identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03C5112FFFEDFFA8FF02FC10FD085537.text	03C5112FFFEDFFA8FF02FC10FD085537.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Culicoides Latreille 1809	<div><p>Genus Culicoides Latreille, 1809</p><p>Type species: Ceratopogon punctatus Meigen, 1804</p></div>	https://treatment.plazi.org/id/03C5112FFFEDFFA8FF02FC10FD085537	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Szadziewski, Ryszard;Dominiak, Patrycja;Sontag, Elżbieta;Krzemiński, Wiesław;Wang, Bo;Szwedo, Jacek	Szadziewski, Ryszard, Dominiak, Patrycja, Sontag, Elżbieta, Krzemiński, Wiesław, Wang, Bo, Szwedo, Jacek (2019): Haematophagous biting midges of the extant genus Culicoides Latreille (Diptera: Ceratopogonidae) evolved during the mid-Cretaceous. Zootaxa 4688 (4): 535-548, DOI: 10.11646/zootaxa.4688.4.5
03C5112FFFEEFFABFF02F95CFA3C56FC.text	03C5112FFFEEFFABFF02F95CFA3C56FC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Culicoides in Burmese	<div><p>Key to females of Culicoides in Burmese amber</p><p>1. Palpal segment 3 slender (Figs 3B, 6D)................................................................... 2</p><p>- Palpal segment 3 swollen (Figs 4B, 5B)................................................................... 3</p><p>2. Flagellomeres 2–4 cylindrical (Fig. 3C). Entire wing membrane with macrotrichia. Proboscis 2.1 x longer than eye height................................................................ C. bojarskii Szadziewski &amp; Dominiak, sp. nov.</p><p>- Flagellomeres 2–4 spherical (Fig. 6C). Wing membrane with macrotrichia on distal 1/3. Proboscis 1.25 x longer than eye height...................................................... C. myanmaricus Szadziewski &amp; Dominiak, sp. nov.</p><p>3. Palpal segment 3 with distinct sensory pit (Fig. 4B). Flagellomeres 2–8 long, cylindrical (Fig. 4B). Entire wing membrane except basal radial cell with macrotrichia............................ C. burmiticus Szadziewski &amp; Dominiak, sp. nov.</p><p>- Palpal segment 3 without sensory pit (Fig. 5B). Flagellomeres 2–8 short, spherical to subcylindrical (Fig. 5C). Wing membrane with macrotrichia on distal portion................................. C. ellenbergeri Szadziewski &amp; Dominiak, sp. nov.</p></div>	https://treatment.plazi.org/id/03C5112FFFEEFFABFF02F95CFA3C56FC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Szadziewski, Ryszard;Dominiak, Patrycja;Sontag, Elżbieta;Krzemiński, Wiesław;Wang, Bo;Szwedo, Jacek	Szadziewski, Ryszard, Dominiak, Patrycja, Sontag, Elżbieta, Krzemiński, Wiesław, Wang, Bo, Szwedo, Jacek (2019): Haematophagous biting midges of the extant genus Culicoides Latreille (Diptera: Ceratopogonidae) evolved during the mid-Cretaceous. Zootaxa 4688 (4): 535-548, DOI: 10.11646/zootaxa.4688.4.5
03C5112FFFEFFFADFF02FA67FC195019.text	03C5112FFFEFFFADFF02FA67FC195019.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Culicoides bojarskii Szadziewski	<div><p>Culicoides bojarskii Szadziewski &amp; Dominiak, sp. nov.</p><p>Figs 3 A–C</p><p>urn:lsid:zoobank.org:act: FBC637E3-9205-4CCB-91DA-88115D9203A7</p><p>Diagnosis. This new species is distinguished from other Cretaceous Culicoides by the following female characters: antennal flagellum with cylindrical flagellomeres, 2–8 elongate, 9–13 greatly elongate, only flagellomere 1 with sensilla coeloconica; entire wing membrane with macrotrichia; palpal segment 3 slender without sensory pit; proboscis 2.1 x longer than eye height. Male unknown.</p><p>Description. Female (Fig. 3A). Head-body length (minus antenna) ca. 1.0 mm. Head with greatly elongate proboscis (length 0.49 mm), ca. 2.1 x longer than eye height (Fig. 3B). Eyes moderately separated. Antennal flagellum (Fig. 3C) length 0.51 mm, with cylindrical flagellomeres, 2–8 elongate, 9–13 greatly elongate, flagellomere 1 with single sensillum coeloconicum; antennal ratio 1.02. Palpus (Fig. 3B) 5-segmented, slender, ca. 0.25 mm long; segment 3 greatly elongate (ca. 0.10 mm), without sensory pit. Wing (Fig. 3A) length 0.68 mm, costal vein 0.48 mm; costal ratio 0.71; entire membrane including basal radial cell covered with macrotrichia. Hind tarsomere 1 swollen; tibial comb with 3 spines; TR (2) 2.3, TR (3) 2.1. Abdominal sternite 8 with distinct caudomedian excavation.</p><p>Male. Unknown.</p><p>Material examined. Holotype female, NIGP 171164 (BA02 14253). Burmese amber, Hukawng Valley, Myan- mar. Syninclusions: Scatopsidae 1 male, Copeognatha 1, Hemiptera, 1. Deposited in the collections of the Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing.</p><p>Etymology. This new species is dedicated to PhD student Błażej Bojarski, Museum of Amber Inclusions, Uni- versity of Gdańsk, for preparing the inclusions for entomological study.</p></div>	https://treatment.plazi.org/id/03C5112FFFEFFFADFF02FA67FC195019	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Szadziewski, Ryszard;Dominiak, Patrycja;Sontag, Elżbieta;Krzemiński, Wiesław;Wang, Bo;Szwedo, Jacek	Szadziewski, Ryszard, Dominiak, Patrycja, Sontag, Elżbieta, Krzemiński, Wiesław, Wang, Bo, Szwedo, Jacek (2019): Haematophagous biting midges of the extant genus Culicoides Latreille (Diptera: Ceratopogonidae) evolved during the mid-Cretaceous. Zootaxa 4688 (4): 535-548, DOI: 10.11646/zootaxa.4688.4.5
03C5112FFFE8FFACFF02FAF0FF7251D1.text	03C5112FFFE8FFACFF02FAF0FF7251D1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Culicoides burmiticus Szadziewski	<div><p>Culicoides burmiticus Szadziewski &amp; Dominiak, sp. nov.</p><p>Figs 4 A–B</p><p>urn:lsid:zoobank.org:act: 8F5AD5B1-78A5-491A-8036-FD4EC46EAD2A</p><p>Diagnosis. This new species is distinguished from other Cretaceous Culicoides by the following female characters: antennal flagellum with cylindrical flagellomeres, flagellomeres 2–8 much shorter than elongate 9–13, only flagellomere 1 with single sensillum coeloconicum; wing membrane except basal radial cell with macrotrichia; palpal segment 3 stout with distinct, deep sensory pit; proboscis moderately short. Male unkown.</p><p>Description. Female (Fig. 4A). Very well preserved, all structures present and well visible. Head-body length 1.03 mm. Eyes narrowly separated. Antennal flagellum (Fig. 4B) length 0.54 mm; flagellomere 1 with single sensillum coeloconicum, flagellomeres 2–13 cylindrical without sensilla coecolonica; antennal ratio 1.44. Palpus (Fig. 4B) 5-segmented; segment 3 stout 0.045 mm long, with deep sensory pit. Wing (Fig. 4A) length measured from basal arculus 0.70 mm; costal ratio 0.69; wing membrane except basal radial cell with macrotrichia. Legs slender, hind femur most massive; hind tibial comb with 3 spines; tarsomeres 4 cylindrical; tarsal ratios similar on all legs, TR (1) 2.2, TR (2) 2.5, TR (3) 2.3; claws small, equal-size.</p><p>Male. Unknown.</p><p>Material examined. Holotype female, BUB 1745, Burmese amber, Hukawng Valley, Myanmar. Syninclusions. Collembola 2, Acari 1, Empididae 1 female, Ceratopogonidae: Leptoconops 3 females. Deposited in Museum of Natural History, PAS Kraków.</p><p>Etymology. This new species name refers to Burma (Myanmar), the country of origin of the amber in Southeast Asia.</p></div>	https://treatment.plazi.org/id/03C5112FFFE8FFACFF02FAF0FF7251D1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Szadziewski, Ryszard;Dominiak, Patrycja;Sontag, Elżbieta;Krzemiński, Wiesław;Wang, Bo;Szwedo, Jacek	Szadziewski, Ryszard, Dominiak, Patrycja, Sontag, Elżbieta, Krzemiński, Wiesław, Wang, Bo, Szwedo, Jacek (2019): Haematophagous biting midges of the extant genus Culicoides Latreille (Diptera: Ceratopogonidae) evolved during the mid-Cretaceous. Zootaxa 4688 (4): 535-548, DOI: 10.11646/zootaxa.4688.4.5
03C5112FFFE9FFAFFF02FA82FAEC51D0.text	03C5112FFFE9FFAFFF02FA82FAEC51D0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Culicoides ellenbergeri Szadziewski	<div><p>Culicoides ellenbergeri Szadziewski &amp; Dominiak, sp. nov.</p><p>Figs 5 A–C</p><p>urn:lsid:zoobank.org:act: C343228E-064A-4344-A63B-C748F54A468D</p><p>Diagnosis. This new species is distinguished from other Cretaceous Culicoides by the following combination of female characters: antennal flagellum with flagellomeres 2–8 spherical, much shorter than the more elongate 9–13, only flagellomere 1 with 7 sensilla coeloconica; distal portion of wing membrane with macrotrichia; palpal segment 3 greatly swollen without sensory pit, with several distinct capitate sensilla; proboscis short. Male unknown.</p><p>Description. Female (Fig. 5A). Poorly preserved, distorted, abdomen missing. Eyes bare, separation not visible. Antennal flagellum (Fig. 5C) length 0.32 mm; flagellomere 1 with 7 sensilla coeloconica, flagellomeres 2–13 without sensilla coeloconica, but with distinct sensilla basiconica; flagellomeres 2–8 more or less spherical, much shorter than the more elongate 9–13; antennal ratio 1.13. Palpus (Fig. 5B) 5-segmented; palpal segment 3 greatly swollen without sensory pit, with several distinct capitate sensilla on ventral surface. Wing (Fig. 5A) length 0.61 mm; costal ratio 0.69; distal portion of wing with macrotrichia in cells r 3, m 1 and m 2; basal radial and cubital cells without macrotrichia. Scutellum with about 8 long marginal setae. Tarsomere 1 of hind leg slender; TR (1) 2.1, TR (2) 2.0, TR (3) 1.8.</p><p>Male. Unknown.</p><p>Material examined. Holotype female, MAI 5666. Burmese amber, Hukawng Valley, Myanmar; purchased by Sieghard Ellenberger from Kassel, Germany. Deposited in Museum of Amber Inclusions, University of Gdańsk.</p><p>Etymology. This new species is dedicated to Sieghard Ellenberger of Kassel, Germany, a generous collector who purchased this biting midge and donated it to the Museum of Amber Inclusions, University of Gdańsk.</p></div>	https://treatment.plazi.org/id/03C5112FFFE9FFAFFF02FA82FAEC51D0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Szadziewski, Ryszard;Dominiak, Patrycja;Sontag, Elżbieta;Krzemiński, Wiesław;Wang, Bo;Szwedo, Jacek	Szadziewski, Ryszard, Dominiak, Patrycja, Sontag, Elżbieta, Krzemiński, Wiesław, Wang, Bo, Szwedo, Jacek (2019): Haematophagous biting midges of the extant genus Culicoides Latreille (Diptera: Ceratopogonidae) evolved during the mid-Cretaceous. Zootaxa 4688 (4): 535-548, DOI: 10.11646/zootaxa.4688.4.5
03C5112FFFEAFFAEFF02FA9CFA3351B5.text	03C5112FFFEAFFAEFF02FA9CFA3351B5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Culicoides myanmaricus Szadziewski	<div><p>Culicoides myanmaricus Szadziewski &amp; Dominiak, sp. nov.</p><p>Figs 6 A–D</p><p>urn:lsid:zoobank.org:act: 4671B5EB-F1B5-4F9C-830D-40E4965AA00A</p><p>Diagnosis. This species is distinguished from other Cretaceous Culicoides by the following combination of female characters: antennal flagellomere 1 with sensilla coeloconica, flagellomeres 2–8 subcylindrical to cylindrical, gradually increasing in length distally; palpal segment 3 slender, greatly elongate, without sensory pit; proboscis moderately elongate; wing membrane with macrotrichia on distal 1/3. Male unknown.</p><p>Description. Female (Fig. 6A). Head-body length minus antenna 1.15 mm. Proboscis ca. 0.25 mm long, 1.25 x longer than eye height. Eyes broadly separated (Fig. 6B). Antennal flagellum (Fig. 6C) ca. 0.39 mm long; first flagellomere with sensilla coeloconica; flagellomeres 2–13 gradually increasing in length; antennal ratio ca. 1.08. Palpus (Fig. 6D) 5-segmented, very slender, greatly elongate; total length 0.21 mm; segment 3 greatly elongate, without sensory pit, length 0.10 mm. Wing (Fig. 6A) length 0.63 mm; membrane with macrotrichia only on distal 1/3; basal radial cell without macrotrichia; costal ratio 0.81. Tarsomere 1 of hind leg slender; TR (1) 2.1, TR (3) 1.7. Abdominal sternite 8 with distinct caudomedian excavation.</p><p>Male. Unknown.</p><p>Material examined. Holotype female, NIGP 171165 (BA02 881). Burmese amber, Hukawng Valley, Myanmar. Syninclusion Hemiptera: Fulgoroidea of 3 rd or 4 th instar. Deposited in the collections of the Nanjing Institute of Ge- ology and Palaeontology, Chinese Academy of Sciences, Nanjing.</p><p>Etymology. The species name refers to Myanmar (Burma) in Southeast Asia where Burmese amber is mined.</p></div>	https://treatment.plazi.org/id/03C5112FFFEAFFAEFF02FA9CFA3351B5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Szadziewski, Ryszard;Dominiak, Patrycja;Sontag, Elżbieta;Krzemiński, Wiesław;Wang, Bo;Szwedo, Jacek	Szadziewski, Ryszard, Dominiak, Patrycja, Sontag, Elżbieta, Krzemiński, Wiesław, Wang, Bo, Szwedo, Jacek (2019): Haematophagous biting midges of the extant genus Culicoides Latreille (Diptera: Ceratopogonidae) evolved during the mid-Cretaceous. Zootaxa 4688 (4): 535-548, DOI: 10.11646/zootaxa.4688.4.5
