identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03CFA3486B67FFC1FF1EFBBA5EE0F9C0.text	03CFA3486B67FFC1FF1EFBBA5EE0F9C0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala coffea Filippini, Galante, and Mico	<div><p>Anomala coffea Filippini, Galante, and Micó, new species</p> <p>(Figs. 1, 9, 17, 25, 34)</p> <p>Type Material. Holotype: ♂ "Est. Pitilla, 700 m, 9 km S Sta. Cecilia, P. N. Guanacaste, Prov. Guanacaste, Costa Rica, C. Moraga 8 a 24 ago 1991, L-N 330200,380200 / INBIOCRI000409517" deposited at MNCR. Paratypes (6): 1♀ " <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-85.42777&amp;materialsCitation.latitude=10.990556" title="Search Plazi for locations around (long -85.42777/lat 10.990556)">Estac. Pitilla</a>, 700 m, 9 Km S. Santa Cecilia, Guanac. Pr. COSTA RICA Jul 1988 GNP Biodiversity Survey W85 25 40", N10 59 26" / INB0003315377"; 1♂ " Est. Pitilla, 700 m, 9 km S Sta. Cecilia, P. N. Guanacaste, Prov. Guan. COSTA RICA, Tp Malaise, 1990. L N 330200,380200 / INBIOCRI000304012"; 1♂ " Est. Pitilla, 700 m, 9 km S Sta. Cecilia, P. N. Guanacaste, Prov. Guanacaste, Costa Rica, C. Moraga, 2 a 15 may 1992, L N 330200,380200 / INBIOCRI000405396"; 1♂ " Est. Pitilla, 700 m, 9 km S Sta. Cecilia, P. N. Guanacaste, Prov. Guanacaste, Costa Rica, K. Taylor, 31 mar - 29 abr 1992, L- N 330200,380200 / INBIOCRI000523773"; 1♀ " Est. Pitilla, 700 m, 9 km S Sta. Cecilia, P. N. Guanacaste, Prov. Guan. COSTA RICA, C. Moraga, 6-28 Ene</p> <p>1992, L- N 330200,380200 / CEUA00106195 "; 1♂ "Est. Pitilla, 700 m, 9km S Sta. Cecilia, P.N. Guanacaste, Prov. Guanacaste, Costa Rica, 22 oct a 2 nov 1992, C. Moraga L-N 330200,380200 / CEUA00106194 ". Four paratypes are deposited at MNCR, and two paratypes are deposited at Universidad de Alicante.</p> <p>Holotype. Male (Fig. 1). Body oval. Length 8.99 mm, width 5.19 mm. Head and pronotum dark brown. Scutellum reddish brown. Elytra dark brown but lighter than pronotum. Pygidium ochre. Legs orange on femur, reddish brown on tibia and tarsus. Venter orange on sternum, abdominal sternites ochre. Head: Clypeus trapezoidal, short, surface densely punctate-reticulate, convex; anterior margin curved, clypeus thick in frontal view; width/length ratio 2.50. Frons sparsely punctate, flat. Ocular canthus long, thin, with rounded apex. Ratio of interocular width/width of eye 2.84. Ratio antennal funiculus/club 0.74. Pronotum: Subtrapezoidal. Ratio width/length 1.69. Lateral margins angulated at 1/3 of pronotal length from the anterior margins. Anterior angles acute, posterior angles obtuse. Basal margin sinuate, subapical bead obliterated at middle. Surface with deep, dense punctures. Scutellum: Subtriangular, with rounded sides, and acute apex. Ratio width/length 1.19. Surface sparsely punctate. Elytra: With costae defined by regular rows of punctures. Subsutural interstice with 1–2 rows of sparse punctures, 2 nd –4 th interstices with 1 irregular, secondary stria. Pygidium: Sparsely punctate with long setae on posterior margin. Ratio width/length 1.77. Venter: Space between mesocoxae narrow, flat. Mesometasternal suture well-defined at base of mesocoxae. Abdominal sternites with 1 row of setae and 4–5 rows of punctures per sternite. Last sternite strigate, with subapical bead thin and sinuate, apical membrane thin. Legs: Protibia (Fig. 9) tridentate, apical tooth long and curved, middle tooth above internal apex of protibia, triangular in shape, acute, basal tooth sharp. Metatibia stout, fusiform; ratio length/width 2.81. First external carina with row of sparse setae, surface punctate. External protarsal claw strongly curved, internal claw bifurcate, with upper branch slightly shorter and 2/3 width of the lower one, inferior margin curved. Male genitalia: Parameres slender (Fig. 17), with squared, protruding apex in lateral view, ventral margin sinuate. Endophallus (Fig. 25) with 1 long, glabrous sacculus. Ejaculatory duct opening frontally on a ventral inflation.</p> <p>Female. Similar to male except protibia (Fig. 9) with wider teeth; last sternite with curved apex.</p> <p>Variation. Six paratypes. Dorsal surface varies from dark reddish brown to dark brown; pygidium and venter from orange to ochre. Body length 8.99–10.17 mm, body width 5.19–5.76 mm. Clypeal width/length ratio 2.50–2.93. Ratio of interocular</p> <p>6) A. robiginosa; 7) A. strigodermoides; 8) A. unilineata. Scale bar = 5 mm.</p> <p>width/width of eye 2.84–3.19. Ratio of antennal funiculus/club 0.73–0.79. Pronotal width/length ratio 1.69–1.72. Scutellar width/length ratio 1.13–1.33. Pygidial width/length ratio 1.68–1.82. Metatibial length/width ratio 2.60–2.81.</p> <p>Etymology. From the generic name of coffee plants, to be treated as a noun in apposition, for its resemblance in shape and color to toasted coffee beans, a famous Costa Rican product.</p> <p>Distribution. This species is found in the humid forest on the northwest side of the Guanacaste mountain range, Costa Rica, at 700 m elevation (Fig. 34).</p> <p>Diagnosis. This species is recognized by its small size, color homogeneously dark brown, elytra with defined costae and interstices with rows of punctures, and tridentate protibia. The small size and brown color is similar to that of species of Anomala in the subgenus Bucaphallanus Ramírez-Ponce and Morón, 2012, but A. coffea does not share any diagnostic character with any of its species and has more defined punctation on the elytra.</p> </div>	https://treatment.plazi.org/id/03CFA3486B67FFC1FF1EFBBA5EE0F9C0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Filippini, V.;Galante, E.;Micó, E.	Filippini, V., Galante, E., Micó, E. (2015): Descriptions of New Species ofAnomalaSamouelle (Coleoptera: Scarabaeidae: Rutelinae) from Costa Rica. The Coleopterists Bulletin 69 (3): 463-476, DOI: 10.1649/0010-065x-69.3.463, URL: http://dx.doi.org/10.1649/0010-065x-69.3.463
03CFA3486B64FFC4FD74F9D8586CFC92.text	03CFA3486B64FFC4FD74F9D8586CFC92.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala cyclops Filippini, Galante and Mico	<div><p>Anomala cyclops Filippini, Galante and Micó, new species</p> <p>(Figs. 2, 10, 18, 26, 35)</p> <p>Type Material. Holotype: ♂ "Cerro El Hacha,</p> <p>300 m, 12 km SE de La Cruz, Prov. Guanacaste,</p> <p>Costa Rica, 25 jun 1992, III curso Parataxon. L-N 329200,368000 / INBIOCRI000755645" deposited at MNCR. Paratypes (9): 1♂ "Est. Sta. Rosa, 300 m, P. N. Sta. Rosa, Prov. Guanacaste, Costa Rica, 3 a 12 jun 1992, III Curso Parataxon. L- N 313000,359800" / INBIOCRI000458888"; 1♀ "Est. Sta. Rosa, 300 m, P. N. Guanacaste, Prov. Guan, COSTA RICA. D. H. Janzen &amp; W. Hallwachs, Jun 1991, L- N 313000,359800 / INBIOCRI 000590279; 1♂ 1♀ "Est. Sta. Rosa, 300 m, P. N. Sta. Rosa, Prov. Guanacaste, Costa Rica, 1 a 12 jun 1992, A. M. Mora, L-N 313000,359800" / INBIOCRI000912716 and CEUA00106197; 1♀ "Los Almendros, P. N. Guanacaste, Prov. Guana., COSTA RICA. 12–31 May 1993, E. Lopez, L- N 334800, 369800 / INBIOCRI001168088"; 1♂ "Fca. Jenny, 30 km N de Liberia, P. N. Guanacaste, Prov. Guan., COSTA RICA. 25–29 May 1993. E. Araya. L-N 316200,364400 / INBIOCRI001183834"; 2♂ "Finca Jenny, 30 km N de Liberia, P. N. Guanacaste, Prov. Guan., COSTA RICA. 20–24 May 1993. E. Araya. L-N 316200,364400" / INBIOCRI 001184178 and CEUA00106196; 1♀ "Est. Murcielago, 8 km SO. de Cuajiniquil, P. N. Guanacaste, Prov. Guana, COSTA RICA. 100 m. 3 May 1994, C. Cano, L N 320300_347200 # 2907 / INBIOCRI001891273". Seven paratypes are deposited at MNCR, and two paratypes are deposited at Universidad de Alicante.</p> <p>Holotype. Male (Fig. 2). Body oval. Length 9.82 mm, width 5.09 mm. Head reddish brown. Pronotum ochre with a pentagonal, dark brown macula on disc, with apex pointing posteriorly,</p> <p>and a small macula on lateral foveae. Scutellum dark brown. Elytra ochre with dark brown sides, expanding at base, on median and apical parts of suture. Pygidium and venter ochre. Legs ochre with reddish tarsi and apical part of tibiae. Head: Clypeus trapezoidal, surface densely punctatereticulate, flat. Anterior margin straight, clypeus thin in frontal view, width/length ratio 2.42. Frons punctate, less densely than clypeus, with a triangular concavity. Ocular canthus long, thin, with rounded apex. Ratio of interocular width/width of eye 2.9. Ratio antennal funiculus/club 0.85. Pronotum: Subtrapezoidal. Ratio width/length 1.75. Lateral margins angulated at 1/3 of pronotal length from the anterior margins. Anterior angles quadrate, posterior angles obtuse. Basal margin sinuate, subapical bead complete. Surface with deep, dense punctures. Scutellum: Subtriangular, with rounded sides and acute apex. Ratio width/ length 1.4. Surface sparsely punctate. Elytra: With costae defined by regular rows of punctures. Subsutural interstice with 2 rows of sparse punctures, 2 nd –4 th interstices with 1 irregular, secondary stria. Pygidium: Sparsely punctate with long setae on posterior margin. Ratio width/length 1.52. Venter: Space between mesocoxae narrow, slightly convex. Mesometasternal suture welldefined at base of mesocoxae. Abdominal sternites</p> <p>35) Distribution of A. cyclops, A. mesosticta, and A. strigodermoides; 36) Distribution of A. divisa and A. unilineata.</p> <p>with 1 row of setae and 3–5 rows of punctures per sternite. Last sternite strigate, with subapical bead thin and sinuate, apical membrane thin. Legs: Protibia (Fig. 10) tridentate, apical tooth long and curved, middle tooth below internal apex of protibia, triangular in shape, acute, basal tooth sharp. Metatibia fusiform. Ratio length/width 3.16. First external carina well-developed, surface punctate. Internal protarsal claw bifurcate, with upper branch slightly shorter and half width of the lower one, inferior margin sinuate. Male genitalia: Parameres with wide apex in lateral view (Fig. 18), ventral margin sinuate. Endophallus (Fig. 26) with 1 long, glabrous sacculus. Ejaculatory duct opening frontally on a lateral inflation.</p> <p>Female. Similar to male except protibia (Fig. 10) with wider teeth; last sternite with curved apex.</p> <p>Variation. Nine paratypes. Head varies from orange to dark reddish brown; presence of a complete or incomplete reddish brown transversal band on elytra or median expansion on suture; pygidium and venter vary from ochre to light brown. Body length 8.44–10.29 mm, body width 4.41–5.37 mm. Clypeal width/length ratio 2.11–2.54. Ratio of interocular width/width of eye 2.50–2.90. Ratio antennal funiculus/club 0.67–0.85. Pronotal width/ length ratio 1.67–1.78. Scutellar width/length ratio 1.10–1.43. Pygidial width/length ratio 1.48–1.68. Metatibial length/width ratio 2.88–3.28.</p> <p>Etymology. From the Latin noun Cyclops, -opis, a one-eyed mythological creature, to be treated as a noun in apposition, for the single macula on the pronotum.</p> <p>Distribution. Anomala cyclops inhabits the dry forest in northeastern Costa Rica, from 100 to 300 m elevation (Fig. 35).</p> <p>Diagnosis. This species is recognized by its medium–small size, ochre color with a median dark macula on the pronotum and a transverse band on the elytra, tridentate protibia, parameres with a wide apex in lateral view and sinuate ventral margin. Species with a similar pronotal pattern and size are Anomala boliviana Ohaus, 1897 (Bolivia), Anomala flavilla centralis Bates, 1888 (Mexico), Anomala centralis LeConte, 1863 (Mexico), and Anomala chevrolati Bates, 1888 (Mexico). However, all these species have a bidentate protibia, whereas A. cyclops has a tridentate protibia. Moreover, A. boliviana has maculae on the elytra and pygidium, ochre elytral suture, first interstice of elytra wider with 3–4 irregular rows of punctures, and a larger pronotal macula. Anomala flavilla centralis has only the elytral suture pigmented, and the parameres are long with an acute and curved apex and the ventral margin acute and pointing posteriorly in lateral view. Anomala centralis has a triangular pronotal macula, ochre elytral suture, and elytra with darker shading on the lateral margins. Anomala chevrolati has an irregular reddish pronotal macula, a wide, dark elytral suture homogeneously wide on the apical half, denser punctures on the elytra, and finer secondary punctures on the elytral surface. Anomala bruchiana Ohaus, 1911 has a tridentate protibia and a small pronotal macula, but it has a triangular shape, ochre elytra with a narrow, dark elytral suture, and slender, long parameres with an acute apex and curved ventral margin. Anomala hoepfneri Bates, 1888 has a similar elytral pattern but with a large, irregular pronotal macula, bidentate protibia, and wide and densely punctured elytral interstices.</p> </div>	https://treatment.plazi.org/id/03CFA3486B64FFC4FD74F9D8586CFC92	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Filippini, V.;Galante, E.;Micó, E.	Filippini, V., Galante, E., Micó, E. (2015): Descriptions of New Species ofAnomalaSamouelle (Coleoptera: Scarabaeidae: Rutelinae) from Costa Rica. The Coleopterists Bulletin 69 (3): 463-476, DOI: 10.1649/0010-065x-69.3.463, URL: http://dx.doi.org/10.1649/0010-065x-69.3.463
03CFA3486B61FFCBFF1CFC965863FDA1.text	03CFA3486B61FFCBFF1CFC965863FDA1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala divisa Filippini, Galante, and Mico	<div><p>Anomala divisa Filippini, Galante, and Micó, new species</p> <p>(Figs. 3, 11, 19, 27, 36)</p> <p>Type Material. Holotype: ♂ " Alajuela. Costa Rica Cinco Esquinas de Carrizal, 1800m 16 abril de 1988 Col: G. Barrantes / INBIOCRI002517128" deposited at MNCR. Paratypes (6): 2♂ " Zarcero, Alfaro Ruiz, 1700m Alajuela Pr. COSTA RICA. Apr - May 1989. A. Del Valle &amp; A. Solis, L- N 240500, 493500" / INBIOCRI000022463 and CEUA00106199; 3♂ 1♀ " Est. Biol. Las Alturas 1500 m, Coto Brus, Prov. Puntarenas, Costa Rica, F. Araya, 23 mar a 2 may 1992, L- S 322500,591300" / CEUA00106198, INBIOCRI000909434, INBIO CRI000916198 and INBIOCRI000909406. Four paratypes are deposited at MNCR, and two paratypes are deposited at Universidad de Alicante.</p> <p>Holotype. Male (Fig. 3). Body oval. Length 12.87 mm, width 6.47 mm. Head reddish brown with ochre apical fringe and clypeus. Pronotum ochre with 2 large, trapezoidal, reddish brown maculae with bronze luster touching anterior and posterior margins. Scutellum ochre with thin, reddish sides. Elytra ochre with reddish suture. Pygidium ochre. Legs ochre with light to reddish brown apical part of tibia and tarsus. Venter reddish to dark brown, sternum with sides ochre. Head: Clypeus trapezoidal, anterior angles widely curved, surface densely punctate-reticulate, convex; anterior margin straight, thin in frontal view, ratio width/length 1.99. Frons densely punctate, flat. Ocular canthus long, thin, with acute apex. Ratio of interocular width/width of eye 3.52. Ratio antennal funiculus/club 0.79. Pronotum: Subtrapezoidal. Ratio width/length 1.81. Lateral margins angulated at 1/3 of pronotal length from the anterior margins. Anterior angles quadrate, posterior angles quadrate. Basal margin sinuate, subapical bead complete. Surface with deep, dense punctures. Scutellum: Subtriangular, with rounded sides and acute apex. Ratio width/length 1.56. Surface sparsely punctate. Elytra: Costae defined by regular rows of punctures. Subsutural interstice with 2–4 rows of sparse punctures, 2 nd –4 th interstices with 1–2 irregular, secondary stria(e). Surface covered with secondary punctures. Pygidium: Finely granulated with long setae on posterior margin. Ratio width/length 1.57. Venter: Space between mesocoxae narrow, slightly convex. Mesometasternal suture well-defined at base of mesocoxae. Abdominal sternites with 1 row of setae (2 rows on second sternite) and 5–7 rows of punctures per sternite. Last sternite strigate, with subapical bead thin and sinuate, apical membrane wide. Legs: Protibia (Fig. 11) bidentate, apical tooth long and curved, basal tooth below internal apex of protibia triangular, acute. Metatibia slender, fusiform, length/width ratio 3.39. First external carina with a row of sparse setae, surface punctate. Internal protarsal claw bifurcate, with upper branch of same length and half width of the lower one; inferior margin with a sharp angle at half length. Male genitalia: Parameres with blunt apex in lateral view (Fig. 19), ventral angle defined. Endophallus (Fig. 27) with 1 ventral and 1 dorsal inflation. On dorsal inflation, a long curved, claw and a patch of sclerotized, thick setae; ejaculatory duct opening located on ventral inflation. Median lobes very thin and long, a dorsally located sclerotized plate between their apices.</p> <p>Female. Similar to male except protibia (Fig. 11) with wider teeth; internal protarsal claw narrower with curved inferior margin; last sternite with curved apex.</p> <p>Variation. Six paratypes. Maculae on pronotum can fuse into one large macula with a bronze or green luster. Venter of female specimen is ochre. Claw on endophallus can have a bifurcated tip. Body length 12.04–13.56 mm, body width 6.47–7.18 mm. Clypeal width/length ratio 1.85–2.01. Ratio of interocular width/width of eye 3.42–3.93. Ratio of antennal funiculus/club 0.70–0.79. Pronotal width/length ratio 1.79–1.87. Scutellar width/length ratio 1.22–1.56. Pygidial width/length ratio 1.57–1.66. Metatibial length/ width ratio 3.16–3.61.</p> <p>Etymology. From the past participle of the Latin verb divido, to divide. The ochre median line on the pronotum and dark suture on the elytra visually divide in half the dorsal aspect of this species.</p> <p>Distribution. Anomala divisa occurs in the Central and Talamanca mountain ranges of Costa Rica, from 1,500 to 1,800 m elevation (Fig. 36).</p> <p>Diagnosis. This species is recognized by the presence of two large, trapezoidal, dark maculae on the pronotum and the parameres with blunt apex and defined ventral angle. Similar species, also present in Costa Rica, are Anomala jansoni Ohaus, 1897 and Anomala quiche Ohaus, 1897. The former species is slightly larger, with smaller maculae on the pronotum and long, slender, curved parameres in lateral view. The latter species has smaller and more separated maculae on the pronotum, maculae on the elytral calli, and parameres with a wider and blunter apex (aedeagi illustrated in Filippini et al. 2015a).</p> </div>	https://treatment.plazi.org/id/03CFA3486B61FFCBFF1CFC965863FDA1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Filippini, V.;Galante, E.;Micó, E.	Filippini, V., Galante, E., Micó, E. (2015): Descriptions of New Species ofAnomalaSamouelle (Coleoptera: Scarabaeidae: Rutelinae) from Costa Rica. The Coleopterists Bulletin 69 (3): 463-476, DOI: 10.1649/0010-065x-69.3.463, URL: http://dx.doi.org/10.1649/0010-065x-69.3.463
03CFA3486B6EFFCAFF08FD66580CFE9B.text	03CFA3486B6EFFCAFF08FD66580CFE9B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala mersa Filippini, Galante, and Mico	<div><p>Anomala mersa Filippini, Galante, and Micó, new species</p> <p>(Figs. 4, 12, 20, 28, 34)</p> <p>Type Material. Holotype: ♂ " COSTA RICA, Prov. Guanacaste, Bagaces, P.N. Palo Verde, Sector Palo Verde. 0–50m. 29 NOV-15 DIC 1999. W. Porras. Manual L_N_260952_385020 #57243 / INB0003151362" deposited at MNCR. Paratypes (4): 4♂ " COSTA RICA, Prov. Guanacaste, Bagaces, P.N. Palo Verde, Sector Palo Verde. 0–50m. 29 NOV-15 DIC 1999. W. Porras. Manual L_N_ 260952_385020 #57243" / CEUA00106201, INB0003151360, INB0003151358 and INB000 3151412. Three paratypes are deposited at MNCR and one at Universidad de Alicante.</p> <p>Holotype. Male (Fig. 4). Body oval. Length 9.31 mm, width 4.73 mm. Head reddish brown. Pronotum dark brown with wide, ochre sides, with green luster. Scutellum brown. Elytra dark brown with large ochre maculae on basal part and 2 small, ochre maculae at apex. Pygidium light brown with ochre apex and sides at base. Legs ochre with reddish brown tarsus, metatibia, and apical part of pro- and mesotibiae. Venter ochre on sternum, with darker median suture, and reddish brown abdominal sternites. Head: Clypeus trapezoidal, surface densely punctate-reticulate, flat. Anterior margin straight, clypeus thin in frontal view, width/ length ratio 1.98. Frons densely punctate, convex on suture with clypeus. Ocular canthus long, thin, with rounded apex. Ratio of interocular width/ width of eye 3.46. Ratio antennal funiculus/club 0.71. Pronotum: Subtrapezoidal, basal portions of lateral margins nearly parallel. Ratio width/ length 1.66. Lateral margins angulated at 1/3 of pronotal length from the anterior margins. Anterior angles quadrate, posterior angles quadrate. Basal margin sinuate, subapical bead complete. Surface with dense punctures. Scutellum: Subtriangular, with rounded sides and acute apex. Ratio width/ length 1.24. Surface densely punctate. Elytra: With striae defined by regular rows of punctures. Subsutural interstice with 1–2 rows of sparse punctures, 2 nd –4 th interstices with 1 irregular, secondary stria. Surface covered with secondary punctures. Pygidium: Strigate with long setae on posterior margin. Ratio width/length 1.60. Venter: Space between mesocoxae narrow, slightly convex. Mesometasternal suture well-defined at base of mesocoxae. Abdominal sternites with 1 row of setae and 5–6 rows of punctures per sternite. Last sternite strigate, with subapical bead thin and sinuate, apical membrane thin. Legs: Protibia (Fig. 12) bidentate, apical tooth long and curved, basal tooth below internal apex of protibia triangular, sharp. Metatibia slightly narrower subapically. Ratio length/ width 2.92. First external carina with a row of sparse setae. Surface rugose. Internal protarsal claw bifurcate, with upper branch slightly shorter and 2/3 width of lower one, inferior margin curved. Male genitalia: Parameres with blunt apex in lateral view (Fig. 20), ventral angle acute. Endophallus (Fig. 28) with a long sacculus covered with short setae, a patch of sclerotized spines at base. Ejaculatory duct opening on lateral inflation.</p> <p>Female. Unknown.</p> <p>Variation. Four paratypes. Posterior margin of pronotum dark or with two ochre median maculae, size of ochre maculae on elytra variable, pygidium varies from ochre to ochre with light brown maculae on lateral depressions to light brown; venter varies from ochre with brown bands on abdominal sternites to mainly brown. Body length 8.93–9.31 mm, body width 4.73–5.00 mm. Clypeal width/length ratio 1.80–2.02. Ratio of interocular width/width of eye 3.13–3.61. Ratio of antennal funiculus/club 0.57–0.71. Pronotal width/length ratio 1.66–1.70. Scutellar width/length ratio 1.21–1.37. Pygidial width/length ratio 1.54–1.62. Metatibial length/ width ratio 2.82–3.11.</p> <p>Etymology. From the past participle of the Latin verb mergo, dipped, for the black apical portion of the elytra, which appears as if it was dipped into black dye.</p> <p>Distribution. Anomala mersa inhabits the dry forest of Parque Nacional Palo Verde on the northern side of Golfo de Nicoya, Costa Rica, from 0 to 50 m elevation (Fig. 34).</p> <p>Diagnosis. This species is recognized by its small size, elytra dark brown with large, ochre maculae on the basal part, and parameres with blunt apex in lateral view and ventral angle acute. The particular elytral pattern easily sets this species apart from other Anomala. Species with similar patterns are dark specimens of Anomala pseudoeucoma Filippini, Micó, and Galante, 2013 (Costa Rica) and Anomala nigrosellata Ohaus, 1905 (Venezuela), which are much larger in size. The former species has the entire surface covered in dense setae and longer parameres with an acute apex and sinuate ventral margin. The latter species has a squared, dark macula on the elytra, hence the lateral elytral margins are ochre, coarser punctures on the elytra, and narrow parameres with a sinuate ventral margin.</p> </div>	https://treatment.plazi.org/id/03CFA3486B6EFFCAFF08FD66580CFE9B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Filippini, V.;Galante, E.;Micó, E.	Filippini, V., Galante, E., Micó, E. (2015): Descriptions of New Species ofAnomalaSamouelle (Coleoptera: Scarabaeidae: Rutelinae) from Costa Rica. The Coleopterists Bulletin 69 (3): 463-476, DOI: 10.1649/0010-065x-69.3.463, URL: http://dx.doi.org/10.1649/0010-065x-69.3.463
03CFA3486B6FFFC9FF13FE6F5B5EFE9B.text	03CFA3486B6FFFC9FF13FE6F5B5EFE9B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala mesosticta Filippini, Galante, and Mico	<div><p>Anomala mesosticta Filippini, Galante, and Micó, new species</p> <p>(Figs. 5, 13, 21, 29, 35)</p> <p>Type Material. Holotype: ♂ " Los Arbolitos, Prov. Hered. COSTA RICA. 30m. 20–27 MAR 1993. F. Araya. L_N_291400_536100 #1952 / INBIOCRI 001677111" deposited at MNCR. Paratypes (9): 2♂ "La Virgen de Sarapiquí, prov. Here., COSTA RICA. 9 a 30 mar 1993. M. Ortiz. L_N_263950_521050" / INBIOCRI001299837 and CEUA00106193; 2♂ " Amubri, Prov. Limon, COSTA RICA. 70 m. 08–27 nov 1993, G. M. Gallardo, L S 385500_ 578100 # 2461" / CEUA00106192 and INBIOCRI 001957606; 1♀ " Limon, Costa Rica Reserva Hitoy- Cerere Rio Cerere 100 msnm 24 marzo 1987 Angel Solis / INBIOCRI002517679"; 1♀ " Rio Sardinas, 10 m, R.N.F.S. Barra del Colorado, Prov. Limon, COSTA RICA. 18 a 30 feb 1993. F. Araya, L-N- 291500, 564700 / INBIOCRI001297396"; 1♂ "R.B. Hitoy Cerere, Valle La Estrella, Prov. Limon, COSTA RICA. 100– 200m. 9–14 May 1994, G. Carballo, L N 643400_184600 # 2856 / INBIOCRI001798864"; 1♂ " Amubri, Prov. Limon, COSTA RICA. 70 m. 12–31 Oct 1993. G. Gallardo, L S 385500_578000 # 2407 / INBIOCRI001644531"; 1♂ " Cuatro Esquinas, P. N. Tortuguero, Prov. Limon, COSTA RICA. 0 m, April 1989, R. Aguilar &amp; J. Solano, L- N 280000_ 590500 / INBIOCRI000084822". Seven paratypes are deposited at MNCR, and two paratypes are deposited at Universidad de Alicante.</p> <p>Holotype. Male (Fig. 5). Body oval. Length 10.88 mm, width 5.52 mm. Head reddish brown. Pronotum dark brown on disc, with sides and basal median part ochre. Scutellum dark brown with reddish sides. Elytra ochre with dark brown margins, expanding on median and apical parts of suture; punctures of striae pigmented. Pygidium, legs, and venter ochre. Head: Clypeus trapezoidal, surface densely punctate-reticulate, convex. Anterior margin straight, clypeus thin in frontal view, width/length ratio 2.24. Frons punctate, less densely than clypeus, with a triangular concavity. Ocular canthus long, thin, with rounded apex. Ratio of interocular width/width of eye 2.99. Ratio antennal funiculus/ club 0.73. Pronotum: Subtrapezoidal. Ratio width/ length 1.7. Lateral margins angulated at 1/3 of pronotal length from the anterior margins. Anterior angles quadrate, posterior angles obtuse. Basal margin sinuate, subapical bead complete. Surface with deep dense punctures. Scutellum: Subtriangular, with rounded sides and acute apex. Ratio width/length 1.38. Surface densely punctate. Elytra: With costae defined by regular rows of punctures. Subsutural interstice with 2–4 rows of sparse punctures, 2 nd –4 th interstices with 1 irregular, secondary stria. Surface covered with secondary punctures. Pygidium: With coalescing punctures and long setae on posterior margin. Ratio width/length 1.66. Venter: Space between mesocoxae narrow, slightly convex. Mesometasternal suture well-defined at base of mesocoxae. Abdominal sternites with 1 row of setae and 3–5 rows of punctures per sternite. Sixth abdominal sternite slightly raised, as a bump. Last sternite punctate, with subapical bead thick and sinuate, apical membrane wide. Legs: Protibia (Fig. 13) tridentate, apical tooth long and curved, middle tooth below internal apex of protibia triangular, acute, basal tooth scarcely developed. Metatibia fusiform. Ratio length/ width 3.23. First external carina well-developed. Surface punctate. Internal protarsal claw bifurcate, with upper branch slightly shorter, as wide as the width of the lower one, inferior margin curved. Male genitalia: Parameres short with wide, blunt apex in lateral view (Fig. 21), ventral margin sinuate. Endophallus (Fig. 29) composed of a long sacculus, with a ridge of sclerotized spines at about half the length and a stripe of sclerotized setae at base. Ejaculatory opening lateral, at apex of an inflation.</p> <p>Female. Similar to male except protibia (Fig. 13) with longer and wider teeth, last sternite with curved apex.</p> <p>Variation. Nine paratypes. Head and pronotal disc vary from reddish to dark brown; extension of dark maculae on pronotum variable to almost dark with thin, ochre sides; dark lateral bands and median expansion on elytra variable in width; venter varies from ochre to light brown. Body length 10.26–11.76 mm, body width 5.52–6.38 mm. Clypeal width/length ratio 2.00–2.24. Ratio of interocular width/width of eye 2.91–3.05. Ratio of antennal funiculus/club 0.73–0.87. Pronotal width/ length ratio 1.62–1.72. Scutellar width/length ratio 1.33–1.49. Pygidial width/length ratio 1.59–1.73. Metatibial length/width ratio 2.79–3.23.</p> <p>Etymology. From the Greek prefix méso s, middle, and adjective stiktós, maculated, for the central macula on the elytra.</p> <p>Distribution. Anomala mesosticta occurs in the Caribbean lowlands of Costa Rica, from 0 to 200 m elevation (Fig. 35).</p> <p>Diagnosis. This species is recognized by its ochre elytra with dark brown margins expanding medially, tridentate protibia, and parameres short with a wide blunt apex. This species has an elytral pattern similar to that of Anomala denticollis Bates, 1888 (Guatemala), and A. hoepfneri. The former species has sinuate lateral margins on the pronotum and black maculae on the calli, which are not present in A. mesosticta. The latter species is larger, has dark maculae on the basal calli and light-colored lateral elytral margins, and wide interstices on the elytra with several irregular rows of punctures.</p> </div>	https://treatment.plazi.org/id/03CFA3486B6FFFC9FF13FE6F5B5EFE9B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Filippini, V.;Galante, E.;Micó, E.	Filippini, V., Galante, E., Micó, E. (2015): Descriptions of New Species ofAnomalaSamouelle (Coleoptera: Scarabaeidae: Rutelinae) from Costa Rica. The Coleopterists Bulletin 69 (3): 463-476, DOI: 10.1649/0010-065x-69.3.463, URL: http://dx.doi.org/10.1649/0010-065x-69.3.463
03CFA3486B6CFFC9FF7EFE6F5981FAF4.text	03CFA3486B6CFFC9FF7EFE6F5981FAF4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala robiginosa Filippini, Galante, and Mico	<div><p>Anomala robiginosa Filippini, Galante, and Micó new species</p> <p>(Figs. 6, 14, 22, 30, 34)</p> <p>Type material. Holotype: ♂ " Zarcero, Alfaro Ruiz, 1700m Alajuela Pr. COSTA RICA. Apr - May 1989. A. Del Valle &amp; A. Solis, L- N 240500_493500 / INBIOCRI002517719" deposited at MNCR. Paratypes (3): 1♀ " Santa Rosa National Park Guana. Prov. Costa Rica 18–20 May 1979 D. H. Janzen / INBIOCRI000111925"; 1♂ " ALAJUELA, COSTA RICA Zarcero, 1700 m 23-V-87 COL: A. SOLIS B. / CEUA00106202 "; 1♂ " Alajuela. Costa Rica. Zarcero, 1700 m 23 may 1987 Angel Solís / INBIOCRI002517717". Two paratypes are deposited at MNCR, and one paratype is deposited at Universidad de Alicante.</p> <p>Holotype. Male (Fig. 6). Body oval. Length 11.23 mm, width 5.75 mm. Head, pronotum, and scutellum bronze-brown. Elytra ochre with suture and punctures reddish brown. Pygidium, legs, and venter orange-brown. Head: Clypeus trapezoidal, surface densely punctate-reticulate, convex. Anterior margin straight, thin in frontal view, width/ length ratio 2.09. Frons densely punctate, with a parabolic concavity. Ocular canthus long, thin, with rounded apex. Ratio of interocular width/width of eye 2.48. Ratio of antennal funiculus/club 0.7. Pronotum: Trapezoidal. Ratio width/length 1.84. Lateral margins angulated at 1/3 of pronotal length from the anterior margins. Anterior angles quadrate, posterior angles obtuse. Basal margin sinuate, obliterated at middle. Surface with deep, dense punctures. Scutellum: Subtriangular, with straight lateral margins and acute apex. Ratio width/length 1.27. Surface densely punctate. Elytra: With costae defined by rows of sparse, coalescing punctures. Subsutural interstice with 1–2 rows of sparse punctures, 2 nd –4 th interstices with 1 irregular, secondary stria. Pygidium: Finely granulated, with long setae on posterior margin. Ratio width/length 1.65. Venter: Space between mesocoxae narrow, slightly convex. Mesometasternal suture welldefined at base of mesocoxae. Abdominal sternites with 1 row of setae (2 rows on second sternite) and 3–4 rows of punctures per sternite. Last sternite punctate, with subapical bead thick and sinuate, apical membrane wide. Legs: Protibia (Fig. 14) bidentate, apical tooth long and straight, basal tooth below internal apex of protibia triangular, sharp, obtuse. Metatibia slightly narrower sub- apically. Ratio length/width 2.98. First external carina with row of sparse setae. Surface punctate above second external carina and rugose below. Internal protarsal claw bifurcate, with upper branch slightly shorter and 1/3 width of lower one. Inferior margin with a sharp angle at half length. Male genitalia: Parameres slender (Fig. 22), inferior margin sinuate. Apical portion of ventral plate membranous. Endophallus (Fig. 30) developing dorsally with respect to median lobes; sclerotized, flat plate present posteriorly at base, above which a rounded, hairy inflation is present; ejaculatory duct opening located frontally.</p> <p>Female. Similar to male except protibia (Fig. 14) with longer and wider first tooth; internal protarsal claw narrower with curve inferior margin; last sternite with curved apex.</p> <p>Variation. Three paratypes. Head, pronotum, and venter vary from orange to dark brown; presence of small dark maculae on basal third of elytra. Body length 10.68–11.49 mm, body width 5.75– 6.44 mm. Clypeal width/length ratio 2.01–2.33. Ratio of interocular width/width of eye 2.24–2.75. Ratio of antennal funiculus/club 0.67–0.74. Pronotal width/length ratio 1.72–1.84. Scutellar width/length ratio 1.24–1.37. Pygidial width/length ratio 1.65–1.82. Metatibial length/width ratio 2.77–3.03.</p> <p>Etymology. From the Latin adjective robiginosus, -a, -um, rusty, for the rust-colored elytral maculae.</p> <p>Distribution. Anomala robiginosa inhabits the dry forest in the northwestern Guanacaste lowlands and Tilarán and northwestern extreme of Central mountain ranges of Costa Rica, from 500 to 1,700 m elevation (Fig. 34).</p> <p>Diagnosis. Pronotum bronze-brown, elytra with pigmented punctures, parameres slender, apex squared in lateral view, endophallus developing dorsally with respect to the median lobes. This species is very similar to A. sticticoptera, but it can be distinguished by the shape of the aedeagus. In A. sticticoptera, the parameres are shorter, with a narrow apex, and the ventral angle is acute and protruding posteriorly (Fig. 33).</p> </div>	https://treatment.plazi.org/id/03CFA3486B6CFFC9FF7EFE6F5981FAF4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Filippini, V.;Galante, E.;Micó, E.	Filippini, V., Galante, E., Micó, E. (2015): Descriptions of New Species ofAnomalaSamouelle (Coleoptera: Scarabaeidae: Rutelinae) from Costa Rica. The Coleopterists Bulletin 69 (3): 463-476, DOI: 10.1649/0010-065x-69.3.463, URL: http://dx.doi.org/10.1649/0010-065x-69.3.463
03CFA3486B6CFFC8FD6AFAB459B6FBBE.text	03CFA3486B6CFFC8FD6AFAB459B6FBBE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala strigodermoides Filippini, Galante, and Mico	<div><p>Anomala strigodermoides Filippini, Galante, and Micó, new species</p> <p>(Figs. 7, 15, 23, 31, 35)</p> <p>Type Material. Holotype: ♂ " COSTA RICA. Prov. Alajuela. Bijagua. Albergue Heliconias. 800-900m. 24–26 MAR 2006. B. Gamboa, M. Moraga, J. A. Azofeifa. Tp. Luz. L_N_299800_ 423800 #85743 / INB0004012910" deposited at MNCR. Paratypes (6): 1♀ " COSTA RICA. Prov. Cartago. Reserva Biol. El Copal. 900–1000 m. 24 ABR 2006. B. Gamboa, M. Moraga, E. Navarro. Tp. Luz. L_N_196916_563695 #86066 / INB0004012683"; 1♀ " COSTA RICA.Prov. Alajuela.Bijagua.Albergue</p> <p>Heliconias. 800- 900m. 24–26 MAR 2006.B. Gamboa, M. Moraga, J. A. Azofeifa. Tp. Luz. L_N_299800_ 423800 #85743 / INB0004012911"; 1♀ " COSTA RICA. Prov. Alajuela. Bijagua. Albergue Heliconias. 800- 900m. 24–26 MAR 2006. B. Gamboa, M. Moraga, J. A. Azofeifa. Tp. Luz. L_N_299800_423800 #85743 / INB0004012912"; 1♀ " COSTA RICA. Prov. Alajuela. Guatuso. P.N. Volcán Tenorio. Punto 1: Catarata Río Buenavista. 800m. 5 MAY 2008. J. A. Azofeifa. Tp de Luz 1. L_N_298380_427850 #93735 / CEUA00106207 "; 1♂ " P.N. Volcán Tenorio. Catarata Río Buenavista. Alajuela, Costa Rica. 780m 15/IV/2010. L. M. Zumbado, J.A.Azofeifa. / CEUA00106208 "; 1♂ " P.N. Volcán Tenorio. Catarata Río Buenavista. Alajuela, Costa Rica. 780m 15/IV/2010. L. M. Zumbado, J.A.Azofeifa. / CEUA00106209 ". Three paratypes are deposited at MNCR, and three paratypes are deposited at Universidad de Alicante.</p> <p>Holotype. Male (Fig. 7). Body elongate. Length 9.04 mm, width 4.89 mm. Head, pronotum, and scutellum metallic green; elytra ochre with 3 pairs of brown maculae along basal margin and a brown, wavy, median, transverse band. Rows of punctures lightly pigmented. Entire surface with green luster. Pygidium, legs, and venter brown with green luster. Head: Clypeus trapezoidal, surface densely punctate-reticulate, flat. Anterior margin straight, ratio width/length 1.9. Frons densely punctate, flat. Ocular canthus long and thin, with acute apex. Ratio of interocular width/width of eye 2.76. Ratio of antennal funiculus/club 0.64. Pronotum: Trapezoidal, long, narrower than elytra. Width 1.26 times length. Lateral margins angulated at 1/3 of pronotal length from anterior margins. Anterior angles acute, posterior angles obtuse. Basal margin thin, sinuate, complete. Surface irregular, with coalescing punctures. Scutellum: Subtriangular, surface sparsely punctate. Ratio width/length 1.24. Elytra: With striae defined by regular rows of punctures. Surface covered with secondary punctures. Marginal membrane complete, very thin. Pygidium: Finely granulated, with short setae on disc and long setae at hind margin. Ratio width/length 1.87. Venter: Space between mesocoxae narrow, slightly convex. Mesometasternal suture welldefined at base of mesocoxae. Abdominal sternites with 2–3 rows of setae and 3–5 rows of punctures per sternite. Last sternite strigate, with apical margin thin and curved, protruding membrane thin. Legs: Protibia bidentate (Fig. 15); apical tooth long and curved, basal tooth above internal apex of protibia, triangular, acute. Metatibia slen- der, fusiform. Ratio length/width 2.72. First external carina well-developed. Surface punctate above second external carina and rugose below. External protarsal claw curved; internal claw bifurcate, with upper branch of same length and 2/3 width of the lower one; inferior margin with a sharp angle near base. Male genitalia: Parameres with squared apex in lateral view (Fig. 23). Endophallus (Fig. 31) with a wide sacculus, ejaculatory duct opening at basal lateral inflation.</p> <p>Female. Similar to male except protibia with wider apical tooth; internal protarsal claw narrower with curved inferior margin.</p> <p>Variation. Six paratypes. Elytra with or without green luster; maculae on elytra vary from reddish to dark brown, may also be present on apex; venter varies from reddish to brown. Body length 8.17–9.64 mm, body width 4.42–4.89 mm. Clypeal width/length ratio 1.89–2.08. Ratio of interocular width/width of eye 2.26–3.04. Ratio of antennal funiculus/club 0.60–0.74. Pronotal width/length ratio 1.26–1.32. Scutellar width/length ratio 1.14–1.33. Pygidial width/length ratio 1.87 (n = 1). Metatibial width/length ratio 2.49–2.82.</p> <p>Etymology. From the name of genus Strigoderma and suffix -oides, for its general resemblance with species of Strigoderma Burmeister.</p> <p>Distribution. This species is known from the Guanacaste and Talamanca mountain ranges of Costa Rica, from 750 to 1,050 m elevation (Fig. 35).</p> <p>Diagnosis. The long and narrow pronotum with a coarse surface readily separates this species from other Neotropical Anomala. Another species with a relatively narrow pronotum is Anomala repressa Ohaus, 1908 (Mexico), which is of similar size. However, in A. repressa, the pronotum is wider than long, with a sinuate margin, the entire surface is covered with setae, and the elytra have a peculiar pattern with two light-colored maculae on a black to reddish basal color.</p> </div>	https://treatment.plazi.org/id/03CFA3486B6CFFC8FD6AFAB459B6FBBE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Filippini, V.;Galante, E.;Micó, E.	Filippini, V., Galante, E., Micó, E. (2015): Descriptions of New Species ofAnomalaSamouelle (Coleoptera: Scarabaeidae: Rutelinae) from Costa Rica. The Coleopterists Bulletin 69 (3): 463-476, DOI: 10.1649/0010-065x-69.3.463, URL: http://dx.doi.org/10.1649/0010-065x-69.3.463
03CFA3486B6DFFCFFD7AFB4A5E76FE6B.text	03CFA3486B6DFFCFFD7AFB4A5E76FE6B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala unilineata Filippini, Galante, and Mico	<div><p>Anomala unilineata Filippini, Galante, and Micó, new species</p> <p>(Figs. 8, 16, 24, 32, 36)</p> <p>Type Material. H o l o t y p e: ♂ "S a nt a R o s a National Park Guanacaste Prov. COSTA RICA. 17–19 May 1980 DH Janzen &amp; W. Hallwachs / INBIOCRI001117054" deposited at MNCR. Paratype: 1♂ " Santa Rosa National Park Guanacaste Province Costa Rica. D. H. Janzen 5–12 March 1978 / INBIOCRI001118447", deposited at MNCR.</p> <p>Holotype. Male (Fig. 8). Body oval. Length 8.52 mm, width 4.27 mm. Head and pronotum dark bronze-brown with green luster. Scutellum dark bronze-brown. Elytra ochre with dark brown suture and median wavy band. Pygidium reddish brown. Legs from yellowish to dark brown. Venter reddish brown. Head: Clypeus trapezoidal, with anterior angles widely curved, surface densely punctate-reticulate, convex. Anterior margin straight, thin in frontal view, width/length ratio 2.03. Frons punctate, less densely than clypeus, with a triangular concavity. Ocular canthus long, thin, with acute apex. Ratio of interocular width/width of eye 2.71. Ratio of antennal funiculus/club 0.7. Pronotum: Subtrapezoidal. Ratio width/length 1.57. Lateral margins regularly convex. Anterior angles acute, posterior angles obtuse. Basal margin complete. Surface with fine, dense punctures. Scutellum: Subtriangular, with rounded sides and blunt apex. Ratio width/length 1.16. Surface sparsely punctate. Elytra: With costae defined by regular rows of punctures. Subsutural interstice with 1–2 rows of sparse punctures, 2 nd –4 th interstices with 1 irregular, secondary stria. Pygidium: Strigate with long setae on posterior margin. Ratio width/length 1.64. Venter: Space between mesocoxae narrow, flat. Mesometasternal suture well-defined at base of mesocoxae. Abdominal sternites with 1 row of setae and 2–4 rows of punctures per sternite. Last sternite strigate, with subapical bead thin and sinuate, apical membrane thin. Legs: Protibia (Fig. 16) bidentate, apical tooth long and curved, basal tooth below internal apex of protibia triangular, sharp, acute. Metatibia stout, fusiform. Ratio length/width 3.27. First external carina well-developed. Surface rugose. External protarsal claw strongly curved, internal claw bifurcate, with upper branch of same length and half width of the lower one, inferior margin sinuate. Male genitalia: Parameres with acute blunt apex in lateral view (Fig. 24). Endophallus (Fig. 32) elongate, dorsally covered with setae, and with 2 ventral patches of sclerotized setae. Sclerotized, flat plate present on ventral inflation, where ejaculatory duct opening is located.</p> <p>Female. Unknown.</p> <p>Variation. One paratype. Head, pronotum, and venter vary from reddish to dark brown. Body length 7.74–8.52 mm, body width 4.27–4.28 mm. Clypeal width/length ratio 2.03–2.07. Ratio of interocular width/width of eye 2.71–2.83. Ratio of antennal funiculus/club 0.70–0.73. Pronotal width/length ratio 1.57–1.60. Scutellar width/ length ratio 1.16–1.22. Metatibial length/width ratio 2.96–3.27.</p> <p>Etymology. From the Latin adjectives unus, -a, -um, one, and lineatus, -a, -um, striped, for the single band across the elytra.</p> <p>Distribution. This species occurs in the dry forest in northeastern Costa Rica, from 220 to 300 m elevation (Fig. 36).</p> <p>Diagnosis. This species is recognized by its small size, pronotum dark bronze-brown, elytra ochre with dark brown, median, wavy band, and endophallus elongated with two ventral patches of sclerotized setae.</p> <p>Another species with one transverse band on the elytra is Anomala donovani Stephens, 1830, but it is larger, with a black pronotum, pigmented punctures on the elytra, and shaded apical portion of the elytra. Other small species, like Anomala discoidalis Bates, 1888, Anomala retusicollis Bates, 1888, Anomala tessellatipennis Blanchard, 1851, and Anomala undulata (Melsheimer, 1844) have the elytral pattern formed by isolated maculae, not forming a continuous band, and differently shaped parameres and endophalli.</p> </div>	https://treatment.plazi.org/id/03CFA3486B6DFFCFFD7AFB4A5E76FE6B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Filippini, V.;Galante, E.;Micó, E.	Filippini, V., Galante, E., Micó, E. (2015): Descriptions of New Species ofAnomalaSamouelle (Coleoptera: Scarabaeidae: Rutelinae) from Costa Rica. The Coleopterists Bulletin 69 (3): 463-476, DOI: 10.1649/0010-065x-69.3.463, URL: http://dx.doi.org/10.1649/0010-065x-69.3.463
