taxonID	type	description	language	source
03C187F69F24FF97FC66FC31FA180D33.taxon	diagnosis	Diagnosis (modified from MARSHALLet al. 2018). Metanotum entirely concealed at dorsal midline (except in Platypedia Uhler, 1888 and Tibicina Kolenati, 1857). Fore wing veins CuP and 1 A unfused (fused in Platypedia and Neoplatypedia Davis, 1920, and partially fused in Gibbocicada gen. nov.). Hind wing veins RP and Munfused at their bases. Male operculum with distinctive S-shape and deeply concave lateral margins (except in Gibbocicada gen. nov.), and with distal margins not reaching distal marginsof tympanal cavities. Abdominal timbal cavity lacking timbal covers or with a partial turned-back rim. Pygofer with distal shoulder undeveloped; pygofer upper lobe usually absent (present in Selymbriini Moulds & Marshall, 2018). Uncus exceedingly long and non-retractable within pygofer. Claspers absent. Aedeagus with ventrobasal pocket present; aedeagus restrained by tubular encapsulation below uncus. Apical part of theca with apair ofleaf-like lateral lobes (exceptin Gibbocicada gen. nov.).	en	Ruschel, Tatiana Petersen (2018): Gibbocicada brasiliana, new genus and new species from Brazil and a key for the genera of Tibicinini (Hemiptera: Auchenorrhyncha: Cicadidae). Acta Entomologica Musei Nationalis Pragae 58 (2): 559-566, DOI: 10.2478/aemnp-2018-0047
03C187F69F24FF91FC5EF9A6FD72084C.taxon	type_taxon	Type genus. Tibicina Kolenati, 1857 (type species Cicada haematodes Scopoli, 1763).	en	Ruschel, Tatiana Petersen (2018): Gibbocicada brasiliana, new genus and new species from Brazil and a key for the genera of Tibicinini (Hemiptera: Auchenorrhyncha: Cicadidae). Acta Entomologica Musei Nationalis Pragae 58 (2): 559-566, DOI: 10.2478/aemnp-2018-0047
03C187F69F24FF91FC5EF9A6FD72084C.taxon	diagnosis	Diagnosis (modified from DISTANT 1905 and LEE 2012). Headincluding eyes narrower than the lateral angle of pronotalcollar and narrowerthan mesonotum. Supra-antennal platesnot meeting the eyes laterally (except in Okanagodes Davis, 1919). Pronotum longer than head, generally more or less distinctly narrowed anteriorly. Mesonotum wider than pronotum (not including the pronotal collar). Mesonotal stridulatory apparatus absent. Male operculum not reaching posterior margins of sternite II. Wings hyaline (tinged in yellowish brownin Subtibicina Lee, 2012); base of wings usually reddish or yellowish orange; veins of fore wings fuscousblack (only in Subpsaltria Chen, 1943). Fore wings: somewhat talc-like and obscurely wrinkled, their greatest width always more than one third of their length; marginal area present. Hind wing first cubital cell width at distal end about equal to second cubital cell. Abdomen more or less cylindricaland attenuated posteriorly. Timbal organs present. Uncus usually very prolonged and prominent. Pygofer upper lobes absent. Claspers absent. Theca not laterally expanded or distally ornamented.	en	Ruschel, Tatiana Petersen (2018): Gibbocicada brasiliana, new genus and new species from Brazil and a key for the genera of Tibicinini (Hemiptera: Auchenorrhyncha: Cicadidae). Acta Entomologica Musei Nationalis Pragae 58 (2): 559-566, DOI: 10.2478/aemnp-2018-0047
03C187F69F24FF91FC5EF9A6FD72084C.taxon	distribution	Distribution. Holarctic, Orientaland Neotropical (Fig. 1).	en	Ruschel, Tatiana Petersen (2018): Gibbocicada brasiliana, new genus and new species from Brazil and a key for the genera of Tibicinini (Hemiptera: Auchenorrhyncha: Cicadidae). Acta Entomologica Musei Nationalis Pragae 58 (2): 559-566, DOI: 10.2478/aemnp-2018-0047
03C187F69F24FF91FC5EF9A6FD72084C.taxon	description	Key to the genera of Tibicinini	en	Ruschel, Tatiana Petersen (2018): Gibbocicada brasiliana, new genus and new species from Brazil and a key for the genera of Tibicinini (Hemiptera: Auchenorrhyncha: Cicadidae). Acta Entomologica Musei Nationalis Pragae 58 (2): 559-566, DOI: 10.2478/aemnp-2018-0047
03C187F69F27FF93FEF5FD2BFDFE092C.taxon	description	(Figs 2 – 18)	en	Ruschel, Tatiana Petersen (2018): Gibbocicada brasiliana, new genus and new species from Brazil and a key for the genera of Tibicinini (Hemiptera: Auchenorrhyncha: Cicadidae). Acta Entomologica Musei Nationalis Pragae 58 (2): 559-566, DOI: 10.2478/aemnp-2018-0047
03C187F69F27FF93FEF5FD2BFDFE092C.taxon	type_taxon	Type species. Gibbocicada brasiliana sp. nov., here designated.	en	Ruschel, Tatiana Petersen (2018): Gibbocicada brasiliana, new genus and new species from Brazil and a key for the genera of Tibicinini (Hemiptera: Auchenorrhyncha: Cicadidae). Acta Entomologica Musei Nationalis Pragae 58 (2): 559-566, DOI: 10.2478/aemnp-2018-0047
03C187F69F27FF93FEF5FD2BFDFE092C.taxon	diagnosis	Diagnosis. The new genus can be distinguished from all other genera in the Tibicinini by the following combination of characters: head short, three times shorter than pronotoum in dorsal view (Fig. 6); lateral ocelli almost reaching the anterior margin of pronotum in dorsal view (Fig. 6); anterior margin of lateral lobe of pronotum reaching the eyes (Fig. 6); pronotum and mesonotum pronounced dorsally in lateral view (Fig. 5); dorsal midline of pronotal collar very broad, as wide as the diameter of eye (Fig. 6); lateral angle of pronotal collar laterally pronounced (Fig. 6); meracanthus large, of the same length as operculum (Fig. 9); fore wings with the median and cubitus anterior veins rising from the same point in the basal cell (Fig. 3); tergite 1 fully covered by cruciform elevation; uncus posteriorly developed (Fig. 13); basal lobe long, almostreaching the uncus (Fig. 13). The new genus can be distinguished from Okanagodes by the distance between the lateral ocellus and eyes greater than diameter of ocellus, and by the postclypeus short, slightly prominent relative to the anterior margin of head in dorsal view; from Clidophleps by the medial cell notmuch narrower than radial cell; from Okanagana by the head (including eyes) broader than the pronotum; from Paharia, Subpsaltria, and Tibicina by the pronotum long, three timeslonger than head, the aedeagus not elongated nor curved anteriorly; from Subtibicina by the fore wings being broad (less than three times the length), not tinged with yellowish brown, and by the uncus being arched and not flat; and from Tibicinoides by the base of wings colorless, and the medial cell of fore wings almost twice longer than ulnar cell 3.	en	Ruschel, Tatiana Petersen (2018): Gibbocicada brasiliana, new genus and new species from Brazil and a key for the genera of Tibicinini (Hemiptera: Auchenorrhyncha: Cicadidae). Acta Entomologica Musei Nationalis Pragae 58 (2): 559-566, DOI: 10.2478/aemnp-2018-0047
03C187F69F27FF93FEF5FD2BFDFE092C.taxon	description	Description. Head (Fig. 6) (including eyes) wide, slightly broader than pronotum (excluding the pronotal collar), and narrower than mesonotum; head short, three times shorter than pronotoum in dorsal view. Lateral ocelli widelyseparated and almost reaching the anteriormargin of pronotum. Vertex wide, the distance between the lateral ocellus and eyes greater than diameter of ocellus. Supra-antennal plates not prominent and not meeting eye. Postclypeus short and narrow in dorsal view, its width equal to the distance between the lateral ocelli; slightly prominent relative to the anterior margin of head in dorsal view; rectangular in ventral view (Fig. 7) and obtuse in lateral view. Pronotum and mesonotum pronounced dorsally in lateral view (Fig. 5). Pronotum (Fig. 6) long, three times longer than head, not depressed laterally; anterior margin of lateral lobe of pronotum reaching the eyes; pronotal collar broad in dorsal view, as wide as diameter of eye; lateral angle of pronotal collar laterally pronounced, exceeding the lateral margin of mesonotum; lateralpart of pronotalcollar narrow relative to lateral angle. Mesonotal stridulatory apparatus absent. Operculum (Fig. 9) broad, the lateral andposterior margins slightly convex, and distal margins not reaching thedistal margins of tympanal cavities. Meracanthus (Fig. 9) large, of the same length as operculum. Basisternum 3 flat, covered by the meso- and metacoxae. Cruciform elevation with the central and lateral areas flat. Profemora armed with two spines, the primary one the longest and parallel to the posterior margin of femur, and the secondary one sharp and straight. Tarsi three-segmented. Fore wings (Fig. 2) hyaline; broad, width almost three times the length; with eight apical cells; the median and cubitus anterior veins rising from the same point in the basal cell; medial cell almost twice longer than ulnar cell 3 and not much narrower than radial cell; hind wings (Fig. 2) about half the length of fore wings, with six apical cells. Male timbal covers absent (Fig. 10). Tergite 1 fully covered by cruciform elevation. Sternite II (Fig. 8) distant from metacoxae. Uncus longer than sternite VIII, elevated and posteriorly developed. Basal lobe of pygofer long, almost reaching the uncus (Fig. 13). Aedeagus not elongated nor curved anteriorly (Fig. 12).	en	Ruschel, Tatiana Petersen (2018): Gibbocicada brasiliana, new genus and new species from Brazil and a key for the genera of Tibicinini (Hemiptera: Auchenorrhyncha: Cicadidae). Acta Entomologica Musei Nationalis Pragae 58 (2): 559-566, DOI: 10.2478/aemnp-2018-0047
03C187F69F27FF93FEF5FD2BFDFE092C.taxon	etymology	Etymology. The name is derived from the Latin adjective gibbus (= humped) and the noun cicada (= cicada). Itrefers to the pronotum and mesonotum which are both pronounced dorsally in lateral view. The gendre is feminine.	en	Ruschel, Tatiana Petersen (2018): Gibbocicada brasiliana, new genus and new species from Brazil and a key for the genera of Tibicinini (Hemiptera: Auchenorrhyncha: Cicadidae). Acta Entomologica Musei Nationalis Pragae 58 (2): 559-566, DOI: 10.2478/aemnp-2018-0047
03C187F69F27FF93FEF5FD2BFDFE092C.taxon	discussion	Comments. Based on the diagnosis of the subfamily proposed by MARSHALL et al. (2018), Gibbocicada gen. nov. clearly belongs to the Tibicininae by the metanotum concealed at dorsal midline (Fig. 2); the hind wing veins RP and Munfused at their bases (Figs 3, 4); the distal marginsof male operculum not reachingthe distal margins of tympanal cavities and basallynot extendingbeyond meracanthus, but notin S-shape (Fig. 9); males lackingtimbal covers (Fig. 10); pygofer distal shoulder undeveloped; pygofer upper lobe absent; uncus very long and not retractable within pygofer; claspers absent; and the presence of a ventrobasalpocket (Figs 12 – 14). The forewing veins CuP and 1 A are unfused in most genera of Tibicininae, but fused in Platypedia Uhler, 1888 and Neoplatypedia Davis, 1920 (Platypediini) (MARSHALLet al. 2018). Gibbocicada gen. nov. presents these veins partially fused, i. e. thebasal third of CuP and 1 A are unfused (Fig. 3). Among all tribes of Tibicininae, Gibbocicada gen. nov. is more similar to the genera classified in Tibicinini, with the pronotum more or less distinctly narrowed anteriorly (Fig. 6); fore wings talc-like and obscurely wrinkled, their greatest breadth more than one third of their length (Fig. 2); abdomen subcylindrical (Fig. 2); and uncus longer than sternite VIII (DISTANT 1905). LEE (2012) listed other characteristics shared by the genera of Tibicinini that Ialso observed in Gibbocicada gen. nov.: hind wing first cubital cell width at distal end about equal to second cubital cell; male operculum not reaching posterior margins of sternite II (Fig. 9); pygofer upper lobes absent, claspers absent and uncus very long (Figs 13, 14). The new genus cannot be placed in the other tribes of Tibicininae for the lack of their diagnostic characters, i. e. Tettigadini because the species in the tribe are characterized by the lateral margins of the pronotum dilated and usually medially angulate (DISTANT 1905), and by a mesonotal stridulatory apparatus (JACOBI 1907, BOULARD 1976, MOULDS 2005); Selymbrini because they have supra-antennal plates almostreaching the eye, a rounded postclypeus, hind coxae without lateral protruberance and a very short theca laterally expanded and distally ornamented (MARSHALL et al. 2018); Chilecicadini Sanborn, 2014 because they havethe anteriorpronotum about as wide as the posterior pronotum, the lateral angle of the pronotal collar anteriorto thehind margin, and the partiallyexcavated lateral margin of the pronotal collar (SANBORN 2014); and Platypediini Kato, 1932 because the tribe includes only species without timbal organs (MOULDS 2005).	en	Ruschel, Tatiana Petersen (2018): Gibbocicada brasiliana, new genus and new species from Brazil and a key for the genera of Tibicinini (Hemiptera: Auchenorrhyncha: Cicadidae). Acta Entomologica Musei Nationalis Pragae 58 (2): 559-566, DOI: 10.2478/aemnp-2018-0047
03C187F69F20FF93FEA2FDD3FBF60F2C.taxon	description	(Figs 2 – 18)	en	Ruschel, Tatiana Petersen (2018): Gibbocicada brasiliana, new genus and new species from Brazil and a key for the genera of Tibicinini (Hemiptera: Auchenorrhyncha: Cicadidae). Acta Entomologica Musei Nationalis Pragae 58 (2): 559-566, DOI: 10.2478/aemnp-2018-0047
03C187F69F20FF93FEA2FDD3FBF60F2C.taxon	materials_examined	Type locality. Brazil, Ceará state, Mucambo municipality, Carquejo village, ca. 3 ° 52 ′ S, 40 ° 44 ′ W. Type material. HOLOTYPE:, ʻBRAZIL / Carquejo / Est Ceará / Dirings // IV 1960 ʼ (MZSP, dry-mounted ona pin; Fig. 2). PARATYPES: 2 1 ♀, same data as the holotype (MZSP).	en	Ruschel, Tatiana Petersen (2018): Gibbocicada brasiliana, new genus and new species from Brazil and a key for the genera of Tibicinini (Hemiptera: Auchenorrhyncha: Cicadidae). Acta Entomologica Musei Nationalis Pragae 58 (2): 559-566, DOI: 10.2478/aemnp-2018-0047
03C187F69F20FF93FEA2FDD3FBF60F2C.taxon	description	Description. Male. Bodycoloration greenish yellow (Fig. 2), only the head with black markings. Head. Vertex with transverse black band departing from each eye, surrounding the ocelli and extending to the posterior area behind the eyes. Eyes rounded, slightly projecting laterally beyond anterior angles of pronotum. Lateral ocelli widely separated and laterally projected, higher than median ocellus in frontal view (Fig. 6). Postclypeus unstained, with ten transverse grooves. Central sulcus shallow and slender. Anteclypeus and carina tawny, apex half-moon shaped. Lorum with several silver setae (Fig. 7). Labium long, reaching the metacoxae (Fig. 8). Thorax. Pronotum (Fig. 6) with lateral and sub-lateral lobes with wrinkles. Mesonotum (Fig. 2) with submedian andlateral sigillae slightly stained in pale green. Operculum (Fig. 9) broad, widely spacedbythe metacoxae, and laterally directed; posterior margin slightly convex, lateral margin very short. Meracanthus large, coveringthe internal angle of operculum in dorsal view. Proximal margins of metacoxae forming acute projections (Fig. 8). Wings hyaline. Fore wings (Figs 2 – 3): basal cell slightly smoky, twice as long as wide; posterior vein ofbasal cell arched; cubitus anterior vein slightly concave; cubital cell wider than clavus; medial cell twice as long as the ulnar cell 3, almost half of the length of fore wing; radius anterior vein closely aligned with subcosta for its length, divergent at apex; median vein long, reaching half thelength of radial cellproximal to M 3 + 4 fork; apical cell 1 longer than any other apical cell. Abdomen (Fig. 2) subcylindrical, fusiform, short, the length equal to the combined length of head and thorax in dorsal view. Timbals elongate laterally, posteriorly directed, anterior margin of the same length as anterior margin of tergite 1 (Fig. 10). Sternite VII subrectangular, with posterior margin concave and slightly emarginated; sternite VIII triangular (Fig. 11). Genitalia (Figs 12 – 14). Pygofer distalshoulder undeveloped. Basal lobe of pygofer long, reaching uncus. Uncus long with setae, tapering to apex, thelateralmargins folding to aedeagus. Aedeagus broad, taperingto apex. Theca with ventral branches, apex of theca beak-shaped in lateral view, dorsally grooved and bifurcated on the anterior margin, with wrinkles. Vesica originating in a fissure from basal plate to the distal third of the theca. Apex of theca with two slender sclerotized branches closely spaced on the fissure of vesica. Female (Figs 15 – 18). Same characteristics as in the male, but body a little more yellowish (Fig. 15). Operculum similar to that of the male, with the lateral margin very short and posterior margin slightly convex (Fig. 16). Meracanthus large, longer than operculum, covering the internal angle of operculum in dorsal view (Fig. 16). Sternite VII longer than any other, posterior margin with an obtuse middle groove reaching more than half of sternite, formingtwo lobes (Fig. 17). Lateral margins of abdominal segment 9 touching medially (Fig. 17). Ovipositor sheath shorter than tip of dorsal beak (Fig. 18). Measurements (in mm). Males (n = 3, values for the holotypein parentheses) lb 23.0 – 25.1 (25.1); wh 6.21 – 6.66 (6.48); lh 1.08 – 1.26 (1.17); wp 7.56 – 8.28 (8.28); lp 3.33 – 3.96 (3.96); wm 6.48 – 6.93 (6.93); lm 4.23 – 4.77 (4.77); lfw 31.1 – 34.6 (34.6); wfw 11.3 – 12.9 (12.9); lhw 18.3 – 20.3 (20.3); whw 7.7 – 8.8 (8.6). Female (n = 1): lb 26.8; wh 6.66; lh 1.26; wp 8.46; lp 3.78; wm 7.2; lm 5.13; lfw 34; wfw 13; lhw 19.6; whw 8.8.	en	Ruschel, Tatiana Petersen (2018): Gibbocicada brasiliana, new genus and new species from Brazil and a key for the genera of Tibicinini (Hemiptera: Auchenorrhyncha: Cicadidae). Acta Entomologica Musei Nationalis Pragae 58 (2): 559-566, DOI: 10.2478/aemnp-2018-0047
03C187F69F20FF93FEA2FDD3FBF60F2C.taxon	etymology	Etymology. The name refers to the country of the type locality. Adjective.	en	Ruschel, Tatiana Petersen (2018): Gibbocicada brasiliana, new genus and new species from Brazil and a key for the genera of Tibicinini (Hemiptera: Auchenorrhyncha: Cicadidae). Acta Entomologica Musei Nationalis Pragae 58 (2): 559-566, DOI: 10.2478/aemnp-2018-0047
03C187F69F20FF93FEA2FDD3FBF60F2C.taxon	distribution	Distribution. Brazil (Ceará).	en	Ruschel, Tatiana Petersen (2018): Gibbocicada brasiliana, new genus and new species from Brazil and a key for the genera of Tibicinini (Hemiptera: Auchenorrhyncha: Cicadidae). Acta Entomologica Musei Nationalis Pragae 58 (2): 559-566, DOI: 10.2478/aemnp-2018-0047
