taxonID	type	description	language	source
03CCCC27FFDAFFB9FF1FF9D3CECEFC92.taxon	description	The alignment of all Rhyacophila COI gene sequences generated in our lab was performed and edited manually using MEGA X (Kumar et al. 2018). Fine adjustments (for individual, undefined nucleotides) were made by eye, as the COI does not present any gaps. We undertook phylogenetic analyses using Maximum Likelihood (ML) and Bayesian inference (BI) methods in the CIPRES Science Gateway v. 3.3 online server (Miller et al. 2010). We ran jModeltest v. 2.1.9 (Darriba et al. 2012; Guindon & Gascuel 2003) on the partitioned data to identify the best-fitting substitution model that we used in the ML and BI tree inferences. We performed phylogenetic inference under ML in RAxML-HPC BlackBox (Stamatakis 2014), using 10,000 bootstrap replicates to assess node support. We performed BI in MrBayes- 3.2.7 (Ronquist et al. 2012) using the best-fitting substitution model and setting lset nst = mixed rates = invgamma, for 5 × 106 generations, and reconstructed Majority Rule trees (with 50 % threshold, made with Geneious Prime ® 2022.0.1) after discarding the first 25 % of the generations as burn-in. We visualized the best tree using FigTree v 1.4.4. (Rambaut 2019), and we labelled the nodes with posterior probabilities. We calculated distances among and within species using the maximum composite likelihood and p-distance models with MEGA X. We deposited nucleotide sequences in GenBank [accession numbers and Tissues and DNA Collection numbers (MNCN: ADN) in Table 1, and in the dataset by Valladolid et al. 2025].	en	Valladolid, Maria, Arauzo, Mercedes, Coppa, Gennaro, Reding, Jean-Paul G., Dorda, Beatriz A., Rey, Isabel (2025): The Rhyacophila fasciata Species Complex (Trichoptera: Rhyacophilidae) in France and Switzerland, with description of a new species, Rhyacophila caussensis Valladolid & Coppa sp. nov. from France, based on morphological, genetic and ecological evidence. Zootaxa 5692 (3): 401-445, DOI: 10.11646/zootaxa.5692.3.1, URL: https://doi.org/10.11646/zootaxa.5692.3.1
03CCCC27FFDBFFB9FF1FF9F6C908F8BE.taxon	etymology	Derivatio nominis: Derived from the name of the area where the new species has been found, the Grands Causses, a set of high calcareous plateaus that extend across the south of the Massif central of France. Material Examined: Holotype ♂: FRANCE (CAU 2), river La Sorgue, source, Cornus, Aveyron, 43.877 ºN, 3.205 ºE, 574 m a. s. l., 20 / vii / 2013, leg. G. Coppa [T 5587 (OPIE)]. Paratypes: FRANCE, 1 ♂, river La Sorgue, source, Cornus, Aveyron, 20 / vii / 2013, (G. Coppa) [T 5587 (OPIE)]; 1 ♂ + 1 ♀, river Le Cernon, Saint Rome de Cernon, Aveyron, 21 / vii / 2013, (G. Coppa) [T 5580 (OPIE)].	en	Valladolid, Maria, Arauzo, Mercedes, Coppa, Gennaro, Reding, Jean-Paul G., Dorda, Beatriz A., Rey, Isabel (2025): The Rhyacophila fasciata Species Complex (Trichoptera: Rhyacophilidae) in France and Switzerland, with description of a new species, Rhyacophila caussensis Valladolid & Coppa sp. nov. from France, based on morphological, genetic and ecological evidence. Zootaxa 5692 (3): 401-445, DOI: 10.11646/zootaxa.5692.3.1, URL: https://doi.org/10.11646/zootaxa.5692.3.1
03CCCC27FFDDFFBFFF1FFF73CE78FC4E.taxon	description	MALE Genitalia (Fig. 9, 13): 2 nd segment of each inferior appendage (Fig. 9 A, 13 a) with posterior edge slightly concave in central area, ventral edge straight, 2 times longer that dorsal edge; apicodorsal vertex of 2 nd segment ~ 135 ° and round, apicoventral angle ~ 50 °, projecting as thick lobe tapered to round apex. Parameres in ventral view curved posteromesad in distal half (Fig. 9 BV, p). In lateral view (Figs 9 BL, 13 d) each constricted at short distance beyond base, dilated at middle, with dorsal and ventral margins rounded and almost parallel; two rows of thick spines on midventral margin, pointing outward; midlateral surface with spicules or setae, from middle posteroventral to posterodorsal edges, bigger from ventral to middle area, smaller from middle to dorsal area. Aedeagus (phallicata) in lateral view (Fig. 9 CL) with anterodorsal margin and posterior corner of dorsal concavity hooked anterad, apical edge of aedeagus straight or slightly convex; ventral lobe of aedeagus in ventral view almost circular, with pointed apex (Fig. 9 BV, vl), in lateral view vertical edge straight, posteroventral angle round, almost 90 º (Fig. 9 CL). In dorsal view posterior edges of lateroventral lobes of phallus apically triangular, apicolateral margins convex (Fig. 9 CD, lvl). Apicodorsal lobe of segment IX in dorsal view oval (Figs 9 DD, al; 13 b), with small apicomesal excision; preanal appendages (Fig. 9 DD, pa) elongate rectangular, 4 / 5 as long as apicodorsal lobe, round subapically and posterior edges triangular. In ventral view, apical band U-shaped (Fig. 9 DV, ab), longer than wide, its posterior arms parallel and round apically, its non-sclerotized mesal area U-shaped (Fig. 9 DV, va), anal sclerites triangular, posterior edges concave (Fig. 9 DV, as).	en	Valladolid, Maria, Arauzo, Mercedes, Coppa, Gennaro, Reding, Jean-Paul G., Dorda, Beatriz A., Rey, Isabel (2025): The Rhyacophila fasciata Species Complex (Trichoptera: Rhyacophilidae) in France and Switzerland, with description of a new species, Rhyacophila caussensis Valladolid & Coppa sp. nov. from France, based on morphological, genetic and ecological evidence. Zootaxa 5692 (3): 401-445, DOI: 10.11646/zootaxa.5692.3.1, URL: https://doi.org/10.11646/zootaxa.5692.3.1
03CCCC27FFDDFFA2FF1FFABBCEFFFE8E.taxon	description	MALE Genitalia (Figs 11, 14): 2 nd segment of each inferior appendage (Figs 11 A, 14 a) with posterior edge slightly convex in dorsal and concave in ventral area respectively, ventral edge straight or slightly concave, less than 2 times longer that dorsal edge; apicodorsal vertex of 2 nd segment ~ 110 ° and round, apicoventral angle ~ 35 °, projecting as thick lobe tapered to round apex. Parameres in ventral view curved posteromesad in distal half (Figs 11 BV, p; 14 c). In lateral view (Figs 11 BL, 14 d) each constricted 1 / 3 distance beyond base, dilated at middle, with semicircular dorsal margin, convex ventral margin; along midventral posterior margin three long, thick spines; midlateral surface with thin spicules or setae of the same size, from middle posteroventral to posterodorsal edges. Aedeagus (phallicata) in lateral view (Fig. 11 CL) with anterodorsal margin and posterior corner of dorsal concavity hooked anterad, apical edge of aedeagus mostly triangular; ventral lobe of aedeagus in ventral view triangular (Fig. 11 BV, vl), with bifid apex, in lateral view vertical edge deeply concave, with triangular projection in posteroventral edge, posteroventral angle less than 90 º (Fig. 11 CL). In dorsal view posterior edges of lateroventral lobes of phallus apically straight or slightly convex, apicolateral margins straight or slightly convex (Fig. 11 CD, lvl). Apicodorsal lobe of segment IX in dorsal view oval, slightly dilated subapicolaterally (Figs 11 DD, al; 14 b), with small apicomesal excision; preanal appendages (Fig. 11 DD, pa) round, 4 / 5 long than apicodorsal lobe, posterior edges convex. In ventral view, apical band pentagonal shaped (Fig. 11 DV, ab), almost as long as wide, its posterior arms converging posterad and round apically, its non-sclerotized mesal area oval (Fig. 11 DV, va), anal sclerites triangular, posterior edges concave (Fig. 11 DV, as).	en	Valladolid, Maria, Arauzo, Mercedes, Coppa, Gennaro, Reding, Jean-Paul G., Dorda, Beatriz A., Rey, Isabel (2025): The Rhyacophila fasciata Species Complex (Trichoptera: Rhyacophilidae) in France and Switzerland, with description of a new species, Rhyacophila caussensis Valladolid & Coppa sp. nov. from France, based on morphological, genetic and ecological evidence. Zootaxa 5692 (3): 401-445, DOI: 10.11646/zootaxa.5692.3.1, URL: https://doi.org/10.11646/zootaxa.5692.3.1
03CCCC27FFC5FFA7FF1FFEAECF41FBB2.taxon	description	MALE Biometrics: Lengths: Body (distance from front of head to distal edge of segment IX): 7.8 – 13.0 mm (x ̄ = 10.3, n = 27), forewing 10.1 – 15.8 mm (x ̄ = 12.8, n = 27), hind wing 9.1 – 14.2 mm (x ̄ = 11.3, n = 27). Genitalia (Figs 19, 23): 2 nd segment of each inferior appendage (Figs 19 A, 23 a) with posterior edge slightly concave in dorsal area, ventral edge straight or slightly concave, ~ 2.2 times longer that dorsal edge; apicodorsal vertex of 2 nd segment ~ 120 º and angled, apicoventral angle ~ 40 °, projecting as thick lobe tapering to apex. Parameres in ventral view curved posteromesad in distal half (Figs 19 BV, p; 23 c). In lateral view (Figs 19 BL, 23 d) each constricted 1 / 3 distance beyond base, dilated at middle, with round dorsal and ventral margins; without spines along midventral margin; midlateral surface with few long and thick spines on ventral area and fine spicules or setae from middle to ventral edge, absent in middle anteroventral and posterodorsal edges. Aedeagus (phallicata) in lateral view (Fig. 19 CL) with anterodorsal margin and posterior corner of dorsal concavity hooked anterad, apical edge of aedeagus straight projected posteroventrad in small ventral apex (Fig 19 CL); ventral lobe of aedeagus semicircular, with small posterior apex (Fig 19 BV, vl), in lateral view vertical edge straight, small projection in posteroventral angle (Fig. 19 CL). In dorsal view posterior edges of lateroventral lobes of phallus apically straight or slightly convex, apicolateral margins round (Fig 19 CD, lvl). Apicodorsal lobe of segment IX in dorsal view dilated subapicolaterally (Figs 19 DD, al; 23 b), with small apicomesal excision; preanal appendages (Fig 19 DD, pa) round, 4 / 5 as long as apicodorsal lobe, round subapically and posterior edges convex. In ventral view, apical band (Fig 19 DV, ab) V-shaped, longer than wide, its posterior arms converging, parallel and round apically, its non-sclerotized mesal area slightly oval, narrow (Fig 19 DV, va), anal sclerites triangular, posterior edges concave (Fig 19 DV, as).	en	Valladolid, Maria, Arauzo, Mercedes, Coppa, Gennaro, Reding, Jean-Paul G., Dorda, Beatriz A., Rey, Isabel (2025): The Rhyacophila fasciata Species Complex (Trichoptera: Rhyacophilidae) in France and Switzerland, with description of a new species, Rhyacophila caussensis Valladolid & Coppa sp. nov. from France, based on morphological, genetic and ecological evidence. Zootaxa 5692 (3): 401-445, DOI: 10.11646/zootaxa.5692.3.1, URL: https://doi.org/10.11646/zootaxa.5692.3.1
03CCCC27FFC5FFAAFF1FF946CD62FD66.taxon	description	MALE Biometrics: Body length (distance from front of head to distal edge of segment IX): 9.2 – 10.0 mm (x ̄ = 9.6, n = 3), forewing 11.3 – 12.7 mm (x ̄ = 12.0, n = 3), hind wing 10.2 – 11.4 mm (x ̄ = 10.6, n = 3). Genitalia (Figs 21, 24): 2 nd segment of each inferior appendage (Figs 21 A, 24 a) with posterior edge slightly convex in central area, ventral edge straight or slightly convex, ~ 2.1 times longer that dorsal edge; apicodorsal vertex of 2 nd segment ~ 135 ° and angled, apicoventral angle ~ 45 °, projecting as thick lobe tapering to round apex. Parameres in ventral view curved posteromesad in distal half (Figs 21 BV, p; 24 c). In lateral view (Figs 21 BL, 24 d) each constricted 1 / 3 distance beyond base, dilated at middle, with round dorsal and straight or slightly convex ventral margins respectively; along posterior midventral margin two rows with three long, thick spines, pointing outward; midlateral surface with spicules or setae, from middle posteroventral to posterodorsal edges, decreasing on size from ventral to dorsal edges. Aedeagus (phallicata) in lateral view (Fig. 21 CL) with anterodorsal margin and posterior corner of dorsal concavity hooked anterad, apical edge of aedeagus straight or slightly convex; ventral lobe of aedeagus in ventral view triangular (Fig. 21 BV, vl), in lateral view vertical edge convex, posteroventral angle almost 90 º (Fig. 21 CL). In dorsal view posterior edges of lateroventral lobes of phallus apically straight or slightly convex, apicolateral margins round (Fig. 21 CD, lvl). Apicodorsal lobe of segment IX in dorsal view almost circular, slightly dilated subapicolaterally (Figs 21 DD, al; 24 b), with small apicomesal excision in some specimens; preanal appendages (Fig. 21 DD, pa) rectangular, 3 / 4 as long as apicodorsal lobe, pointed subapically and posterior edges convex. In ventral view, apical band diamond shaped (Fig. 21 DV, ab), almost as long as wide, its posterior arms convergent posterad, rounded apically, its non-sclerotized mesal area oval (Fig. 21 DV, va), anal sclerites triangular, posterior edges straight or slightly concave (Fig. 19 DV, as).	en	Valladolid, Maria, Arauzo, Mercedes, Coppa, Gennaro, Reding, Jean-Paul G., Dorda, Beatriz A., Rey, Isabel (2025): The Rhyacophila fasciata Species Complex (Trichoptera: Rhyacophilidae) in France and Switzerland, with description of a new species, Rhyacophila caussensis Valladolid & Coppa sp. nov. from France, based on morphological, genetic and ecological evidence. Zootaxa 5692 (3): 401-445, DOI: 10.11646/zootaxa.5692.3.1, URL: https://doi.org/10.11646/zootaxa.5692.3.1
