identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03C287AAFFFD0B05FF357869FE4EFE80.text	03C287AAFFFD0B05FF357869FE4EFE80.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Actinocyclus cuneiformis (Wallich) F. Gomez, Lu Wang & Senjie Lin 1860	<div><p>Actinocyclus cuneiformis (Wallich) F. Gómez, Lu Wang &amp; Senjie Lin Figs 1, 2</p><p>Basionym.:  Hemidiscus cuneiformis Wallich</p><p>Ref. illus.: Hustedt 1927 –1930, p. 904, fig. 542d; Ricard 1987, Figs 30–34; the images in Round et al. 1990, pp. 192–193, are also evidently this species; Gómez et al. 2017, figs 1–23</p><p>Samples: GU66F-4</p><p>Dimensions: long axis 71–74 µm, short axis 41–45 µm</p><p>Diagnostics: Distinctively shaped, lens-shaped in valve view but the frustules wedge-shaped (cuneiform) because the valve faces are at an angle to one another. Areolae with a series of radiating patterns around the margin, especially on the more strongly curved side. Large rimoportulae visible in the oblique view. A pseudonodulus is present on the flatter side.</p><p>Comments: Two valves observed in one sample. The ventral curvatures of  A. cuneiformis vary from straight to sinuous and Hustedt (1927 – 1931) combined several older Eudotia species as varieties. Although Round et al. (1990) say that the valves are cuneiform, it seems to us that Wallich’s (1860) epithet refers to the shape of the frustule (described by Round et al. 1990 as “shaped like orange segments”), rather than to the shape of the valve. The genus  Hemidiscus was subsumed into  Actinocyclus by Gómez et al. (2017); they did not discuss any varieties of  H. cuneiformis but their images, like ours, conform to  var. ventricosa (Castracane) Hustedt. We have therefore omitted any varietal designation.</p></div>	https://treatment.plazi.org/id/03C287AAFFFD0B05FF357869FE4EFE80	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobban, Christopher S.;Witkowski, Andrzej	Lobban, Christopher S., Witkowski, Andrzej (2023): Marine benthic diatoms of Guam: new records, Dictyoneis apapae sp. nov., and updates to the checklist. Micronesica 2023 (2): 1-75, DOI: 10.5281/zenodo.12118784
03C287AAFFFE0B05FF357C5DFD38FD23.text	03C287AAFFFE0B05FF357C5DFD38FD23.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amphora spectabilis Gregory 1857	<div><p>Amphora spectabilis Gregory Figs 3, 4</p><p>Ref. illus.: Levkov 2009, p. 265, pl. 114, figs 1–5, pl. 252, figs 1–5 (SEM).</p><p>Samples: GU55B-5</p><p>Dimensions: Length 30 µm, width 12 µm, stria density 8 in 10 µm (dorsal) and 13 in 10 µm (ventral)</p><p>Diagnostics: More strongly and coarsely striated on the dorsal surface, dorsal striae often biseriate and bifurcating towards the sternum, ventral striae uniseriate.</p><p>Comment: Our specimens (seen also in Yap and Palau) are much shorter than the 68–129 µm given by Levkov (2009). A Palauan specimen (Fig. 4) seen in SEM shows that the striae are mostly biseriate, the bifurcations resulting from the two rows diverging; these images conform to the specimens shown by Levkov in SEM.</p></div>	https://treatment.plazi.org/id/03C287AAFFFE0B05FF357C5DFD38FD23	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobban, Christopher S.;Witkowski, Andrzej	Lobban, Christopher S., Witkowski, Andrzej (2023): Marine benthic diatoms of Guam: new records, Dictyoneis apapae sp. nov., and updates to the checklist. Micronesica 2023 (2): 1-75, DOI: 10.5281/zenodo.12118784
03C287AAFFFE0B04FF357E09FAE1FE7D.text	03C287AAFFFE0B04FF357E09FAE1FE7D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amphora subhyalina Podzorski & Hakansson	<div><p>Amphora subhyalina Podzorski &amp; Håkansson Figs 5–7</p><p>Syn.:  Amphora insulana Stepanek &amp; Kociolek</p><p>Ref. illus.: Podzorski &amp; Håkansson 1987, p. 52; pl. 15, fig. 10, pl. 51, figs 7, 8; Stepanek &amp; Kociolek 2018, p. 14, pl. 5, figs 13–16, pl. 8, figs 1–4.</p><p>Samples: GU44Y-13, GU44Z-15, GU52Q-1a, GU66G-2, GU68A-2</p><p>Dimensions: Length 27–45 µm, width 7–8 µm; dorsal striae 36 in 10 µm in the middle increasing to 39 toward the apices, ventral striae 38 in 10 µm.</p><p>Diagnostics: Valves and copulae frequently forming a hyaline oval pattern in LM. In SEM, more prominent elongated areolae on either side of the dorsal rib with the rest of the dorsal striae comprising small areolae; prominent conopea at the central raphe endings.</p><p>Comments: As far as we can tell from the SEM illustrations of valves in Podzorski &amp; Håkansson (1987) and Stepanek &amp; Kociolek (2018), the same species is being described and therefore the former name has priority. Although clearly shown in SEM by Stepanek &amp; Kociolek (2018), they did not include a written description of ultrastructure. Podzorski &amp; Håkansson (1987), although showing SEMs, also did not specify stria densities. Our SEM images show dorsal striae of irregular transapical slits, ending with a long, prominent slit at valve margin, mantle sharply delimited by a ridge and bearing a similar series of prominent slits (Figs 6, 7). Small central area on dorsal side of central nodule, and central striae somewhat more prominent. Ventral valve face with single row of slit-like areolae, interrupted by a conopeum developed ventrally and the ventrally deflected raphe. Internally, as shown in Stepanek &amp; Kociolek (2018, pl. 8, fig. 4) the proximal raphe endings are straight. The girdle bands have a single line of slits along the narrow ventral part and faint striae on the wide dorsal part (Fig. 7 arrow). The lack of a ventral raphe ledge is indicative of  Halamphora rather than  Amphora, as summarized by Stepanek &amp; Kociolek (2018, pp. 8–9 vs 28–29), but the broad girdle bands are indicative of  Amphora . Neither Podzorski &amp; Håkansson (1987) nor Stepanek &amp; Kociolek (2018) showed girdle bands. Their materials were, respectively, from Palawan (Philippines) and Florida Keys.</p><p>These fairly large, hyaline frustules resemble  Halamphora hyalina (Kützing) Rimet &amp; R.Jahn (Rimet et al. 2018) and  Halamphora pseudohyalina (Simonsen) Stepanek &amp; Kociolek (2018) (neither yet recorded from our region) in the presence of the distinct central area and a ventral surface;  H. pseudohyalina has fine dorsal striae 28–30 in 10 µm barely distinguished in LM and ventral striae ca 65 in 10 µm, whereas as  H. hyalina and  A. vaughanii Giffen have clearly visible striae, 20–23 in 10 µm and 28–30 in 10 µm, respectively.  Amphora hyalina, reported from Yap by Navarro &amp; Lobban (2009, figs 97, 98), has an exposed, dorsally deflected raphe with a conopeum extending dorsally on the dorsal raphe ledge; the striae are finely biseriate, the areolae consistently small, apicallyelongated slits.  Amphora vaughanii was recorded from Guam (Lobban et al. 2012, pl. 56, figs 7–9), it has biseriate striae of very fine pores on the dorsal surface and a line of very closely spaced slits on the mantle, conopea absent; its position in  Halamphora or  Amphora has not yet been considered.</p></div>	https://treatment.plazi.org/id/03C287AAFFFE0B04FF357E09FAE1FE7D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobban, Christopher S.;Witkowski, Andrzej	Lobban, Christopher S., Witkowski, Andrzej (2023): Marine benthic diatoms of Guam: new records, Dictyoneis apapae sp. nov., and updates to the checklist. Micronesica 2023 (2): 1-75, DOI: 10.5281/zenodo.12118784
03C287AAFFFF0B04FF357CCBFCA0FD34.text	03C287AAFFFF0B04FF357CCBFCA0FD34.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Auricula densistriata K. Osada (Osada 1997	<div><p>Auricula densistriata Osada Figs 8, 9</p><p>Lobban et al. (2012) reported  A. complexa (Gregory) Cleve from Guam, but J.S. Park et al. (2018) noted that the valve size and stria density of the voucher specimens match  Auricula densistriata K. Osada (Osada 1997) . Stria density for  A. complexa is 15–20 in 10 µm (Peragallo &amp; Peragallo 1897 – 1908), but Lobban et al. reported 26–28 in 10 µm. In SEM (Figs 8, 9) we measured 27 striae.  Auricula complexa species was correctly reported from Federated States of Micronesia — Yap by Navarro &amp; Lobban (2009) and Chuuk by Park et al. (2022) —and may well be present in Guam but we have no evidence of it, so for now remove it from the flora.</p></div>	https://treatment.plazi.org/id/03C287AAFFFF0B04FF357CCBFCA0FD34	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobban, Christopher S.;Witkowski, Andrzej	Lobban, Christopher S., Witkowski, Andrzej (2023): Marine benthic diatoms of Guam: new records, Dictyoneis apapae sp. nov., and updates to the checklist. Micronesica 2023 (2): 1-75, DOI: 10.5281/zenodo.12118784
03C287AAFFFF0B04FF357E12FEA8FB13.text	03C287AAFFFF0B04FF357E12FEA8FB13.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bacillaria J.F.Gmelin 1788	<div><p>“  Bacillaria Group B” sensu Schmid var. tumidula Figs 10–14</p><p>Previous Micronesian record: J.S. Park et al. 2022, p. 45, figs 144, 145</p><p>Samples: GU21AM-2, GU21AN-1; GU52J-3, GU52K-4, GU52S-2; GU58G-4D</p><p>Diagnostics: Colonial, as in  B. paxillifera, the “slide rule” diatom, but inflated in the middle. Uniseriate striae with external costae.</p><p>Comment: As noted by Lobban et al. (2012) for the nominate form, marine forms were orphaned when Jahn &amp; Schmid (2007) redefined  B. paxillifera as freshwater/brackish, naming several new species, but excluding three marine groups distinguished by Schmid (2007). Our assignment of this variety to Group B, based on the nominate form, has not been verified and remains in limbo until Jahn completes her analysis. The short cristae at the central area are characteristic but we have images of specimens without them and without costae; further progress awaits completion of work on this group.</p></div>	https://treatment.plazi.org/id/03C287AAFFFF0B04FF357E12FEA8FB13	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobban, Christopher S.;Witkowski, Andrzej	Lobban, Christopher S., Witkowski, Andrzej (2023): Marine benthic diatoms of Guam: new records, Dictyoneis apapae sp. nov., and updates to the checklist. Micronesica 2023 (2): 1-75, DOI: 10.5281/zenodo.12118784
03C287AAFFFF0B02FF357839FEC0FE1F.text	03C287AAFFFF0B02FF357839FEC0FE1F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Berkeleya hyalina (Round & M. E. Brooks) Cox	<div><p>Berkeleya cf. hyalina (Round &amp; M.E.Brooks) Cox Figs 15–25</p><p>Ref. illus.: Round &amp; Brooks 1973, figs 1–4, 6–16; Cox 1975a, figs 7, 33, 35; Cox 1975b, figs 9, 18, 22, 23; Chastain &amp; Stewart 1985, figs 1–14; Lobban 1985, figs 3–5; Witkowski et al. 2000, p. 156, pl.62, fig. 22</p><p>Samples: GU77A-2</p><p>Observations: Colonial (tube-dwelling) species forming wide, branched colonies with many narrow tubes bundled together, each narrow tube containing a single file of cells (Figs 15–17); plastids Hshaped (Fig. 18). Valves hyaline, 45–51 µm long, 6 µm wide, striae parallel except convergent near apices, 37 in 10 µm. Cells linear-lanceolate with broadly rounded ends (Fig. 19). In SEM, striae parallel except radiate near the apices, areolae quadrangular except a row of larger areolae along one side of the raphe (Figs 20, 21). Central raphe endings separated by 7.1–8.6 µm, curved slightly away from the side with the larger areolae.</p><p>Comments: These microscopic colonies are very much smaller than those in southern California but that is not unusual for Guam tube-dwelling diatoms (so far only  Homoeocladia [ Nitzschia]  martiana C.Agardh colonies found at seaweed size; Lobban &amp; Tsuda 1993). Our cells appear to differ from  B. hyalina in larger size, wider separation of central raphe endings and shape of the apex, from  B. fragilis in shape and longer central area, and from  B. micans, which is longer but narrower with coarser striae and very long central area (Table 1). However, variability in some of these characters warrants caution.</p><p>First, there are discrepancies between Chastain &amp; Stewart’s (1985) California populations compared to type material, some of them noted by those authors, i.e., the stria density, the shape of the apex, and the extent of the axial row of enlarged areolae. The difference in stria density might be significant but Chastain and Stewart’s statement is broad and imprecise (“approximately 32–38” in 10 µm), and Round &amp; Brooks’ is even more vague (“more than 40”). Lobban (1985) in a floristic survey including southern California, reported striae of  B. hyalina to be “ca. 45 in 10 µm… visible only in the electron microscope” but no SEM was published. Perhaps a more significant difference lies in the hemilanceolate depression formed by the very long areolae in San Diego populations, which Chastain &amp; Stewart noted as a “signal feature,” though saying that “the row of enlarged pores… appears to be somewhat more prominent” than in the type material. All species of  Berkeleya in Table 1 are reported in the literature to have a row of enlarged areolae along one side of the raphe, so what is distinctive in the San Diego populations is the width and the depression. Round &amp; Brooks (1973, figs 7, 8, 15) noted that the larger pores appeared to be in a “slight depression;” the pores are trapezoidal, wider toward the axis. Chastain &amp; Stewart (1985, figs 6, 10) show a single row of larger areolae in their TEM (fig. 6) but in the SEM (fig. 10) most in the widest part are divided. We have seen such depressed zones in a few samples that we could not identify to species (Figs 24, 25) and even, rarely, in the present colony. In all these, the distance between central raphe endings was &lt;7 µm but perhaps not significantly different from the majority of the colony (in the two specimens in Figs 24, 25 = 4.2 and 6.2 µm but in Guam specimens up to 6.7 µm). This brings into question whether the depressed area is a consistent enough character to use for taxonomy, and how much distance between central raphe is needed to separate species. Chastain &amp; Stewart noted the great difference in latitude and water temperature between Togo (type locality) and San Diego, to which we might add, the different ocean basins.</p><p>Second, there are the differences between our material and both of the other two populations which require additional material from Guam and re-examination of type material (or better, more samples from Togo as well!). At this point both the Guam and Togo populations can be considered inadequately studied. The biggest problem with the Togo material is the vague stria density given by Round &amp; Brooks; we tried counting it from the published images (Round &amp; Brooks 1973, figs 8, 15) and got answers very much higher than the&gt;40 given, but only magnifications are given for the images and perhaps the plates were reduced in publication without correction of the captions. Thus, we have identified our specimens only as “  cf. hyalina ” and shown their morphology for future reference.</p></div>	https://treatment.plazi.org/id/03C287AAFFFF0B02FF357839FEC0FE1F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobban, Christopher S.;Witkowski, Andrzej	Lobban, Christopher S., Witkowski, Andrzej (2023): Marine benthic diatoms of Guam: new records, Dictyoneis apapae sp. nov., and updates to the checklist. Micronesica 2023 (2): 1-75, DOI: 10.5281/zenodo.12118784
03C287AAFFF90B02FF357CDCFEA4FB47.text	03C287AAFFF90B02FF357CDCFEA4FB47.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Biddulphiella tridens (Ehrenberg) Ashworth & Sims	<div><p>Biddulphiella tridens (Ehrenberg) Ashworth &amp; Sims Figs 26–29</p><p>Syn.:  Biddulphia tridens (Ehrenberg) Ehrenberg</p><p>Biddulphia tuomeyi (J.W. Bailey) Roper</p><p>Ref. illus.: Schmidt et al. 1874–1959, pl. 118, figs 1–7 and pl. 119, figs 1–6; Peragallo &amp; Peragallo 1897 –1908, pl. 94, fig. 2; Hustedt 1927 –1930, fig. 491; Navarro 1981, fig. 15; Lobban &amp; Jordan 2010, fig. 5q, 5r (all as  B. tuomeyi); López Fuerte et al. 2010, pl. 12, figs 6–7; Ashworth et al. 2013, figs 5b, 5i; Sims et al. 2023, fig. 13</p><p>Samples: GU7Y-4</p><p>Dimensions: Length 42–63 µm, width 27–30 µm</p><p>Diagnostics: Deeply sculpted valve in girdle view, central dome flanked by two smaller ones that abut or are themselves separated from the tall pseudocellus-bearing horns on the two poles. Many spines on the domes and the central dome with two rimoportulae with very long external tubes. The deep grooves (sulci) (Fig. 29) were used to distinguish Biddulphiella from  Biddulphia (Sims et al. 2023) .</p><p>Comments: Differs from  Biddulphia biddulphiana (J.E.Smith) Boyer (Navarro &amp; Lobban 2009, figs 28–31) in the more elongate, rostrate valve outline and number of domes. VanLandingham (1968, p. 562) commented on which specific epithet had priority and listed the extensive synonymy under  B. tridens .</p></div>	https://treatment.plazi.org/id/03C287AAFFF90B02FF357CDCFEA4FB47	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobban, Christopher S.;Witkowski, Andrzej	Lobban, Christopher S., Witkowski, Andrzej (2023): Marine benthic diatoms of Guam: new records, Dictyoneis apapae sp. nov., and updates to the checklist. Micronesica 2023 (2): 1-75, DOI: 10.5281/zenodo.12118784
03C287AAFFF90B02FF3579E5FE7FF9D9.text	03C287AAFFF90B02FF3579E5FE7FF9D9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Biremis ambigua (Cleve) D. G. Mann	<div><p>Biremis ambigua (Cleve) D.G.Mann Figs 30, 31</p><p>Ref. illus.: Cox 1990, figs 19–28; Sabbe et al. 1995, figs 34, 35, 38, 51, 52; Witkowski et al. 2000, p. 158, pl. 155, figs 2–6; Tatarek &amp; Wiktor 2005</p><p>Samples: GU52O-4</p><p>Dimensions: Length 38–54 µm, width ca. 6 µm; striae 9–10 in 10 µm</p><p>Diagnostics: Linear, symmetrical valve, similar in size and stria density to  B. ridicula (see below), which is strongly asymmetrical.</p><p>Comments:  Biremis is characterized by alveolate striae. Several smaller species have been observed but not yet identified.</p></div>	https://treatment.plazi.org/id/03C287AAFFF90B02FF3579E5FE7FF9D9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobban, Christopher S.;Witkowski, Andrzej	Lobban, Christopher S., Witkowski, Andrzej (2023): Marine benthic diatoms of Guam: new records, Dictyoneis apapae sp. nov., and updates to the checklist. Micronesica 2023 (2): 1-75, DOI: 10.5281/zenodo.12118784
03C287AAFFF90B01FF357B6EFD68FD4F.text	03C287AAFFF90B01FF357B6EFD68FD4F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Biremis ridicula (Giffen) D. G. Mann	<div><p>Biremis ridicula (Giffen) D.G. Mann Figs 32–37</p><p>Ref. illus.: Giffen 1976, figs 10–13 (as  Amphora ridicula); Round et al. 1990 pp. 548–549 (figs c, d, but not identified to species); Witkowski et al. 2000, pl. 154, figs 14–16</p><p>Samples: GU52O-4</p><p>Dimensions: Length 46–74 µm, width of valve 9 µm, width of frustule in girdle view 24 µm; striae 7 in 10 µm</p><p>Diagnostics:  Amphora -like frustule, asymmetrical (Figs 32, 35). Valve asymmetrical (Figs 34, 36), lanceolate, with short, broad striae along each margin and the raphe strongly displaced to the “ventral” striae, leaving strongly asymmetrical axial area. Striae alveolate (Fig. 37).</p><p>Comments: This species is similar in size and stria density to  Biremis ambigua (see above), but that species is symmetrical. The relatively large frustules are shaped like  Amphora arenaria Donkin, with which it co-occurred in this sample from calcareous sand, but the striae are very distinctive. Round et al. (1990) described the areolae in this genus as consisting of two incompletely separated chambers, closed internally by a cribrum, and opening externally by slit-like foramina, one at each end of the areola, resulting in two longitudinal lines that are the most evident feature in external SEM views. Witkowski et al. (2000) mention this species being reported only from South Africa and the North Sea (Europe), populations for which there is no SEM yet; a comparison needs to be made to confirm the identity of the Guam specimens.</p></div>	https://treatment.plazi.org/id/03C287AAFFF90B01FF357B6EFD68FD4F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobban, Christopher S.;Witkowski, Andrzej	Lobban, Christopher S., Witkowski, Andrzej (2023): Marine benthic diatoms of Guam: new records, Dictyoneis apapae sp. nov., and updates to the checklist. Micronesica 2023 (2): 1-75, DOI: 10.5281/zenodo.12118784
03C287AAFFFA0B01FF357F9DFB31F99E.text	03C287AAFFFA0B01FF357F9DFB31F99E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Campylodiscus robertsianus Greville 1863	<div><p>Campylodiscus robertsianus Greville Fig. 38</p><p>Refs.: Greville 1863, p. 14, pl. I, fig. 5; Schmidt et al. 1874–1959, pl. 17, figs 8, 9; pl. 207, fig. 22; Williams 1988, pl. 27, figs 3, 4</p><p>Samples: GU74B-4</p><p>Dimensions: Diam. 100 µm</p><p>Diagnostics: Lines of large, paired oval pores running along lines from the margin to the central area.</p><p>Comments: Greville noted that the most similar species is  C. diplostictus G.Norman ex Greville (Williams 1988, pl. 26, figs 5, 6), but that species is twice the diameter and has many interpolated short “striae” not present in our specimen. Despite some vague topography in the central space, compared to completely hyaline in Greville’s (1863) drawing, we are satisfied that we have  C. robertsianus . The character of these pores (called punctae or granules by Peragallo &amp; Peragallo 1897 –1908) seems not to have been studied in SEM.</p><p>Arrangement of pores distinguish  C. robertsianus from species with scattered pores, such as  Coronia daemeliana (Grunow) Ruck &amp; Guiry, already transferred from  Campylodiscus .  Campylodiscus robertsianus is one of several taxa present in the Guam flora and lacking infundibula that most likely qualify for transfer to  Coronia but were not studied by Ruck et al. (2016a, b, Ruck &amp; Guiry 2016), i.e.,  Campylodiscus decorus var. pinnatus (the species was transferred but not the variety),  C. humilis and  C. brightwellii in our flora. Park et al. (2018) already transferred the first of these to  Coronia decora var. pinnata (Peragallo) Lobban &amp; JoonS.Park, and we are tempted to propose here the transfer of  C. robertsianus . However, Ruck &amp; Guiry (2016) gave no unambiguous morphological diagnosis for  Coronia, relying instead on sequence data. Nomenclaturally, while a  variety must go with its parent species when the species is transferred, transferring  C. robertsianus would require a decision that this species accords with the new genus, which is uncertain in the absence of molecular data. This species was described from Queensland, Australia and has been reported from Samoa. Greville (1863) delighted in it as “one of the most exquisite species of this charming genus.” We have only this single specimen from Guam and a fragment from Yap, both in LM.</p></div>	https://treatment.plazi.org/id/03C287AAFFFA0B01FF357F9DFB31F99E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobban, Christopher S.;Witkowski, Andrzej	Lobban, Christopher S., Witkowski, Andrzej (2023): Marine benthic diatoms of Guam: new records, Dictyoneis apapae sp. nov., and updates to the checklist. Micronesica 2023 (2): 1-75, DOI: 10.5281/zenodo.12118784
03C287AAFFFA0B00FF357BACFC74FEE6.text	03C287AAFFFA0B00FF357BACFC74FEE6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cocconeis carinata Riaux-Gobin, Ector & Witkowski 2019	<div><p>Cocconeis carinata Riaux-Gobin, Ector &amp; Witkowski Figs 39–41</p><p>Ref. illus.: Riaux-Gobin et al. 2019, figs 1–35</p><p>Samples: GU70A-2A, GU68A-2, inter alia</p><p>Dimensions: Length 8 µm, width 5 µm; stria density 26 in 10 µm on both valves.</p><p>Diagnostics: Sternum valve with crista marginalis dividing the striae. Striae have single transapically elongate areola distal to crista and 2–3 areolae proximal.</p><p>Comments: Abundant in GU70A-2A. This species was recently described from Indian Ocean and Pacific Ocean sites, including the Marquesas I., French Polynesia.</p></div>	https://treatment.plazi.org/id/03C287AAFFFA0B00FF357BACFC74FEE6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobban, Christopher S.;Witkowski, Andrzej	Lobban, Christopher S., Witkowski, Andrzej (2023): Marine benthic diatoms of Guam: new records, Dictyoneis apapae sp. nov., and updates to the checklist. Micronesica 2023 (2): 1-75, DOI: 10.5281/zenodo.12118784
03C287AAFFFB0B00FF357C44FB29FCF6.text	03C287AAFFFB0B00FF357C44FB29FCF6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Coscinodiscus hauckii Cleve	<div><p>Coscinodiscus hauckii Cleve Figs 42, 43</p><p>Ref. illus.: Hustedt 1927 –1930, p. 388, fig. 200b</p><p>Samples: GU52O-4, inter alia</p><p>Dimensions: Diam. 27–30 µm, stria density at margin 21-22 in 10 µm.</p><p>Diagnosis: Ring of short striae surrounding flat central zone with or without short spines.</p><p>Comments: The stria density given by Hustedt is ca. 18 in 10 µm. Hein et al. (2008, p. 18, pl. 2, fig. 2) reported  C. hauckii var. mesoleia, which has a clear center, with 28 striae in 10 µm. It is not clear from Hustedt’s text about the nominate  variety whether he thought the unregelmässig punktiertem Mittelfeld was perforated, because he also used punktiert in reference to the short striae. The  variety may not be justified, given that the difference in the central zone of the nominate  variety comprises only small papillae, and there is only a slight difference in stria densities between the forms (20 in the variety). Our stria density is higher, and Hein et al.’s still higher. Our SEM shows that this species lacks rimoportulae, so it does not belong in  Coscinodiscus (see Round et al. 1990).</p></div>	https://treatment.plazi.org/id/03C287AAFFFB0B00FF357C44FB29FCF6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobban, Christopher S.;Witkowski, Andrzej	Lobban, Christopher S., Witkowski, Andrzej (2023): Marine benthic diatoms of Guam: new records, Dictyoneis apapae sp. nov., and updates to the checklist. Micronesica 2023 (2): 1-75, DOI: 10.5281/zenodo.12118784
03C287AAFFFB0B0FFF357E53FEF3FB15.text	03C287AAFFFB0B0FFF357E53FEF3FB15.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dictyoneis apapae Lobban & Witkowski 2023	<div><p>Dictyoneis apapae Lobban,  sp. nov. Figs 44–50, 52–55, 57, 58</p><p>Diagnosis: Differing from  Dictyoneis marginata in the broadly lanceolate outline, absence of field of small pseudoloculi around the central area, and smaller outer openings of the pseudoloculi, especially along the margin.</p><p>Holotype: Specimen at 10.2 mm E and 6.0 mm S of the mark on slide 3056, deposited at Diatom Collection,  
Academy of 
Natural Sciences of Drexel University, Philadelphia, accession number GC20106. Fig. 44.</p><p>Type locality: Guam, Apra Harbor, Scuba Beach, 13°27.840', 144°39.360', scarce in biofilm on calcareous sand ca. 10 m deep, in communities with  Arcuatasigma spp. (Lobban &amp; Reid 2018),  Progonoia spp. (Lobban 2015b), and several species listed below. Sample GU52X-5b, 10 May 2015, coll. C.S. Lobban &amp; M. Schefter</p><p>Etymology: Apapa, genitive noun in apposition, for the type locality in Apra Harbor, not far from Cabras Island. Ayong (2023) explains that, “Apra is a corruption of the Chamorro word ‘ apapa ’ which means ‘low.’ Apapa is the original name for what is now known as Cabras Island.”</p><p>Morphology: Valves broadly lanceolate (Figs 44–46), 94–115 µm long, 22–26 µm wide, outer pseudoloculi in rows giving apparent stria density, especially in LM, of 8–12 in 10 µm, (Figs 44, 45) but true striae measured on interior views 24–26 in 10 µm (Figs 52, 55). External raphe slits straight, bordered by thin ridges of silica (Fig. 48, arrow), terminal endings deflected in opposite directions. Inner layer of pores completely overlain by pseudoloculate framework forming larger pores, except in a variable zone near the sternum and at the apices where the outer openings are also small (Figs 48, 49 and cf. fracture wall, Fig. 54). The outermost areolae are very large and in LM give the impression of chambers (cf.  Mastogloia) (Figs 44, 45) but are simply part of the outer layer (Figs 46, 47,49). There is also a long groove on each side in the outer layer near the apex (Figs 47, 50, 57). Figs 50 and 51 compare exterior features of the new species with a specimen of  D. marginata (F.W.Lewis) Cleve from Georgia. Interior surface shows regular striae opening by transapical slits (Figs 52–55); many short striae and a few longer ones are interpolated on the mantle forming a denser fringe (32 striae in 10 µm), followed by a hyaline border (Fig. 53). Raphe bordered by thick ribs, central and terminal endings simple, the latter with a very small helictoglossa. Wall structure (Fig. 54) shows deep pseudoloculate outer network over the basal striate layer. Figs 55 and 56 compare the similar internal surfaces of the new species with  D. marginata . Girdle bands (Figs 47, 57, 58): copulae open: valvocopula with deep notch at the closed end filled by ligula on 3 rd copula (Fig 57, arrow), with two rows of pores along advalvar edge, scattered pores/pits elsewhere; two other copulae with similar structure.</p><p>Additional records: GUAM: GU52V-1! (biofilm); YAP, FSM: Y-D2 (seagrass leaves)</p><p>Registration: http://phycobank.org/104437</p><p>Comments:  Dictyoneis apapae differs markedly from  D. marginata [Hustedt 1931 –1959, p. 576, fig. 1009; Montgomery 1978 pl. 81C–F; Round et al. 1990 p. 468 (implicitly showing  D. marginata); Hein et al. 2008, p. 51, pl. 26, fig 2, pl. 27 fig. 1] but there seem to be additional discrepancies among the images in the literature. The apical groove in Round et al. (1990) is long like that in  D. apapae, and they also note that the external raphe slits are bordered by thin ridges of silica, which can be seen in our images of  D. apapae but not in our  D. marginata specimens (Figs 50 vs. 51). Moreover, Round et al. (1990, p. 469, fig. h) show thick transverse costae and note that “the small poroids of the inner layer are difficult to distinguish because of the strong development of the ribs,” a description that does not fit  D. apapae (Fig. 54) or even our Georgia specimens (Fig. 56). These seem to be small discrepancies compared to the differences in the central area, marginal pore size and constriction (the latter two used by Cleve 1890 as taxonomic criteria); we are therefore confident that the Guam species cannot be included in  D. marginata, but not so confident in our identification of the  D. marginata specimen.</p><p>When Cleve (1890) erected the genus, he provided a key to the known species, of which only two were lanceolate, and only one of those had marginal pores larger than the ones on the valve face, that was  D. thrumii Cleve, described in the same article. However, he gave no drawings and the description, based on a single specimen from a sea cucumber gut in China, mentions little of the structure. His valve was larger than our specimens, 150 µm long, 32 µm wide. Montgomery (1978, pl. 81A, B) showed an unnamed lanceolate species from Florida with broad semilanceolate zones of smaller pseudoloculi on each side of the raphe, clearly different from ours. There is need for further study of this genus, and the assertion by Round et al. (1990, p. 468) that, “only the type [species] is recorded at all frequently” should not be a license for assigning every constricted specimen to  D. marginata .</p></div>	https://treatment.plazi.org/id/03C287AAFFFB0B0FFF357E53FEF3FB15	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobban, Christopher S.;Witkowski, Andrzej	Lobban, Christopher S., Witkowski, Andrzej (2023): Marine benthic diatoms of Guam: new records, Dictyoneis apapae sp. nov., and updates to the checklist. Micronesica 2023 (2): 1-75, DOI: 10.5281/zenodo.12118784
03C287AAFFF40B0FFF357832FB8CF882.text	03C287AAFFF40B0FFF357832FB8CF882.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hyalosynedra al-turkii J. S. M. Sabir & E. C. Theriot	<div><p>Hyalosynedra cf. al-turkii J.S.M.Sabir &amp; E.C.Theriot</p><p>Ref. illus.: Navarro &amp; Lobban 2009, figs 48–51 (as  Hyalosynedra laevigata); Sabir et al. (2018), figs 27–32</p><p>Samples: GU50D, GU52U-1</p><p>Dimensions: 35–40 µm long, 4–5 µm wide, 40 biseriate striae in 10 µm.</p><p>Diagnostics: Apex rounded; sternum lanceolate, narrow; ocellulimbus 6–8 pores wide.</p><p>Comments: This species is separated from  H. prasadii by the dimensions according to the key in Sabir et al. (2018). Like that species, it has a lanceolate sternum, in contrast to other species in which the sternum is linear. The images published by Navarro &amp; Lobban (2009) can be better identified as Sabir &amp; Theriot’s new species, although the stria density is higher than their specimens.  Hyalosynedra prasadii, or something like it, has also been noted in a few samples, but both species were scarce and detailed comparison with Red Sea and Florida species is not yet possible and, because of the fine discriminations needed, it is possible that these species are regional endemics and that our western Pacific species are different. We make this provisional record now only to point out that the record in Navarro &amp; Lobbban (2009) is not  H. laevigata . This genus needs more detailed study in our region but  H. laevigata is present in our flora, see below.</p></div>	https://treatment.plazi.org/id/03C287AAFFF40B0FFF357832FB8CF882	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobban, Christopher S.;Witkowski, Andrzej	Lobban, Christopher S., Witkowski, Andrzej (2023): Marine benthic diatoms of Guam: new records, Dictyoneis apapae sp. nov., and updates to the checklist. Micronesica 2023 (2): 1-75, DOI: 10.5281/zenodo.12118784
03C287AAFFF50B0EFF357DCEFB64FDAC.text	03C287AAFFF50B0EFF357DCEFB64FDAC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hyalosynedra laevigata (Grunow) Williams & Round 1986	<div><p>Hyalosynedra laevigata (Grunow) Williams &amp; Round Figs 59–62</p><p>Ref. illus.: Sabir et al. 2018, p. 7, figs 2–16. Non Navarro &amp; Lobban 2009, figs 48–51.</p><p>Samples: GU44Z-15, GU58G-4A</p><p>Dimensions: length 114–132 µm, width 5–6 µm.</p><p>Diagnostics: Long, lanceolate, hyaline cells, apices rounded to weakly capitate; linear midrib visible in LM; pseudocellus with three rows of pores; rimoportulae asymmetrical.</p><p>Comments: Differing from  H. lanceolata Belando, Jiménez &amp; Aboal, which has a broad, lanceolate sternum. It is so similar in size and shape to  Stricosus cardinalii E.C.Theriot &amp; Lobban (see below) that the two taxa can scarcely be distinguished in LM. In SEM, the character of the rimoportulae and the depth of the pseudocellus separate them. See also comments above on  H. cf. al-turkii .</p></div>	https://treatment.plazi.org/id/03C287AAFFF50B0EFF357DCEFB64FDAC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobban, Christopher S.;Witkowski, Andrzej	Lobban, Christopher S., Witkowski, Andrzej (2023): Marine benthic diatoms of Guam: new records, Dictyoneis apapae sp. nov., and updates to the checklist. Micronesica 2023 (2): 1-75, DOI: 10.5281/zenodo.12118784
03C287AAFFF50B0EFF357FB9FCF5FB82.text	03C287AAFFF50B0EFF357FB9FCF5FB82.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Licmophora complanata Lobban	<div><p>Licmophora complanata Lobban Figs 63, 64</p><p>Ref. illus.: Lobban 2021b, figs 1–25.</p><p>Samples: GU44AR-3</p><p>Dimensions: Length 135–143 µm, striae in 10 µm 9–10, increasing to 10–12 near apex.</p><p>Diagnostics: The apically oriented areolae, the triangular profile of the valve, always lying in girdle view, and the broad valvocopula with deep septum, together with apically elongated areolae, make this species unique so far even in LM.</p><p>Comments: Two specimens, including a complete frustule were found in LM.  The apically elongated areolae are evident in both images; the three inflated areas are suggested by the focus and the short, intercalated striae (Fig. 63, black arrowheads), and the path of the midrib and septum are shown by arrow and arrowhead in Fig. 63. The septum is very unusual in this species (Lobban 2021, fig. 19). Previously known only from Majuro, Marshall Islands.</p></div>	https://treatment.plazi.org/id/03C287AAFFF50B0EFF357FB9FCF5FB82	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobban, Christopher S.;Witkowski, Andrzej	Lobban, Christopher S., Witkowski, Andrzej (2023): Marine benthic diatoms of Guam: new records, Dictyoneis apapae sp. nov., and updates to the checklist. Micronesica 2023 (2): 1-75, DOI: 10.5281/zenodo.12118784
03C287AAFFF50B0EFF3579A8FC30F944.text	03C287AAFFF50B0EFF3579A8FC30F944.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mastogloia ovata Grunow 1860	<div><p>Mastogloia ovata Grunow Figs 65, 66</p><p>Ref. illus.: Hustedt 1931 –1959, p. 476, fig. 895</p><p>Lobban et al. 2012, p. 281, pl. 35, figs 1, 2 (as “  M. pseudolatecostata ”)</p><p>Lobban et al. 2012, p. 279, pl. 33, figs 7, 8 (as “  M. ovata ”)</p><p>Samples: GU44T-1, GU44 AJ, GU44AU-1; GU52P-7; GU70A-2A</p><p>Our specimen published as voucher for  M. pseudolatecostata must be rejected because the stria density is too high (21–24 vs 12–14 in 10 µm). It is a better match for  M. ovata than the voucher specimen cited in the same paper. SEM images of such specimens are shown in Figs 65, 66. A specimen from Bikar Atoll, Marshall Islands that matches  M. pseudolatecostata is shown in Fig 67; this species not yet identified in Guam. The specimen claimed as a voucher for  M. ovata in Lobban et al. 2012 less clearly matches the description in Hustedt (1931 –1959) and close to Hein et al. (2008, pl. 40, fig. 3) “  Mastogloia cf. ovata . ”</p><p>Comments: The inner margin of  M. ovata has indentations at each cross wall (Fig. 66), that of  M. pseudolatecostata in the middle of each partectum (Fig. 67).</p></div>	https://treatment.plazi.org/id/03C287AAFFF50B0EFF3579A8FC30F944	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobban, Christopher S.;Witkowski, Andrzej	Lobban, Christopher S., Witkowski, Andrzej (2023): Marine benthic diatoms of Guam: new records, Dictyoneis apapae sp. nov., and updates to the checklist. Micronesica 2023 (2): 1-75, DOI: 10.5281/zenodo.12118784
03C287AAFFF50B0DFF357BE2FDDEFB53.text	03C287AAFFF50B0DFF357BE2FDDEFB53.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mastogloia pisciculus Cleve 1893	<div><p>Mastogloia cf. pisciculus Cleve Figs 68–70</p><p>Ref. illus.: Cleve 1893; Hustedt 1931 –1959, p. 473, fig. 990; Stephens &amp; Gibson 1980, p. 228, figs 31–36; Pennesi et al. 2012. p. 1254, figs 7A–H; Loir &amp; Novarino 2013, p. 42, pl. 18c</p><p>Samples: GU52X-5</p><p>Dimensions: Length 30–35 µm, width 14 µm, striae radiate, 25 in 10 µm</p><p>Diagnostics: Broadly lanceolate, rostrate, depressed area not strongly differentiated but a longitudinal line visible in the middle of it (Figs 68, 69 arrows). Chambers quadrate to elongate with convex inner edges, not reaching apices. In SEM external view (Fig. 69) uniform areolae in the outer zone, transapically elongated areolae in the depressed zone, irregular rib between them; raphe thickened. Internally, outer zone with transapical costae, inner zone smooth. Areolae opening internally, by quadrate set of small pores; outer zone of striae with costate virgae, inner zone with areolae in irregular longitudinal line, sporadically two lines (Fig. 70 arrow). Partecta 3 in 10 µm, in middle portion only, apically elongated with convex inner margins and scattered pores on the abvalvar surface.</p><p>Comments: Uncommon in this biofilm sample. Specimens were smaller than Cleve’s and at low end of range in Hustedt. Hustedt (1931 –1959), Stephens &amp; Gibson (1980) and Loir &amp; Novarino (2013) all show much variation. Moreover, there are considerable discrepancies between the various descriptions, leading us to qualify our identification with “cf.” First, Hustedt (1931 –1959: 558) said that although the forms he found (only near Seychelles) differed from Cleve’s (1893) description (specimens from Florida), e.g. in stria density, he decided not to separate them. Stephens &amp; Gibson reported stria densities in line with type material in Cleve (1893) and Boyer (1927). Specimens in Pennesi et al. were from Suluwesi, Indonesia. The SEM studies by Stephens &amp; Gibson (1980) and Pennesi et al. (2012) are not completely comparable because partecta found by the latter were all broken. The partecta are square to apically lengthened with convex inner margins. Stephens &amp; Gibson showed them to have delicate silica plaques with perforations; partecta in our specimens seem longer than in these other reports and we saw pores but no plaques (Fig. 70). Stephens &amp; Gibson did not observe the edge of the mantle, where we and Pennesi et al. saw biseriate striae. Internally, there is a single line of quadripunctate foramina under the depressed area shown in both Stephens &amp; Gibson and Pennesi et al. but in our specimens, there was a much looser row, sometimes double. Stephens &amp; Gibson (1980; Florida) considered whether their specimens might match  M. arabica Hendey, or if that might be synonymous with  M. pisciculus . Establishing boundaries in these disparate populations of rare taxa will require more work; for now, we present our SEM observations to define what we found here.</p></div>	https://treatment.plazi.org/id/03C287AAFFF50B0DFF357BE2FDDEFB53	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobban, Christopher S.;Witkowski, Andrzej	Lobban, Christopher S., Witkowski, Andrzej (2023): Marine benthic diatoms of Guam: new records, Dictyoneis apapae sp. nov., and updates to the checklist. Micronesica 2023 (2): 1-75, DOI: 10.5281/zenodo.12118784
03C287AAFFF60B0DFF3579F8FB7FF8D8.text	03C287AAFFF60B0DFF3579F8FB7FF8D8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Moreneis hexagona J. Park, Koh & Witkowski 2012	<div><p>Moreneis cf. hexagona J. Park, Koh &amp; Witkowski Figs 71–74</p><p>Ref. illus.: J. Park et al. 2012, p. 192, fig. 6; J. S. Park et al. 2018, p. 108, fig. 26.</p><p>Samples: GU 52O-4, GU 44AS-4</p><p>Dimensions: Length 31–32 µm, width 13 µm; striae 15 in 10 µm, areolae 16 in 10 µm.</p><p>Diagnostics: Central and terminal raphe endings deflected in same direction, central endings in opposite directions (Fig. 73), distinguish the genus from  Petroneis and  Navicula . Species based on shape, dimensions, and stria density. Valve rostrate; striae radiate, some shorter ones in center.</p><p>Comments: Rare specimens match the description of  M. hexagona, which is smaller and more finely striated than  M. angulata (42–48 µm x 16–16.5 µm, striae 11 in 10 µm, areolae 12 in 10 µm). J. Park et al. (2012) noted a single areola at the Voigt faults (Fig. 72, arrows) and larger areolae adjacent to axial area, which are present in our specimen. Our SEM images are the first for this species and show the areolae to be c-shaped (Fig. 73).  Moreneis hexagona is distinguished from several other species by shape and stria density. J. S. Park et al. (2018) reported  M. cf. hexagona from Chuuk with some hesitation because of a size difference. Present in biofilm with another  Moreneis sp. in GU 52O-4, but also found in a gathering of  Halimeda in GU 44AS-4. The other species does not match any of the species in this newly erected genus but it may have been described under  Navicula .</p></div>	https://treatment.plazi.org/id/03C287AAFFF60B0DFF3579F8FB7FF8D8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobban, Christopher S.;Witkowski, Andrzej	Lobban, Christopher S., Witkowski, Andrzej (2023): Marine benthic diatoms of Guam: new records, Dictyoneis apapae sp. nov., and updates to the checklist. Micronesica 2023 (2): 1-75, DOI: 10.5281/zenodo.12118784
03C287AAFFF70B0CFF357DCEFD49FBD2.text	03C287AAFFF70B0CFF357DCEFD49FBD2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nitzschia incurva Grunow 1878	<div><p>Nitzschia incurva Grunow Figs 75–78</p><p>Syn.:  Nitzschia lorenziana Grunow in Cleve &amp; Grunow (nom. illeg.)</p><p>Ref. illus.: Navarro 1982, pl. 35, fig. 4; Poulin et al. 1990, p. 87, figs 67, 68, 71, 72 (SEM); Witkowski et al. 2000, p. 392, pl. 210, figs 24, 25, pl. 211, fig. 3, pl. 212, figs 1–3 (all as  N. lorenziana)</p><p>Samples: GU58G-4A, GU44I-1, GU44Z-15</p><p>Dimensions: Length 18.5 µm, width 7.4 µm; striae 20 in 10 µm.</p><p>Diagnostics: Weakly sigmoid in girdle view, keel eccentric except in long-rostrate apices, striation bold in LM because of external costae between the striae; central striae biseriate towards to keel, triseriate towards the margin (Fig. 78), becoming rings of pores with central row of pores and diminishing towards apex (Fig. 77). Eccentric keels of the two valves are on opposite sides of the frustule (nitzschioid, not hantzschioid) (Figs 77, 78). Keel canal is closed except for circular fenestrae (Fig. 77).</p><p>Comments: Guiry (2017) explains why  lorenziana is illegitimate and provides the earlier synonym. The ultrastructure of our specimens clearly matches SEM images in Poulin et al. (1990). It is apparently a brackish/estuarine species, with wide temperature tolerance (from Gulf of St. Lawrence to Florida + Guam). Abundant in Vietnam and Indonesia (Witkowski, unpubl.). Lopez-Fuerte et al. (2010: p. 63, pl. 37, fig. 21) show  var. subtilis Grunow from Mexico and list the nominate variety.  Nitzschia incurva somewhat resembles  N. gaoi Bing Liu, S. Blanco &amp; B.Q. Huang (2019), but that has small conopeal canals and uniseriate striae. Lobban &amp; Ashworth (2022b) did not transfer  N. gaoi or their own  N. maiae Lobban, Ashworth, Calaor &amp; E.C.Theriot to  Homoeocladia because of maintaining a narrow definition of the morphology of  Homoeocladia and noted some other species in the region with doubtful formation of a conopeal canal that also have T-shaped external costae (one shown in Lobban 2023a, figs 4F, G).</p></div>	https://treatment.plazi.org/id/03C287AAFFF70B0CFF357DCEFD49FBD2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobban, Christopher S.;Witkowski, Andrzej	Lobban, Christopher S., Witkowski, Andrzej (2023): Marine benthic diatoms of Guam: new records, Dictyoneis apapae sp. nov., and updates to the checklist. Micronesica 2023 (2): 1-75, DOI: 10.5281/zenodo.12118784
03C287AAFFF70B0CFF357978FBDFFA4A.text	03C287AAFFF70B0CFF357978FBDFFA4A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Olifantiella mascarenica Riaux-Gobin & Compere	<div><p>Olifantiella mascarenica Riaux-Gobin &amp; Compère Figs 79, 80</p><p>Ref. illus.: Riaux-Gobin &amp; Compère 2009, figs 6–22; Riaux-Gobin 2015, figs 24, 25</p><p>Sample: GU52G-A</p><p>Dimensions: Valve 9.3 µm long x 2.3 µm wide; striae 50 in 10 µm.</p><p>Diagnostics: This species differs from  O. pilosella Riaux-Gobin in the shape of the buciniportula, which here is a single trumpet.</p><p>Comments:  Olifantiella spp. are barely discernable in LM, and the details are at the limit even of the desktop SEM. These images were taken at U. Texas on a full-sized SEM.</p></div>	https://treatment.plazi.org/id/03C287AAFFF70B0CFF357978FBDFFA4A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobban, Christopher S.;Witkowski, Andrzej	Lobban, Christopher S., Witkowski, Andrzej (2023): Marine benthic diatoms of Guam: new records, Dictyoneis apapae sp. nov., and updates to the checklist. Micronesica 2023 (2): 1-75, DOI: 10.5281/zenodo.12118784
03C287AAFFF70B0CFF3578E0FB77F9BD.text	03C287AAFFF70B0CFF3578E0FB77F9BD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Parlibellus biblos (Cleve) Cox	<div><p>Parlibellus biblos (Cleve) Cox Figs 81–84</p><p>Ref. illus: Hustedt 1931 –1959, p. 703, fig. 1178 (as  Stauroneis biblos [Cleve] Hustedt); Cox 1988, p. 23; Lobban et al. 2012, p. 296, pl. 52, figs 4, 5) [as  Stauroneis retrostauron (Mann) Meister]</p><p>Comments: J.S. Park et al. (2022, p. 14) made the case for recognizing this taxon as  Parlibellus biblos .  Stauroneis retrostauron must be deleted from the species list for Guam for now.</p></div>	https://treatment.plazi.org/id/03C287AAFFF70B0CFF3578E0FB77F9BD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobban, Christopher S.;Witkowski, Andrzej	Lobban, Christopher S., Witkowski, Andrzej (2023): Marine benthic diatoms of Guam: new records, Dictyoneis apapae sp. nov., and updates to the checklist. Micronesica 2023 (2): 1-75, DOI: 10.5281/zenodo.12118784
03C287AAFFF70B0BFF357B8AFAADFE30.text	03C287AAFFF70B0BFF357B8AFAADFE30.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Plagiogramma minor (W. Gregory) Chunlian Li, Ashworth & Witkowski	<div><p>Plagiogramma minor (W.Gregory) Chunlian Li, Ashworth &amp; Witkowski Figs 85, 86</p><p>Basionym:  Denticula minor W.Gregory</p><p>Ref. illus.: Kaczmarska et al. 2017, p. 27, figs 13A–D.</p><p>Samples: GU52O-4, GU52X-5 (biofilm on calcareous sand)</p><p>Dimensions: Length 11–12 µm, width 5 µm; striae 17 in 10 µm</p><p>Diagnostics: rhomboidal valves with central elevation and two elevated apical pore fields; large flaps on the four most-apical spines.</p><p>Comments: Kaczmarska et al. (2017) compared their specimens to similar species and concluded that there was no match. Our specimens are very close to their fig. 13A. In their nomenclatural note, Kaczmarska et al. claimed that the correct epithet is minus, since  Dimeregramma is a neuter noun, and Li et al. (2020, p. 13) used this transferring the species to  Plagiogramma (also neuter). However, minus is an adverb and the correct adjective here is  minor, which has the same ending across all genders in the nominative singular (Stearn 1973), as used by Gregory (1857) in the basionym.</p></div>	https://treatment.plazi.org/id/03C287AAFFF70B0BFF357B8AFAADFE30	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobban, Christopher S.;Witkowski, Andrzej	Lobban, Christopher S., Witkowski, Andrzej (2023): Marine benthic diatoms of Guam: new records, Dictyoneis apapae sp. nov., and updates to the checklist. Micronesica 2023 (2): 1-75, DOI: 10.5281/zenodo.12118784
03C287AAFFF00B0BFF357F16FA83FC1F.text	03C287AAFFF00B0BFF357F16FA83FC1F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pleurosigma simulacrum Lobban & Sterrenburg	<div><p>Pleurosigma simulacrum Lobban &amp; Sterrenburg</p><p>Ref.: Lobban 2021a, p. 253, figs 91–102.</p><p>Here we just want to note that this species, described from Guam and Yap and differentiated from  P. intermedium W.Smith, is also similar in shape to  P. acus A. Mann (1925, p. 133, pl. 29, figs 4, 5), described from the Philippines as being very similar to  P. intermedium . Stidolph (2002) pointed out that  P. acus has the diagonal striae at ca 80º to one another, contrasting with  P. intermedium and  P. simulacrum .  Pleurosigma simulacrum and  P. acus both lack the calcar slit at the apex, which is present in  P. intermedium . Mann also noted that the striae of  P. acus are much coarser than other straight species (14, vs 26 in 10 µm for  P. simulacrum). Thus, there are at least four straight, narrow species of  Pleurosigma, the fourth being  P. patagonicum Ferrario &amp; Sar (Sar et al. 2012) (Table 2). We have checked all images we attributed to  P. simulacrum, now including specimens from Saipan, Chuuk and Palau (Lobban, unpublished), and confirmed that they were correctly identified.</p></div>	https://treatment.plazi.org/id/03C287AAFFF00B0BFF357F16FA83FC1F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobban, Christopher S.;Witkowski, Andrzej	Lobban, Christopher S., Witkowski, Andrzej (2023): Marine benthic diatoms of Guam: new records, Dictyoneis apapae sp. nov., and updates to the checklist. Micronesica 2023 (2): 1-75, DOI: 10.5281/zenodo.12118784
03C287AAFFF00B0BFF35792CFE7BF925.text	03C287AAFFF00B0BFF35792CFE7BF925.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Proschkinia complanata (Grunow) D. G. Mann	<div><p>Proschkinia complanata (Grunow) D.G.Mann Figs 87–90</p><p>Ref. illus.: Lobban et al. 2012, p. 296, pl. 1, figs 4–6 and pl. 53, figs 1–4 [as  P. complanatoides (Hustedt) Karayeva]; Kim et al. 2020, p. 960, figs 2, S2</p><p>Samples: GU44Z-15, GU52Q-10a, GU62A-7, GU66G-5, GU68F-8</p><p>Dimensions: Length 32–48 μm, width 6–8 μm; striae 30–32 in 10 μm</p><p>Comments: Majewska et al. (2019, p. 625), working on  Proschkinia spp. epizoic on sea turtles, described a new species  P. torquata Bosak, Van de Vijver &amp; Majewska, which they compared with  P. complanatoides . They suggested that our Guam specimens might be a new species because they saw no widening of the central area (GU44Z-15: Fig. 90) and the internal fistula was not like  P. complanatoides . However, Kim et al. (2020) studied Grunow’s specimens of  P. complanata with SEM and showed a fistula matching ours, also matching in the presence of a flap over the external fistula opening (Figs 87, 88) and a raised rib on the raphe internally for some distance from the central nodule (Fig. 90). One short stria on the side with the fistula (Fig. 88), and a silica flap over the fistula opening; central striae on opposite sometimes further apart (Figs 87, 88 vs. 90). Comparing our specimens to  P. torquata, the fistula has a similar series of spheres, and the external opening is also covered by a flap, but  P. torquata has longitudinal silica strips over the central part of the valve face and the apices are rounded rather than acute. Kim et al. (2020, table 4) listed Guam specimens with  P. complanata not  P. complanatoides and their images of authentic (including type) material convince us that there are no differences that would warrant a new species. We have seen this species across Micronesia, and it seems so far to be the only species present. We remove  P. complanatoides from the Guam flora.</p></div>	https://treatment.plazi.org/id/03C287AAFFF00B0BFF35792CFE7BF925	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobban, Christopher S.;Witkowski, Andrzej	Lobban, Christopher S., Witkowski, Andrzej (2023): Marine benthic diatoms of Guam: new records, Dictyoneis apapae sp. nov., and updates to the checklist. Micronesica 2023 (2): 1-75, DOI: 10.5281/zenodo.12118784
03C287AAFFF20B09FF357DCEFDE3FD4E.text	03C287AAFFF20B09FF357DCEFDE3FD4E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudictyota bicornis (Cleve) P. A. Sims & D. M. Williams	<div><p>Pseudictyota bicornis (Cleve) P.A.Sims &amp; D.M.Williams Figs 91–93</p><p>Ref. illus.: Cleve 1878, fig. 30; Ashworth et al. 2013, fig 7b; Sims et al. 2018, figs 133–138</p><p>Sample: GU52U-1</p><p>Dimensions: 51 µm long (on the axis with the ocelli), 42 µm wide</p><p>Diagnostics: Quadrate, false pseudoloculate valve, having two elevations with ocelli and two unelevated points with rimoportulae.</p><p>Comments: Differing from  Pseudictyota dubia (Brightwell) P.A.Sims &amp; D.M.Williams (Navarro &amp; Lobban 2009, figs 21–24), which has six angles, three with ocelli and three with rimoportulae; and from  P. reticulata (Roper) P.A.Sims &amp; D.M.Williams (Lobban et al. 2012, pl. 3, figs 3, 4, as  Triceratium dictyotum P.A.Sims &amp; R.Ross), which is broadly lanceolate with two angles each with both an ocellus and a rimoportula.</p></div>	https://treatment.plazi.org/id/03C287AAFFF20B09FF357DCEFDE3FD4E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobban, Christopher S.;Witkowski, Andrzej	Lobban, Christopher S., Witkowski, Andrzej (2023): Marine benthic diatoms of Guam: new records, Dictyoneis apapae sp. nov., and updates to the checklist. Micronesica 2023 (2): 1-75, DOI: 10.5281/zenodo.12118784
03C287AAFFF20B09FF357F9BFE17FA90.text	03C287AAFFF20B09FF357F9BFE17FA90.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Serratifera andersonii Chunlian Li, Dabek & Wachnika	<div><p>Serratifera andersonii Chunlian Li, Dąbek &amp; Wachnika Figs 94, 95</p><p>Ref. illus.: C. Li et al. 2018, p. 63, figs 360–373</p><p>Sample: GU26A</p><p>Dimensions: Length 5.5–9.2 µm, width 2.3 µm, striae 15 in 10 µm</p><p>Diagnostics: Small, clavate (heteropolar), with a clear pore field at one pole, simple, stick-like spines arising from areolae, whereas in  S. rhombica there are “accessory” spines flanking the larger central spines (C. Li et al. 2018).</p><p>Comments: Many small  Fragilariaceae have been studied recently, incorporating much debate on generic boundaries.  Serratifera Ashworth, Chunlian Li &amp; Witkowski (in C. Li et al. 2016) was established largely on molecular grounds and emended on morphological grounds by the same authors (in C. Li et al. 2018). Morales et al. (2019: 274) argued against the new genus and for inclusion of its species in the older genus  Pseudostaurosira D.M.Williams &amp; Round. Witkowski did not accept the argument (personal communication during ms. preparation), so we have presented this and the next species under their original names. Morales et al. transferred them to  Pseudostaurosira andersonii (S.Sato, Chunlian Li &amp; Witkowski) E.A.Morales, Novais &amp; C.E.Wetzel, and  P. rhombica (S.Sato, Chunlian Li &amp; Witkowski) E.A.Morales, Novais &amp; C.E.Wetzel, respectively. The senior author cannot contribute to this debate. All small Fragillariaceae are rare in our Guam samples to date, probably because the frustules are washing in from their primary habitats (mostly sediments and biofilms).</p></div>	https://treatment.plazi.org/id/03C287AAFFF20B09FF357F9BFE17FA90	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobban, Christopher S.;Witkowski, Andrzej	Lobban, Christopher S., Witkowski, Andrzej (2023): Marine benthic diatoms of Guam: new records, Dictyoneis apapae sp. nov., and updates to the checklist. Micronesica 2023 (2): 1-75, DOI: 10.5281/zenodo.12118784
03C287AAFFF20B09FF3578B6FDF2F9B4.text	03C287AAFFF20B09FF3578B6FDF2F9B4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Serratifera rhombica S. Sato, Chunlian Li & Witkowski 2018	<div><p>Serratifera rhombica S.Sato, Chunlian Li &amp; Witkowski Figs 96, 97</p><p>Ref. illus.: C. Li et al. 2018, p. 81, figs 431–436</p><p>Sample: GU52P-9</p><p>Dimensions: Length 8 µm, width 2.7 µm, striae 13 in 10 µm</p><p>Diagnostics: Small, rhomboidal (slightly heteropolar). Spines have accessory spines alongside, but these are not clear in our images.</p></div>	https://treatment.plazi.org/id/03C287AAFFF20B09FF3578B6FDF2F9B4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobban, Christopher S.;Witkowski, Andrzej	Lobban, Christopher S., Witkowski, Andrzej (2023): Marine benthic diatoms of Guam: new records, Dictyoneis apapae sp. nov., and updates to the checklist. Micronesica 2023 (2): 1-75, DOI: 10.5281/zenodo.12118784
03C287AAFFF20B08FF357B91FB95FCE0.text	03C287AAFFF20B08FF357B91FB95FCE0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Stricosus harrisonii E. C. Theriot & Lobban	<div><p>Stricosus harrisonii E.C.Theriot &amp; Lobban Figs 98–103</p><p>Ref. illus.: Sabir et al. 2018, figs 95–98</p><p>Samples: GU44U-2, GU44Z-15, GU44BV-3, GU52P-9, GU52Q-1a.</p><p>Description of wild material: Plastids relatively large, numerous (Figs 98, 99), valves 113–193 µm long, 7 µm wide, 40 striae in 10 µm (Figs 100, 101), ocellulimbus 8–9 pores tall (Fig. 102), usually two spines (Fig. 102), rimoportula symmetrical, as for the genus (Fig. 103).</p><p>Comments: Although the record of this species was established in 2018, we wish to add these further observations.  Stricosus harrisonii is the only linear species of  Stricosus found so far in Guam. It was described as 88–152 µm long, relatively broad (5–9 µm), with a narrow, linear sternum, 38–41 biseriate striae in 10 µm, and with a single spine over the ocellulimbus (Sabir et al. 2018). In a variety of wild samples, however, there were two spines, as shown here. Also described from the same material was  S. cardinalii, (Sabir et al. 2018, figs 99–103), which usually has two spines over the ocellulimbus. While  S. harrisonii is essentially linear, the shape and size of  S. cardinalii is very much like  Hyalosynedra laevigata (see above). In SEM,  S. cardinalii is distinguished by the symmetrical rimoportulae and deep ocellulimbi characteristic of  Stricosus .  Stricosus harrisonii and  S. cardinallii from Guam are distinct from similar large species in Florida/Caribbean ( S. navarroensis E.C.Theriot &amp; Ashworth,  S. blumbergii E.C.Theriot &amp; Ashworth) and the Red Sea ( S. alfageehii Sabir &amp; E.C.Theriot,  S. madanii Sabir &amp; E.C.Theriot) (Sabir et al. 2018), possibly suggesting regional endemicity. Belando et al. (2018) identified the sequence labeled GU44AI as being  H. laevigata but it is actually  Stricosus harrisonii, which was published concurrently.</p></div>	https://treatment.plazi.org/id/03C287AAFFF20B08FF357B91FB95FCE0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobban, Christopher S.;Witkowski, Andrzej	Lobban, Christopher S., Witkowski, Andrzej (2023): Marine benthic diatoms of Guam: new records, Dictyoneis apapae sp. nov., and updates to the checklist. Micronesica 2023 (2): 1-75, DOI: 10.5281/zenodo.12118784
03C287AAFFF30B08FF357E45FD32FAF0.text	03C287AAFFF30B08FF357E45FD32FAF0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Thalassiosira nanolineata (A. Mann) Fryxell & Hasle 1977	<div><p>Thalassiosira nanolineata (A. Mann) Fryxell &amp; Hasle Figs 104–106</p><p>Ref. illus.: Mann 1925, p. 68, pl. 14, fig. 4; Fryxell &amp; Hasle 1977 p. 32, figs. 74–80; Sar et al. 2002 Table 1; Y. Li et al. 2013, p. 99, figs 89–94; J.S. Park et al. 2016, p. 414, fig. 28</p><p>Samples: GU44Z-15, GU52P-9</p><p>Dimensions: 12.8–19.8 diam., areolae 8–9 in 10 µm</p><p>Diagnostics: Areolae linear, not radial, single row of fultoportulae around the margin, their external part ending in a cuplike structure with a central post. Rimoportula inserted between two fultoportulae (rather than replacing one). Two central fultoportulae.</p><p>Comments: Although the areolar density is lower than most records (Y. Li et al. 2013 the exception), these specimens seem to agree with the modern concept of the species originally described from the Philippines by Mann (1925). The surveys of  Thalassiosira diversity in Argentina (Sar et al. 2002), Guangdong Province, China (Y. Li et al. 2013), and South Korea (J.S. Park et al. 2016) are all very helpful in identifying the species here.</p></div>	https://treatment.plazi.org/id/03C287AAFFF30B08FF357E45FD32FAF0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobban, Christopher S.;Witkowski, Andrzej	Lobban, Christopher S., Witkowski, Andrzej (2023): Marine benthic diatoms of Guam: new records, Dictyoneis apapae sp. nov., and updates to the checklist. Micronesica 2023 (2): 1-75, DOI: 10.5281/zenodo.12118784
03C287AAFFF30B08FF357856FEFFF8A2.text	03C287AAFFF30B08FF357856FEFFF8A2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tryblionella jelineckii (Grunow) D. G. Mann	<div><p>Tryblionella cf. jelineckii (Grunow) D.G.Mann Figs 107–110</p><p>Ref. illus.: Peragallo &amp; Peragallo 1897 –1908 p. 268, pl. 69, fig 19, Hein et al. 2008, p. 84, pl. 58, fig. 19 (both as  Nitzschia jelineckii Grunow); J.S. Park et al. 2022, p. 20, fig. 58.</p><p>Samples: GU52X-5</p><p>Dimensions: Length 121–158 µm, width 22 µm; striae 12–14 in 10 µm, fibulae 7 in 10 µm.</p><p>Diagnostics: Large panduriform species with prominent external costae running the width of the cell.</p><p>Comments: Similar to  T. plana (W.Smith) Pelletan [Peragallo &amp; Peragallo 1897 –1908 p. 270, pl. 70, fig. 18; Sims 1996, pl. 194, fig 2; Lopez-Fuerte et al. 2010, pl. 38, fig. 1 (all as  Nitzschia plana W.Smith)], but that has 19–20 striae in 10 µm. Not clear from literature that either species has external costae. Montgomery (1978) has two images (pl. 160C and 161D) labeled as “  Nitzschia jelinecki ” but neither has external costae. Peragallo &amp; Peragallo (1897 –1908) commented that they had only seen it in “exotic” samples, the one figured from Java; type was from the Nicobar Islands.  This species needs much more study before we can say whether the Guam specimens belong in it or elsewhere.</p></div>	https://treatment.plazi.org/id/03C287AAFFF30B08FF357856FEFFF8A2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobban, Christopher S.;Witkowski, Andrzej	Lobban, Christopher S., Witkowski, Andrzej (2023): Marine benthic diatoms of Guam: new records, Dictyoneis apapae sp. nov., and updates to the checklist. Micronesica 2023 (2): 1-75, DOI: 10.5281/zenodo.12118784
03C287AAFFEC0B17FF357C38FAB0FC54.text	03C287AAFFEC0B17FF357C38FAB0FC54.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Vikingea gibbocalyx (Brun) Witkowski, Lange-Bertalot & Metzltin	<div><p>Vikingea gibbocalyx (Brun) Witkowski, Lange-Bertalot &amp; Metzltin Figs 111–113</p><p>Ref. illus.: Witkowki et al. 2000, p. 125, pl. 55, figs 8–16, pl. 56, figs 1–3, 5, pl. 57, figs 1–4; Hein et al. 2008, p. 40, pl. 16, fig. 25; Van de Vijver et al. 2009, figs 1–34, Riaux-Gobin et al. 2011, p. 40, pl. 81, figs 1–3, pl.82, figs 1–6</p><p>Dimensions: Length 25–28 µm, width 17–20 µm; striae 9 in 10 µm (RV), 7–8 in 10 µm (SV)</p><p>Samples: GU52X-5, GU52N-7, GU52P-3</p><p>Diagnostics: The raised apices of the RV are diagnostic of the genus in LM, the large circular central area diagnostic of the species in contrast to  V. florifera Riaux-Gobin, O.E.Romero, Compère &amp; A.Y.Al-Handel (2011), which has a broad fascia.</p><p>Comments: These specimens are in the lower end of the size range compared to most literature (see van de Vijver et al. 2009, table 1). There are two troubling differences between our specimens and those shown in the recent literature. First, comparison of the internal SV in our Fig. 112 with figs 16, 19 in van de Vijver et al. (2009) and similar images in Witkowski et al. (2000) and Riaux-Gobin et al. (2011) shows absence in our specimen of the elliptical opening and slit in the marginal ridges; this may be significant even though it was also absent in some specimens by van de Vijver et al. (2009, fig.20). Secondly, Van de Vijver et al. (2009: 271, fig. 27, arrow) show small costae extending from the RV margin into the first row of areolae, a feature clearly absent from the external RV shown in Fig. 113. This is yet another genus with strong potential for additional species to be found.</p></div>	https://treatment.plazi.org/id/03C287AAFFEC0B17FF357C38FAB0FC54	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobban, Christopher S.;Witkowski, Andrzej	Lobban, Christopher S., Witkowski, Andrzej (2023): Marine benthic diatoms of Guam: new records, Dictyoneis apapae sp. nov., and updates to the checklist. Micronesica 2023 (2): 1-75, DOI: 10.5281/zenodo.12118784
03C287AAFFE60B1DFF357C4DFEA8FDF4.text	03C287AAFFE60B1DFF357C4DFEA8FDF4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Actinocyclus tenuissimus Cleve 1878	<div><p>Actinocyclus tenuissimus Cleve</p><p>Reported by Lobban et al. (2012), still a valid species but in view of discussion in J.S. Park et al. (2022), we reconsidered the specimens. While the size and stria density are in the range of  A. tenuissimus, the central area and instinct interfascicular rows are characteristic of  A. subtilis (Gregory) Ralfs, to which we now refer all our Guam specimens (see also Lobban &amp; Tharngan in prep.).</p></div>	https://treatment.plazi.org/id/03C287AAFFE60B1DFF357C4DFEA8FDF4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobban, Christopher S.;Witkowski, Andrzej	Lobban, Christopher S., Witkowski, Andrzej (2023): Marine benthic diatoms of Guam: new records, Dictyoneis apapae sp. nov., and updates to the checklist. Micronesica 2023 (2): 1-75, DOI: 10.5281/zenodo.12118784
03C287AAFFE60B1DFF357F51FAE1FD1A.text	03C287AAFFE60B1DFF357F51FAE1FD1A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ardissonea fulgens (Greville) Grunow ex De Toni 1892	<div><p>Ardissonea fulgens (Greville) Grunow ex De Toni</p><p>Reported by Lobban et al. (2012), still a valid species but Lobban et al. (2022) showed that it differs from the species in our flora and renamed our species  Ardissoneopsis fulgicans Lobban &amp; Ashworth.</p></div>	https://treatment.plazi.org/id/03C287AAFFE60B1DFF357F51FAE1FD1A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobban, Christopher S.;Witkowski, Andrzej	Lobban, Christopher S., Witkowski, Andrzej (2023): Marine benthic diatoms of Guam: new records, Dictyoneis apapae sp. nov., and updates to the checklist. Micronesica 2023 (2): 1-75, DOI: 10.5281/zenodo.12118784
03C287AAFFE60B1DFF357E30FE7AFC5B.text	03C287AAFFE60B1DFF357E30FE7AFC5B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ardissonea fulgens (Greville) Grunow ex De Toni 1892	<div><p>Ardissonea fulgens var. (Lobarzewsky) Rabenhorst</p><p>Reported by Lobban et al. (2012), Lobarzewsky’s diatom proved to be a valid but dubious species and not what we have here, so Lobban et al. (2022) renamed our species  Ardissoneopsis appressata Lobban &amp; Ashworth.</p></div>	https://treatment.plazi.org/id/03C287AAFFE60B1DFF357E30FE7AFC5B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobban, Christopher S.;Witkowski, Andrzej	Lobban, Christopher S., Witkowski, Andrzej (2023): Marine benthic diatoms of Guam: new records, Dictyoneis apapae sp. nov., and updates to the checklist. Micronesica 2023 (2): 1-75, DOI: 10.5281/zenodo.12118784
03C287AAFFE60B1DFF357EF0FB0BFBD8.text	03C287AAFFE60B1DFF357EF0FB0BFBD8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Auricula complexa (Gregory) Cleve	<div><p>Auricula complexa (Gregory) Cleve</p><p>Still a valid species but not what we have here; see  Auricula densistriata above.</p></div>	https://treatment.plazi.org/id/03C287AAFFE60B1DFF357EF0FB0BFBD8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobban, Christopher S.;Witkowski, Andrzej	Lobban, Christopher S., Witkowski, Andrzej (2023): Marine benthic diatoms of Guam: new records, Dictyoneis apapae sp. nov., and updates to the checklist. Micronesica 2023 (2): 1-75, DOI: 10.5281/zenodo.12118784
03C287AAFFE60B1DFF35796EFCE7FB7F.text	03C287AAFFE60B1DFF35796EFCE7FB7F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Climacosphenia moniligera Ehrenberg.	<div><p>Climacosphenia moniligera Ehrenberg.</p><p>Reported by Navarro &amp; Lobban (2009). Still a valid species but Lobban et al. (2022) show that the curved  Climacosphenia in Guam is  C. scimiter A.Mann.</p></div>	https://treatment.plazi.org/id/03C287AAFFE60B1DFF35796EFCE7FB7F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobban, Christopher S.;Witkowski, Andrzej	Lobban, Christopher S., Witkowski, Andrzej (2023): Marine benthic diatoms of Guam: new records, Dictyoneis apapae sp. nov., and updates to the checklist. Micronesica 2023 (2): 1-75, DOI: 10.5281/zenodo.12118784
03C287AAFFE60B1DFF3579CCFC9BFA9E.text	03C287AAFFE60B1DFF3579CCFC9BFA9E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Haslea howeana (Hagelstein) Giffin	<div><p>Haslea howeana (Hagelstein) Giffin</p><p>Possibly still a valid species but Lobban et al. (2020) recognized our material as  Navicula tsukamotoi (Sterrenburg et al.) Yuhang Li et Kuidong Xu</p></div>	https://treatment.plazi.org/id/03C287AAFFE60B1DFF3579CCFC9BFA9E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobban, Christopher S.;Witkowski, Andrzej	Lobban, Christopher S., Witkowski, Andrzej (2023): Marine benthic diatoms of Guam: new records, Dictyoneis apapae sp. nov., and updates to the checklist. Micronesica 2023 (2): 1-75, DOI: 10.5281/zenodo.12118784
03C287AAFFE60B1DFF3578ABFED8F99A.text	03C287AAFFE60B1DFF3578ABFED8F99A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lyrella hennedyi (W. Smith) Stickle & D. G. Mann	<div><p>Lyrella hennedyi (W.Smith) Stickle &amp; D.G.Mann (including var. granulosa Grunow)</p><p>Reconsideration of the literature to identify specimens in Yap lead to a better identification of specimens reported in Lobban et al. (2012) as  L. clavata Gregory (Lobban &amp; Tharngan in prep.), based on Hustedt’s (1961–1966) key, differentiating groups on the basis of rostrate ends, including  clavata, from those with rounded apices, including  hennedyi . Stria densities are similar in these two species.</p></div>	https://treatment.plazi.org/id/03C287AAFFE60B1DFF3578ABFED8F99A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobban, Christopher S.;Witkowski, Andrzej	Lobban, Christopher S., Witkowski, Andrzej (2023): Marine benthic diatoms of Guam: new records, Dictyoneis apapae sp. nov., and updates to the checklist. Micronesica 2023 (2): 1-75, DOI: 10.5281/zenodo.12118784
03C287AAFFE60B1DFF357BB0FC1EF8B0.text	03C287AAFFE60B1DFF357BB0FC1EF8B0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nanofrustulum shiloi (J. J. Lee, Reimer & McEnery) Round, Hallsteinsen & Paasche 1999	<div><p>Nanofrustulum shiloi (J.J.Lee, Reimer &amp; McEnery) Round, Hallsteinsen &amp; Paasche</p><p>This species was reported from Guam by Lobban &amp; Navarro (2009) (figs. 40–43) but based on recent work by C.Li et al. (2018), the specimens illustrated are not this species or genus and cannot be identified to any of the other genera described there. These specimens must await further study. We do not have any other specimens so far that can be identified as  N. shiloi, although we anticipate it is present in the flora. The images remain posted in ProtistCentral under the name GU7B-UnID (http://www.protistcentral.org/Taxa/get/taxa_id/586261).</p></div>	https://treatment.plazi.org/id/03C287AAFFE60B1DFF357BB0FC1EF8B0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobban, Christopher S.;Witkowski, Andrzej	Lobban, Christopher S., Witkowski, Andrzej (2023): Marine benthic diatoms of Guam: new records, Dictyoneis apapae sp. nov., and updates to the checklist. Micronesica 2023 (2): 1-75, DOI: 10.5281/zenodo.12118784
03C287AAFFE70B1CFF357C38FB2CFE90.text	03C287AAFFE70B1CFF357C38FB2CFE90.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Proschkinia complanatoides (Hustedt) Karayeva	<div><p>Proschkinia complanatoides (Hustedt) Karayeva</p><p>Still a valid species but not what we have here; see  Proschkinia complanata above.</p></div>	https://treatment.plazi.org/id/03C287AAFFE70B1CFF357C38FB2CFE90	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobban, Christopher S.;Witkowski, Andrzej	Lobban, Christopher S., Witkowski, Andrzej (2023): Marine benthic diatoms of Guam: new records, Dictyoneis apapae sp. nov., and updates to the checklist. Micronesica 2023 (2): 1-75, DOI: 10.5281/zenodo.12118784
03C287AAFFE70B1CFF357CB5FBF2FE15.text	03C287AAFFE70B1CFF357CB5FBF2FE15.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Stauroneis retrostauron (Mann) Meister 1937	<div><p>Stauroneis retrostauron (Mann) Meister</p><p>Still a valid species but not what we have here; see  Parlibellus biblos above.</p></div>	https://treatment.plazi.org/id/03C287AAFFE70B1CFF357CB5FBF2FE15	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobban, Christopher S.;Witkowski, Andrzej	Lobban, Christopher S., Witkowski, Andrzej (2023): Marine benthic diatoms of Guam: new records, Dictyoneis apapae sp. nov., and updates to the checklist. Micronesica 2023 (2): 1-75, DOI: 10.5281/zenodo.12118784
03C287AAFFE70B1CFF357F32FB1EFD56.text	03C287AAFFE70B1CFF357F32FB1EFD56.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Synedra bacillaris (Grunow) Hustedt 1932	<div><p>Synedra bacillaris (Grunow) Hustedt</p><p>Reported by Lobban et al. (2012). Lobban et al. (2022) renamed that species  Grunowago bacillaris (Grunow) Lobban &amp; Ashworth but were unable to establish its presence in Guam, so established a new species,  Grunowago pacifica Lobban &amp; Ashworth, for the Guam material.</p></div>	https://treatment.plazi.org/id/03C287AAFFE70B1CFF357F32FB1EFD56	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobban, Christopher S.;Witkowski, Andrzej	Lobban, Christopher S., Witkowski, Andrzej (2023): Marine benthic diatoms of Guam: new records, Dictyoneis apapae sp. nov., and updates to the checklist. Micronesica 2023 (2): 1-75, DOI: 10.5281/zenodo.12118784
03C287AAFFE20B19FF357EDCFAE1FBA6.text	03C287AAFFE20B19FF357EDCFAE1FBA6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Campylodiscus giffenii (M. H. Giffen) Lobban	<div><p>Campylodiscus giffenii (M.H. Giffen) Lobban &amp; JoonS.Park in Park et al. (2022)</p><p>Synonym:  Surirella scalaris M.H. Giffen</p><p>Synonym:  Campylodiscus scalaris (M.H. Giffen) Lobban &amp; Joon S.Park in Park et al. (2018). Illegit.</p></div>	https://treatment.plazi.org/id/03C287AAFFE20B19FF357EDCFAE1FBA6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobban, Christopher S.;Witkowski, Andrzej	Lobban, Christopher S., Witkowski, Andrzej (2023): Marine benthic diatoms of Guam: new records, Dictyoneis apapae sp. nov., and updates to the checklist. Micronesica 2023 (2): 1-75, DOI: 10.5281/zenodo.12118784
03C287AAFFE20B19FF357F27FD5BFD87.text	03C287AAFFE20B19FF357F27FD5BFD87.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Campylodiscus neofastuosus (Ehrenberg) Ruck & Nakov	<div><p>Campylodiscus neofastuosus (Ehrenberg) Ruck &amp; Nakov</p><p>Synonym:  Surirella fastuosa Ehrenberg</p></div>	https://treatment.plazi.org/id/03C287AAFFE20B19FF357F27FD5BFD87	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobban, Christopher S.;Witkowski, Andrzej	Lobban, Christopher S., Witkowski, Andrzej (2023): Marine benthic diatoms of Guam: new records, Dictyoneis apapae sp. nov., and updates to the checklist. Micronesica 2023 (2): 1-75, DOI: 10.5281/zenodo.12118784
03C287AAFFE20B19FF357FA5FC99FD04.text	03C287AAFFE20B19FF357FA5FC99FD04.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Coronia ambigua (Greville) Ruck & Guiry	<div><p>Coronia ambigua (Greville) Ruck &amp; Guiry</p><p>Synonym:  Campylodiscus ambiguus Greville.</p></div>	https://treatment.plazi.org/id/03C287AAFFE20B19FF357FA5FC99FD04	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobban, Christopher S.;Witkowski, Andrzej	Lobban, Christopher S., Witkowski, Andrzej (2023): Marine benthic diatoms of Guam: new records, Dictyoneis apapae sp. nov., and updates to the checklist. Micronesica 2023 (2): 1-75, DOI: 10.5281/zenodo.12118784
03C287AAFFE20B19FF357E22FD61FC89.text	03C287AAFFE20B19FF357E22FD61FC89.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Coronia decora (Brebisson) Ruck & Guiry	<div><p>Coronia decora (Brébisson) Ruck &amp; Guiry</p><p>Synonym:  Campylodiscus decorus Brébisson</p></div>	https://treatment.plazi.org/id/03C287AAFFE20B19FF357E22FD61FC89	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobban, Christopher S.;Witkowski, Andrzej	Lobban, Christopher S., Witkowski, Andrzej (2023): Marine benthic diatoms of Guam: new records, Dictyoneis apapae sp. nov., and updates to the checklist. Micronesica 2023 (2): 1-75, DOI: 10.5281/zenodo.12118784
03C287AAFFE20B19FF357E5FFC23FC0F.text	03C287AAFFE20B19FF357E5FFC23FC0F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Coronia decora var. pinnata (Peragallo) Lobban 1888	<div><p>Coronia decora var. pinnata (Peragallo) Lobban &amp; JoonS.Park in Park et al. (2018)</p><p>Synonym:  Campylodiscus decorus var. pinnatus H. Peragallo</p></div>	https://treatment.plazi.org/id/03C287AAFFE20B19FF357E5FFC23FC0F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobban, Christopher S.;Witkowski, Andrzej	Lobban, Christopher S., Witkowski, Andrzej (2023): Marine benthic diatoms of Guam: new records, Dictyoneis apapae sp. nov., and updates to the checklist. Micronesica 2023 (2): 1-75, DOI: 10.5281/zenodo.12118784
03C287AAFFE20B19FF357983FD2EFB2A.text	03C287AAFFE20B19FF357983FD2EFB2A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Disymmetria excentrica (Lobban) Lobban 2023	<div><p>Disymmetria excentrica (Lobban) Lobban 2023b</p><p>Synonym:  Lauderia excentrica Lobban</p></div>	https://treatment.plazi.org/id/03C287AAFFE20B19FF357983FD2EFB2A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobban, Christopher S.;Witkowski, Andrzej	Lobban, Christopher S., Witkowski, Andrzej (2023): Marine benthic diatoms of Guam: new records, Dictyoneis apapae sp. nov., and updates to the checklist. Micronesica 2023 (2): 1-75, DOI: 10.5281/zenodo.12118784
03C287AAFFE20B19FF357897FD6CFA37.text	03C287AAFFE20B19FF357897FD6CFA37.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epithemia guettingeri (Krammer) Lobban	<div><p>Epithemia guettingeri (Krammer) Lobban &amp; JoonS. Park in Park et al. (2018)</p><p>Synonym:  Rhopalodia guettingerii Krammer</p></div>	https://treatment.plazi.org/id/03C287AAFFE20B19FF357897FD6CFA37	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobban, Christopher S.;Witkowski, Andrzej	Lobban, Christopher S., Witkowski, Andrzej (2023): Marine benthic diatoms of Guam: new records, Dictyoneis apapae sp. nov., and updates to the checklist. Micronesica 2023 (2): 1-75, DOI: 10.5281/zenodo.12118784
03C287AAFFE20B19FF357B14FCF3F9B4.text	03C287AAFFE20B19FF357B14FCF3F9B4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epithemia musculus Kutzing	<div><p>Epithemia musculus Kützing</p><p>Synonym:  Rhopalodia musculus (Kützing) O.F.Müller</p></div>	https://treatment.plazi.org/id/03C287AAFFE20B19FF357B14FCF3F9B4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobban, Christopher S.;Witkowski, Andrzej	Lobban, Christopher S., Witkowski, Andrzej (2023): Marine benthic diatoms of Guam: new records, Dictyoneis apapae sp. nov., and updates to the checklist. Micronesica 2023 (2): 1-75, DOI: 10.5281/zenodo.12118784
03C287AAFFE20B19FF357B92FBD2F8DB.text	03C287AAFFE20B19FF357B92FBD2F8DB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Epithemia pacifica Krammer 2018	<div><p>Epithemia pacifica (Krammer in Lange-Bertalot &amp; Krammer) Lobban &amp; JoonS.Park in Park et al. (2018)</p><p>Synonym:  Rhopalodia pacifica Krammer in Lange-Bertalot &amp; Krammer.</p></div>	https://treatment.plazi.org/id/03C287AAFFE20B19FF357B92FBD2F8DB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobban, Christopher S.;Witkowski, Andrzej	Lobban, Christopher S., Witkowski, Andrzej (2023): Marine benthic diatoms of Guam: new records, Dictyoneis apapae sp. nov., and updates to the checklist. Micronesica 2023 (2): 1-75, DOI: 10.5281/zenodo.12118784
03C287AAFFE20B18FF357A70FD7BFEFE.text	03C287AAFFE20B18FF357A70FD7BFEFE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Florella intermedia (Grunow) Lobban, Navarro & TM Schuster 2021	<div><p>Florella intermedia (Grunow) Lobban, Navarro &amp; TM Schuster 2021</p><p>Synonyms:  Striatella intermedia Grunow</p><p>Florella portoricensis Navarro</p></div>	https://treatment.plazi.org/id/03C287AAFFE20B18FF357A70FD7BFEFE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobban, Christopher S.;Witkowski, Andrzej	Lobban, Christopher S., Witkowski, Andrzej (2023): Marine benthic diatoms of Guam: new records, Dictyoneis apapae sp. nov., and updates to the checklist. Micronesica 2023 (2): 1-75, DOI: 10.5281/zenodo.12118784
03C287AAFFE30B18FF357C4CFD5FFE15.text	03C287AAFFE30B18FF357C4CFD5FFE15.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Florella lindigiana (Grunow) Lobban, Navarro & TM Schuster 2021	<div><p>Florella lindigiana (Grunow) Lobban, Navarro &amp; TM Schuster 2021</p><p>Synonyms:  Striatella lindigiana Grunow</p><p>Florella pascuensis Navarro</p></div>	https://treatment.plazi.org/id/03C287AAFFE30B18FF357C4CFD5FFE15	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobban, Christopher S.;Witkowski, Andrzej	Lobban, Christopher S., Witkowski, Andrzej (2023): Marine benthic diatoms of Guam: new records, Dictyoneis apapae sp. nov., and updates to the checklist. Micronesica 2023 (2): 1-75, DOI: 10.5281/zenodo.12118784
03C287AAFFE30B18FF357F32FB9FFD66.text	03C287AAFFE30B18FF357F32FB9FFD66.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Luticola tropica Levkov, Metzeltin & Pavlov 2013	<div><p>Luticola tropica Levkov, Metzeltin &amp; Pavlov 2013</p><p>Synonym:  Luticola inserata var. undulata (Hustedt) Navarro &amp; Lobban 2009</p><p>Refs: Levkov et al. 2013, Rybak et al. 2021</p><p>Brackish species reported from Guam by Navarro &amp; Lobban (2009)</p></div>	https://treatment.plazi.org/id/03C287AAFFE30B18FF357F32FB9FFD66	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobban, Christopher S.;Witkowski, Andrzej	Lobban, Christopher S., Witkowski, Andrzej (2023): Marine benthic diatoms of Guam: new records, Dictyoneis apapae sp. nov., and updates to the checklist. Micronesica 2023 (2): 1-75, DOI: 10.5281/zenodo.12118784
03C287AAFFE30B18FF357FC3FD15FCEB.text	03C287AAFFE30B18FF357FC3FD15FCEB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Neofragilaria anomala (Giffen) Witkowski & Dabek	<div><p>Neofragilaria anomala (Giffen) Witkowski &amp; Dąbek (in C.Li et al. 2015)</p><p>Synonym:  Neofragilaria nicobarica</p></div>	https://treatment.plazi.org/id/03C287AAFFE30B18FF357FC3FD15FCEB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobban, Christopher S.;Witkowski, Andrzej	Lobban, Christopher S., Witkowski, Andrzej (2023): Marine benthic diatoms of Guam: new records, Dictyoneis apapae sp. nov., and updates to the checklist. Micronesica 2023 (2): 1-75, DOI: 10.5281/zenodo.12118784
03C287AAFFE30B18FF357E40FCC5FC23.text	03C287AAFFE30B18FF357E40FCC5FC23.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Perissonoe crucifera Kitton 1987	<div><p>Perissonoë crucifera (Kitton in Prichard) Desikachary, Gowthaman, Hema, A.K.S.K. Prasad &amp; Prema (1987)</p><p>Syn.:  Perissonoë cruciata (Janisch &amp; Rabenhorst) G.W.Andrews &amp; Stoelzel</p><p>Reported by Lobban et al. (2012) under the synonym.</p></div>	https://treatment.plazi.org/id/03C287AAFFE30B18FF357E40FCC5FC23	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobban, Christopher S.;Witkowski, Andrzej	Lobban, Christopher S., Witkowski, Andrzej (2023): Marine benthic diatoms of Guam: new records, Dictyoneis apapae sp. nov., and updates to the checklist. Micronesica 2023 (2): 1-75, DOI: 10.5281/zenodo.12118784
03C287AAFFE30B18FF357909FDCCFB7C.text	03C287AAFFE30B18FF357909FDCCFB7C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Plagiogramma porcipellis Ashworth & Chunlian Li 2020	<div><p>Plagiogramma porcipellis Ashworth &amp; Chunlian Li in C.Li et al. 2020</p><p>Synonym:  Plagiogramma staurophoron (Gregory) Heiberg (as given in Lobban et al. 2012, p. 254, pl. 10, figs 3–5) and possibly the same as “  Plagiogramma cf. pulchellum Greville ” reported from Chuuk by J.S.Park et al. (2018)</p></div>	https://treatment.plazi.org/id/03C287AAFFE30B18FF357909FDCCFB7C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobban, Christopher S.;Witkowski, Andrzej	Lobban, Christopher S., Witkowski, Andrzej (2023): Marine benthic diatoms of Guam: new records, Dictyoneis apapae sp. nov., and updates to the checklist. Micronesica 2023 (2): 1-75, DOI: 10.5281/zenodo.12118784
03C287AAFFE30B18FF357B2EFC55F9B7.text	03C287AAFFE30B18FF357B2EFC55F9B7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudictyota bicornis (Cleve) PA Sims & DM Williams	<div><p>Pseudictyota bicornis (Cleve) PA Sims &amp; DM Williams in Sims et al. 2018</p><p>Synonym:  Triceratium bicorne Cleve</p><p>Sims et al. 2018: 38, figs 139–43 (as ‘Pseudodictyota bicorne’)</p></div>	https://treatment.plazi.org/id/03C287AAFFE30B18FF357B2EFC55F9B7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobban, Christopher S.;Witkowski, Andrzej	Lobban, Christopher S., Witkowski, Andrzej (2023): Marine benthic diatoms of Guam: new records, Dictyoneis apapae sp. nov., and updates to the checklist. Micronesica 2023 (2): 1-75, DOI: 10.5281/zenodo.12118784
03C287AAFFE30B18FF3579CAFB9BFA9B.text	03C287AAFFE30B18FF3579CAFB9BFA9B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudictyota dubia (Brightwell) PA Sims & DM Williams	<div><p>Pseudictyota dubia (Brightwell) PA Sims &amp; DM Williams in Sims et al. 2018</p><p>Synonym:  Triceratium dubium Brightwell</p><p>Sims et al. 2018, pl. 37, figs 133–138 (as ‘Pseudodictyota dubium’)</p></div>	https://treatment.plazi.org/id/03C287AAFFE30B18FF3579CAFB9BFA9B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobban, Christopher S.;Witkowski, Andrzej	Lobban, Christopher S., Witkowski, Andrzej (2023): Marine benthic diatoms of Guam: new records, Dictyoneis apapae sp. nov., and updates to the checklist. Micronesica 2023 (2): 1-75, DOI: 10.5281/zenodo.12118784
03C287AAFFE30B18FF3578B0FCE3FA19.text	03C287AAFFE30B18FF3578B0FCE3FA19.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudictyota reticulata (Roper) PA Sims & DM Williams	<div><p>Pseudictyota reticulata (Roper) PA Sims &amp; DM Williams in Sims et al. 2018</p><p>Synonym:  Triceratium dictyotum (Roper) Sims &amp; Ross.</p></div>	https://treatment.plazi.org/id/03C287AAFFE30B18FF3578B0FCE3FA19	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobban, Christopher S.;Witkowski, Andrzej	Lobban, Christopher S., Witkowski, Andrzej (2023): Marine benthic diatoms of Guam: new records, Dictyoneis apapae sp. nov., and updates to the checklist. Micronesica 2023 (2): 1-75, DOI: 10.5281/zenodo.12118784
03C287AAFFE30B27FF357A0AFD82FE7D.text	03C287AAFFE30B27FF357A0AFD82FE7D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Schizostauron trachyderma (F. Meister) Gorecka & Riaux-Gobin Our	<div><p>Schizostauron cf. trachyderma (F.Meister) Górecka &amp; Riaux-Gobin</p><p>Our species identified as  Achnanthes citronella is a species of  Schizostauron but we can so far only propose it as  Schizostauron cf. trachyderma (F.Meister) Górecka &amp; Riaux-Gobin. The difficulties in deciding whether it is that species,  S. kajotkei Dabek, Górecka &amp; Witkowski, or neither was discussed with reference to Chuuk specimens by J.S.Park et al. (2022), and the SEM images we have to date for this species do not resolve the question. See Riaux-Gobin et al. (2015), Davidovich et al. (2017) and Górecka et al. (2021).</p></div>	https://treatment.plazi.org/id/03C287AAFFE30B27FF357A0AFD82FE7D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobban, Christopher S.;Witkowski, Andrzej	Lobban, Christopher S., Witkowski, Andrzej (2023): Marine benthic diatoms of Guam: new records, Dictyoneis apapae sp. nov., and updates to the checklist. Micronesica 2023 (2): 1-75, DOI: 10.5281/zenodo.12118784
03C287AAFFDC0B27FF357CCBFD2DFDE2.text	03C287AAFFDC0B27FF357CCBFD2DFDE2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tetramphora decussata (Grunow 1877) Stepanek & Kociolek 2016	<div><p>Tetramphora decussata (Grunow) Stepanek &amp; Kociolek 2016</p><p>Synonym:  Amphora decussata Grunow</p></div>	https://treatment.plazi.org/id/03C287AAFFDC0B27FF357CCBFD2DFDE2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobban, Christopher S.;Witkowski, Andrzej	Lobban, Christopher S., Witkowski, Andrzej (2023): Marine benthic diatoms of Guam: new records, Dictyoneis apapae sp. nov., and updates to the checklist. Micronesica 2023 (2): 1-75, DOI: 10.5281/zenodo.12118784
03C287AAFFDC0B27FF357F48FCCAFD60.text	03C287AAFFDC0B27FF357F48FCCAFD60.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tetramphora intermedia (Cleve 1895) Stepanek & Kociolek 2016	<div><p>Tetramphora intermedia (Cleve) Stepanek &amp; Kociolek 2016</p><p>Synonym:  Amphora rhombica var. intermedia Cleve</p></div>	https://treatment.plazi.org/id/03C287AAFFDC0B27FF357F48FCCAFD60	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Lobban, Christopher S.;Witkowski, Andrzej	Lobban, Christopher S., Witkowski, Andrzej (2023): Marine benthic diatoms of Guam: new records, Dictyoneis apapae sp. nov., and updates to the checklist. Micronesica 2023 (2): 1-75, DOI: 10.5281/zenodo.12118784
