identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03C587AAFFFAFF9CFF6FFE750CDEFBAD.text	03C587AAFFFAFF9CFF6FFE750CDEFBAD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Stenotanais Bird & Holdich 1984	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Genus  Stenotanais Bird &amp; Holdich, 1984</p>
            <p>Diagnosis. (Modified after Larsen 2005) Female. Body elongate (11.5–13 times as long as wide). Pereonites longer than wide, with straight lateral edges. Pleon short (15–20 % of total body length). Antennule shorter than carapace, with four articles. Antenna with six articles. Mandibular molar tapering with terminal spines. Maxillule with eight to ten spiniform terminal setae. Maxillipedal endite with medial lobes, spiniform lateral corners and robust distal seta. Cheliped carpus prominent and often with large carpal shield. Chela robust. Fixed finger with two robust (as wide as a pereopod unguis) ventral setae. Pereopods without coxae. Pereopods 1–3 basis wider than on pereopods 4–6; merus and carpus with robust spiniform setae. Pereopods 4–6 dactylus and unguis not fused; dactylus with double-row of small spines. Pleopods short and broad, with plumose or simple setae. Uropods prominent, often as long as antennae: endopod with two articles; exopod with one or two articles; specialized setae with a wide and flat basis present on either exo- or endopod.</p>
            <p>Male. Antennule thicker than in female, functional mouthparts present (see remarks on new species).</p>
            <p> Type species:  Stenotanais crassiseta Bird &amp; Holdich, 1984 . </p>
            <p>Gender of generic name: Masculine.</p>
            <p> Species currently assigned to this genus:  Stenotanais arenasi n.sp. :  Stenotanais crassiseta Bird &amp; Holdich, 1984 ;  Stenotanais hamicauda Bird &amp; Holdich, 1984 ;  Stenotanais macrodactylus Larsen, 2005 . </p>
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	https://treatment.plazi.org/id/03C587AAFFFAFF9CFF6FFE750CDEFBAD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Larsen, Kim	Larsen, Kim (2011): The tanaidacean assemblage from the Central Pacific Manganese Nodule Province. II. The genera Stenotanais and Typhlotanais (Crustacea). Zootaxa 3088: 39-53, DOI: 10.5281/zenodo.279029
03C587AAFFFAFF99FF6FFB1A0CC0F809.text	03C587AAFFFAFF99FF6FFB1A0CC0F809.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Stenotanais arenasi	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Stenotanais arenasi n.sp.</p>
            <p>(Fig. 1 – 2)</p>
            <p>Material examined. Holotype, male, dissected, (MNHN-Ta1033). St. CAROT-B, no 22, 06/06-2004. 14˚03.10’N, 130˚07.74’W. 4954 meters. No sediment date available.</p>
            <p>Diagnosis. Male. Antennule article 4 only marginally longer than article 3, with fusion line. Cheliped carpal shield absent. Uropod basal article shorter than pleotelson and endopod; exopod biarticulated, shorter than endopod article 1, with one basally thick and flat seta. Female. Unknown.</p>
            <p>Etymology. The species is named after Dr. Francisco Arenas who organises and leads the fieldwork collection expeditions at CIIMAR.</p>
            <p>Description. Adult male, body length. 3.4 mm.</p>
            <p>Body (fig. 1A) elongate, 13 times as long as wide. Carapace elongate, as long as pereonite 2, not wider than pereonites. Pereon with pereonites all longer than wide. Pereonite 1 somewhat shorter than pereonite 2. Pereonite 2 and 4 subequal, longer than other pereonites. Pereonite 3 as long as pereonite 5. Pereonite 6 shortest. Pleon short (17% of total body length). All pleonites subequal, with pleopods. Pleotelson with blunt apex, longer than last two pleonites combined.</p>
            <p>Antennule (fig. 1B) shorter than carapace. Articles 1 and 2 twice as wide as article 4. Article 1 not much longer than article 2, with one simple and four setulose outer distal setae. Article 2 longer than articles 3 and 4 combined, almost square, with one robust outer and one small inner distal setae. Article 3 narrower than articles 1 and 2, marginally shorter than article 4, with one robust inner distal seta and outer row of setules. Article 4 about 0.25 times as long as article 1, with proximal fusion line (possibly an artefact), with six setae of which five are robust and one distal aesthetasc.</p>
            <p>Antenna (fig. 1C) 0.85 times as long as antennule. Article 1 fused to cephalothorax, wider than following articles, naked. Article 2 with one dorsodistal seta arising from a wide dorsal projection. Article 3 about as long as article 2, with one dorsodistal seta arising from a narrower dorsal projection. Article 4 longer than other articles, with two simple and two small setulated distal setae. Article 5 longer than article 3, with one simple distal seta. Article 6 smallest, decreasing in width at midlenght, with one robust seta at midlenght and three robust distal setae.</p>
            <p>Mouthparts. Labrum (fig. 1D) rounded, with distal setules. Mandibular molar tapering, with terminal spines. Left mandible (fig. 1E) incisor broad with three denticles; lacinia mobilis shaped like a broad spine. Right mandible (fig. 1F) incisor narrower than on left mandible with outer process. Labium (fig. 1G) outer corners with small, sparsely setulose process. Maxillule (fig. 1H) endite inner margin with small spines, with ten apical spiniform setae of which at least three are serrated; palp with two apical setae. Maxilla (fig. 1I) ovoid and featureless, but large relative to maxilliped. Maxilliped (fig. 1J) basis fused medially, with one seta at palp insertion; endites almost as wide as basis, divided into three lobe-like structures, outer corners with spiniform process. Palp article 1 without setae but with many scales; article 2 with one outer and three simple inner setae and scattered scales; article 3 with two larger bipinnate and three smaller simple inner setae; article 4 with four larger bipinnate and one smaller simple inner setae and one simple outer seta. Epignath (fig. 1K) naked.</p>
            <p>Cheliped (fig. 2A) basis with slender proximal part, attached to cephalothorax by elongated sclerite wrapped beneath the cephalothorax, naked. Merus prominent (half as long as carpus) with one ventral seta. Carpus marginally longer than propodus including fixed finger, without ventral carpal shield, with two ventral setae and one small dorsodistal seta. Propodus robust with two smaller simple and on larger bipinnate inner setae, with one robust (as thick as a pereopodal unguis) outer seta at dactylus insertion. Fixed finger with two robust (as thick as a peropodal unguis) ventral setae and three robust (as thick as a pereopodal unguis) setae on inner margin, inner margin also with relatively weak processes. Dactylus naked.</p>
            <p>Pereopod 1 (fig. 2B) Basis wider than those of pereopods 4–6, naked. Ischium with one ventral seta. Merus triangular, with one long (almost as long as carpus) spiniform seta. Carpus as long as merus, with two long distal spiniform setae and distal scales. Propodus with one spiniform ventral seta and one robust dorsal seta and dorsodistal spine, with distal scales. Dactylus and unguis of almost equal length, combined shorter than propodus, and not fused.</p>
            <p>Pereopod 2 (fig. 2C) broken. Carpus with two short and one long (longer than half of propodus) spiniform distal setae. Propodus with ventral subdistal spiniform seta but without dorsal seta, with distal and ventral setules/ scales.</p>
            <p>Pereopod 3 (fig. 2D) as pereopod 1 except: merus with one additional simple distal seta. Carpus with additional distal spiniform seta. Propodus with one additional simple distal seta.</p>
            <p>Pereopod 4 (fig. 2E) basis naked. Ischium with two setae. Merus with two spiniform distal setae. Carpus with two short and one long (longer than half of propodus) spiniform distal setae, with dorsodistal bone-shaped seta. Propodus with three spiniform distal setae. Dactylus and unguis combined longer than propodus and not fused. Dactylus with row sof ventral spines.</p>
            <p>Pereopod 5 (fig. 2F) as pereopod 4.</p>
            <p>Pereopod 6 (fig. 2G) as pereopod 5 except: basis with one setulated dorsoproximal seta. Ischium naked. Propodus with only two spiniform ventrodistal setae and three robust dorsodistal setae.</p>
            <p>Pleopods (fig. 1L) basal article naked. Endopod with 11 inner plumose setae, large gap between most proximal and other setae. Exopod with five plumose inner setae and one outer seta, with gap between most proximal and basal article.</p>
            <p>Uropods (fig. 1M) as long as antennule. Basal article longer than exopod but much shorter than pleotelson, naked. Endopod slender, with two articles of subequal length; article 1 with two distal setae; article 2 with one subdistal simple, four simple distal, and one setulated distal setae. Exopod slender, with two articles of subequal length; article 1 with one simple distal seta; article 2 with one simple and one specialized distal setae with flat and wide basis.</p>
            <p> Remarks. This species agrees well with most diagnostic characters of  Stenotanais : the long pereonites 1–5, the wide basis of the anterior pereopods, possession of thick uropodal setae, structures of maxilliped endite distal margin, and the thick propodal setae of the cheliped. However, the lack of a carpal shield on the chelipeds is in sharp contrast to other species of  Stenotanais . Also the mandibular molars, although tapering, are wider than seen in other species and the antennule article 4 shorter than in other species. These conflicting characters suggest that this species is early derived from  Stenotanais or may not belong to this genus at all, and genus affiliation should be considered preliminary. </p>
            <p>The wide antennule articles 1 and 2 leads me to believe this specimen is a male, although no penes could be observed. The thick antennule article is a character usually associated with males in the Tanaidomorpha (Lang 1968, Larsen 2002, Larsen &amp; Heard 2004). Also the subdivision (fusion line) of the 4th article, albeit incomplete and possibly an artifact, is indicative of the male gender (Larsen &amp; Shimomura 2007, 2009).</p>
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	https://treatment.plazi.org/id/03C587AAFFFAFF99FF6FFB1A0CC0F809	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Larsen, Kim	Larsen, Kim (2011): The tanaidacean assemblage from the Central Pacific Manganese Nodule Province. II. The genera Stenotanais and Typhlotanais (Crustacea). Zootaxa 3088: 39-53, DOI: 10.5281/zenodo.279029
03C587AAFFFEFF98FF6FFF100DF9FE25.text	03C587AAFFFEFF98FF6FFF100DF9FE25.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Stenotanais	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Key to  Stenotanais</p>
            <p> 1. Uropod basal article longer than pleotelson....................................................  S. macrodactylus</p>
            <p>- Uropod basal article shorter than pleotelson................................................................ 2</p>
            <p>2. Uropodal exopod bi-articulate, not reaching beyond first endopod article.......................................... 3</p>
            <p> - Uropodal exopod uni-articulate and oar-shaped, reaching at least to 0.75 of entire endopod length..........  S. hamicauda</p>
            <p> 3. Antennule article 4 almost twice as long as article 3. Cheliped carpal shield present. Endopod with one flat and wide seta dis- tally. Exopod without flat and wide setae.........................................................  S. crassiseta</p>
            <p> - Antennule article 4 only marginally longer than article 3. Cheliped carpal shield absent. Endopod without flat and wide seta. Exopod with one flat and wide seta................................................................  S. arenasi</p>
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	https://treatment.plazi.org/id/03C587AAFFFEFF98FF6FFF100DF9FE25	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Larsen, Kim	Larsen, Kim (2011): The tanaidacean assemblage from the Central Pacific Manganese Nodule Province. II. The genera Stenotanais and Typhlotanais (Crustacea). Zootaxa 3088: 39-53, DOI: 10.5281/zenodo.279029
03C587AAFFFEFF98FF6FFDD50BF4FACB.text	03C587AAFFFEFF98FF6FFDD50BF4FACB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Typhlotanais sensu subsp. lato G.O. Sars 1882	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Genus  
Typhlotanais 
sensu lato
G.O. Sars, 1882
</p>
            <p>Diagnosis. See Błażewicz-Paszkowycz (2007).</p>
            <p> Type species:  T. aequiremis (Lilljeborg, 1864) . </p>
            <p>Gender of generic name: Feminine.</p>
            <p> Species currently assigned to this genus: subgenus Monsmerinx Sieg, 1986;  T. brevicornis (Lilljeborg, 1864) ;  T. parvus Sieg, 1986 ; subgenus  Typhlotanais G.O. Sars, 1882 ;  T. aequiremis (Lilljeborg, 1864) ;  T. andeepae Blazewicz-Paszkowycz, 2007 ;  T. angstromensis Blazewicz-Paszkowycz &amp; Bamber, 2009 ;  T. assimilis G.O. Sars, 1882 ;  T. brachyurus Beddard, 1886 ;  T. compactus Kudinova-Pasternak, 1966 ;  T. cornutus (G.O. Sars, 1879) ;  T. crassus Dojiri &amp; Sieg, 1997 ;  T. dubius Tzareva, 1982 ;  T. eximius Hansen, 1913 ;  T. finmarchicus G.O. Sars, 1882 ;  T. froufesae n.sp. ;  T. grahami Blazewicz-Paszkowycz, 2004 ;  T. greenwichensis Shiino, 1970 ;  T. gruzovi Tzareva, 1982 ;  T. inaequipes Hansen, 1913 ;  T. incognitus Larsen et al., 2006 ;  T. inermis Hansen, 1913 ;  T. irregularis Hansen, 1913 ;  T. kerguelenensis Beddard, 1886 ;  T. kussakini Kudinova-Pasternak, 1970 ;  T. longicephala Kudinova-Pasternak, 1970 ;  T. longimanus Dollfus, 1897 ;  T. magnus Kudinova-Pasternak, 1990 ;  T. mananensis Wallace, 1919 ;  T. messinensis G.O. Sars, 1882 ;  T. mucronatus Hansen, 1913 ;  T. ohtsukae Larsen &amp; Shimomura, 2007 ;  T. parangularis Kudinova-Pasternak, 1975 ;  T. peculiaris Lang, 1968 ;  T. penicillatus G.O. Sars, 1882 ;  T. pereosetulosa n.sp. ;  T. plebejus Hansen, 1913 ;  T. plicatus Kudinova-Pasternak, 1993 ;  T. proctagon Tattersall, 1904 ;  T. profundus Hansen, 1913 ;  T. rectus Kudinova-Pasternak, 1966 ;  T. setosus Kudinova-Pasternak, 1966 ;  T. simplex Kudinova-Pasternak, 1984 ;  T. squamiger Blazewicz-Paszkowycz, 2007 ;  T. solidus Hansen, 1913 ;  T. spinicauda Hansen, 1913 ;  T. spinipes Kudinova-Pasternak, 1982 ;  T. tenuicornis G.O. Sars, 1882 ;  T. tenuimanus (Lilljeborg, 1864) ;  T. trispinosus Hansen, 1913 ;  T. variabilis Hansen, 1913 ;  T. williamsae Dojiri &amp; Sieg, 1997 . </p>
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	https://treatment.plazi.org/id/03C587AAFFFEFF98FF6FFDD50BF4FACB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Larsen, Kim	Larsen, Kim (2011): The tanaidacean assemblage from the Central Pacific Manganese Nodule Province. II. The genera Stenotanais and Typhlotanais (Crustacea). Zootaxa 3088: 39-53, DOI: 10.5281/zenodo.279029
03C587AAFFFEFF97FF6FFA050D51FE0B.text	03C587AAFFFEFF97FF6FFA050D51FE0B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Typhlotanais froufesae	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Typhlotanais froufesae n. sp.</p>
            <p>(Figs 3–5)</p>
            <p>Material examined. Holotype, non-ovigerous female, dissected, (MNHN-Ta1034). CAROT-B, no 24, 04/06- 2004. 14˚02.96’N, 130°08.18’W, 4909 m.</p>
            <p>Diagnosis. Antennule as long as carapace. Antenna article 1–3 with multiple setules; article 4 long (about five times the length of article 3). Pereopod 1 coxa with anterior process. Cheliped carpus without row of small dorsal setae. Pereopods 1–3 basis without multiple setules. Pereopods 2–3 merus and carpus with scales at distal margin. Pereopods 4–6 prickly tubercle comparably small (smaller than half of carpus) and surrounded by spines. Uropod endopod biarticulated; exopod uniarticulated.</p>
            <p> Etymology. The species is named after Dr. Elsa Froufe who conducts the genetic analysis of the  Tanaidacea at CIIMAR. </p>
            <p>Description. Female (body and appendages of holotype). Body length 1.8 mm</p>
            <p>Body (Fig. 3 A) long (12 times as long as wide). Cephalothorax longer than wide, with slightly rounded lateral margins. Pereonites all wider than long; pereonite1 shortest, almost twice as wide as long, half as long as pereonite 2; pereonites 2–4 subequal. Pereonite 5 shorter than pereonite 4. Pereonite 6 marginally longer than pereonite 1. Pleon 25% total body length. Pleonites 1–5 subequal. Pleotelson as long as combined length of last two pleonites, with rounded apex.</p>
            <p>Antennule (Fig. 3 B) as long as cephalothorax. Article 1 thick (just over twice as long as wide, with several proximal and medial setulated setae; distally with one simples and three setulated setae. Article 2 shorter than wide, with two simple distal setae. Article 3 long (more than three times as long as article 2), with apical spur-like process, two aesthestascs, four simple and one setulose distal setae.</p>
            <p>Antenna (Fig. 3 C) about 75% as long as antennule. Article 1 with scattered short thin setae. Article 2 with multiple setules, more than twice as long as article 3. Article 3 about as long as article 1, with few setules. Article 4 more than twice as long as article 5, with two simple and multiple setulated distal setae. Article 5 marginally longer than article 2, with one simple distal seta. Article 6 minute, with three terminal setae and one aesthetasc.</p>
            <p>Mouthparts. Labrum (Fig. 3 D) with rounded and setulose apex, without spines. Mandibles (Figs E–F) molar well developed, longer than incisor with clearly demarcated terminal spines. Left mandible (Fig. 3 F) incisor bluntly bifurcate; lacinia mobilis well developed and longer than incisor, with blunt apex. Right mandible (Fig. 3 E) incisor weakly bifid, with outer crenulations. Labium (Fig. 3 G) with flat, sparsely setulose distal corners. Maxillule (Fig. 3 H) endite with eight spiniform distal setae, palp as long as endite, with two apical setae Maxilla (Fig. 3 H) ovoid and featureless. Maxilliped (Fig. 5 A): basis fused medially, with long (as long as endite) distal seta near palp insertion; endite with one simple seta and serrated outer distal corners, with inner articulated tubercle outer projection and setules on corners; palp article1 with multiple setules or scales; article 2, with three bipinnate distal setae on inner margin and proximal setulations, with one smaller seta on outer margin; article 3 with four bipinnate setae on inner margin; article 4, with one simple seta outer and five bipinnate distal inner setae. Epignath (Fig. 3 I) naked tapering into a pointed apex.</p>
            <p>Cheliped (Fig. 4 A) basis attached via small triangular sclerite, distal part larger than proximal part, shorter than carpus, with one sub-dorsomedial seta near carpus connection. Merus less than half as long as carpus, with one ventral seta. Carpus almost three times longer than wide, with two simple ventromedial seta, one small stout ventrodistal seta, and one short dorsoproximal seta, with proximal scales. Propodus (including fixed finger) slender, marginally shorter than carpus, with one seta at dactylus insertion. Fixed finger with two ventral and three inner setae, inner margin smooth. Dactylus with dorsoproximal seta.</p>
            <p>Pereopod 1 (Fig. 4 B) coxa with anterior directed process with one seta. Basis longer than three succeeding articles combined, with one simple and one setulose dorsoproximal setae. Ischium with one ventral seta. Merus elongated, shorter than carpus, with one simple ventrodistal seta. Carpus only marginally shorter than propodus, with five distal setae, one thicker than others. Propodus only marginally longer than dactylus and unguis combined, with two dorso-subdistal setae, one distal seta, apical spine and one short ventrodistal spiniform seta. Dactylus less than half as long as unguis; both naked.</p>
            <p>Pereopod 2 (Fig. 4 C) coxa projection much smaller than on pereopod 1, apparently naked. Basis naked. Ischium with one ventral seta. Merus shorter than carpus, with prominent scales on distal margin, with three distal setae of which one is thicker than the others. Carpus with prominent scales on distal margin, with three dorsodistal setae of which one is thicker than the others, with one short ventrodistal spiniform seta. Propodus more than twice as long as dactylus and unguis combined, with two dorso-subdistal setae, apical spine and one short ventrodistal spiniform seta. Dactylus with one proximal seta, less than half as long as unguis.</p>
            <p>Pereopod 3 (Fig. 4 D) shorter than, but similar to pereopod 2 except: coxa projection with seta. Basis with dorsoproximal setulated seta. Dactylus with proximal seta.</p>
            <p>Pereopod 4 (Fig. 4 E) basis more robust than on pereopod 1–3, about 2.5 times longer than wide, apparently naked. Ischium with one ventral seta. Merus with scattered scales and two spiniform distal setae. Carpus apparently with one bone-shaped and one dorsodistal spiniform setae and with clearly defined prickly tubercle surrounded by row of minute spines. Propodus with two ventro-subdistal spiniform and one robust distal setae. Dactylus three times as long as unguis; both combined shorter than propodus, with row of small ventral spines.</p>
            <p>Pereopod 5 (Fig. 4 F) similar to pereopod 4 except: basis with one ventro-subdistal setulated seta. Ischium with two ventral setae. Propodus with ventral setules and scales and dorsomedial setulated seta.</p>
            <p>Pereopod 6 (Fig. 4 G) similar to as pereopod 5 except propodus with four distal and one subdistal spiniform setae but without dorsomedial setulated seta.</p>
            <p>Pleopods (Fig. 5 B) no differences between the five pairs were observed (although most were in a bad condition). Basal article elongated and naked; exopod with one plumose seta on inner margin and ten plumose setae on outer margin, pronounced gap between proximal and other setae; endopod with 15 plumose setae on outer margin, pronounced gap between proximal and other setae.</p>
            <p>Uropod (Fig. 3 J) basal article almost square. Exopod slender, 75% as long as endopod, uniarticulated with one small proximal seta; distally with two dissimilar length distal setae of which one is longer than exopod. Endopod slender, article 1 almost twice as long as article 2, with two simple and three setulated distal setae, article 2 with two short setulated and four simple distal setae of which two are longer than endopod.</p>
            <p> Remarks. The groupings by Błażewicz-Paszkowycz (2007) of typhlotanaid species, while not taxonomic entities, are very helpful for narrowing down identifications. The coxa projection of the pereopod 1 and the long maxilliped basis setae indicate that this species may belong to the ‘ greenwichensis’ group. However, the ‘  greenwichensis ’ group is also defined (Błażewicz-Paszkowycz 2007) by a suite of other characters: 1) Lack of gap between the cheliped basis and pereonite 1. 2) Row of short setae on the cheliped carpus. The new species clearly demonstrates a gap between the cheliped basis and pereonite 1, but it has no row of short seta on the cheliped carpus. Since another species,  T. mimosis , also displays a coxal projection on pereopods 1–3 but nevertheless is assigned to the ‘  mixtus ’ group (Błażewicz-Paszkowycz 2007), the distinctions between these groups are not clear. Therefore comparisons are made with the species from both these ‘groups’ while abstaining from assigning the new species to a specific group. </p>
            <p> The ‘ greenwichensis’ group currently consists of only two named species (  T. greenwichensis and  T. messinensis ). The new species differs from  T. greenwichensis by the much shorter antenna article 4, by lacking the row of small dorsal setae on the chelipeds carpus.  T. greenwichensis is also recorded from a much more shallow-water habitat of 79 meters (Shiino, 1970).  Typhlotanais froufesae n.sp. differs from  T. messinensis by the antennule being as long as the carapace and by lacking the row of small dorsal setae on the chelipeds carpus. The third ‘species’ in the greenwichensis’ group (‘?  T. greenwichensis ‘  [sic] sensu Błażewicz-Paszkowycz, 2007:107 ) might well be a separate, cryptic species rather than the ‘true’  T. greenwichensis but can be separated from  T. froufesae on the same characters as the original  T. greenwichensis . </p>
            <p> The ‘ mixtus’ group, also constitutes of only two named species (  T. mixtus Hansen, 1913 and  T. mimosis Błażewicz-Paszkowycz, 2007 ). From  T. mixtus , the new species differs by the pereopod coxa projection and from  T. mimosis by lacking the long dorsal setae on the first pereonite. Also, the comparatively small prickly tubercles are not in concord with the ‘ mixtus’ group definition. However, the ‘ mixtus’ group definition contains other conflicting characters, such as the pereopods 4–5 dactylus and unguis combined being almost as long as propodus while it is later stated that these are actually longer than the propodus (Błażewicz-Paszkowycz 2007:108). These conflicts suggest that many of these typhlotanaid ‘groups’ at present do not represent monophyletic lines. </p>
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	https://treatment.plazi.org/id/03C587AAFFFEFF97FF6FFA050D51FE0B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Larsen, Kim	Larsen, Kim (2011): The tanaidacean assemblage from the Central Pacific Manganese Nodule Province. II. The genera Stenotanais and Typhlotanais (Crustacea). Zootaxa 3088: 39-53, DOI: 10.5281/zenodo.279029
03C587AAFFF1FF90FF6FFDB8097DFA78.text	03C587AAFFF1FF90FF6FFDB8097DFA78.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Typhlotanais pereosetulosa	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Typhlotanais pereosetulosa n. sp.</p>
            <p>(Figs 6–7)</p>
            <p>Material examined. Holotype, female, dissected (MNHN-Ta 1035). Station Zone Ouest # 30, 12/06-2004, 09°33.81’N, 150°0.98’W, 5051 meters.</p>
            <p>Diagnosis. Pereonite 2 longer than other pereonites. Antennule as long as carapace. Antenna article 2 and 3 with multiple setules; antenna article 4 short (about three times the length of article 3). Labrum lateral edges with spines. Cheliped carpus without row of small dorsal setae. Pereopod 1 coxa without anterior process. Pereopod 1–3 basis with multiple setules. Pereopods 2–3 merus and carpus without serrated distal margin. Pereopods 4–6 prickly tubercle comparatively small (smaller than half of carpus) not surrounded by spines. Distal pleopods with fewer setae than anterior pleopods. Uropodal endo- and exopod biarticulated.</p>
            <p>Etymology. The species is named after the diagnostic character of the setation of the bases of pereopods 1–3.</p>
            <p>Description. Female (body and appendages of holotype). Body length 1.6 mm.</p>
            <p>Body (Fig. 6 A) elongated, 7 times as long as wide. Cephalothorax longer than wide, with slightly rounded lateral margins. Pereonites all except pereonite 2, wider than long; pereonites1 and 6 subequal, shorter than other pereonites, almost twice as wide as long, longer than half of pereonite 2. Pereonite 2 longest, square. Pereonites 3 and 5 subequal. Pereonite 5 shorter than pereonite 4. Pleon 25% total body length. Pleonites 1–5 subequal. Pleotelson as long as combined length of last two pleonites, with flat apex.</p>
            <p>Antennule (Fig. 6 B) longer than cephalothorax. Article 1 slender, four times as long as wide, with two simple medial setae, one simple and two setulated distal setae. Article 2 shorter than 0.25 times of article 1, with two simple distal setae. Article 3 twice as long as article 2, with apical process and seven distal setae of which two are longer than article.</p>
            <p>Antenna (Fig. 6 C) about 60% as long as antennule. Article 1 wider than other articles, with scattered short thin setae. Article 2 with multiple setules on all margins, less than twice as long as article 3. Article 3 shorter than article 1, with multiple setules and one simple distal seta. Article 4 twice as long as article 2, with sparse setulation and three distal setae of which one is longer than rest of antenna. Article 5 marginally longer than article 2, with one simple distal seta. Article 6 minute, with four terminal setae.</p>
            <p>Mouthparts: Labrum (Fig. 6 D) with flat and setulose apex, lateral margins with conspicuous spines. Mandibles (Figs 6 E–F) molar well developed, longer than incisor with distal spiniform denticles. Left mandible (Fig. 6 F) incisor bifurcate; lacinia mobilis well developed and larger than incisor, with pointed outer corner. Right mandible (Fig. 6 E) incisor blunt. Labium (Fig. 6 G) with flat, sparsely setulose distal corners. Maxillule (Fig. 6 H) endite with eight spiniform distal setae, palp not recovered. Maxilla (Fig. 6 I) small, ovoid and featureless. Maxilliped (Fig. 6 J): basis fused medially, with long distal seta near palp insertion; endite with two simple setae and serrated outer distal corners, no tubercles; palp article 1 naked; article 2 with three bipinnate distal setae on inner margin and proximal setulations, with one seta on outer margin; article 3 with four bipinnate setae on inner margin; article 4 less than half as wide as other articles, with one simple seta outer and five bipinnate distal inner setae. Epignath not recovered.</p>
            <p>Cheliped (Fig. 7 A) basis attached via triangular sclerite, distal part larger than proximal part, shorter than carpus, with one seta near carpus. Merus half as long as carpus, with one ventral seta. Carpus as long as propodus inclusive fixed finger, with two simple ventromedial setae, one small stout ventrodistal seta, and one short dorsodistal seta. Propodus (including fixed finger) more than twice as long as wide, with one seta at dactylus insertion. Fixed finger with two ventral and three inner setae, inner margin smooth. Dactylus naked.</p>
            <p>Pereopod 1 (Fig. 7 B) coxa without projections, with single seta. Basis six times as long as wide, with several setules on dorsal margin, with one dorsoproximal setulose seta. Ischium with one ventral seta. Merus marginally shorter than carpus, with one simple distal setae on each margin. Carpus sub-rectangular, longer than half of propodus, with two dorsodistal and two ventrodistal setae. Propodus less than twice as long as dactylus and unguis combined, with three dorso-subdistal setae, apical spine and one ventrodistal seta. Dactylus less than half as long as unguis, with single seta. Unguis with spatulated apex</p>
            <p>Pereopod 2 (Fig. 7 C) similar to pereopod 1 except dactylus and unguis shorter and unguis without spatulated apex.</p>
            <p>Pereopod 3 (Fig. 7 D) similar to pereopod 2.</p>
            <p>Pereopod 4 (Fig. 7 E) basis only marginally more robust than on pereopod 1–3, about 3 times longer than wide, apparently naked. Ischium with one ventral seta. Merus with proximal scales and two spiniform ventrodistal setae.</p>
            <p>Carpus apparently with one dorsodistal spiniform seta and with prickly tubercle surrounded by spines. Propodus with two ventrodistal and one dorsodistal spiniform setae. Dactylus three times as long as unguis; both combined shorter than propodus.</p>
            <p>Pereopod 5 (Fig. 7 F) similar to pereopod 4 except: merus with one ventrodistal spiniform seta. Carpus with prickly tubercle surrounded by spines and one spiniform seta. Propodus with ventral setules and scales.</p>
            <p>Pereopod 6 (Fig. 7 G) similar to pereopod 4: Carpus with prickly tubercle surrounded by spines and one dorsodistal spiniform setae. Propodus with ventral setules and scales, two ventrodistal and three dorsodistal spiniform setae. Dactylus with row of ventral spines</p>
            <p>Pleopods 1–3 (Fig. 6 K) basal article naked; exopod with one plumose seta on inner margin and 12 plumose setae on outer margin, pronounced gap between proximal and other setae; endopod with 13 plumose setae on outer margin, pronounced gap between proximal and other setae.</p>
            <p>Pleopods 4–5 (Fig. 6 L), basal article naked; exopod with one plumose seta on inner margin and with only five plumose setae on outer margin, pronounced gap between proximal and other setae; endopod with 12 plumose setae on outer margin; pronounced gap between proximal and other setae.</p>
            <p>Uropod (Fig. 6 M) basal article 0.4 times as long as wide. Rami biarticulated, slender. Exopod 75% as long as endopod, article 1 with one small seta; article 2 with two dissimilar length distal setae of which one is longer than exopod. Endopod article 1 with two simple distal setae, article 2 with four distal setae of which two are longer than endopod.</p>
            <p> Remarks. The new species key out to the ‘ mixtus’ group (Hansen 1913) using the key of Błażewicz-Paszkowycz (2007:25) but differs by the pereopods 4 and 5 merus prickly tubercles being smaller (smaller than half of carpus) and the combined length of dactylus and unguis clearly shorter than propodus. The new species differ from  T. mimosis by lacking the long dorsal setae on the first pereonite. </p>
            <p> The new species is unique in having multiple small short setae on the pereopod 1 and 2 bases. Multiple small setae on the basis of the first pereopods are also recorded from  T. aequiremis , and apparently only for pereopod 2 and; these are larger than in  T. pereosetulosa which also differ in the number of uropod articles as well as several other characters. </p>
            <p> The labrum spines is a somewhat unusual character but have been recorded from  Paratyphlotanais alveolus Błażewicz-Paszkowycz (2007) but the uropodal endopod distal article of  T. pereosetulosa is not shorter than the proximal and thus cannot be place in that genus. </p>
            <p> The dissimilar structure of the pleopods is another interesting feature that has not been examined in sufficient detail among the  Tanaidacea . Dissimilarity between pleopods have been observed in other tanaidaceanss to a greater (  Sphaeromapseudes - see Larsen (2011b) or lesser extent (  Zeuxo - Larsen et al. research in progress); also Błażewicz-Paszkowycz (2007) illustrates and even diagnoses dissimilar pleopods in  Meromonakantha natatoris although not in the same specimen. However, most tanaidacean species are described as with “all pleopods similar” and it is doubtful if these appendages have been studied in sufficient detail in previous descriptions (even those by this author). </p>
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	https://treatment.plazi.org/id/03C587AAFFF1FF90FF6FFDB8097DFA78	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Larsen, Kim	Larsen, Kim (2011): The tanaidacean assemblage from the Central Pacific Manganese Nodule Province. II. The genera Stenotanais and Typhlotanais (Crustacea). Zootaxa 3088: 39-53, DOI: 10.5281/zenodo.279029
