taxonID	type	description	language	source
03C5112FFFEDFFA8FF02FC10FD085537.taxon	materials_examined	Type species: Ceratopogon punctatus Meigen, 1804	en	Szadziewski, Ryszard, Dominiak, Patrycja, Sontag, Elżbieta, Krzemiński, Wiesław, Wang, Bo, Szwedo, Jacek (2019): Haematophagous biting midges of the extant genus Culicoides Latreille (Diptera: Ceratopogonidae) evolved during the mid-Cretaceous. Zootaxa 4688 (4): 535-548, DOI: 10.11646/zootaxa.4688.4.5
03C5112FFFEFFFADFF02FA67FC195019.taxon	description	Figs 3 A – C urn: lsid: zoobank. org: act: FBC 637 E 3 - 9205 - 4 CCB- 91 DA- 88115 D 9203 A 7	en	Szadziewski, Ryszard, Dominiak, Patrycja, Sontag, Elżbieta, Krzemiński, Wiesław, Wang, Bo, Szwedo, Jacek (2019): Haematophagous biting midges of the extant genus Culicoides Latreille (Diptera: Ceratopogonidae) evolved during the mid-Cretaceous. Zootaxa 4688 (4): 535-548, DOI: 10.11646/zootaxa.4688.4.5
03C5112FFFEFFFADFF02FA67FC195019.taxon	diagnosis	Diagnosis. This new species is distinguished from other Cretaceous Culicoides by the following female characters: antennal flagellum with cylindrical flagellomeres, 2 – 8 elongate, 9 – 13 greatly elongate, only flagellomere 1 with sensilla coeloconica; entire wing membrane with macrotrichia; palpal segment 3 slender without sensory pit; proboscis 2.1 x longer than eye height. Male unknown.	en	Szadziewski, Ryszard, Dominiak, Patrycja, Sontag, Elżbieta, Krzemiński, Wiesław, Wang, Bo, Szwedo, Jacek (2019): Haematophagous biting midges of the extant genus Culicoides Latreille (Diptera: Ceratopogonidae) evolved during the mid-Cretaceous. Zootaxa 4688 (4): 535-548, DOI: 10.11646/zootaxa.4688.4.5
03C5112FFFEFFFADFF02FA67FC195019.taxon	description	Description. Female (Fig. 3 A). Head-body length (minus antenna) ca. 1.0 mm. Head with greatly elongate proboscis (length 0.49 mm), ca. 2.1 x longer than eye height (Fig. 3 B). Eyes moderately separated. Antennal flagellum (Fig. 3 C) length 0.51 mm, with cylindrical flagellomeres, 2 – 8 elongate, 9 – 13 greatly elongate, flagellomere 1 with single sensillum coeloconicum; antennal ratio 1.02. Palpus (Fig. 3 B) 5 - segmented, slender, ca. 0.25 mm long; segment 3 greatly elongate (ca. 0.10 mm), without sensory pit. Wing (Fig. 3 A) length 0.68 mm, costal vein 0.48 mm; costal ratio 0.71; entire membrane including basal radial cell covered with macrotrichia. Hind tarsomere 1 swollen; tibial comb with 3 spines; TR (2) 2.3, TR (3) 2.1. Abdominal sternite 8 with distinct caudomedian excavation. Male. Unknown.	en	Szadziewski, Ryszard, Dominiak, Patrycja, Sontag, Elżbieta, Krzemiński, Wiesław, Wang, Bo, Szwedo, Jacek (2019): Haematophagous biting midges of the extant genus Culicoides Latreille (Diptera: Ceratopogonidae) evolved during the mid-Cretaceous. Zootaxa 4688 (4): 535-548, DOI: 10.11646/zootaxa.4688.4.5
03C5112FFFEFFFADFF02FA67FC195019.taxon	materials_examined	Material examined. Holotype female, NIGP 171164 (BA 02 14253). Burmese amber, Hukawng Valley, Myan- mar. Syninclusions: Scatopsidae 1 male, Copeognatha 1, Hemiptera, 1. Deposited in the collections of the Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing.	en	Szadziewski, Ryszard, Dominiak, Patrycja, Sontag, Elżbieta, Krzemiński, Wiesław, Wang, Bo, Szwedo, Jacek (2019): Haematophagous biting midges of the extant genus Culicoides Latreille (Diptera: Ceratopogonidae) evolved during the mid-Cretaceous. Zootaxa 4688 (4): 535-548, DOI: 10.11646/zootaxa.4688.4.5
03C5112FFFEFFFADFF02FA67FC195019.taxon	etymology	Etymology. This new species is dedicated to PhD student Błażej Bojarski, Museum of Amber Inclusions, Uni- versity of Gdańsk, for preparing the inclusions for entomological study.	en	Szadziewski, Ryszard, Dominiak, Patrycja, Sontag, Elżbieta, Krzemiński, Wiesław, Wang, Bo, Szwedo, Jacek (2019): Haematophagous biting midges of the extant genus Culicoides Latreille (Diptera: Ceratopogonidae) evolved during the mid-Cretaceous. Zootaxa 4688 (4): 535-548, DOI: 10.11646/zootaxa.4688.4.5
03C5112FFFE8FFACFF02FAF0FF7251D1.taxon	description	Figs 4 A – B urn: lsid: zoobank. org: act: 8 F 5 AD 5 B 1 - 78 A 5 - 491 A- 8036 - FD 4 EC 46 EAD 2 A	en	Szadziewski, Ryszard, Dominiak, Patrycja, Sontag, Elżbieta, Krzemiński, Wiesław, Wang, Bo, Szwedo, Jacek (2019): Haematophagous biting midges of the extant genus Culicoides Latreille (Diptera: Ceratopogonidae) evolved during the mid-Cretaceous. Zootaxa 4688 (4): 535-548, DOI: 10.11646/zootaxa.4688.4.5
03C5112FFFE8FFACFF02FAF0FF7251D1.taxon	diagnosis	Diagnosis. This new species is distinguished from other Cretaceous Culicoides by the following female characters: antennal flagellum with cylindrical flagellomeres, flagellomeres 2 – 8 much shorter than elongate 9 – 13, only flagellomere 1 with single sensillum coeloconicum; wing membrane except basal radial cell with macrotrichia; palpal segment 3 stout with distinct, deep sensory pit; proboscis moderately short. Male unkown.	en	Szadziewski, Ryszard, Dominiak, Patrycja, Sontag, Elżbieta, Krzemiński, Wiesław, Wang, Bo, Szwedo, Jacek (2019): Haematophagous biting midges of the extant genus Culicoides Latreille (Diptera: Ceratopogonidae) evolved during the mid-Cretaceous. Zootaxa 4688 (4): 535-548, DOI: 10.11646/zootaxa.4688.4.5
03C5112FFFE8FFACFF02FAF0FF7251D1.taxon	description	Description. Female (Fig. 4 A). Very well preserved, all structures present and well visible. Head-body length 1.03 mm. Eyes narrowly separated. Antennal flagellum (Fig. 4 B) length 0.54 mm; flagellomere 1 with single sensillum coeloconicum, flagellomeres 2 – 13 cylindrical without sensilla coecolonica; antennal ratio 1.44. Palpus (Fig. 4 B) 5 - segmented; segment 3 stout 0.045 mm long, with deep sensory pit. Wing (Fig. 4 A) length measured from basal arculus 0.70 mm; costal ratio 0.69; wing membrane except basal radial cell with macrotrichia. Legs slender, hind femur most massive; hind tibial comb with 3 spines; tarsomeres 4 cylindrical; tarsal ratios similar on all legs, TR (1) 2.2, TR (2) 2.5, TR (3) 2.3; claws small, equal-size. Male. Unknown.	en	Szadziewski, Ryszard, Dominiak, Patrycja, Sontag, Elżbieta, Krzemiński, Wiesław, Wang, Bo, Szwedo, Jacek (2019): Haematophagous biting midges of the extant genus Culicoides Latreille (Diptera: Ceratopogonidae) evolved during the mid-Cretaceous. Zootaxa 4688 (4): 535-548, DOI: 10.11646/zootaxa.4688.4.5
03C5112FFFE8FFACFF02FAF0FF7251D1.taxon	materials_examined	Material examined. Holotype female, BUB 1745, Burmese amber, Hukawng Valley, Myanmar. Syninclusions. Collembola 2, Acari 1, Empididae 1 female, Ceratopogonidae: Leptoconops 3 females. Deposited in Museum of Natural History, PAS Kraków.	en	Szadziewski, Ryszard, Dominiak, Patrycja, Sontag, Elżbieta, Krzemiński, Wiesław, Wang, Bo, Szwedo, Jacek (2019): Haematophagous biting midges of the extant genus Culicoides Latreille (Diptera: Ceratopogonidae) evolved during the mid-Cretaceous. Zootaxa 4688 (4): 535-548, DOI: 10.11646/zootaxa.4688.4.5
03C5112FFFE8FFACFF02FAF0FF7251D1.taxon	etymology	Etymology. This new species name refers to Burma (Myanmar), the country of origin of the amber in Southeast Asia.	en	Szadziewski, Ryszard, Dominiak, Patrycja, Sontag, Elżbieta, Krzemiński, Wiesław, Wang, Bo, Szwedo, Jacek (2019): Haematophagous biting midges of the extant genus Culicoides Latreille (Diptera: Ceratopogonidae) evolved during the mid-Cretaceous. Zootaxa 4688 (4): 535-548, DOI: 10.11646/zootaxa.4688.4.5
03C5112FFFE9FFAFFF02FA82FAEC51D0.taxon	description	Figs 5 A – C urn: lsid: zoobank. org: act: C 343228 E- 064 A- 4344 - A 63 B-C 748 F 54 A 468 D	en	Szadziewski, Ryszard, Dominiak, Patrycja, Sontag, Elżbieta, Krzemiński, Wiesław, Wang, Bo, Szwedo, Jacek (2019): Haematophagous biting midges of the extant genus Culicoides Latreille (Diptera: Ceratopogonidae) evolved during the mid-Cretaceous. Zootaxa 4688 (4): 535-548, DOI: 10.11646/zootaxa.4688.4.5
03C5112FFFE9FFAFFF02FA82FAEC51D0.taxon	diagnosis	Diagnosis. This new species is distinguished from other Cretaceous Culicoides by the following combination of female characters: antennal flagellum with flagellomeres 2 – 8 spherical, much shorter than the more elongate 9 – 13, only flagellomere 1 with 7 sensilla coeloconica; distal portion of wing membrane with macrotrichia; palpal segment 3 greatly swollen without sensory pit, with several distinct capitate sensilla; proboscis short. Male unknown.	en	Szadziewski, Ryszard, Dominiak, Patrycja, Sontag, Elżbieta, Krzemiński, Wiesław, Wang, Bo, Szwedo, Jacek (2019): Haematophagous biting midges of the extant genus Culicoides Latreille (Diptera: Ceratopogonidae) evolved during the mid-Cretaceous. Zootaxa 4688 (4): 535-548, DOI: 10.11646/zootaxa.4688.4.5
03C5112FFFE9FFAFFF02FA82FAEC51D0.taxon	description	Description. Female (Fig. 5 A). Poorly preserved, distorted, abdomen missing. Eyes bare, separation not visible. Antennal flagellum (Fig. 5 C) length 0.32 mm; flagellomere 1 with 7 sensilla coeloconica, flagellomeres 2 – 13 without sensilla coeloconica, but with distinct sensilla basiconica; flagellomeres 2 – 8 more or less spherical, much shorter than the more elongate 9 – 13; antennal ratio 1.13. Palpus (Fig. 5 B) 5 - segmented; palpal segment 3 greatly swollen without sensory pit, with several distinct capitate sensilla on ventral surface. Wing (Fig. 5 A) length 0.61 mm; costal ratio 0.69; distal portion of wing with macrotrichia in cells r 3, m 1 and m 2; basal radial and cubital cells without macrotrichia. Scutellum with about 8 long marginal setae. Tarsomere 1 of hind leg slender; TR (1) 2.1, TR (2) 2.0, TR (3) 1.8. Male. Unknown.	en	Szadziewski, Ryszard, Dominiak, Patrycja, Sontag, Elżbieta, Krzemiński, Wiesław, Wang, Bo, Szwedo, Jacek (2019): Haematophagous biting midges of the extant genus Culicoides Latreille (Diptera: Ceratopogonidae) evolved during the mid-Cretaceous. Zootaxa 4688 (4): 535-548, DOI: 10.11646/zootaxa.4688.4.5
03C5112FFFE9FFAFFF02FA82FAEC51D0.taxon	materials_examined	Material examined. Holotype female, MAI 5666. Burmese amber, Hukawng Valley, Myanmar; purchased by Sieghard Ellenberger from Kassel, Germany. Deposited in Museum of Amber Inclusions, University of Gdańsk.	en	Szadziewski, Ryszard, Dominiak, Patrycja, Sontag, Elżbieta, Krzemiński, Wiesław, Wang, Bo, Szwedo, Jacek (2019): Haematophagous biting midges of the extant genus Culicoides Latreille (Diptera: Ceratopogonidae) evolved during the mid-Cretaceous. Zootaxa 4688 (4): 535-548, DOI: 10.11646/zootaxa.4688.4.5
03C5112FFFE9FFAFFF02FA82FAEC51D0.taxon	etymology	Etymology. This new species is dedicated to Sieghard Ellenberger of Kassel, Germany, a generous collector who purchased this biting midge and donated it to the Museum of Amber Inclusions, University of Gdańsk.	en	Szadziewski, Ryszard, Dominiak, Patrycja, Sontag, Elżbieta, Krzemiński, Wiesław, Wang, Bo, Szwedo, Jacek (2019): Haematophagous biting midges of the extant genus Culicoides Latreille (Diptera: Ceratopogonidae) evolved during the mid-Cretaceous. Zootaxa 4688 (4): 535-548, DOI: 10.11646/zootaxa.4688.4.5
03C5112FFFEAFFAEFF02FA9CFA3351B5.taxon	description	Figs 6 A – D urn: lsid: zoobank. org: act: 4671 B 5 EB-F 1 B 5 - 4 F 9 C- 830 D- 40 E 4965 AA 00 A	en	Szadziewski, Ryszard, Dominiak, Patrycja, Sontag, Elżbieta, Krzemiński, Wiesław, Wang, Bo, Szwedo, Jacek (2019): Haematophagous biting midges of the extant genus Culicoides Latreille (Diptera: Ceratopogonidae) evolved during the mid-Cretaceous. Zootaxa 4688 (4): 535-548, DOI: 10.11646/zootaxa.4688.4.5
03C5112FFFEAFFAEFF02FA9CFA3351B5.taxon	diagnosis	Diagnosis. This species is distinguished from other Cretaceous Culicoides by the following combination of female characters: antennal flagellomere 1 with sensilla coeloconica, flagellomeres 2 – 8 subcylindrical to cylindrical, gradually increasing in length distally; palpal segment 3 slender, greatly elongate, without sensory pit; proboscis moderately elongate; wing membrane with macrotrichia on distal 1 / 3. Male unknown.	en	Szadziewski, Ryszard, Dominiak, Patrycja, Sontag, Elżbieta, Krzemiński, Wiesław, Wang, Bo, Szwedo, Jacek (2019): Haematophagous biting midges of the extant genus Culicoides Latreille (Diptera: Ceratopogonidae) evolved during the mid-Cretaceous. Zootaxa 4688 (4): 535-548, DOI: 10.11646/zootaxa.4688.4.5
03C5112FFFEAFFAEFF02FA9CFA3351B5.taxon	description	Description. Female (Fig. 6 A). Head-body length minus antenna 1.15 mm. Proboscis ca. 0.25 mm long, 1.25 x longer than eye height. Eyes broadly separated (Fig. 6 B). Antennal flagellum (Fig. 6 C) ca. 0.39 mm long; first flagellomere with sensilla coeloconica; flagellomeres 2 – 13 gradually increasing in length; antennal ratio ca. 1.08. Palpus (Fig. 6 D) 5 - segmented, very slender, greatly elongate; total length 0.21 mm; segment 3 greatly elongate, without sensory pit, length 0.10 mm. Wing (Fig. 6 A) length 0.63 mm; membrane with macrotrichia only on distal 1 / 3; basal radial cell without macrotrichia; costal ratio 0.81. Tarsomere 1 of hind leg slender; TR (1) 2.1, TR (3) 1.7. Abdominal sternite 8 with distinct caudomedian excavation. Male. Unknown.	en	Szadziewski, Ryszard, Dominiak, Patrycja, Sontag, Elżbieta, Krzemiński, Wiesław, Wang, Bo, Szwedo, Jacek (2019): Haematophagous biting midges of the extant genus Culicoides Latreille (Diptera: Ceratopogonidae) evolved during the mid-Cretaceous. Zootaxa 4688 (4): 535-548, DOI: 10.11646/zootaxa.4688.4.5
03C5112FFFEAFFAEFF02FA9CFA3351B5.taxon	materials_examined	Material examined. Holotype female, NIGP 171165 (BA 02 881). Burmese amber, Hukawng Valley, Myanmar. Syninclusion Hemiptera: Fulgoroidea of 3 rd or 4 th instar. Deposited in the collections of the Nanjing Institute of Ge- ology and Palaeontology, Chinese Academy of Sciences, Nanjing.	en	Szadziewski, Ryszard, Dominiak, Patrycja, Sontag, Elżbieta, Krzemiński, Wiesław, Wang, Bo, Szwedo, Jacek (2019): Haematophagous biting midges of the extant genus Culicoides Latreille (Diptera: Ceratopogonidae) evolved during the mid-Cretaceous. Zootaxa 4688 (4): 535-548, DOI: 10.11646/zootaxa.4688.4.5
03C5112FFFEAFFAEFF02FA9CFA3351B5.taxon	etymology	Etymology. The species name refers to Myanmar (Burma) in Southeast Asia where Burmese amber is mined.	en	Szadziewski, Ryszard, Dominiak, Patrycja, Sontag, Elżbieta, Krzemiński, Wiesław, Wang, Bo, Szwedo, Jacek (2019): Haematophagous biting midges of the extant genus Culicoides Latreille (Diptera: Ceratopogonidae) evolved during the mid-Cretaceous. Zootaxa 4688 (4): 535-548, DOI: 10.11646/zootaxa.4688.4.5
