identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03CF3F6EFFD7FFB0FF79F8A9FE93DDE6.text	03CF3F6EFFD7FFB0FF79F8A9FE93DDE6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cyana artemis Volynkin & Černý 2019	<div><p>Cyana artemis Volynkin &amp; Černý, sp. n.</p><p>(Figs 1–4, 23, 24, 35)</p><p>urn:lsid:zoobank.org:act: B035E3CB-7A67-459D-926E-F33B582A7B2E</p><p>Type material. Holotype (Figs 1, 23): ♂, “N-Vietnam [ North Vietnam], 1400m, Mai-chau, Urwald [primary forest], 40 km SE Moc-chau, 20.50’ N 104.50 ’E, 07.–15.IV.1995, leg. Sinjaev &amp; EinhSamml. [local collectors]”, slide MWM 34524 Volynkin (MWM/ ZSM).</p><p>Paratypes: CHINA: 1 ♂, China, Guangxi, 35 km NNE Nanning, 10 km S Silong, 800 m, 25–26.III.1999, leg. G. Müller, slide MWM 34537 (♂) Volynkin (Coll. MWM/ ZSM) ; THAILAND: 5 ♂, 1 ♀, Thailand, Changwat Nan, 30 km E Pua, 1700 m, 22–23.IX.1999, leg. A. Szabó &amp; Z. Czere, slides MWM 34527 (♂), MWM 34553 (♂) Volynkin (Coll. MWM/ ZSM) ; 2 ♂, N Thailand, Chiang Mai, Fang, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.04222&amp;materialsCitation.latitude=19.903055" title="Search Plazi for locations around (long 99.04222/lat 19.903055)">Doi Ang Khang</a>, 1425m, 19°54’11’’N, 99°2’32’’E, 25–26.IV.2006, leg. K. Černý (Coll. CKC) ; 1 ♂, same locality, but 5. VI.2006, leg. T. Ihle (Coll. CKC); 8 ♂, 5 ♀, N Thailand, Chiang Mai, Fang, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.04222&amp;materialsCitation.latitude=19.903055" title="Search Plazi for locations around (long 99.04222/lat 19.903055)">Doi Ang Khang</a>, 1425m, 19°54’11’’N, 99°2’32’’E, 23–27.VII.2006, leg. T. Ihle, slides AV3814 ♂, AV3815 ♀, AV3816 ♀ Volynkin ( Coll. CKC) ; 1 ♀, N Thailand, Chiang Mai, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.915&amp;materialsCitation.latitude=18.8" title="Search Plazi for locations around (long 98.915/lat 18.8)">Doi Suthep</a>, 1085m, 18°48’N, 98°54’54’’E, 8. VI.2005, leg. K. Černý (Coll. CKC) ; 1 ♂, W Thailand, Tak, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.94006&amp;materialsCitation.latitude=16.75615" title="Search Plazi for locations around (long 98.94006/lat 16.75615)">Doi Mussoe</a>, 812 m, 16°45.369’N, 98°56.404’E, 18–21.IX.2007, leg. T. Ihle (Coll. CKC) ; LAOS: 1 ♂, Laos, prov. Luang Prabang, Pak Yaeng Noi, 1200 m, 25–27.V.2013, leg. T. Ihle (Coll. CKC) ; VIETNAM: 6 ♂, 2 ♀, same data as in the holotype, slides ZSM Arct. 119/2017 ♂, MWM 34525 (♀) Volynkin ( Coll. MWM/ ZSM) ; 1 ♂, North Vietnam, Tam Dao, 60 km NW Hanoi, 950 m (21.34’ N 105.20 ’E), 17.X.1994, leg. V. Sinjaev (Coll. MWM/ ZSM) ; 1 ♂, same locality, but 1–15.XI.1992, sec. forest, leg. Sinajev &amp; Simonov, ex coll. Schintlmeister, slide MWM 34531 (♂) Volynkin (Coll. MWM/ ZSM); 1 ♂, N Vietnam, 1600 m, Mt. Fan-si-pan (North), Cha-pa, primary forest (22.17’ N 103.44 ’E), 20– 30.IV.1995, leg. Sinjaev &amp; local collectors, slide MWM 34526 (♂) Volynkin (Coll. MWM/ ZSM) .</p><p>Diagnosis. The closest relative of the new species is C. moelleri (Figs 5–8) known from Nepal and northern and northeastern India. Cyana artemis sp. n. can be distinguished from it by its slightly broader transverse lines. In addition, the new species is somewhat larger than C. moelleri in general, and usually has somewhat larger black spots on forewing. Females of C. artemis also differ from those of C. moelleri by the stronger curved outer margin of the postmedial line. The genital capsules of the two species are very similar and the valva apex of C. artemis sp. n. (Figs 23, 24) is only narrower and the uncus is slightly longer than those of C. moelleri respectively (Fig. 25). However, the clear distinctive characters are expressed in the aedeagus and vesica structure. The aedeagus of C. artemis sp. n. is broader distally than that of C. moelleri, and its subapical area of short dentation is significantly broader. The vesica of C. artemis sp. n. is broader than that of C. moelleri, its dorsal diverticulum is much longer and broader and has several short but broad subdiverticula subbasally; in addition, vesica of C. artemis sp. n. bears two large, round clusters of numerous robust spinules, whereas in C. moelleri there are a long lateral band and a narrow subapical bunch of spinules. The female genitalia of C. artemis sp. n. (Fig. 35) differ from those of C. moelleri (Fig. 36) by the broader ductus bursae with shorter anterior sclerotised section, narrower sclerotised area of the posterior section of the corpus bursae, and slightly larger signum bursae.</p><p>Description. Adult (Figs 1–4). Forewing length 15–17 mm in males (16.5 mm in the holotype) and 17–18 mm in females. Antennae of both sexes weakly ciliate. Head white. Thorax white, with two large orange spots medially; tegulae and patagiae with broad orange margins. Abdomen white with slight admixture of ochreous scales distally. Forewing ground color white; transverse lines orange; subbasal line wavy, broadened at costa; antemedial line wavy medially; postmedial line angled and strongly narrowed in the cell; subterminal line broad, semilunar-like curved; male with two large black spots in the cell: round inner and reniform outer; two black strokes at costa opposite the R angle of the cell. Female with three round spots in the cell. Cilia white. Hindwing white; discal spot diffuse, gray, semilunar. Male genitalia (Figs 23, 24). Uncus dorso-ventrally flattened, short, narrowly arrowhead-shaped, its basal third connected to tuba analis. Tuba analis broad; scaphium not developed; subscaphium as broad setose field. Tegumen narrow, shorter than valvae. Juxta moderately sclerotised, trapezoidal, with round short lower concavity. Vinculum robust, U-shaped. Valva broad and elongated; costal margin slightly curved medially; valva apex broad, slightly narrowed distally and rounded apically; sacculus narrow, heavily sclerotised, its distal process robust, slightly curved dorsally, pointed apically, shorter than distal lobe of valva. Aedeagus large, strongly broadened distally, with heavily sclerotised subapical area bearing numerous small denticles. Vesica globular, with several diverticula: two short and membranous diverticula medially; long and curved, membranous dorsal diverticulum with a small subbasal subdiverticulum; globular granulated diverticulum with a small membranous subdiverticulum subapically; a round cluster of robust spinules subapically; and a round subbasal cluster of small spinules ventrally. Female genitalia (Fig. 35). Ovipositor short, broadly conical. Papillae anales trapezoidal with rounded angles. Apophyses thin; apophyses anteriores approximately two times shorter than apophyses posteriores. Ostium bursae broad. Postvaginal plate moderately sclerotised, lens-shaped, rounded anteriorly. Ductus bursae short, its posterior section membranous, and anterior section heavily sclerotised, with longitudinal wrinkles. Posterior section of corpus bursae narrowed, heavily sclerotised, wrinkled laterally at base of appendix bursae. Anterior section of corpus bursae broad, globular, weakly sclerotised, with round dentate signum medially. Basal section of appendix bursae narrow, heavily sclerotised, situated dorsally; distal section of appendix bursae membranous, saccate, situated laterally.</p><p>Distribution. Cyana artemis sp. n. is widely distributed in SE China (Guangxi Prov.), W and N Thailand (provinces Tak, Chiang Mai and Nan), N Laos (Luang Prabang Prov.) and N Vietnam (Hoa Binh, Lao Cai and Vinh Phuc). The record of C. moelleri for Thailand (Černý &amp; Pinratana 2009) belongs to C. artemis sp. n.</p><p>Etymology. In the ancient Greek mythology, Artemis is the goddess of the hunt, the wilderness, wild animals and chastity.</p></div>	https://treatment.plazi.org/id/03CF3F6EFFD7FFB0FF79F8A9FE93DDE6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Volynkin, Anton V.;Černý, Karel	Volynkin, Anton V., Černý, Karel (2019): Three new species of Cyana Walker, 1854 from Indochina and China (Lepidoptera, Erebidae, Arctiinae, Lithosiini). Zootaxa 4586 (2): 338-350, DOI: 10.11646/zootaxa.4586.2.8
03CF3F6EFFD5FFB1FF79FDB1FDE1DDC3.text	03CF3F6EFFD5FFB1FF79FDB1FDE1DDC3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cyana butyracea Volynkin & Černý 2019	<div><p>Cyana butyracea Volynkin &amp; Černý, sp. n.</p><p>(Figs 9, 10, 26, 27)</p><p>urn:lsid:zoobank.org:act: 37BA6B72-4CD0-4131-9B6F-BFFF92B5B2BF</p><p>Type material. Holotype (Figs 9, 26): ♂, “ Nord-Vietnam [North Vietnam], Tam Dao (Sek. Wald [secondary forest]), 60 km NW Hanoi, 1200 m (21.34’ N 105.20 ’E), 1.–5.V.1993, leg. Sinajev &amp; Simonov”, slide ZSM Arct. 112/2017 ♂ Volynkin (Coll. MWM/ ZSM).</p><p>Paratypes: VIETNAM: 1 ♂, same data as in the holotype ( Coll. MWM/ ZSM) ; 2 ♂, N Vietnam, 16[00]– 1800 m, Mt. Fan-si-pan ( West), Cha-pa, sec. forest, 22.20’ N 103.40 ’E (cult.) IV.1995, leg. Sinjaev &amp; local collectors (Coll. MWM/ ZSM) ; 2 ♂, S Vietnam, Kon Tum Prov., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=107.7&amp;materialsCitation.latitude=15.116667" title="Search Plazi for locations around (long 107.7/lat 15.116667)">Dac Glei</a>, 15°07’N 107°42’E, h= 700 m, 8.VIII.1996, leg. Siniaev &amp; Afonin (Coll. MWM/ ZSM) ; 4 ♂, South Vietnam, Bao Loc (sec. forest), Rung Cat Tien, 1500 m (11.32’ N 107.48 ’E), 10.–20.XII.1992, leg. Sinajev &amp; Simonov”, slide ZSM Arct. 113/2017 ♂ Volynkin (Coll. MWM/ ZSM).</p><p>Diagnosis. Cyana butyracea sp. n. (Figs 9, 10) is similar externally to the narrow-lined specimens of C. intercomma (widespread in Malay Peninsula, Thailand, Laos, Cambodia and Yunnan Prov. of China) (Černý &amp; Pinratana 2009; Bayarsaikhan &amp; Bae 2016) (Figs 11–16) and C. moelleri (Figs 5–8), but can easily be distinguished from them by its pale butter-yellow forewing coloration and small discal spot. The forewing pattern of the new species also resembles that of C. bacilla Fang, 1992 (Figs 17–18) known from SE China and Laos, but in the latter the forewing coloration is white, the forewing pattern elements are red (those are dark yellow in C. butyracea sp. n.), the antemedial line is straight (that is slightly wavy in C. butyracea sp. n.), the hindwing is pale red (that is pale butter-yellow in C. butyracea sp. n.), and the male genitalia structure is principally different (Figs 31, 38). The male genitalia structure of the new species (Figs 26, 27) is most similar to those of C. intercomma (Figs 28–30) by the shape of valva and the vesica structure, but differs by its slightly broader valva, slightly larger sclerotised crest of the medial costal plate of valva, and weaker subapical dentation of the aedeagus. In comparison to that of C. intercomma, the vesica of C. butyracea sp. n. lacks subapical and basal ventral clusters of spinules; in addition, the dorsal diverticulum of C. butyracea sp. n. bears longer cluster of spinules stretching along its inner margin, whereas in C. intercomma that is bunch-like and situated apically.</p><p>Description. Adult (Figs 9, 10). Forewing length 14–15 mm in males (14 mm in the holotype). Male antennae weakly ciliate. Head white. Thorax white, tegulae and patagiae encircled by orange hairs. Abdomen white with slight admixture of ochreous scales distally. Forewing ground color pale butter-yellow; transverse lines orange; subbasal line wavy, broadened at costa; antemedial line wavy; postmedial line oblique, strongly narrowed opposite the cell, with admixture of black scales near costa; subterminal line broad, crescent-like curved; two large black spots in the cell: round inner and reniform outer; orange stroke on costa opposite the outer reniform black spot. Cilia pale butter-yellow. Hindwing ligth butter-yellow; discal spot absent. Male genitalia (Figs 26, 27). Uncus dorso-ventrally flattened, short, narrowly arrowhead-shaped, its basal third connected to tuba analis. Tuba analis broad; scaphium not developed; subscaphium as broad setose field. Tegumen narrow, shorter than valvae. Juxta moderately sclerotised, trapezoidal, with round short lower concavity. Vinculum robust, U-shaped. Valva broad and elongated; costal margin slightly curved medially; medial costal plate with a long and very short plate-like sclerotised crest; valva apex broad and rounded; sacculus narrow, heavily sclerotised, its distal process robust, slightly curved dorsally, pointed apically, shorter than distal lobe of valva. Aedeagus tube-like, straight, with heavily sclerotised subapical area bearing numerous small denticles. Vesica globular, membranous, with several diverticula: four short and granulated diverticula medially; long and finger-shaped, membranous dorsal diverticulum with a long cluster of robust spinules laterally; in medial part of vesica a small spinula may be present.</p><p>Female unknown.</p><p>Distribution. Cyana butyracea sp. n. is to date, found in North and South Vietnam (Provinces Vinh Phuc, Lao Cai, Kon Tum and Dong Nai).</p><p>Etymology. ‘Butyracea’ means ‘buttery’ in Latin. The specific epithet refers to the unusual pale butter-yellow forewing coloration of the new species.</p></div>	https://treatment.plazi.org/id/03CF3F6EFFD5FFB1FF79FDB1FDE1DDC3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Volynkin, Anton V.;Černý, Karel	Volynkin, Anton V., Černý, Karel (2019): Three new species of Cyana Walker, 1854 from Indochina and China (Lepidoptera, Erebidae, Arctiinae, Lithosiini). Zootaxa 4586 (2): 338-350, DOI: 10.11646/zootaxa.4586.2.8
03CF3F6EFFD4FFB6FF79FDD5FD92DFDB.text	03CF3F6EFFD4FFB6FF79FDD5FD92DFDB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cyana kucheri Volynkin & Černý 2019	<div><p>Cyana kucheri Volynkin &amp; Černý, sp. n.</p><p>(Figs 19, 20, 32, 33, 39)</p><p>urn:lsid:zoobank.org:act: 5A5202E2-DB33-4AB8-9146-BDA4ED814923</p><p>Type material. Holotype (Figs 19, 32): ♂, “N-Vietnam [ North Vietnam], 1400 m, Mai-chau, Urwald [primary forest], 40 km SE Moc-chau, 20.50’ N 104.50 ’E, 07.–15.IV.1995, leg. Sinjaev &amp; EinhSamml. [local collectors]”, slide ZSM Arct. 78/2017 ♂ Volynkin (Coll. MWM/ ZSM).</p><p>Paratypes: 17 ♂, 1 ♀, same data as in the holotype, slides ZSM Arct. 77/2017 ♂, ZSM Arct. 79/2017 ♀, Volynkin (Coll. MWM/ ZSM) .</p><p>Diagnosis. Cyana kucheri sp. n. (Figs 19, 20) is an unmistakable species characterized by its forewing pattern resembling that of C. artemis sp. n. (Figs 1–4), C. moelleri (Figs 5–8) and C. intercomma (Figs 11–16), and hindwing being pale rosy orange. The male genital capsule of C. kucheri sp. n. (Figs 32, 33) differs clearly from those of C. artemis sp. n. (Figs 23, 24), C. moelleri (Fig. 25) and C. intercomma (Figs 28–30) by its more elongated and narrower valva with a longer and strongly curved distal saccular process. The vesica structure of the new species is more similar to that of C. intercomma by presence of the elongated dorsal diverticulum bearing a bunch of spinules apically, but in C. kucheri sp. n. that diverticulum is longer and the spinules on its apex are much weaker than those in C. intercomma; distal cluster of spinules is broad and round in C. kucheri sp. n., whereas in C. intercomma that is band-like; vesica of C. kucheri sp. n. has a broad subbasal ventral diverticulum with a broad cluster of short and thin spinules, whereas vesica of C. intercomma lacks a subbasal ventral diverticulum but has a basal dorsal bunch of robust spinules. In addition, the aedeagus of C. intercomma has a heavily sclerotised subapical transverse band covered with small but robust dens, whereas in C. kucheri sp. n. the aedeagus lacks subapical sclerotization. The male genitalia structure of C. kucheri sp. n. (Figs 32, 33) is most similar to that of C. gazella (Fig. 34). The genital capsule of the new species differs from that of C. gazella by its less robust vinculum, narrower and slightly longer valva with a smaller medial crest, and more robust distal saccular process. Compared to that of C. gazella, the vesica of C. kucheri sp. n. has a much broader medial section with a much broader distal diverticulum; dorsal diverticulum is much narrower and longer in C. kucheri sp. n. and bears a narrower cluster of thinner spinules than that of C. gazella; in addition, in C. kucheri sp. n. the subapical cluster of spinules is slightly broader and the ventral subbasal cluster of spinules is much larger than those of C. gazella . The female genitalia of C. kucheri sp. n. (Fig. 39) differ clearly from those of C. artemis sp. n. (Fig. 35), C. moelleri (Fig. 36) and C. intercomma (Fig. 37) by their narrow ductus bursae without sclerotization, much smaller and strongly rugose sclerotised posterior section of corpus bursae, elliptical signum bursae (those are round in the three other species), and much larger, elongated and curved appendix bursae, whereas in the three other species appendices bursae are much smaller and globular or saccate. The female genitalia structure of C. kucheri sp. n. (Fig. 39) is most similar to that of C. gazella (Fig. 40). However, it can be easily distinguished by its much larger appendix bursae and longer signum bursae; in addition, the corpus bursae of C. kucheri sp. n. is weakly sclerotised, while that of C. gazella has a heavily sclerotised area laterally.</p><p>Description. Adult (Figs 19, 20). Forewing length 16.5– 17 mm in males (16.5 mm in the holotype) and 20 mm in female. Antennae of both sexes ciliate. Head white. Thorax white, tegulae and patagiae with broad orange margins. Abdomen pale red with slight admixture of ochreous scales. Forewing ground color white with slight admixture of pale red scales. Transverse lines orange; subbasal line broad, slightly wavy, broadened at costa; antemedial line broad, its outer margin straight, while inner margin with a broad concavity in the cell. Postmedial line broad posteriorly and strongly broadened in the cell and at costa. Subterminal line short, broadly crescent-like, expressed only in the anterior part of wing. Two large black spots in the cell: inner spot reniform, outer spot rounded, situated more posteriorly. Black stroke on postmedial line near costa. Hindwing pale rosy orange, discal spot absent. Male genitalia (Figs 32, 33). Uncus short, narrow, distally tapered, its basal half connected to tuba analis. Tuba analis broad, subscaphium as a broad setose field. Tegumen narrow, shorter than valva. Juxta trapezoidal, with deep trigonal lower concavity. Vinculum short but robust, U-shaped. Valva elongated, broadened medially, its costal margin strongly curved medially; distal section of valva strongly elongated, slightly narrowed distally, rounded apically. Sacculus heavily sclerotised, narrow; distal saccular process robust, somewhat shorter than distal section of valva, slightly curved dorsally, pointed apically. Aedeagus elongated, broadened medially, narrowed apically. Medial part of vesica globular; subbasal dorsal diverticulum as long as aedeagus, slightly narrowed distally, bears a cluster of numerous small spinules; two globular granulated diverticula medially; subbasal cluster of small spinules ventrally; broad, round cluster of large spinules subapically. Female genitalia (Fig. 39). Ovipositor short, broadly conical. Papillae anales trapezoidal with rounded angles. Apophyses thin; apophyses anteriores approximately two times shorter than apophyses posteriores. Ostium bursae moderately broad. Ductus bursae short, membranous, wrinkled anteriorly. Posterior section of corpus bursae conically narrowed, heavily sclerotised. Anterior section of corpus bursae globular, membranous, with a large, elliptical, weakly dentate signum. Basal section of appendix bursae broadly conical, heavily sclerotised, strongly wrinkled; distal part of appendix bursae membranous, elongated, much longer than corpus bursae, crescent-like curved, connected to the sclerotised basal part by a narrow membranous isthmus.</p><p>Distribution. Cyana kucheri sp. n. is to date, only known from its type locality, Mai Chau district of the Hoa Binh Province in the northern Vietnam.</p><p>Etymology. The species name is dedicated to Artem A. Kucher (Barnaul, Russia), an excellent veterinarian and a senior author’s family friend.</p></div>	https://treatment.plazi.org/id/03CF3F6EFFD4FFB6FF79FDD5FD92DFDB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Volynkin, Anton V.;Černý, Karel	Volynkin, Anton V., Černý, Karel (2019): Three new species of Cyana Walker, 1854 from Indochina and China (Lepidoptera, Erebidae, Arctiinae, Lithosiini). Zootaxa 4586 (2): 338-350, DOI: 10.11646/zootaxa.4586.2.8
