taxonID	type	description	language	source
03CE87B3FFC5FFA68DD0FE955CC87BE6.taxon	description	(Figs 1 – 2, 5 – 8, 13 – 17, 23 – 26, 31 – 34) http: // zoobank. org / urn: lsid: zoobank. org: act: 974 B 23 EA-E 128 - 4 B 10 - A 8 C 7 - E 3 AA 5 E 9855 F 1	en	Giusto, Carlo (2021): Squamapion gardinii n. sp. from the Mediterranean Region and notes on some species belonging to the genus Squamapion Bokor, 1923 (Coleoptera, Apionidae). Zootaxa 4941 (2): 217-238, DOI: 10.11646/zootaxa.4941.2.4
03CE87B3FFC5FFA68DD0FE955CC87BE6.taxon	materials_examined	Type locality. Genova-Quinto al Mare: Rio San Pietro (Genoa Province, Liguria, Italy).	en	Giusto, Carlo (2021): Squamapion gardinii n. sp. from the Mediterranean Region and notes on some species belonging to the genus Squamapion Bokor, 1923 (Coleoptera, Apionidae). Zootaxa 4941 (2): 217-238, DOI: 10.11646/zootaxa.4941.2.4
03CE87B3FFC5FFA68DD0FE955CC87BE6.taxon	diagnosis	Diagnosis. A Squamapion species belonging to the “ S. vicinum group ” as defined by Wanat (1997). It differs from all other Palaearctic Squamapion in having the following combination of characters: stocky general appear- ance; legs robust; vestiture dense, especially on sides of body, composed of narrowly lanceolate white scales, on elytral intervals arranged in 2 rows; rostrum long and thin, particularly in female; mesorostral dilatation dentiform in both sexes.	en	Giusto, Carlo (2021): Squamapion gardinii n. sp. from the Mediterranean Region and notes on some species belonging to the genus Squamapion Bokor, 1923 (Coleoptera, Apionidae). Zootaxa 4941 (2): 217-238, DOI: 10.11646/zootaxa.4941.2.4
03CE87B3FFC5FFA68DD0FE955CC87BE6.taxon	materials_examined	Type material. Holotype (Ƌ): ITALY — Liguria: Genoa Province: “ Genova-Quinto al / Mare: Rio San Pietro / 100 m ca. 6. V. 2020 / C. Giusto leg. ” // “ on Thymus vulgaris L. ” (BMNH). Paratypes (329 ƋƋ, 410 ♀♀): FRANCE — Provence-Alpes-Côte d’Azur: Alpes-Maritimes Department: Gourdon, env. S, 700 m a. s. l., 21. V. 2013, C. Giusto, G. Gardini & S. Zoia leg., on Thymus vulgaris L. (1 ♀ CG); Tende, Vallon du Réfrei, 30. X. 2015, H. Pierotti leg. (1 ♀ LD); — Bouches-du-Rhône Department: Aix-en-Provence, 3. XI. 1968, G. Bartoli leg., on Thymus vulgaris L. (4 ♀♀ MSNG); — Hautes-Alpes Department: Garde-Colombe, Saint-Genis: Gorges du Riou, 7. VII. 2015, R. Casalini leg. (1 Ƌ, 1 ♀ CG; 1 Ƌ, 1 ♀ RC); — Var Department: Callas, env. N, 460 m a. s. l., 23. IV. 2011, Ch. Germann leg. (1 ♀ CGE). ITALY — Abruzzo: Pescara Province: Popoli, Sorgenti del Pescara, 250 – 300 m a. s. l., 22. VI. 2017, C. Giusto & G. Gardini leg., on Thymus vulgaris L. (1 Ƌ CG); — Emilia-Romagna: Piacenza Province: Ottone, Losso, 550 m a. s. l., 21. VI. 2020, C. Giusto leg., on Thymus vulgaris L. (1 ♀); — Liguria: Genoa Province: Bogliasco, 9. VI. 1961, G. Bartoli leg. (7 ƋƋ, 2 ♀♀ MSNG); Bogliasco, Bric Gianesi, slopes S, 600 m a. s. l., 7. I. 2018, C. Giusto leg., on Thymus vulgaris L. (14 ♀♀ CG); Bogliasco, Monte Santa Croce, slopes S, 500 m a. s. l., 12. I. 2020, C. Giusto leg., on Thymus vulgaris L. (3 ♀♀ CG); Camogli, Monte di Portofino, slopes W, 11. IV. 1968, G. Bartoli leg., on Thymus vulgaris L. (1 ♀ MSNG); Camogli, Monte Tocco, 470 m a. s. l., 25. V. 2020, C. Giusto leg., on Thymus vulgaris L. (3 ƋƋ CG); Camogli, San Rocco, Mortola, env. E, 350 m a. s. l., 25. V. 2020, C. Giusto leg., on Thymus vulgaris L. (4 ƋƋ, 1 ♀ CG); Chiavari, Monte Castello, slopes S, 250 m a. s. l., 27. III. 1984, C. Giusto leg., on Thymus vulgaris L. (1 ♀ CG); Crocefieschi, 20. IX. 1905, A. Dodero leg. (2 ♀♀ MSNG); Crocefieschi, 700 m a. s. l., 16. II. 2020, C. Giusto leg., on Thymus vulgaris L. (10 ♀♀ CG); Davagna, Canate di Marsiglia env., 450 m a. s. l., 25. IV. 2016, C. Giusto leg., on Thymus vulgaris L. (25 ƋƋ, 10 ♀♀ CG); Genova, VI. 1911, C. Mancini leg. (1 Ƌ, 1 ♀ MSNG); Genova-Apparizione, 250 m a. s. l., 30. V. 2020, C. Giusto leg., on Thymus vulgaris L. (3 ƋƋ, 1 ♀ CG); Genova-Ba- vari env., 300 m a. s. l., 31. V. 2013, C. Giusto, G. Gardini & S. Zoia leg., on Thymus vulgaris L. (27 ƋƋ, 19 ♀♀ CG); Genova-Bavari, VI. 1907, C. Mancini leg. (1 ♀ MSNG); Genova-Bavari env., 300 m a. s. l., 7. VI. 2017, C. Giusto & G. Gardini leg., on Thymus vulgaris L. (10 ƋƋ, 4 ♀♀ CG); Genova-Bavari env., 250 m a. s. l., 22. V. 2020, C. Gi- usto leg., on Thymus vulgaris L. (13 ƋƋ, 10 ♀♀ CG); Genova-Chiappeto, 2. VI. 1953, F. Capra leg. (1 ♀ MSNG); Genova-Granarolo, 21. VII. 1961, G. Bartoli leg. (3 ƋƋ, 5 ♀♀ MSNG); Genova-Granarolo, 4. IV. 1967, G. Bartoli leg., on Thymus sp. (2 ♀♀ MSNG); Genova-Molassana, 21. V. 1944, G. Binaghi leg., on Thymus vulgaris L. (11 ƋƋ, 7 ♀♀ MSNG); Genova-Molassana, 28. III. 1976, G. Gardini leg. (1 ♀ MSNG); Genova-Molassana, 11. V. 1982, C. Giusto leg., on Thymus vulgaris L. (5 ƋƋ CG); Genova-Molassana, 16. VIII. 1962, L. Briganti leg. (1 ♀ CG); Genova-Molassana, V. 1944, A. Baliani leg. (1 Ƌ CG); Genova-Quezzi, 19. XI. 1967, G. Bartoli leg., on Thymus vulgaris L. (5 ♀♀ MSNG); Genova-Quezzi, 29. V. 1963, G. Bartoli leg., on Thymus sp. (1 ♀ MSNG); Genova-Quinto al Mare, Rio San Pietro, 200 m a. s. l., 15. III. 2020, C. Giusto leg., on Thymus vulgaris L. (3 ♀♀ CG); Genova-Quinto al Mare, Rio San Pietro, 70 m a. s. l., 28. IV. 2019, C. Giusto leg., on Thymus vulgaris L. (13 ƋƋ, 11 ♀♀ CG); Genova- Quinto al Mare, Rio San Pietro, 100 m a. s. l., 6. V. 2020, C. Giusto leg., on Thymus vulgaris L. (3 ƋƋ, 3 ♀♀ BMNH; 39 ƋƋ, 43 ♀♀ CG; 3 ƋƋ, 3 ♀♀ CMNC; 3 ƋƋ, 3 ♀♀ EC; 3 ƋƋ, 3 ♀♀ MNHN; 3 ƋƋ, 3 ♀♀ MSNG; 3 ƋƋ, 3 ♀♀ MSNM; 3 ƋƋ, 3 ♀♀ MW; 3 ƋƋ, 3 ♀♀ NMW; 3 ƋƋ, 3 ♀♀ RC; 3 ƋƋ, 3 ♀♀ SDEI; 3 ƋƋ, 3 ♀♀ ZMHB); Genova-Righi, 30. IV. 1976, G. Bartoli leg., on Thymus vulgaris L. (1 Ƌ, 1 ♀ MSNG); Genova-Righi, 5. VI. 1980, G. Bartoli leg., on Thymus vulgaris L. (1 ♀ MSNG); Genova-San Desiderio, 16. IV. 1961, G. Bartoli leg., on Thymus sp. (6 ♀♀ MSNG); Genova-San Desiderio, 8. IV. 1962, G. Bartoli leg., on Thymus sp. (4 ♀♀ MSNG); Genova, Monte Fasce, 830 m a. s. l., 11. VII. 2020, C. Giusto leg., on Thymus vulgaris L. (1 ♀ CG); Genova, Monte Fasce, slopes S, 700 m a. s. l., 11. VII. 2020, C. Giusto leg., on Thymus vulgaris L. (2 ƋƋ, 1 ♀ CG); Genova, Monte Fasce, slopes S, 300 m a. s. l., 26. X. 2014, C. Giusto leg., on Thymus vulgaris L. (5 ƋƋ, 20 ♀♀ CG); Genova, Monte Fasce, slopes W, 300 m a. s. l., 18. XI. 2017, C. Giusto leg., on Thymus vulgaris L. (2 ♀♀ CG); Genova, Monte Fasce, slopes S, 400 m a. s. l., 6. V. 2020, C. Giusto leg., on Thymus vulgaris L. (14 ƋƋ CG; 3 ƋƋ, 3 ♀♀ CGE; 3 ƋƋ, 3 ♀♀ FS; 3 ƋƋ, 3 ♀♀ KS; 3 ƋƋ, 3 ♀♀ LD; 3 ƋƋ, 3 ♀♀ MK; 3 ƋƋ, 3 ♀♀ TAUI; 3 ƋƋ, 3 ♀♀ TLFN; 3 ƋƋ, 3 ♀♀ ZMUH); Genova, Monte Fasce, 16. IX. 1967, G. Bartoli leg., on Thymus vulgaris L. (1 ♀ MSNG); Genova, Monte Fasce, 25. VIII. 1884, Caneva leg. (1 ♀ MSNG – Apion atomarium, det. J. Desbrochers des Loges); Genova, Monte Gazzo, slopes SE, 200 – 300 m a. s. l., 1. I. 2019, C. Giusto leg., on Thymus vulgaris L. (13 ♀♀ CG); Genova, Monte Ratti, slopes S, 500 m a. s. l., 14. I. 2018, C. Giusto leg., on Thymus vulgaris L. (5 ♀♀ CG); Montoggio, Creto, 11. VII. 1961, G. Bartoli leg. (1 Ƌ MSNG); Montoggio, Creto, 4. X. 1968, G. Bartoli leg., on Thymus vulgaris L. (3 ♀♀ MSNG); Montoggio, Creto, 600 m a. s. l., 16. II. 2020, C. Giusto leg., on Thymus vulgaris L. (4 ♀♀ CG); Montoggio, Creto, 600 m a. s. l., 28. IX. 2014, C. Giusto leg., on Thymus vulgaris L. (1 Ƌ, 2 ♀♀ CG); Montoggio, Creto, env. E, 650 m a. s. l., 22. V. 2020, C. Giusto, G. Gardini & P. Gardini leg., on Thymus vulgaris L. (7 ƋƋ, 10 ♀♀ CG); Rapallo, Monte Caravagli, slopes, 400 m a. s. l., 16. I. 1993, C. Giusto leg., on Thymus vulgaris L. (2 ♀♀ CG); Rapallo, Monte Pegge, slopes S, 700 m a. s. l., 1. I. 2020, C. Giusto leg., on Thymus vulgaris L. (4 ♀♀ CG); Rapallo, Passo della Crocetta, 550 m a. s. l., 20. VI. 1992, C. Giusto leg., on Thymus vulgaris L. (1 Ƌ, 4 ♀♀ CG); Sestri Levante, Trigoso, 75 m a. sl., 1. XI. 2020, C. Giusto leg., on Thymus vulgaris L. (4 ♀♀ CG); Sori, 9. III. 1969, G. Bartoli leg., on Thymus vulgaris L. (1 ♀ MSNG); Sori, Canepa env., 350 m a. s. l., 9. VI. 1991, C. Giusto leg., on Thymus vulgaris L. (17 ƋƋ, 10 ♀♀ CG); Sori, Canepa env., 350 – 400 m a. s. l., 31. XII. 2016, C. Giusto leg., on Thymus vulgaris L. (9 ♀♀ CG); Vobbia, 470 m a. s. l., 16. II. 2020, C. Giusto leg., on Thymus vulgaris L. (2 ♀♀ CG); — Imperia Province: Ventimiglia, Capo Mortola, 50 m a. s. l., 10. V. 2019, C. Giusto leg., on Thymus vulgaris L. (1 Ƌ CG); Ventimiglia, Villatella, 400 – 500 m a. s. l., 10. V. 2019, C. Giusto leg., on Thymus vulgaris L. (2 ♀♀ CG); — La Spezia Province: Deiva Marina, Ca’ Marcone env., 475 m a. s. l., 1. XI. 2020, C. Giusto leg., on Thymus vulgaris L. (1 Ƌ, 2 ♀♀ CG); Deiva Marina, Passo del Bracco, 7. VI. 2006, L. Diotti leg. (1 Ƌ, 3 ♀♀ LD); Portovenere, Monte Muzzerone, slopes NW, 220 m a. s. l., 22. II. 2020, C. Giusto leg., on Thymus vulgaris L. (5 ♀♀ CG); — Savona Province: Albenga, 25. IV. 1901, A. Dodero leg. (1 ♀ MSNG); Albenga, Bastia, 31. III. 1968, G. Binaghi leg. (1 ♀ MSNG); Albissola Marina, 20. X. 1926, F. Capra leg. (1 ♀ MSNG); Bergeggi, Monte Mao, 400 m a. s. l., 10. IV. 2017, C. Giusto, G. Gardini & A. Trotta leg., on Thymus vulgaris L. (1 ♀ CG); Boissano env., 150 m a. s. l., 15. V. 1994, C. Giusto leg., on Thymus vulgaris L. (12 ƋƋ, 8 ♀♀ CG); Celle Ligure, Bric dei Corvi, 250 m a. s. l., 7. V. 1995, C. Giusto leg., on Thymus vulgaris L. (2 ƋƋ CG); Cisano sul Neva, Poggio Croce Ceresa, 700 m a. s. l., 10. VII. 2002, C. Giusto leg., on Thymus vulgaris L. (2 ƋƋ CG); Finale Ligure, Finalmarina, 22. V. 1899, A. Dodero leg. (14 ƋƋ, 5 ♀♀ MSNG); Noli, Capo Noli, 3. IV. 1966, G. Bartoli leg., on Thymus sp. (1 ♀ MSNG); Toirano, Salto del Lupo, 180 – 200 m a. s. l., 15. VI. 2016, C. Giusto leg., on Thymus vulgaris L. (6 ƋƋ, 11 ♀♀ CG); Villanova d’Albenga, Coasco, 54 m a. s. l., 21. XI. 2015, C. Giusto leg., on Thymus vulgaris L. (5 ♀♀ CG); — Piedmont: Alessandria Province: Dernice env., 21. IV. 2016, L. Diotti leg. (1 ♀ CG); Dernice, Costa di Montebore, 680 m a. s. l., 21. VI. 2020, C. Giusto leg., on Thymus vulgaris L. (8 ƋƋ, 8 ♀♀ CG); Dernice, Montebore, 20. I. 2018, L. Diotti leg. (1 ♀ CG; 1 ♀ LD); Dernice, Montebore, 29. VIII. 2014, L. Diotti leg. (2 ♀♀ LD); Dernice, Montebore, 28. IX. 2016, L. Diotti leg. (2 ƋƋ, 2 ♀♀ LD); Dernice, Montebore, 24. VI. 2020, L. Diotti leg. (4 ƋƋ, 8 ♀♀ LD); Dernice, Montebore, 24. VI. 2020, M. Tedeschi leg. (3 ƋƋ, 3 ♀♀ MT). SPAIN — Community of Madrid: Madrid Province: Santa María de la Alameda, La Paradilla, 1,100 m a. s. l., 27. VI. 1991, C. Giusto leg., on Thymus vulgaris L. (2 ♀♀ CG).	en	Giusto, Carlo (2021): Squamapion gardinii n. sp. from the Mediterranean Region and notes on some species belonging to the genus Squamapion Bokor, 1923 (Coleoptera, Apionidae). Zootaxa 4941 (2): 217-238, DOI: 10.11646/zootaxa.4941.2.4
03CE87B3FFC5FFA68DD0FE955CC87BE6.taxon	etymology	Etymology. It is a real pleasure and an honor for me to dedicate the new taxon to Giulio Gardini, pseudoscorpion specialist, friend, mentor and research partner in my first 40 years of entomology.	en	Giusto, Carlo (2021): Squamapion gardinii n. sp. from the Mediterranean Region and notes on some species belonging to the genus Squamapion Bokor, 1923 (Coleoptera, Apionidae). Zootaxa 4941 (2): 217-238, DOI: 10.11646/zootaxa.4941.2.4
03CE87B3FFC5FFA68DD0FE955CC87BE6.taxon	description	Description (Ƌ ♀). Body stocky; Lb: Ƌ 1.20 – 1.77 mm [1.64 mm] (1.67 mm), ♀ 1.30 – 1.79 mm [1.60 mm]. Body integument black; femora black; tibiae and tarsi dark brown to nearly black; in male, antennae testaceous with darker club and sometimes with darkened distal funicular segments; in female, antennae dark brown to nearly black. Vestiture dense, composed of white scales, rarely white-greyish; hair-like on prorostrum and ventrite 5; narrowly lanceolate on metarostrum, pronotal disc, elytra and on body venter; lanceolate, with truncate apex, around eyes, on body sides and on legs; on elytral intervals, scales, only just longer than width of intervals, arranged in two quite irregular rows, partially overlapping; at base of intervals 2 – 4, scales more numerous forming short multiple rows; at base of interval 3, scales, with truncate apex, arranged in a more or less condensed patch; interstrial scales similar to those on intervals, barely separated from each other (Figs 1 – 2). Rostrum rather long; in dorsal view, obtusely dentiform at antennal insertion, narrowed from mesorostrum to the half of prorostrum, then cylindrical up to apex; in lateral view, sometimes feebly bent at antennal insertion and with prorostrum regularly curved, less in male, more in female, narrowing at apex in male or equally high on whole length in female (Figs 5 – 8); rostral surface shagreened, almost matt in male, weakly shining in female; shallowly punctate; in male, up to half prorostrum with sparse, whitish, hair-like scales and then with microscopic scales up to apex; in female almost whole prorostrum clothed with microscopic and sparse scales. Head transverse; eyes, in male, moderately large and regularly convex, in female distinctly smaller; frons flat, strongly punctate and microsculptured with almost indistinct median fovea in some specimens. Antennae (Figs 13 – 14) inserted at basal 0.13 – 0.19 of rostrum, slender and with white and weakly erect setae. Prothorax (Figs 1 – 2) in dorsal view transverse, campanulate, moderately rounded at sides, widest at base with prominent hind angles; disc weakly convex, densely punctate; punctures round, 16 – 22 µm in diameter, separated by less than one third of their diameter; interspaces strongly microreticulate; prescutellar fovea as broad as discal punctures or weakly broader, not longer than three punctures. Scutellum triangular with curved sides, weakly de- pressed on disc and with two rudimentary keels at base, glabrous, microsculptured. Elytra (Figs 1 – 2) in dorsal view subrectangular, with slightly curved sides in male, more rounded in female; intervals 1.4 – 1.8 times as wide as striae, flat, rugosely microsculptured and with two rows of shallow punctures; striae sharply margined with septa between punctures distinctly deepened. Legs robust; in male, hind femora thickened; tibiae straight, relatively short, strongly widened from base to apex; protarsi robust; 1 st segment 1.27 – 1.50 times as long as wide; 2 nd 1.00 – 1.21 times as long as wide, 0.80 – 0.87 times as long as 1 st; 3 rd 0.72 – 0.86 times as long as wide, 0.84 – 0.93 times as long as 2 nd, with narrow lobes; onychium robust, 1.09 – 1.33 times as long as 2 nd. Sternite IX as in Fig. 15. Tegmen (Figs 16 – 17); phallobase Y-shaped with tegminal apodeme slightly longer than forked basal piece; parameral plate with median carinae moderately separated, vanishing just behind the supra- fenestral sclerites; each suprafenestral sclerite bears 7 – 10 pores; apical membranous lobes well developed, united, margined with microchaetae; whole folded margins of parameral plate with very fine and dense longitudinal cari- nae. Penis (Figs 23 – 26) in dorsal view more or less slender, variably narrowed in distal half and ending in a slightly asymmetrical blunt tip; endophallus with a more or less defined M-shaped aggregation of spines in orificial region; basal spines small, distal ones bigger. Female sternite VIII as in Fig. 31. Spermatheca (Fig. 32) C-shaped with cornu more or less recurved; spermathecal gland inserted on a just sketched ramus; nodulus slightly protruding. Ovipositor as in Fig. 33; foretube densely clothed with long microtrichia; gonocoxites (Figs 33 – 34) about 4 times as long as wide; styli inserted latero-apically, cylindrical, 2.25 – 2.42 times as long as wide, with 7 – 8 apical setae.	en	Giusto, Carlo (2021): Squamapion gardinii n. sp. from the Mediterranean Region and notes on some species belonging to the genus Squamapion Bokor, 1923 (Coleoptera, Apionidae). Zootaxa 4941 (2): 217-238, DOI: 10.11646/zootaxa.4941.2.4
03CE87B3FFC5FFA68DD0FE955CC87BE6.taxon	distribution	Distribution. Spain, France, Italy.	en	Giusto, Carlo (2021): Squamapion gardinii n. sp. from the Mediterranean Region and notes on some species belonging to the genus Squamapion Bokor, 1923 (Coleoptera, Apionidae). Zootaxa 4941 (2): 217-238, DOI: 10.11646/zootaxa.4941.2.4
03CE87B3FFC5FFA68DD0FE955CC87BE6.taxon	biology_ecology	Host plant. Thymus vulgaris L. (Figs 40 – 42). Bionomy. Squamapion gardinii prefers the garrigue and the arid meadows of the Mediterranean calcareous or siliceous / calcareous slopes, approximately from sea level (Italy, Ventimiglia, Capo Mortola, 50 m a. s. l.) up to 1,100 m a. s. l. (Spain, Santa María de la Alameda, La Paradilla); it is particularly abundant between 100 and 500 m above sea level (Fig. 39). Plants that host S. gardinii show little damage caused by the adults: small damage is visible on the leaves while it is possible to observe the galls — inside which the insects develop — on the stems (Figs 41 – 45). No galls were found on roots. Adults are present on the host plant all year round, but they are particularly numerous from April to June and again quite frequent from October to December.	en	Giusto, Carlo (2021): Squamapion gardinii n. sp. from the Mediterranean Region and notes on some species belonging to the genus Squamapion Bokor, 1923 (Coleoptera, Apionidae). Zootaxa 4941 (2): 217-238, DOI: 10.11646/zootaxa.4941.2.4
03CE87B3FFC5FFA68DD0FE955CC87BE6.taxon	discussion	Remarks. Some authors had already noted in the past that the southernmost populations of S. atomarium were a little different from those of central Europe. Indeed, it is worth remembering what was written by Sainte-Claire Deville (1924): “ J’ai capturé çà et lâ l’A. atomarium dans des montagnes sèches du Midi de la France où le Thymus serpyllum n’existait pas et où par contre Ie T. vulgaris était abondant; les individus de cette provenance sont plus robustes et leur pubescence est plus apparente ”, observations also quoted later by Hoffmann (1929, 1958). Just two years before, Schatzmayr (1922) had already observed some differences within different populations of S. atomarium underlining the extraordinary body length of the specimens from Montoggio (Liguria, Italy), ” sino a 2,4 mm ”. At first glance, thanks to its dense vestiture, S. gardinii appears whitish whilst, probably due to its stocky body, it seems much larger than it really is. Measurements specified in Table 1, in fact, show that S. gardinii on average is just a little bit larger than S. atomarium and that dimensions of the two species are rather similar (cfr. Figs 1 – 2 with Figs 3 – 4). The analysis of rostral morphology reveals the most useful characters to discriminate S. gardinii from S atomarium: both sexes of the new species show — on average — a longer and thinner rostrum and a dentiform mesorostral dilatation, whereas the rostrum of S. atomarium is thicker and the mesorostral dilatation is dentiform in males and obtusely rounded in females (cfr. Figs 5 – 8 with Figs 9 – 12). Among the other species of Squamapion, only Squamapion solariorum (Wagner, 1908) n. stat. (see below), Squamapion bifarium (Balfour-Browne, 1944), Squamapion kandarense (Hustache, 1946) and Squamapion terraesanctae Friedman & Freidberg, 2007 share with S. gardinii the long and thin rostrum. Despite the small number of specimens examined of some of these species (see Appendix 1) and the fact that their measurements and their ratios overlap abundantly, it is easy to observe that, regardless of the curvature of rostrum — more recurved in S. solariorum, S. bifarium and S. kandarense, less in S. terraesanctae — all their female mesorostra are obtusely rounded in dorsal view. Moreover, their vestiture differs from that of S. gardinii being composed of piliform scales ordered in a single row on elytral intervals, sometimes forming 2 – 3 short confused rows at base of intervals 2 – 4, and often arranged in a poorly condensed patch at base of interval 3. The body outlines of all of them are stout, but less than in S. gardinii; their humeral calli are more prominent, their pronotal surfaces are slightly less densely punctate and the interspaces are microreticulate and weakly shiny. A further, although partial, aid for recognizing these species comes from the knowledge of their host plants: S. atomarium is oligophagous on some Thymus species, mainly on taxa belonging to “ Thymus serpyllum L. ” and “ Thymus pulegioides L. ” groups; also S. gardinii is associated to Thymus, but it is monophagous on Thymus vulgaris L. Squamapion terraesanctae lives on Satureja thymbra L. and Origanum syriacum L. Raf. whilst Mentha longifolia typhoides (Briq.) Briq. (= Mentha longifolia cyprica (Heinr. Braun) Harley) is doubtfully quoted as the host plant of S. bifarium by Friedman & Freidberg (2007). Unfortunately, no data about host plants of S. solariorum and S. kandarense are available, even though their association with Thymus is predictable. The ecology and the phenology of S. gardinii and S. atomarium are also different: the latter species inhabits arid meadows on hills and mountains with calcareous or siliceous soils from 250 to 2,450 m a. s. l. being particularly common from 500 to 1,500 m a. s. l. with adults on the host plants from April to December, but the peak attendance is recorded between June and September (especially in September).	en	Giusto, Carlo (2021): Squamapion gardinii n. sp. from the Mediterranean Region and notes on some species belonging to the genus Squamapion Bokor, 1923 (Coleoptera, Apionidae). Zootaxa 4941 (2): 217-238, DOI: 10.11646/zootaxa.4941.2.4
03CE87B3FFCAFFA48DD0F99A5EA97D79.taxon	description	(Figs 46 – 48)	en	Giusto, Carlo (2021): Squamapion gardinii n. sp. from the Mediterranean Region and notes on some species belonging to the genus Squamapion Bokor, 1923 (Coleoptera, Apionidae). Zootaxa 4941 (2): 217-238, DOI: 10.11646/zootaxa.4941.2.4
03CE87B3FFCAFFA48DD0F99A5EA97D79.taxon	materials_examined	Type locality. Aïn Touta (Aïn Touta District, Batna Province, Algeria).	en	Giusto, Carlo (2021): Squamapion gardinii n. sp. from the Mediterranean Region and notes on some species belonging to the genus Squamapion Bokor, 1923 (Coleoptera, Apionidae). Zootaxa 4941 (2): 217-238, DOI: 10.11646/zootaxa.4941.2.4
03CE87B3FFCAFFA48DD0F99A5EA97D79.taxon	diagnosis	Diagnosis. A Squamapion species belonging to the “ vicinum group ” as defined by Wanat (1997). Among the Mediterranean Squamapion species, it differs in having the following combination of characters: long and slender rostrum with obtusely rounded mesorostrum; parallelsided elytra with well developed humeral calli; vestiture, on elytral intervals, arranged in a single regular row. No close affinity seems to exist with Squamapion consors (Desbrochers des Loges, 1875); female of the latter species mainly differs in having thicker and less curved rostrum (Lr: 0.63 – 0.68 mm [0.65 mm]; Lpr: 0.53 – 0.56 mm [0.54 mm]; Lmtr: 0.12 – 0.14 mm [0.13 mm]; Wra: 0.11 – 0.12 mm [0.12 mm]; Wmsr: 0.15 – 0.18 mm [0.16 mm]; Lr / Wmsr: 3.78 – 4.20 [4.01]; Lr / Wra: 5.42 – 5.73 [5.60]); elytral profile, in dorsal view, distinctly rounded; body vestiture composed of yellow-gold, lanceolate scales; vestiture, on elytral intervals, arranged in 2 – 3 irregular rows. For its relationships with S. kandarense, see the remarks below.	en	Giusto, Carlo (2021): Squamapion gardinii n. sp. from the Mediterranean Region and notes on some species belonging to the genus Squamapion Bokor, 1923 (Coleoptera, Apionidae). Zootaxa 4941 (2): 217-238, DOI: 10.11646/zootaxa.4941.2.4
03CE87B3FFCAFFA48DD0F99A5EA97D79.taxon	materials_examined	Type material examined. Holotype (♀): ALGERIA — Batna: Aïn Touta District: “ Ain-Touta / Algeria / Coll. Vauloger ” // “ Wagner det / consors Dbrs. ♀ ” // “ n. sbsp. / Wagner det. ♀ / Solariorum m. ” (MSNM).	en	Giusto, Carlo (2021): Squamapion gardinii n. sp. from the Mediterranean Region and notes on some species belonging to the genus Squamapion Bokor, 1923 (Coleoptera, Apionidae). Zootaxa 4941 (2): 217-238, DOI: 10.11646/zootaxa.4941.2.4
03CE87B3FFCAFFA48DD0F99A5EA97D79.taxon	description	Redescription (♀). Lb: 1.71 mm. Body integument black; femora black; tibiae and tarsi nearly black; antennae brown. Vestiture composed of cream-white lanceolate scales; on elytral intervals, scales are arranged in a single regular row, partially overlapping, as long as interval width; at base of intervals 2 – 4, few scales are arranged in short multiple confused rows; at base of interval 3 few scales, with truncate apex, are arranged in a loosely condensed patch; interstrial scales similar to those on intervals, barely separated from each other (Fig. 46). Rostrum rather long, thin; in dorsal view obtusely rounded at antennal insertion, just narrowed from meso- rostrum to half of prorostrum, then cylindrical up to apex; in lateral view distinctly and regularly curved, equally high on whole length (Figs 47 – 48); rostrum weakly shagreened, shining in the apical quarter; shallowly punctate; in basal half sparsely clothed with lanceolate scales with truncate apex; in apical half with sparser piliform scales up to apex. Head transverse; eyes rather small, regularly convex; frons flat, strongly punctate and microsculptured with almost indistinct median fovea. Antennae inserted at basal 0.16 of rostrum, slender with white and weakly erect setae. Prothorax (Fig. 46) transverse, campanulate, moderately rounded at sides, widest at base and with prominent posterior angles; disc weakly convex, densely punctate; punctures round, 16 – 23 µm in diameter, separated by less than one third of their diameter; interspaces microreticulate; prescutellar fovea as broad as disc punctures, not longer than three punctures. Scutellum subrounded, weakly depressed on disc and with two rudimentary keels, glabrous, microsculptured. Elytra (Fig. 46), in dorsal view, subrectangular, with slightly curved sides; intervals about 1.5 times as wide as striae, flat, rugosely microsculptured and with one row of shallow punctures; striae sharply mar- gined with septa between punctures distinctly deepened. Tibiae rather slender, feebly widened from base to apex; protibiae feebly outcurved, meso- and metatibiae straight; protarsi with 1 st segment 2 times as long as wide; 2 nd 1.00 times as long as wide, 0.65 times as long as 1 st; 3 rd 0.71 times as long as wide, 0.92 times as long as 2 nd, with narrow lobes; onychium robust, 1.23 times as long as 2 nd. Genitalia not examined.	en	Giusto, Carlo (2021): Squamapion gardinii n. sp. from the Mediterranean Region and notes on some species belonging to the genus Squamapion Bokor, 1923 (Coleoptera, Apionidae). Zootaxa 4941 (2): 217-238, DOI: 10.11646/zootaxa.4941.2.4
03CE87B3FFCAFFA48DD0F99A5EA97D79.taxon	distribution	Distribution. Algeria, Morocco.	en	Giusto, Carlo (2021): Squamapion gardinii n. sp. from the Mediterranean Region and notes on some species belonging to the genus Squamapion Bokor, 1923 (Coleoptera, Apionidae). Zootaxa 4941 (2): 217-238, DOI: 10.11646/zootaxa.4941.2.4
03CE87B3FFCAFFA48DD0F99A5EA97D79.taxon	biology_ecology	Host plant. Unknown.	en	Giusto, Carlo (2021): Squamapion gardinii n. sp. from the Mediterranean Region and notes on some species belonging to the genus Squamapion Bokor, 1923 (Coleoptera, Apionidae). Zootaxa 4941 (2): 217-238, DOI: 10.11646/zootaxa.4941.2.4
03CE87B3FFCAFFA48DD0F99A5EA97D79.taxon	discussion	Remarks. This taxon was described based on a single female as a subspecies of Apion consors and to which, as far as I know, to date only a further specimen from Asni (Morocco, Al Haouz province, Marrakech-Safi region) has been doubtfully attributed (Kocher 1953, 1961). The diagnostic characters and the main differences with the typical S. consors, highlighted above, allow full species status to be proposed for Squamapion solariorum (Wagner, 1908) n. stat. The single examined specimen of S. kandarense fits perfectly with the rough description provided by Hustache (1946) and with the short redescription given by Hoffmann (1957). It differs from the holotype of S. solariorum almost exclusively in having a feebly rounded elytral profile and imperceptibly less slender rostrum (to compare measurements and ratios, see Table 1). In light of this and on the basis of the scanty distributional data, a possible synonymy between the two species cannot be excluded; only the examination of additional material of both sexes can help address this question.	en	Giusto, Carlo (2021): Squamapion gardinii n. sp. from the Mediterranean Region and notes on some species belonging to the genus Squamapion Bokor, 1923 (Coleoptera, Apionidae). Zootaxa 4941 (2): 217-238, DOI: 10.11646/zootaxa.4941.2.4
03CE87B3FFD5FFB88DD0FEE65B5A7955.taxon	materials_examined	BOSNIA AND HERZEGOVINA — Federacija Bosne i Hercegovine: Sarajevo Canton: Trnovo, Bjelašnica, 1901, Leonhard leg. (1 Ƌ, 4 ♀♀ CG). BULGARIA — Blagoevgrad Province: Sandanski, Melnik, 250 m a. s. l., 9. VII. 1998 (1 ♀ CG); — Sofia Province: Samokov, Raduil, 2. VIII. 1970, P. Angelov leg. (1 Ƌ CG); — Stara Zagora Province: Stara Zagora, 24. V. 1977 (1 ♀ CG). CZECH REPUBLIC — Central Bohemian Region: Příbram: Sedlčany, Týnčany, 27. VIII. 2010 (1 Ƌ CG). FRANCE — Provence-Alpes-Côte d’Azur: Alpes-de-Haute-Provence Department: Jausiers, Lans env., 1,500 m a. s. l., 2. VIII. 1998, C. Giusto leg. (2 ƋƋ, 3 ♀♀ CG); Larche, Col de Larche env., 2,000 m a. s. l., 2. XI. 1996, C. Giusto leg. (1 Ƌ CG); — Alpes-Maritimes Department: Tourette-du-Château, Mont Vial, 1,500 m a. s. l., 10. VII. 1999, C. Giusto leg. (1 Ƌ CG); Vence, Col de Vence, 970 m a. s. l., 24. VIII. 1997, C. Giusto leg. (1 ♀ CG); — Hautes-Alpes Department: Garde-Colombe, Gorges du Riou, 7. VII. 2015, R. Casalini leg. (1 Ƌ, 1 ♀ RC); Guillestre env., 1,600 m a. s. l., 2. VIII. 1998, C. Giusto leg. (1 Ƌ, 3 ♀♀ CG); — Auvergne-Rhône-Alpes: Drôme Department: Bouvante, Forêt de Lente, 1,400 m a. s. l., 9. X. 2012, C. Giusto leg. (1 ♀ CG); Chamaloc, Col de Rousset env., 1,400 m a. s. l., 11. VIII. 1996, C. Giusto leg. (5 ƋƋ, 4 ♀♀ CG); Chamaloc, Col de Rousset env., 1,400 m a. s. l., 21. VIII. 1997, C. Giusto leg. (3 ƋƋ, 4 ♀♀ CG); Val-Maravel, La Bâtie Crémezin, 5. VII. 2014, R. Casalini leg. (1 ♀ CG); Vassieuxen-Vercors, Col de la Chau, 1,330 m a. s. l., 22. VIII. 1997, C. Giusto leg. (17 ƋƋ, 12 ♀♀ CG); — Isère Department: La Salette-Fallavaux, Notre-Dame de La Salette env., 1,600 m a. s. l., 23. VIII. 1997, C. Giusto leg. (4 ƋƋ, 2 ♀♀ CG) — Occitanie: Hautes-Pyrénées Department: Cadéac, 29. VII. 1920 (1 Ƌ, 1 ♀ MSNG); Pyrénées-Orientales Department: Prats-de-Mollo-la-Preste, 10. VIII. 1904, A. Dodero leg. (1 Ƌ, 1 ♀ MSNG). GERMANY — Berlin: Berlin, Spandau, Grosser Kienhorst, 8. VIII. 2006, H. Winkelmann & Köck leg. (1 Ƌ, 2 ♀♀ CG). GREECE — Autonomous Region of Mount Athos: Mount Athos, A. Schatzmayr leg. (1 ♀ MSNG); — Central Greece: Phocis: Delphi, Parnass, Paganetti leg. (1 Ƌ, 1 ♀ CG); — Central Macedonia: Serres: Sintiki, Achladohori, 600 m a. s. l., 15. V. 2005, H. Winkelmann leg. (1 ♀ CG); — Western Macedonia: Kozani: Kozani, Mount Askio, 1,900 m a. s. l., 24. VIII. 1994, M. Meregalli leg. (1 Ƌ CG). ITALY — Abruzzo: Chieti Province: Altopiano delle Cinquemiglia, Stazione di Palena, 1,250 m a. s. l., 4. VIII. 2015, R. Casalini leg. (1 Ƌ CG); — L’Aquila Province: Assergi, Monte Cristo, slopes W, Prato Grande, 1,450 m a. s. l., 9. VI. 1993, C. Giusto leg. (1 ♀ CG); Gran Sasso d’Italia, Gran Sasso, VII. 1896, A. Fiori leg. (2 ƋƋ, 2 ♀♀ MSNG); Pacentro, Passo San Leonardo, env. E, 1,300 – 1,400 m a. s. l., 22. VI. 2017, C. Giusto leg. (1 ♀ CG); Piz- zoli, Monte San Franco, slopes S, 1,800 – 2,000 m a. s. l., 10. VI. 1993, C. Giusto leg. (1 ♀ CG); Pizzoli, Monte San Franco, sorgente del San Franco, 1,650 m a. s. l., 7. VII. 1990, E. Colonnelli leg. (1 ♀ CG); Pizzoli, Passo delle Capan- nelle, 1,300 m a. s. l., 21. IX. 1999, C. Giusto leg. (1 ♀ CG); Rocca di Mezzo, Monte Velino, 1.800 m, 12. VII. 1909, G. Binaghi leg. (1 ♀ MSNG); Secinaro, Monte Sirente, Valle Puzzilli, 1,850 m a. s. l., 8. VII. 1990, E. Colonnelli leg. (1 Ƌ CG); — Campania: Caserta Province: San Giorgio Matese, Lago del Matese, 1,100 m a. s. l., 17. V. 2001, F. Angelini leg. (1 ♀ CG); — Emilia-Romagna: Emilia-Romagna, VI. 1974, A. Fiori leg. (1 ♀ MSNG); — Modena Province: Pievepelago, Le Tagliole, 22. VII. 2020, L. Diotti leg. (3 ƋƋ, 3 ♀♀ LD); — Parma Province: Corniglio, Passo di Cirone, 1.255 m, 28. VII. 2008, L. Diotti leg. (1 Ƌ, 2 ♀♀ LD); — Rimini Province: Talamello, Monte Pincio, slopes, 600 – 900 m a. s. l., 10. IX. 1988, G. Platia leg. (1 ♀ CG); — Friuli-Venezia Giulia: Trieste Province: Trieste, Trieste-Opicina, 15. V. 1970, R. Caldara leg. (1 ♀ CG); — Udine Province: Resia, Sella Carnizza, env. E, 900 m a. s. l., 13. VII. 2019, C. Giusto leg., on Thymus sp. cfr. T. pulegioides L. (2 ƋƋ, 1 ♀ CG); Venzone, Monte Plauris, slopes S, Graunes, 1,350 – 1,400 m a. s. l., 11. VII. 2019, C. Giusto leg., on Thymus sp. cfr. T. pulegioides L. (2 ƋƋ, 2 ♀♀ CG); — Lazio: Frosinone Province: Filettino, 21. VII. 1908, R. Gestro leg. (1 ♀ MSNG – Apion atomarium, det. Solari); — Rieti Province: Antrodoco, Monte Giano, Prati di Cinno, 1,450 m a. s. l., 4. VIII. 1990, E. Colonnelli leg. (1 ♀ CG); Micigliano, Monte Terminillo, slopes W Monte i Porcini, Costa Ghiaiosa, 1,800 m a. s. l., 24. VI. 2017, C. Gi- usto leg. (1 ♀ CG); — Liguria: Genoa Province: Casella, VII – IX. 1932, C. Mancini leg. (1 Ƌ, 1 ♀ MSNG); Fontanig- orda, VII. 1897, A. Dodero leg. (1 ♀ MSNG); Genova-Voltri, Fiorino, 5. X. 2019, L. Diotti leg. (2 ƋƋ LD); Genova- Voltri, Fiorino, 1. IX. 2007, L. Diotti leg. (2 ♀♀ LD); Montoggio, Creto, 4. X. 1968, G. Bartoli leg. (1 ♀ MSNG); Montoggio, Monte Alpesisa, slopes W, Gola di Sisa, 750 m a. s. l., 28. IX. 2014, C. Giusto leg., on Thymus sp. cfr. T. pulegioides L. (3 ƋƋ, 6 ♀♀ CG); — Savona Province: Monte Galero, slopes SW, 1,500 – 1,700 m a. s. l., 13. IX. 1997, C. Giusto leg. (1 Ƌ CG); — Lombardy: Bergamo Province: Oltre il Colle, Pizzo Arera, 7. XII. 2011, L. Diotti leg. (2 ƋƋ, 2 ♀♀ LD); — Molise: Isernia Province: Carovilli env., 19 – 22. VIII. 2009, F. Montemurro leg. (3 ♀♀ CG); — Piedmont: Alessandria Province: Cassano Spinola, IX. 1937, G. B. Moro leg. (1 ♀ MSNG); Molare, Olbicella env., 400 m a. s. l., 1. VIII. 1992, C. Giusto leg., on Thymus sp. cfr. T. pulegioides L. (6 ƋƋ, 5 ♀♀ CG); — Biella Province: Biella, Oropa, Santuario, IX. 1927, A. Dodero leg. (2 ƋƋ, 6 ♀♀ MSNG); Piedicavallo, VII – VIII, G. Bartoli leg. (1 Ƌ MSNG); — Cuneo Province: Acceglio, Vallone dell’Infernetto, 4. VIII. 2009, L. Diotti leg. (2 ƋƋ, 3 ♀♀ LD); Alto, Madonna del Lago, 1,000 m a. s. l., 20. IX. 1994, C. Giusto leg. (1 Ƌ CG); Alto, Madonna del Lago, 1,000 m a. s. l., 29. IX. 1996, C. Giusto leg. (2 ƋƋ, 1 ♀ CG); Caprauna, Monte Armetta, 1,700 m a. s. l., 29. VII. 1986, C. Giusto leg. (2 ♀♀ CG); Caprauna, Monte Armetta, 1,700 m a. s. l., 29. VII. 1989, C. Giusto leg. (1 Ƌ CG); Caprauna, Colle di Caprauna env., 1,400 m a. s. l., 8. VIII. 1994, C. Giusto leg. (2 ƋƋ, 1 CG); Chiusa di Pesio, Val Pesio, VIII. 1907, A. Dodero leg. (1 Ƌ, 1 ♀ MSNG); Chiusa di Pesio, Val Pesio, VIII. 1907, A. Dodero leg. (1 ♀ MSNG – Apion atomarium, det. J. Desbrochers des Loges); Chiusa di Pesio, Val Pesio, VIII. 907, R. Gestro leg. (1 Ƌ MSNG); Chiusa di Pesio, Val Pesio, VIII. 907, R. Gestro leg. (2 ƋƋ, 3 ♀♀ MSNG – Apion atomarium, det. Solari); Elva, Colle di Sam- péyre, env. S, 1,800 m a. s. l., 5. VIII. 2007, C. Giusto leg. (1 Ƌ CG); Pamparato, Monte Alpet, slopes, 1,200 m a. s. l., 26. VIII. 1995, C. Giusto leg. (1 Ƌ CG); Pontechianale, Chianale, 1.790 m, 4. IX. 2005, L. Diotti leg. (4 ƋƋ, 1 ♀ LD); Roburent, San Giacomo, 1,000 m a. s. l., 26. VIII. 1995, C. Giusto leg. (1 Ƌ CG); Sambuco, 1,200 m a. s. l., 8. XII. 2016, C. Giusto leg., on Thymus sp. cfr. T. pulegioides L. (1 Ƌ CG); Sampeyre, 1,000 m a. s. l., 13. VIII. 1996, C. Giusto leg. (1 ♀ CG); Sampeyre: Becetto: Pian Ciattiva, 1,900 m 7. VIII. 2020, C. Giusto leg., on Thymus sp. cfr. T. pulegioides L. (2 ƋƋ, 3 ♀♀ CG); Sampeyre: Becetto: Meire Ruà, env., 1,500 – 1,600 m a. s. l., 7. VIII. 2020, C. Giusto leg., on Thymus sp. cfr. T. pulegioides L. (1 ♀ CG); Stroppo, San Martino, 1,300 m a. s. l., 23. VII. 2003, C. Giusto leg. (1 Ƌ CG); Valdieri, Sant’Anna, 8. X. 1967, G. Bartoli leg., on Thymus sp. (1 ♀ MSNG); Vinadio, Vallone di Sant’Anna, 1,500 m a. s. l., 23. VII. 2003, C. Giusto leg. (2 ƋƋ CG); — Turin Province: Meana di Susa, 7. VIII. 1907, A. Dodero leg. (1 ♀ MSNG); Villar Pellice, 4. VIII. 1971, G. Bartoli leg. (1 ♀ MSNG); Villar Pellice, Villar Pellice, 5. VIII. 1963, G. Bartoli leg. (1 ♀ MSNG); Villar Pellice, 6. VIII. 1961, G. Bartoli leg., on Thymus sp. (1 Ƌ MSNG); — Trentino- Alto Adige: Bolzano Province: Brunico, VII. 1934, G. Binaghi leg. (1 ♀ MSNG – Apion atomarium, det. F. Solari); Castelrotto, Siusi allo Sciliar, Schuster leg. (1 Ƌ CG); Rio di Pusteria, Valle di Valles, 960 m a. s. l., 3. VIII. 2014, A. Paladini leg. (4 ƋƋ, 4 ♀♀ CG); — Trento Province: Avio, Monte Baldo, Prà Alpesina, 1,500 m a. s. l., 2. VIII. 2016, C. Giusto leg., on Thymus sp. cfr. T. pulegioides L. (2 ƋƋ CG); Brentonico, Monte Altissimo di Nago, road to Rifugio Chiesa, 9. IX. 2014, L. Diotti leg. (1 Ƌ, 2 ♀♀ CG); Folgaria, 25. VIII. 1937 (1 Ƌ MSNG); Lavarone, 13 – 20. VIII. 1899, A. Dodero leg. (2 ♀♀ MSNG); Valdaora, 24. VIII. 1964, G. Bartoli leg., on Thymus sp. (1 Ƌ, 1 ♀ MSNG); Valdaora, 10. VIII. 1965, G. Bartoli leg. (3 ƋƋ, 1 ♀ MSNG); Valdaora, 12. VIII. 1966, G. Bartoli leg. (2 ƋƋ MSNG); — Tuscany: Pistoia Province: Abetone Cutigliano, Boscolungo, VII – VIII. 1891, A. Dodero leg. (1 Ƌ MSNG); — Veneto: Belluno Province: Voltago Agordino, 850 m, 12. VIII. 1967, I. Mercati leg. (1 ♀ MSNG); Voltago Agordino, 850 m, 25. VIII. 1979, I. Mercati leg. (1 ♀ MSNG); — Verona Province: Erbezzo, Passo delle Fittanze della Sega, 1,350 m a. s. l., 1. VIII. 2016, C. Giusto leg., on Thymus sp. cfr. T. pulegioides L. (1 Ƌ CG). SLOVENIA — Goriška Regija: Nova Gorica, Nova Gorica, Lokve, VIII. 1992, M. Franciscolo leg. (1 Ƌ CG); — Notranjsko-kraška Regija: Cerknica, Cerkniška Dolina, 550 m a. s. l., 18. VI. 1996, C. Giusto leg. (1 ♀ CG). SPAIN — Andalusia: Granada Province: Sierra de Baza, E slopes, Santa Barbara, 2,400 m a. s. l., 24. VI. 1986, M. Meregalli leg. (1 ♀ CG); — Malaga Province: Sierra de Ronda, 850 m a. s. l., 24. IV. 1980, M. Meregalli leg. (1 ♀ CG); — Catalonia: Lleida Province: Sant Llorenç de Morunys, Port del Compte, La Bòfia, 2,450 m a. s. l., 25. VI. 1979, M. Meregalli leg. (1 ♀ CG). SWITZERLAND — Canton of Valais: Goms: Binn, Giesse, 1,480 m a. s. l., 4 – 5. VII. 2015, C. Germann & C. Giusto leg. (3 ƋƋ, 3 ♀♀ CG); Ernen, Ausserbinn, 1,290 m a. s. l., 6. VII. 2015, C. Giusto leg. (6 ƋƋ, 3 ♀♀ CG); — Westlich Raron: Ausserberg, 1,000 m a. s. l., 5. VII. 2015, C. Giusto leg. (1 ♀ CG).	en	Giusto, Carlo (2021): Squamapion gardinii n. sp. from the Mediterranean Region and notes on some species belonging to the genus Squamapion Bokor, 1923 (Coleoptera, Apionidae). Zootaxa 4941 (2): 217-238, DOI: 10.11646/zootaxa.4941.2.4
03CE87B3FFD3FFBF8DD0FF1E5DCD7F39.taxon	materials_examined	CYPRUS — “ Cypro ”, Baudi leg. (2 ♀♀ MSNG); Troódos, Mesa Potamos, Krios River, picnic area, 11. VIII. 2005, L. Friedman & Y. Antoniou leg. (2 ♀♀ CG).	en	Giusto, Carlo (2021): Squamapion gardinii n. sp. from the Mediterranean Region and notes on some species belonging to the genus Squamapion Bokor, 1923 (Coleoptera, Apionidae). Zootaxa 4941 (2): 217-238, DOI: 10.11646/zootaxa.4941.2.4
03CE87B3FFD3FFBF8DD0FEE65BDF7FF5.taxon	materials_examined	MOROCCO — Meknes: Ifrane Province: Ain Leuh, 11. VI. 2013, F. Angelini leg. (1 ♀ CG).	en	Giusto, Carlo (2021): Squamapion gardinii n. sp. from the Mediterranean Region and notes on some species belonging to the genus Squamapion Bokor, 1923 (Coleoptera, Apionidae). Zootaxa 4941 (2): 217-238, DOI: 10.11646/zootaxa.4941.2.4
03CE87B3FFD3FFBF8DD0FD725D057B29.taxon	materials_examined	FRANCE — Auvergne-Rhône-Alpes: Drôme Department: Saint-May, Gorges de Saint-May, 600 m a. s. l., 17. VII. 2015, R. Casalini leg. (2 ƋƋ, 2 ♀♀ CG; 14 ƋƋ, 10 ♀♀ RC); — Occitanie: Aude Department: Aude (12 ♀♀ MSNG); Montagne d’Alaric, 6. X. 1901 (2 ♀♀ MSNG); — Provence-Alpes-Côte d’Azur: Alpes-Maritimes Department: La Brigue, Chapelle Notre-Dame-des-Fontaines, 3. VIII. 2016, L. Diotti leg. (1 ♀ CG); — Vaucluse Department: Beaumont-du-Ventoux, Mont Ventoux, 8. VII. 2015, R. Casalini leg. (5 ƋƋ, 5 ♀♀ CG; 10 ƋƋ, 4 ♀♀ RC). ITALY — Liguria: Imperia Province: Sanremo, Monte Bignone, 4. VIII. 2016, L. Diotti leg. (1 ♀ CG, 1 ♀ LD); Ventimiglia, Passo del Cornà, slopes E, 550 – 650 m a. s. l., 11. V. 2019, C. Giusto leg., on Tyhmus vulgaris L. (1 ♀ CG); — Savona Province: Castelbianco, Monte Peso Grande, slopes S, 1,000 m a. s. l., 5. IX. 1994, C. Giusto leg. (3 ♀♀ CG); SPAIN — Andalusia: Andalusia, Baudi leg. (1 Ƌ, 1 ♀ MSNG); — Malaga Province: Atajate env., 25. IV. 1980, M. Meregalli leg. (2 ♀♀ CG); Sierra de Ronda, 850 m a. s. l., 24. IV. 1980, M. Meregalli leg. (2 ♀♀ CG); — Comunidad de Madrid: Madrid Province: Santa María de la Alameda, La Paradilla, 1,100 m a. s. l., 27. VI. 1991, C. Giusto leg., on Tyhmus vulgaris L. (1 ♀ CG).	en	Giusto, Carlo (2021): Squamapion gardinii n. sp. from the Mediterranean Region and notes on some species belonging to the genus Squamapion Bokor, 1923 (Coleoptera, Apionidae). Zootaxa 4941 (2): 217-238, DOI: 10.11646/zootaxa.4941.2.4
03CE87B3FFD3FFBF8DD0FDAA5A387C8D.taxon	materials_examined	ISRAEL — Central District of Israel: Shoham Park, 130 m a. s. l., 10. IV. 2014, C. Giusto leg., on Origanum syriacum L. (34 ƋƋ, 25 ♀♀ CG); — Jerusalem District: Ramat Razi’el, 11. III. 2001, L. Friedman leg. (1 ♀ Paratype CG).	en	Giusto, Carlo (2021): Squamapion gardinii n. sp. from the Mediterranean Region and notes on some species belonging to the genus Squamapion Bokor, 1923 (Coleoptera, Apionidae). Zootaxa 4941 (2): 217-238, DOI: 10.11646/zootaxa.4941.2.4
03CE87B3FFD3FFBE8DD0FAD65B657A11.taxon	materials_examined	FRANCE — Auvergne-Rhône-Alpes: Drôme Department: Vassieux-en-Vercors, Col de la Chau, 1,330 m a. s. l., 22. VIII. 1997, C. Giusto leg. (5 ♀♀ CG); — Provence-Alpes-Côte d’Azur: Alpes-Maritimes Department: Gourdon, env. S, 700 m a. s. l., 21. V. 2013, C. Giusto, G. Gardini & S. Zoia leg. (1 ♀ CG); — Occitanie: Hérault Department: Montpellier, H. Lavagne leg. (1 ♀ MSNG). ITALY — Emilia-Romagna: Parma Province, Bedonia, Passo del Tomarlo, 20. VII. 2005, L. Diotti leg. (4 ƋƋ, 2 ♀♀ CG); — Reggio Emilia Province: Carpineti, Montelago, 700 m a. s. l., 8. V. 1993, C. Giusto leg. (1 ♀ CG); — Friuli-Venezia Giulia: Udine Province: Resia, Sella Carnizza, env. E, 900 m a. s. l., 13. VII. 2019, C. Giusto leg., on Thymus sp. cfr. T. pulegioides L. (3 ƋƋ, 2 ♀♀ CG); Venzone, Monte Plauris, Malga Confin env., 1,100 – 1,200 m a. s. l., 10. VII. 2019, C. Giusto leg., on Thymus sp. cfr. T. pulegioides L. (3 ƋƋ, 3 ♀♀ CG); Venzone, Monte Plauris, Malga Confin env., 1,400 – 1,500 m a. s. l., 10. VII. 2019, C. Giusto leg., on Thymus sp. cfr. T. pulegioides L. (3 ♀♀ CG); Venzone, Monte Plauris, slopes S, Graunes, 1,350 – 1,400 m a. s. l., 11. VII. 2019, C. Giusto leg., on Thymus sp. cfr. T. pulegioides L. (4 ƋƋ, 1 ♀ CG); Venzone, Stazione di Carnia, 18. X. 1953, Springer leg. (1 Ƌ CG); — Lazio: Rieti Province: Antrodoco: Monte Giano, Prato Lungo, 1,450 m a. s. l., 12. VII. 1990, E. Colonnelli leg. (4 ƋƋ, 3 ♀♀ CG); Micigliano: Monte Terminillo, slopes W Monte i Porcini, Costa Ghiaiosa, 1,800 m a. s. l., 24. VI. 2017, C. Giusto leg. (8 ƋƋ, 3 ♀♀ CG); — Roma Province: Rocca di Papa Monte Cavo, 900 m a. s. l., 28. X. 1988, C. Giusto leg. (1 ♀ CG); — Liguria: Genoa Province: Favale di Malvaro, Passo della Scoglina, 900 m a. s. l., 3. VII. 1993, C. Giusto leg., on Thymus sp. cfr. T. pulegioides L. (1 Ƌ, 1 ♀ CG); — Savona Province: Urbe, Martina Olba, 484 m a. s. l., 1. VI. 1986, C. Giusto leg., on Thymus sp. cfr. T. pulegioides L. (1 ♀ CG); Urbe, Martina Olba, 484 m a. s. l., 10. IX. 1986, C. Gi- usto leg., on Thymus sp. cfr. T. pulegioides L. (4 ♀♀ CG); Urbe, Martina Olba, 484 m a. s. l., 11. VII. 1983, C. Giusto leg., on Thymus sp. cfr. T. pulegioides L. (2 ♀♀ CG); Urbe, Martina Olba, 484 m a. s. l., 13. VIII. 1994, C. Giusto leg., on Thymus sp. cfr. T. pulegioides L. (1 ♀ CG); Urbe, Martina Olba, 484 m a. s. l., 20. VIII. 1983, C. Giusto leg., on Thymus sp. cfr. T. pulegioides L. (1 Ƌ, 1 ♀ CG); Urbe, Martina Olba, 484 m a. s. l., 25. VII. 1983, C. Giusto leg., on Thymus sp. cfr. T. pulegioides L. (2 ƋƋ CG); Urbe, Martina Olba, 484 m a. s. l., 28. VII. 1981, C. Giusto leg., on Thymus sp. cfr. T. pulegioides L. (1 Ƌ CG); — Piedmont: Biella Province: Piedicavallo, VII – VIII, G. Bartoli leg. (2 ƋƋ MSNG); — Cuneo Province: Caprauna, Colle di Caprauna env., 1,400 m a. s. l., 8. VIII. 1994, C. Giusto leg., on Thymus sp. cfr. T. pulegioides L. (5 ƋƋ, 7 ♀♀ CG); Caprauna, Monte Armetta, 1,700 m a. s. l., 29. VII. 1989, C. Giusto leg., on Thymus sp. cfr. T. pulegioides L. (9 ƋƋ, 14 ♀♀ CG); Caprauna, Monte Armetta, 1,700 m a. s. l., 8. VIII. 1994, C. Giusto & G. Gardini leg., on Thymus sp. cfr. T. pulegioides L. (1 Ƌ, 1 ♀ CG); Montaldo Mondovì, Sant’Anna Collarea, 1,000 m a. s. l., 6. VIII. 1995, C. Giusto leg., on Thymus sp. cfr. T. pulegioides L. (7 ƋƋ, 1 ♀ CG); Ormea, Prale env., 800 m a. s. l., 28. VI. 2003, C. Giusto leg., on Thymus sp. cfr. T. pulegioides L. (1 Ƌ, 1 ♀ CG); Sampeyre: Becetto: Pian Ciattiva, 1,900 m 7. VIII. 2020, C. Giusto leg., on Thymus sp. cfr. T. pulegioides L. (6 ƋƋ, 5 ♀♀ CG); Stroppo, San Martino, 1,300 m a. s. l., 23. VII. 2003, C. Giusto leg., on Thymus sp. cfr. T. pulegioides L. (2 ƋƋ, 3 ♀♀ CG); Vinadio, Vallone di Sant’Anna, 1,500 m a. s. l., 23. VII. 2003, C. Giusto leg., on Thymus sp. cfr. T. pulegioides L. (12 ƋƋ, 9 ♀♀ CG) — Turin Province: Val Pellice, Comba Liussa, 15. VIII. 1961, G. Bartoli leg. (4 ƋƋ, 2 ♀♀ MSNG). POLAND — Łódź Voivodeship: Skiernewice District: Rogów, 28. VII. 1976, M. Wanat leg. (1 Ƌ CG); ROMANIA — Arad District: Milova, 200 m a. s. l., 19. IV. 1983, M. Košťál leg. (1 ♀ CG). SLOVAKIA — Košice Region: Sobrance District: Podhorod’, 8. VII. 1980, M. Košťál leg. (1 ♀ CG); — Prešov Region: Snina District: Nová Sedlica, 15. VII. 1979, M. Košťál leg. (1 ♀ CG); SWITZERLAND — Canton of Valais: Goms: Binn, Giesse, camping, 1,500 – 1,550 m a. s. l., 4 – 5. VII. 2015, C. Germann & C. Giusto leg., on Thymus sp. cfr. T. serpyllum L. (11 ƋƋ, 9 ♀♀ CG); Ernen, Ausserbinn, 1,290 m a. s. l., 6. VII. 2015, C. Giusto leg., on Thymus sp. cfr. T. serpyllum L. (7 ƋƋ, 3 ♀♀ CG).	en	Giusto, Carlo (2021): Squamapion gardinii n. sp. from the Mediterranean Region and notes on some species belonging to the genus Squamapion Bokor, 1923 (Coleoptera, Apionidae). Zootaxa 4941 (2): 217-238, DOI: 10.11646/zootaxa.4941.2.4
