B7ECF869.taxon	diagnosis	Diagnosis Dicharax notus and D. omissus are relatively widespread and variable species in northern Thailand and neighbouring central Myanmar. Regarding their conchological variability and identification, see Páll-Gergely et al. (2021).	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A34BFFA9B911FAF2B7ECF869.taxon	materials_examined	Type material examined MYANMAR • 15 syntypes (Fig. 71 A – E); Fort Stedman, Burma; Woodthorpe coll.; NHMUK 1903.7. 1.3065. See also Páll-Gergely et al. (2021). Type locality “ Fort Stedman, Burma ”.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A34AFFABB920FEECB435FDEF.taxon	description	Fig. 71 F – J	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A34AFFABB920FEECB435FDEF.taxon	diagnosis	Diagnosis See under D. notus and Páll-Gergely et al. (2021).	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A34AFFABB920FEECB435FDEF.taxon	materials_examined	Type material examined THAILAND / MYANMAR • 2 syntypes (Fig. 71 F – J); Siam and Shan boundary; Woodthorpe coll.; NHMUK 1903.7. 1.1228. See also Páll-Gergely et al. (2021). Type locality “ Siam and Shan boundary ”.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A349FFABB93FFD93B2E3FAED.taxon	description	Fig. 61 K – O	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A349FFABB93FFD93B2E3FAED.taxon	diagnosis	Diagnosis This species is characterized by a smooth shell (with the exception of widely-spaced R 1 ribs along the suture), a smooth R 2, two low R 3 swellings, a rounded aperture with upper and lower incisions, and a keeled umbilical part of R 2. See also under D. daflaensis and D. ganeshaiahi sp. nov.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A349FFABB93FFD93B2E3FAED.taxon	materials_examined	Type material examined INDIA • 3 syntypes (Fig. 61 K – O); Pachita village, Dafla Hills; Godwin-Austen coll.; NHMUK 1903.7. 1.2614. Type locality “ Pachita village (Camp no. 7 of the Expeditionary Force, 1874 – 75), Dafla Hills, Assam ”.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A349FFAAB912FA9DB498FEEF.taxon	description	Fig. 62 K – O	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A349FFAAB912FA9DB498FEEF.taxon	diagnosis	Diagnosis This species is characterized by a glossy, depressed shell, a finely ribbed R 2, a strongly inflated, long R 3 swelling, and a deep constriction between R 2 and R 3. See also under D. dalingensis, D. lenticulus, D. nattoungensis and D. nongtungensis.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A349FFAAB912FA9DB498FEEF.taxon	materials_examined	Type material examined MYANMAR • 3 probable syntypes (Fig. 62 K – O); Phungdo, Arakan; Blanford coll.; NHMUK 1906.4. 4.178. Type locality “ Phuong do, near Cape Negrais, Arakan ”.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A348FFAAB923FE9CB2DAFBAB.taxon	description	Fig. 70 K – O	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A348FFAAB923FE9CB2DAFBAB.taxon	diagnosis	Diagnosis This species is characterized by a small shell, a smooth R 2, and the nearly smooth transition between R 2 and R 3 (no constriction between the two). See also under D. nattoungensis and D. nongtungensis.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A348FFAAB923FE9CB2DAFBAB.taxon	materials_examined	Type material examined INDIA • 5 syntypes (Fig. 70 K – O); Jawai, Jaintia; NHMUK 1903.7. 1.2688. Type locality “ near Jawai ”, “ on the banks of the Kopili river on the road from Jawai to Asálú, viâ Súfai ”.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A348FFACBADCFBD8B2DBFD35.taxon	description	Fig. 72	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A348FFACBADCFBD8B2DBFD35.taxon	diagnosis	Diagnosis A small species of Dicharax with smooth R 1, having some widely-spaced ribs on the last whorl of R 1, a combined R 2 + R 3 length of ca 150 degrees, an R 3 with a single, low, central swelling, and a moderately fringed peristome.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A348FFACBADCFBD8B2DBFD35.taxon	etymology	Etymology The specific epithet ‘ semivivus ’ (‘ half-alive, almost dead’) refers to the condition of the holotype, which was corroded, but was probably alive when found.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A348FFACBADCFBD8B2DBFD35.taxon	materials_examined	Type material examined Holotype INDIA – Mizoram • empty shell (D: 2.4 mm, H: 1.8 mm, Fig. 72); Blue Mountain, Lawngtlai district; 22.727 ° N, 93.135 ° E; 1400 m a. s. l.; 27 Jan. 2019; N. A. Aravind leg.; Ar 45; NZSI LM 1754.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A348FFACBADCFBD8B2DBFD35.taxon	description	Description Shell small, off-white, semitranslucent; shell surface corroded but the holotype (single available specimen) was probably alive judging from the in situ presence of the operculum; shell outline slightly oval in dorsal view; spire elevated, conical; body whorl rounded; protoconch low, smooth, its boundary not visible due to corrosion consisting of 1.5 whorls; R 1 of ca 1.75 whorls, corroded, nearly smooth, last whorl with widely-spaced, low ribs near suture and inside umbilicus, no ribs reach mid part of whorls; boundary between R 1 and R 2 conspicuous due to the sudden increase in rib density; R 2 with ca 26 low ribs, spaces between the ribs are approximately the width of a single rib or narrower; R 2 + R 3 ca 150 degrees; R 3 slightly longer than R 2; boundary between R 2 and R 3 clearly visible due to a deep constriction; R 3 with a low central swelling, anterior and posterior slopes are declivous; aperture strongly oblique to shell axis, rounded with a shallow upper (parieto-palatal), a deep lower (basal) incision and a fringed palatal part; together with the upper and lower incisions, there are 5 incisions on the peristome; peristome thin overall, boundary between inner and outer peristomes conspicuous; inner peristome slightly protruding; outer peristome only slightly expanded, not reflected, undulated following the fringed inner peristome; umbilicus ca one fourth of shell width, nearly rounded. OPERCULUM. The outer side is smooth, inner side could not be examined. MEASUREMENTS. D: 2.4 mm, H: 1.8 mm.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A348FFACBADCFBD8B2DBFD35.taxon	distribution	Distribution Known only from the Blue Mountain region.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A348FFACBADCFBD8B2DBFD35.taxon	diagnosis	Differential diagnosis The most similar species is Dicharax nitidus (Fig. 55), but it is larger, and has a longer R 3, with a more posteriorly-situated swelling. Dicharax jaintiacus (Fig. 32) is larger, has a higher spire and a narrower umbilicus, a more strongly expanded outer peristome and a more strongly fringed aperture.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A34EFFAFB922FD56B7D8FD23.taxon	description	Fig. 69 F – J	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A34EFFAFB922FD56B7D8FD23.taxon	diagnosis	Diagnosis The combination of the strong, widely-spaced R 1 ribs, the relatively widely-spaced R 2 ribs and the blunt, central R 3 swelling distinguish this species from its congeners.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A34EFFAFB922FD56B7D8FD23.taxon	materials_examined	Type material examined INDIA • 1 syntype of A. theobaldi (Fig. 69 F – J); Khasi Hills, NHMUK 1906.4. 4.60 • 12 syntypes of Alycaeus theobaldi var. diyungensis (Fig. 69 F – J); Diyung Valley, N of Asalu; NHMUK 1903.7. 1.2546 • 1 syntype of A. maosmaiensis (Fig. 69 K – O) separated in a vial with pink wool + 4 additional syntypes; Maosmai, nr Cherrapoonjee, Khasi; NHMUK 20191067. Additional material examined INDIA • 3 shells; Nongphrian, Khasi Hills; Dr Alfred Oberwimmer coll.; NHMW (mixed sample with D. hebes) • 1 shell; same data as for preceding; NHMW 14715 (mixed sample with D. hebes). Type localities “ in montibus Khasi ” (A. theobaldi); “ Diyung Valley, north of Asalu, N. Cachar ” (A. theobaldi var. diyungensis); “ Khasi Hills, near Cherrapunji, at the mouth of the Maosmai cave ” (A. maosmaiensis).	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A34EFFAFB922FD56B7D8FD23.taxon	discussion	Remarks According to the original description of Alycaeus theobaldi var. diyungensis, that variety differs from the typical A. theobaldi in the following traits: “ This Diyung Valley species is a departure from A. theobaldi of the Khasi Hills in having the apex less high and conical, the suture more impressed, the shell more openly umbilicated; costulation next suture stronger ” (Godwin-Austen 1914: 402). However, we do not see any of these differences comparing the type specimens of the two taxa. Therefore, we treat D. theobaldi diyungensis as a junior synonym of D. theobaldi. The type specimens of D. theobaldi, D. theobaldi diyungensis and D. maosmaiensis are similar in shell size and shell shape, while the sculpture of R 1 shows differences. Namely, D. theobaldi and D. theobaldi diyungensis have strong, widely-spaced ribs that are more or less regular at the beginning of R 1, and become irregular and more widely-spaced towards its end, while D. maosmaiensis has a smooth beginning of R 1, and at the end of R 1 the ribs are relatively weak, and slightly denser than in the other two nominal taxa. Nevertheless, these differences are considered part of the intraspecific variability. A similar case (intraspecific variability in R 1 rib density) was shown in the case of Dicharax notus (see Páll-Gergely et al. 2021).	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A34DFF51BAF8FD27B2F5F982.taxon	description	Figs 65 D – F, 66 C – D, 73	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A34DFF51BAF8FD27B2F5F982.taxon	diagnosis	Diagnosis A small species of Dicharax with glossy R 1 and R 3, a combined R 2 + R 3 length of ca 150 degrees, an R 3 with a clearly separated main swelling and a secondary swelling, and a rounded aperture with a slight upper incision.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A34DFF51BAF8FD27B2F5F982.taxon	etymology	Etymology This species is named after Prof. R. Uma Shaanker, Founder Trustee of Ashoka Trust for Research in Ecology and the Environment (ATREE), Bangalore, India. He has contributed immensely to the understanding of ecology, evolution, and conservation of flora and fauna of the Western Ghats, India.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A34DFF51BAF8FD27B2F5F982.taxon	materials_examined	Type material examined Holotype INDIA – Mizoram • empty shell (D: 2.7 mm, H: 2 mm, Fig. 73); Blue Mountain, Lawngtlai district; 22.727 ° N, 93.135 ° E; 1400 m a. s. l.; 27 Jan. 2019; N. A. Aravind leg.; Ar 73; NZSI LM 1742. Paratypes INDIA – Mizoram • 28 empty shells; same data as for holotype; NZSI LM 1743 • 2 empty shells; same data as for holotype; Ar 58; NZSI LM 1744 • 1 empty shell; same data as for holotype; Ar 75; NZSI LM 1907 • 3 empty shells; same data as for holotype; Ar 40; NZSI LM 1745 • 1 empty shell; same data as for holotype; Ar 50; NZSI LM 1746 • 3 empty shells; same data as for holotype; Ar 63; NZSI LM 1747 • 1 empty shell; same data as for holotype; Ar 25; NZSI LM 1748 • 1 empty shell; same data as for holotype; Ar 24; NZSI LM 1749 • 1 empty shell; same data as for holotype; Ar 15; NZSI LM 1750 • 1 empty shell; same data as for holotype; Ar 15; NZSI LM 1908. Additional material examined INDIA – Mizoram • 2 juvenile empty shell (Fig. 65 D – F, 66 C – D); same data as for holotype; NZSI LM 1751 • 1 broken shell; same data as for holotype; Ar 21, NZSI LM 1752 • 3 corroded / broken shells; same data as for holotype; Ar 36, NZSI LM 1753.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A34DFF51BAF8FD27B2F5F982.taxon	description	Description Shell small, light yellowish to orange or ochre, in some specimens with reddish apex; glossy, semitranslucent; shell outline slightly oval in dorsal view; spire slightly elevated, low conical; body whorl rounded; protoconch low, glossy, consisting of 1.25 – 1.5 whorls; R 1 of ca 1.5 whorls, glossy, nearly smooth, with some inconspicuous, dense, low riblets, in some specimens ribs increase in strength on the last ca quarter whorl; spiral striation absent; boundary between R 1 and R 2 conspicuous due to the sudden change to a regularly, densely ribbed surface; R 2 with ca 18 – 22 straight, sharp, lamella-like ribs; R 2 ribs low, blunt, curved towards aperture, spaces between the ribs are approximately the width of a single rib; cross sectional view of R 2 was examined in a specimen (Fig. 66 C – D): anterior crust forms a significantly elevated rib, which has an anterior projection (i. e., the R 2 ribs are curved towards the aperture); anterior crust folds over the much weaker posterior crust; microtunnel wide compared to ribs, its cross sectional view teardrop-shaped; R 2 + R 3 ca 150 degrees; R 3 slightly longer than R 2; boundary between R 2 and R 3 clearly visible due to a deep constriction; R 3 forms a blunt tip towards constriction and possesses two swellings; the upper swelling is visible only on the dorsal side, stops at the centre of the whorl, while the lower swelling (i. e., the one situated closer to the aperture) reaches the ventral side of shell; the dent separating the two swellings is elongated, beginning at the suture; aperture strongly oblique to shell axis, rounded with a very slight upper (parieto-palatal) incision; boundary between inner and outer peristomes conspicuous; inner peristome slightly thickened, protruding; outer peristome more conspicuous, sharp, strongly expanded, and reflected towards umbilicus; umbilicus very narrow, ca 1 / 5 – 1 / 6 of shell width, elongated, oval. OPERCULUM. Outer surface multispiral, without elevated lamina, inner side with an elevated, flat-topped nipple. MEASUREMENTS. D: 2.6 – 3.1 mm, H: 2 – 2.2 mm.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A34DFF51BAF8FD27B2F5F982.taxon	distribution	Distribution Known only from the Blue Mountain region.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A34DFF51BAF8FD27B2F5F982.taxon	diagnosis	Differential diagnosis Dicharax akhaensis is similar in having two swellings and being glossy, but differs from the new species by having more widely-spaced R 1 ribs, denser R 2 ribs (and possibly lower, although the holotype is corroded), a fringed aperture, and a wider umbilicus. Dicharax birugosus is similar in colour, size, sculpture and length of R 1 and R 2, but its R 3 is shorter than that of the new species, the aperture is slightly fringed and has a deep basal incision. Moreover, the two swellings are arranged differently: the main swelling is oblique to the constriction in Dicharax birugosus, and rather parallel in the new species, while the area on which the secondary swelling is situated is much wider in the new species. Dicharax subculmen belongs to the “ Elephant Dicharax ” species group but is somewhat similar to this new species, at least in dorsal view. Nevertheless, the new species has a more depressed shell and a wider umbilicus, a deeper ditch between the primary and secondary R 3 swellings, a longer R 3 with a larger distance between the peristome and the secondary swelling, and a thinner peristome. Dicharax kurzianus has a fringed aperture. Dicharax multirugosus has a more depressed shell, and has three separate R 3 swellings.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3B3FF51B9BFF986B574F8B0.taxon	discussion	Remarks This species group is characterized by a low and blunt R 3 swelling that is situated close to the outer peristome, and a slightly fringed to rounded peristome.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3B3FF50B977F8D1B7F7FD4E.taxon	description	Fig. 74 A – E	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3B3FF50B977F8D1B7F7FD4E.taxon	diagnosis	Diagnosis This species differs from the other species of this species group by the absence of an R 3 swelling.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3B3FF50B977F8D1B7F7FD4E.taxon	materials_examined	Type material examined INDIA • 1 syntype (labelled as holotype) (Fig. 74 A – E); Sikkim; elev. 11000 ft; Dr Stoliczka coll.; NZSI M. 8082. Type locality “ Sikkim, at 11,000 ft ”.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3B2FF50B938FCF1B5F5FAC2.taxon	description	Fig. 74 F – J	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3B2FF50B938FCF1B5F5FAC2.taxon	materials_examined	Type material examined INDIA • holotype (single shell mentioned in the original description: Fig. 74 F – J); Rechila Pk, Sikkim; N. Robert leg.; NHMUK 1903.7. 1.1252. Type locality “ Rechila Peak, Daling District, on Sikhim-Bhutan Frontier, 10,300 feet ”.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3B2FF50B938FCF1B5F5FAC2.taxon	diagnosis	Differential diagnosis Dicharax strangulatus has a shallower constriction between R 2 and R 3, and the R 3 swelling of that species is less steep. See also under D. montanus and D. strigatus.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3B2FF53B91EFA45B7C0FEEC.taxon	description	Fig. 74 K – O	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3B2FF53B91EFA45B7C0FEEC.taxon	materials_examined	Type material examined INDIA • 1 shell (“ possible syntype ”) (Fig. 74 K – O); NHMUK 1856.9. 15.18 • at least 5 shells; Mussoorie, N. W Himalaya, “ figd ”; NHMUK 1903.7. 1.2501 • several shells; NHMUK 1928.7.28.85 - 104 (from general collection). See also newly collected specimens examined in Sajan et al. (2020). Type locality “ Bengalia ”.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3B2FF53B91EFA45B7C0FEEC.taxon	diagnosis	Differential diagnosis See under D. rechilaensis.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3B0FF52BA8CFAB5B7D6F86E.taxon	description	Fig. 75 A – D	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3B0FF52BA8CFAB5B7D6F86E.taxon	diagnosis	Diagnosis Differs from its congeners by the densely ribbed shell, and the keeled body whorl.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3B0FF52BA8CFAB5B7D6F86E.taxon	materials_examined	Type locality “ Samdrup Jongkhar Dzongkhag: 7 km NNW of Daifam (= Jomotshangkha), Toka Phu, 940 m a. s. l., 26 º 57 ’ N 92 º 04 ’ E ” (Bhutan).	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3B0FF52BA8CFAB5B7D6F86E.taxon	diagnosis	Differential diagnosis As mentioned in the original description (Gittenberger et al. 2024), the only species of Alycaeinae in the region with a keeled body whorl is Chamalycaeus sculptilis, which has a beaded peristome. Nevertheless, D. bhucarinatus was described based on the corroded holotype and therefore, the sculpture could not be examined. Thus, it may belong to Chamalycaeus (more likely than Metalycaeus, as mentioned by Gittenberger et al. 2024).	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3B0FF52B947FB44B33BFA95.taxon	discussion	Remarks This species group is characterized by dense ribbing on the entire shell, and rounded aperture.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3B0FF52B924FEECB282FBC7.taxon	description	Fig. 74 P – R	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3B0FF52B924FEECB282FBC7.taxon	materials_examined	Material examined The syntypes should be in the Indian Museum (see Nevill 1878), but they have not been located there; therefore, they are considered lost (Páll-Gergely et al. 2020). Type locality “ Assam ”.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3B0FF52B924FEECB282FBC7.taxon	discussion	Remarks This species may be conspecific with D. rechilaensis, but in the absence of available specimens of D. strigatus, we cannot compare the two species properly. According to the original description of D. strigatus, the shell is 0.15 inch (ca 3.8 mm), while D. rechilaensis is 5 mm (Godwin-Austen 1914). The drawing of D. strigatus in lateral view (Fig. 74 Q) agrees clearly with the photo of D. rechilaensis in the same view (Fig. 74 G). However, the apical view (Fig. 74 R) shows a shallower constriction than in D. rechilaensis (Fig. 74 I). Because of the mismatch of the measurements and the anomalies of the figures, we maintain this as a valid species, until more information becomes available.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3B7FF55BA7CFEECB2DCFC84.taxon	description	Fig. 75 E – H	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3B7FF55BA7CFEECB2DCFC84.taxon	diagnosis	Diagnosis Not similar to any other species of Dicharax. Although no spiral striation was mentioned in the original description, this species rather resembles some species of Metalycaeus, such as M. heudei (Bavay & Dautzenberg, 1900) (see Páll-Gergely et al. 2017). However, the photos in the original description are not of sufficient resolution to see the fine sculpture. We keep this species in Dicharax until we will have the opportunity to examine the holotype or additional material. Nevertheless, this species is not similar to any species of Metalycaeus from the Himalayas.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3B7FF55BA7CFEECB2DCFC84.taxon	materials_examined	Type locality “ Zhemgang Dzongkhag: 19.5 km SE of Tingtibi, 780 m a. s. l., 27 ° 01 ’ N 90 ° 49 ’ E ” (Bhutan).	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3B7FF54BA90FC85B292FE5A.taxon	description	Fig. 75 I – L	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3B7FF54BA90FC85B292FE5A.taxon	diagnosis	Diagnosis The dense ribbing, simple peristome and the morphology of R 3 swelling distinguish this species from all other Alycaeinae from the region.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3B7FF54BA90FC85B292FE5A.taxon	materials_examined	Type locality “ Mongar Dzongkhag: 3 km N of Mongar, 1070 m a. s. l., 27 º 19 ’ N 91 º 13 ’ E ” (Bhutan).	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3B6FF54B901FD04B44EFAE4.taxon	description	Fig. 76	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3B6FF54B901FD04B44EFAE4.taxon	diagnosis	Diagnosis This species can be recognized based on the relatively large, densely ribbed shell, and the strongly oblique aperture.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3B6FF54B901FD04B44EFAE4.taxon	materials_examined	Type material examined MYANMAR • 6 syntypes (Fig. 76); Shan Hills, E of Ava, Burma; Blanford coll.; NHMUK 1906.4. 4.61. Type locality “ The hills east of Mandalay and Ava ”.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3B6FF54B946FDEFB7F8FD07.taxon	discussion	Remarks This species group is characterized by a relatively large, ribbed shell, and a slightly fringed aperture (both from Myanmar).	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3B6FF58B921FA65B7B1FDAA.taxon	description	Figs 77 – 80	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3B6FF58B921FA65B7B1FDAA.taxon	diagnosis	Diagnosis The large shell, widely-spaced ribs, the long R 2, relatively low R 3 swelling and the fringed aperture distinguish this species from its congeners. Based on the lamella-like ribs, this species rather resembles species of Chamalycaeus and Metalycaeus (e. g., M. brahma, see Páll-Gergely et al. 2020), but lacks the spiral striation on its entire shell.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3B6FF58B921FA65B7B1FDAA.taxon	materials_examined	Type material examined MYANMAR • 1 syntype of A. succineus (Fig. 77 A – E); Tangoop Pass, Arakan Hills; NHMUK 1906.4. 4.52 • holotype of A. blanfordi (single specimen mentioned in the original description: Fig. 77 F – J); Chwegalé, Arakan Hills; NHMUK 1906.4. 4.177 • 1 shell, labelled as syntype of A. vulcani; Ava, Burma; MCZ 135705 • 4 shells, possible syntypes of A. vulcani; Puppadoung; ex coll. Theobald; NHMUK 1888.12.4.939 - 942 • 2 shells (possible syntypes of A. vulcani); Puppa Hill, Ava; Blanford leg.; Crosse coll. 1899, Sykes coll. 1954; NHMUK • possible syntypes of A. vulcani (Fig. 78 A – E); Pupa Hills, Burma, extinct volcano; W. T. Blanford coll.; NHMUK 1906.4. 4.49 • possible syntypes of A. vulcani (Fig. 78 F – J); Puppa Ava, Burma; H. F. Blanford coll., ex coll. auctoris; NHMUK 20160325. Additional material examined INDIA – Mizoram • 1 empty shell (Fig. 79); Blue Mountain, Lawngtlai district; 22.727 ° N, 93.135 ° E; 1400 m a. s. l.; 27 Jan. 2019; N. A. Aravind leg.; Ar 70; NZSI LM 1727 • 2 empty shells; same data as for preceding; Ar 26; NZSI LM 1728 • 1 empty shell; same data as for preceding; Ar 22; NZSI LM 1729. Type localities “ in montibus Arakanensibus ” (A. succineus); “ on the upper portion of the isolated peak of Puppa, an extinct volcano lying ca 40 miles E. S. E. of the town of Pu-gán in the territories of the king of Ava ” (A. vulcani); “ Chwegali, Arakan Hills ” [probably Chaingalain Village, Rati Taung: 20 ° 23.8 ’ N 92 ° 39.9 ’ E] (A. blanfordi).	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3B6FF58B921FA65B7B1FDAA.taxon	distribution	Distribution This species is known from the western part of Myanmar and the Indian state of Mizoram (Fig. 80).	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3B6FF58B921FA65B7B1FDAA.taxon	discussion	Remarks The type specimens of Alycaeus blanfordi, A. succineus and A. vulcani are nearly identical. There is no notable difference in shell size, shape, and sculpture. Therefore, they are treated as synonyms, and the oldest name, D. succineus, is valid. Blanford (1863), when describing D. vulcani, compared that species with D. succineus, and wrote that it was more globose than A. succineus. The opposite is true; the syntype of A. succineus is slightly more globular than that of A. vulcani, although the difference represents intraspecific variability. Godwin-Austen (1914) did not compare A. blanfordi with any other species and only noted that the species had very distinctive characters and was worthy of being named. The specimens from the Blue Mountain differ from the ones from the Arakan Mountains (types of A. blanfordi, A. succineus and A. vulcani) by having a glossy R 3, while that of the specimens in the Arakan Mountains are matte with some growth lines. We consider this difference insufficient for species-level distinction.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3B9FF5BB947F9FAB445F972.taxon	discussion	Remarks This species group is not based on morphology, but all species from Myanmar which could not be classified elsewhere are lumped here.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3B9FF5AB93AF917B71FFCEF.taxon	description	Fig. 81 A – J	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3B9FF5AB93AF917B71FFCEF.taxon	diagnosis	Diagnosis R 1 nearly smooth (with only some strong, widely-spaced ribs near the suture), R 3 swelling conical, upper edge of peristome strongly protruding. Based on these traits, this species can be easily distinguished from its congeners.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3B9FF5AB93AF917B71FFCEF.taxon	materials_examined	Type material examined MYANMAR • holotype (single shell mentioned in the original description: Fig. 81 A – E); Ataran, Burma; ex Dr F. Stoliczka coll.; NZSI M. 8073. Additional material examined MYANMAR • 1 shell (Fig. 81 F – J); Kayin State, ca 51 km ESE of Mudon Centre, NE of Htimahto, NE point of Kwooprai Hill; 16 ° 2.396 ′ N, 97 ° 58.200 ′ E; 30 m a. s. l.; 10 Oct. 2018; A. Hunyadi, K. Okubo and J. U. Otani leg.; Coll. HA. Type locality “ Ataran ”.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3B8FF5AB926FC9CB281FA4D.taxon	description	Fig. 81 K – L	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3B8FF5AB926FC9CB281FA4D.taxon	materials_examined	Material examined The type specimens of this species are probably lost (Páll-Gergely et al. 2020). Type locality “ Momeit, Burmah ”.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3B8FF5AB926FC9CB281FA4D.taxon	diagnosis	Differential diagnosis According to the drawings (Fig. 81 K – L), the most similar species is probably D. rugosus, but in that species only the inner surface of the peristome is fringed. See also under D. muspratti.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3B8FF5CB925F9FDB7FAFBD2.taxon	description	Fig. 82	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3B8FF5CB925F9FDB7FAFBD2.taxon	diagnosis	Diagnosis The large shell size, the long R 2 + R 3, and the aperture shape distinguish this species from all other Alycaeinae.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3B8FF5CB925F9FDB7FAFBD2.taxon	materials_examined	Type material examined INDIA • 3 syntypes of A. stoliczkii (Fig. 82 A – E); Naga Hills; NHMUK 1903.7. 1.2622 • 1 syntype of A. peilei (Fig. 82 F – J); Naga Hills; Preston coll.; NHMUK 1915.1. 4.1281. Type localities “ Angaoluo Peak, Nágá Hills at 7,000 feet ”; “ further to the east at Kezakenomih, and at the head of the Lanier River at ca. 5,000 feet where the specimens are much larger ” (A. stoliczkii); “ Naga Hills ” (A. peilei).	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3B8FF5CB925F9FDB7FAFBD2.taxon	diagnosis	Differential diagnosis The most similar species is D. glaber, but D. stoliczkii has a more oval shell (rather rounded in D. glaber), a fringed aperture and a secondary swelling touching the penultimate whorl near the peristome.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3B8FF5CB925F9FDB7FAFBD2.taxon	discussion	Remarks Alycaeus (Charax) peilei is identical with Alycaeus stoliczkii and therefore it is treated as a junior synonym. In the original description of the first species, Preston (1914) did not compare his new species with any other species.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3BEFF5CB969FB70B760F9B4.taxon	type_taxon	Type species	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3BEFF44BAC6F9D5B22DFB82.taxon	description	Figs 83 – 88	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3BEFF44BAC6F9D5B22DFB82.taxon	diagnosis	Diagnosis This species can be recognised based on the relatively large aperture, finely undulated peristome, the elongated basal peristome edge and the blunt centrally located R 3 swelling. We indicate that the Himalayan species of Metalycaeus are not revised in this study, and a comprehensive revision should identify the species-specific traits.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3BEFF44BAC6F9D5B22DFB82.taxon	materials_examined	Type material examined INDIA • 10 syntypes of A. distinctus (Fig. 83 F – J); Naga Hills; NHMUK 1903.7. 1.2619 • 1 syntype of A. kamakiaensis (Fig. 84 A – E); Kamakia Hill, Gowhatty, Assam; NHMUK 1903.7. 1.2705 • holotype (single shell mentioned in the original description: Fig. 83 F – J) of A. rotundatus; Dafla Hills (?); NHMUK 1903.7. 1.2543. MYANMAR • 2 syntypes of A. polygonoma in different vials (Fig. 83 A – E); Tongoop Pass, Arakan Hills; NHMUK 1906.4. 4.51. Additional material examined BANGLADESH • 2 shells (Fig. 88); Chittagong, natural mango garden on a river bank; 22 ° 31.559 ′ N, 92 ° 7.652 ′ E; 27 Sep. 2024; T. Saito and M. S. Shariar leg.; Coll. PGB. INDIA • 2 shells of “ Alycaeus distinctus var. ”; Sadia, Assam; NHMUK 1903.7. 1.2620 • 19 + 6 shells of “ Alycaeus distinctus var. ”; Jatinga Valley, N. Cachar; Godwin-Austen coll.; NHMUK 1903.7. 1.2576 • 1 empty shell (Figs 12 D, 85); Blue Mountain, Lawngtlai district; 22.727 ° N, 93.135 ° E; 1400 m a. s. l.; 27 Jan. 2019; N. A. Aravind leg.; Ar 48; NZSI LM 1762 • 2 empty shells (Figs 86 – 87 [operculum]); same data as for preceding; Ar 71; NZSI LM 1763 • 1 empty shell; same data as for preceding; Ar 43; NZSI LM 1909 • 5 empty shells; same data as for preceding; Ar 46; NZSI LM 1910 • 2 empty shells; same data as for preceding; Ar 51; NZSI LM 1911. Type localities “ in montibus Arakanensibus ” (A. polygonoma); “ Neighbourhood of Asálú, N. Cachar Hills ” (A. distinctus); “ Kamakia Temple Hill near Gowhatty, Assam ” (A. kamakiaensis); “ Dafla hills ” (A. rotundatus); “ Zhemgang Dzongkhag: between Duenmang Tsachu and Gongphu Zero Point, 24 km SE of Zhemgang, 335 m a. s. l., 27 ° 02 ’ N 90 ° 48 ’ E, scree in warm broadleaf forest ” (M. posteriobulbus).	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3BEFF44BAC6F9D5B22DFB82.taxon	distribution	Distribution The new information reveals that M. polygonoma has a larger distribution than previously thought; its area spans from Bhutan and the Dafla Hills (India) to the Arakan Mountains in Myanmar (Fig. 89). Furthermore, this species is reported from Bangladesh for the first time. With these localities, M. polygonoma is one of the few species of Alycaeinae with a large distributional area besides Metalycaeus heudei (Páll-Gergely et al. 2017, 2021), Dicharax fimbriatus and D. cristatus (Möllendorff, 1886) (see Páll-Gergely et al. 2017).	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3BEFF44BAC6F9D5B22DFB82.taxon	discussion	Remarks Type specimens of A. polygonoma, A. distinctus, A. kamakiaensis, A. rotundatus and M. posteriobulbus do not differ in any important conchological characters; therefore, the latter four are junior synonyms of the former. The types of those five species and the newly collected samples from the Blue Mountain show slight variability in shell size, length of R 2, width of umbilicus, and the degree of undulation of the peristome, but all differences are attributed to intraspecific variability.	en	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
