taxonID	type	description	language	source
03C787C3FFDCFFF3C2ABFC14FA57FDB3.taxon	description	(Figs. 1 – 12)	en	Cunha, Eduardo Victor De Paiva, Fernandes, José Antônio Marin (2022): New species of the Hypoxys triangularis group and the first contribution on the morphology of the internal genitalia of Hypoxys Amyot & Serville, 1843 (Hemiptera: Pentatomidae: Edessinae). Zootaxa 5168 (5): 501-520, DOI: 10.11646/zootaxa.5168.5.1
03C787C3FFDCFFF3C2ABFC14FA57FDB3.taxon	diagnosis	Diagnosis. Body uniformly green with corium reddish-brown (Fig. 10) (green with yellow spots in H. triangularis — see Nunes et al. 2020: Fig. 19 E – F), and a median large yellow spot on the ventral surface. Species of the H. triangularis group also lack any transversal black stripe on pronotum and ventral surface, like in the other species groups of Hypoxys, except H. triangularis (see Nunes et al. 2020: Fig. 19 E). Trichobothria with base yellowish (Fig. 10 F). Male genitalia. Paramere (Pa) usually ax-shaped (Fig. 1 B). Inner wall of pygophore laterally swollen (Figs. 3 B – C, 8 — black arrow). Ventral rim (Vr) with expansions slightly developed and rounded (Fig. 4 C). Female genitalia (Fig. 9). Valvifers VIII (Vf 8) with mediolateral crest swollen (black arrow); outer lateral angle of the posterior margin (red arrow) rounded or ending in an acute projection.	en	Cunha, Eduardo Victor De Paiva, Fernandes, José Antônio Marin (2022): New species of the Hypoxys triangularis group and the first contribution on the morphology of the internal genitalia of Hypoxys Amyot & Serville, 1843 (Hemiptera: Pentatomidae: Edessinae). Zootaxa 5168 (5): 501-520, DOI: 10.11646/zootaxa.5168.5.1
03C787C3FFDCFFF3C2ABFC14FA57FDB3.taxon	description	Redescription. Head. Mandibular plates without spot (Figs. 10, 11). Antennae yellow with setae more densely distributed on antennomeres III, IV and V. Thorax. Anterolateral margin of pronotum usually not punctured, except H. subrastratus (see Nunes et al. 2020: Fig. 19 I) and H. saltensis sp. n. (Fig. 10 E). Apex of humeral angle with margin brown (Figs. 10, 11). Pronotum and scutellum usually with brown punctures (Fig. 10 A). Anterior margin of scutellum without yellow stripe, like the one found in some species of H. quadridens and H. oxyacanthus groups. Costal margin of corium with a black spot reaching approximately 1 / 3 of corial extension (Figs. 10, 11). Propleura with concolorous punctures concentrated on posterior half. Evaporatorium slightly whitish (Figs. 10 F, 11 A). Metasternal process with arms of anterior bifurcation receiving fourth and most of third rostral segment (Fig. 10 A). Abdomen. Dorsal surface with brown spots. Connexivum with two barely excavated areas (Fig. 10 C) and usually not associated with dark spot (except H. triangularis). Pseudosutures with or without adjacent dark green spot (Figs. 10 B, 11 D). Male. Pygophore dorsally with brown to reddish brown spot occupying about 1 / 2 of the length of the pygophore; distal part of dorsal surface coarse (Fig. 1 A). Superior process (Pc) of genital cup brown and positioned near to dorsal rim (Fig. 2 C). Paramere directed posterolaterally (Fig. 9); base with a swollen and rounded protuberance (Fig. 9, blue arrow). Proctiger (Proc) with posterior face triangular, elliptical or pentagonal, and bent ventrally (Fig. 8). Female. Valvifers VIII and Valvifers IX (Vf 9) convex (Fig. 2 E). Laterotergites VIII (La 8) with margin black (Fig. 9). Laterotergites IX (La 9) with outer lateral margin sinuous, well projected beyond sclerite uniting laterotergites VIII; almost reaching level of laterotergites VIII (Fig. 4 E). Capsula seminalis (Cs) with two thick projections of equal lengths (Fig. 6 D); projections of capsula seminalis of the same width as pars intermedialis (Pi) (Fig. 6 F).	en	Cunha, Eduardo Victor De Paiva, Fernandes, José Antônio Marin (2022): New species of the Hypoxys triangularis group and the first contribution on the morphology of the internal genitalia of Hypoxys Amyot & Serville, 1843 (Hemiptera: Pentatomidae: Edessinae). Zootaxa 5168 (5): 501-520, DOI: 10.11646/zootaxa.5168.5.1
03C787C3FFDCFFF3C2ABFC14FA57FDB3.taxon	discussion	Comments. Five new species are described into H. triangularis group. Therefore, the new composition of the group is: Hypoxys triangularis; H. capito; H. subrastratus; H. myrtacivorus sp. n.; H. saltensis sp. n.; H. striatus sp. n.; H. curvatus sp. n. and H. ocanensis sp. n. The H. triangularis group has species associated with cultivated plants of the family Myrtaceae Juss. (Angiosperma; Myrtales): H. triangularis associated the Eugenia uniflora L. (brazilian-cherry) and Psidium guajava (guava) (Lima et al. 2010); H. subrastratus associated the Eugenia uniflora (brazilian-cherry) (data base Inaturalist; Fernandes pers. Comm.); H. myrtacivorus sp. n. associated the Myrciaria dubia (Kunth) McVaugh (camu-camu) (Panduro et al. 2001), Eugenia malaccensis L. (red jambo) and Eugenia uniflora L. (see comments of H. myrtacivorus sp. n.). Parameres of this species group are also almost straight in dorsal view and never strongly curved laterally, characteristic shared with Hypoxys balteatus group (see in Mendonça et al. 2021). The internal genitalia of some Edessinae genera were described, but never compared or discussed in a comprehensive way (Pantochlora vivida Stål, 1870, ilustrated — Kumar (1969); Brachystethus Laporte, 1832 — Grazia & Barcellos (2003); Paraedessa Silva & Fernandes, 2013 — Silva et al. (2013); Ascra Say, 1832 — Santos et al. (2015); Plagaedessa Almeida & Fernandes, 2018 — Almeida et al. (2018); Pygoda Amyot & Serville, 1843 — Fernandes et al. (2018); Graziaedessa anastrephae Eger, 2021, only the phallus described — Eger (2021); Edessa ovina group — Fernandes & Silva (2021 )). The internal genitalia of species of Hypoxys have not been describe yet. This is the first attempt to compare and use internal genitalia to characterize the groups of species. Studying the internal genitalia of species belonging to the Hypoxys, we observed some characteristics that support Hypoxys and its groups of species. The female internal genitalia present the following characteristics for Hypoxys: distal part of vesicular area of the ductus receptaculi sclerotized (Fig. 6 E, yellow arrow); base of the capsula seminalis spherical and sclerotized (Fig. 6 D, green arrow); capsula seminalis dilated with thick projections well-developed (Fig. 6 B); posterior annular crest more developed than anterior annular crest (Fig. 6 A). The phallus of Hypoxys species have the phallotheca cylindrical and curved dorsoventrally (Fig. 7). The differences observed between the species belonging to different groups of Hypoxys were: the H. quadridens (Fig. 6 A) and H. oxyacanthus (Fig. 6 B) groups share the projections of capsula seminalis wider and more dilated than the pars intermedialis. In H. triangularis (Fig. 6 D, E, F) and H. balteatus (Fig. 6 C) groups the projections of capsula seminalis are almost as wide as pars intermedialis. Groups H. quadridens and H. oxyacanthus share median portion of pars intermedialis thickened and spherical, while in H. triangularis group this structure is spherical or cylindrical, and in H. balteatus group the same structure is cylindrical. The H. quadridens group shows the base of capsula seminalis conical (Fig. 6 A, red arrow). Hypoxys oxyacanthus group has the globose base of capsula seminalis as wide as the segment X. A comparative study of the internal genitalia in all taxa of Edessinae can help to understand the evolution of the genitalia in the subfamily and add diagnostic characteristics to the genera.	en	Cunha, Eduardo Victor De Paiva, Fernandes, José Antônio Marin (2022): New species of the Hypoxys triangularis group and the first contribution on the morphology of the internal genitalia of Hypoxys Amyot & Serville, 1843 (Hemiptera: Pentatomidae: Edessinae). Zootaxa 5168 (5): 501-520, DOI: 10.11646/zootaxa.5168.5.1
03C787C3FFDCFFF3C2ABFC14FA57FDB3.taxon	distribution	Distribution (Fig. 12). BRAZIL; FRENCH GUIANA; VENEZUELA; PARAGUAY; SURINAME; GUIANA; BOLIVIA; PERU; COLOMBIA, and ARGENTINA.	en	Cunha, Eduardo Victor De Paiva, Fernandes, José Antônio Marin (2022): New species of the Hypoxys triangularis group and the first contribution on the morphology of the internal genitalia of Hypoxys Amyot & Serville, 1843 (Hemiptera: Pentatomidae: Edessinae). Zootaxa 5168 (5): 501-520, DOI: 10.11646/zootaxa.5168.5.1
03C787C3FFDEFFF5C2ABFDC2FC77FD1C.taxon	etymology	Etymology. The specific epithet refers to the feeding plants of this species (family Myrtaceae).	en	Cunha, Eduardo Victor De Paiva, Fernandes, José Antônio Marin (2022): New species of the Hypoxys triangularis group and the first contribution on the morphology of the internal genitalia of Hypoxys Amyot & Serville, 1843 (Hemiptera: Pentatomidae: Edessinae). Zootaxa 5168 (5): 501-520, DOI: 10.11646/zootaxa.5168.5.1
03C787C3FFDEFFF5C2ABFDC2FC77FD1C.taxon	materials_examined	Holotype. PERU. Loreto: 1 ♂ Iquitos, 14. V. 1992, G. Couturier Col. / Host Plant Myrciaria dubia (Myrtaceae) / Isla Mayuy INIAA / IVITA MNHN (EH) 25126 (MNHN). Paratypes (n = 23). No data, 1 ♀ 3093, 72304 (MZUSP); PERU. Loreto: 2 ♂ and 3 ♀ Carretera Iquitos – Nauta, Estacion El Dorado INIAA Unamurú, IX – 2006, J. Vasquez B., sobre Myrciaria dubia (Myrtaceae) “ camu camu ” MNHN (EH) 25130 - 25134 (MNHN); same data 1 ♂ and 1 ♀ (UFPA); 1 ♂ Cocha Supay, Jenaro Herrero, 26. X. 1988, 4 ° 55 ’ S 73 ° 40 ’ W, G. Couturier & H. Inga col., Plante – hote: Myrciaria dubia MNHN (EH) 25127 (MNHN); 1 ♂ Iquitos, 27. V. 1991, G. Couturier Col., Host Plant Eugenia malaccensis L. (Myrtaceae) MNHN (EH) 25128 (MNHN); BRAZIL. Pará: 1 ♂ and 1 ♀ Tucuruí, Rio Tocantins, Acamp. Bagagem, 03. VII. 1984, coleta Noturna (MPEG); 1 ♂ Belém, Bairro Umarizal, XI. 2014 (UFPA); 1 ♀ Belém, ICB — UFPA Campus Guamá, 17. XI. 2021, E. V. P. Cunha col., sobre Eugenia uniflora L., Obs.: indivíduo se alimentando no ramo (UFPA); 1 ♀ Belém, UFPA — Campus Guamá, 09. XII. 2016, E. V. P. Cunha col.; Host Plant Eugenia malaccensis L. (Myrtaceae) (UFPA); 1 ♀ Santarém, E. le Moult. (RMNH); Amazonas: 1 ♂ Igarapé Tacana, Solimões, II. 1962, Cândido & Dyrce (MNRJ); 1 ♀ Benjamin Constant, Rio Javari, II. 1942, Pko. (MNRJ); Rondônia: 2 ♀ 62 km Ariquemes, nr Fazenda Rancho Grande, 8 – 20. XI. 1994, J. E. Eger & L. B. & C. W. O’Brien (JEE); SURINAME. Sipaliwini: 1 ♀ Kayser Gebergte, Exped. H. A. Beatty, 25. XI – 14. XII. 1960, Sergiman Moentari leg. (FMNH); GUIANA. Georgetown: 1 ♀ Belair, Y. VII. 1931, R. T. Potts (USNM); VENEZUELA. Monagas: 1 ♀ Josepín, 50 m, 27. IX. 1955, F. Fernandez y C. J. Rosales (MBUC); COLOMBIA. Meta: 1 ♀ Peralonso, Caño Pachiaquiari, 4. IX. 1965, J. A. Ramos Collector (USNM).	en	Cunha, Eduardo Victor De Paiva, Fernandes, José Antônio Marin (2022): New species of the Hypoxys triangularis group and the first contribution on the morphology of the internal genitalia of Hypoxys Amyot & Serville, 1843 (Hemiptera: Pentatomidae: Edessinae). Zootaxa 5168 (5): 501-520, DOI: 10.11646/zootaxa.5168.5.1
03C787C3FFDEFFF5C2ABFDC2FC77FD1C.taxon	description	Measurements. Total length 11.1 – 15.9; total width 6.2 – 8.5; head length 1.9 – 2.3; head width 2.9 – 3.4; interocular distance: 1.5 – 1.8; length of antennomeres I: 0.7 – 1.0; II: 0.9 – 1.5; III: 1.3 – 1.6; IV: 2,5 – 3,1; V: 2.5 – 3.4; pronotum length 2.4 – 3.8; pronotum width 7.7 – 9.3; scutellum length 5.9 – 7.8; scutellum width 4.4 – 5.4.	en	Cunha, Eduardo Victor De Paiva, Fernandes, José Antônio Marin (2022): New species of the Hypoxys triangularis group and the first contribution on the morphology of the internal genitalia of Hypoxys Amyot & Serville, 1843 (Hemiptera: Pentatomidae: Edessinae). Zootaxa 5168 (5): 501-520, DOI: 10.11646/zootaxa.5168.5.1
03C787C3FFDEFFF5C2ABFDC2FC77FD1C.taxon	diagnosis	Diagnosis. Antennomeres with increasing lengths progressively (I <II <III <IV <V). Anterolateral margin of pronotum unpunctured (Fig. 10 A). Metasternal process with arms of anterior bifurcation robust and receiving all third rostral segment (Fig. 10 B). Pseudosutures of abdomen with adjacent dark green spot (Fig. 10 B). Trichobothria with yellow base (Fig. 10 B). Male genitalia. Dorsal surface with dark spot occupying 1 / 3 of the length of the pygophore (Fig. 1 A). Superior process of the genital cup long and narrow (Fig. 1 B). Paramere with dorsal margin strongly concave; anterior lobe large, subrectangular, reaching dorsal rim; posterior lobe short, rectangular laterally directed (Figs. 1 B, 1 D). Proctiger strongly excavated laterally, excavation and posterior face covered by long and sparse setae; dorsal face tapering posteriorly; posterior face triangular, small, below the level of dorsal face and ventrally bent, lateral areas slightly swollen (Fig. 1 D, 8 D). Inner wall swollen area globose and close to posterolateral angles (Fig. 8 D, black arrow). Ventral rim strongly excavated; bottom of median excavation convex (Fig. 1 C). Phallus (Fig. 7 C). Vesica (vs) half-moon-shaped in lateral view. Phallotheca (ph) slightly curved. Female genitalia (Figs. 1 E). Valvifers VIII with rounded excavation; booth distal angles dentiform (Fig. 9, red arrow); inner angles reaching valvifers IX; mediolateral crest swollen along outer lateral margin (Fig. 9 D, black arrow); mesial borders contiguous forming a spiniform projection (Fig. 9 D). Valvulae IX (Va 9) exposed and sclerotized. Pars intermedialis (pi) spherical medially (Fig. 6 F). Capsula seminalis with three thick projections almost as long as capsula seminalis (Fig. 6 F).	en	Cunha, Eduardo Victor De Paiva, Fernandes, José Antônio Marin (2022): New species of the Hypoxys triangularis group and the first contribution on the morphology of the internal genitalia of Hypoxys Amyot & Serville, 1843 (Hemiptera: Pentatomidae: Edessinae). Zootaxa 5168 (5): 501-520, DOI: 10.11646/zootaxa.5168.5.1
03C787C3FFDEFFF5C2ABFDC2FC77FD1C.taxon	discussion	Comments. This species was collected only on Myrtaceae plants, E. malaccensis (red jambo), E. uniflora (brazilian-cherry) and M. dubia (camu-camu). This species was observed feeding on branches from E. uniflora L., but in M. dubia it was observed feeding on fruits. Hypoxys myrtacivorus sp. n. was considered pest of camu-camu, M. dubia, in Peru, causing a transparent spot with a central red dot (Panduro et al. 2001). Hypoxys myrtacivorus sp. n. had nymphal stages and biological aspects described by Iannacone et al. (2007), identified as Edessa aff. aulacosterna Stål, 1872, but author mentioned that the species probably was new to science. Nevertheless, H. myrtacivorus sp. n. differs from Edessa aulacosterna by several aspects, but two are very conspicuous: humeral angles acute and slightly developed with apex brown to black (not developed, rounded and concolorous in E. aulacosterna); anterior arms of metasternal process swollen and tapering anteriorly (laterally compressed in E. aulacosterna). The shape of the paramere of this species resembles H. curvatus sp. n., but differ because the apex of the posterior lobe is acuminate in H. myrtacivorus sp. n. (in H. curvatus sp. n. the apices of the anterior and posterior lobes are rounded).	en	Cunha, Eduardo Victor De Paiva, Fernandes, José Antônio Marin (2022): New species of the Hypoxys triangularis group and the first contribution on the morphology of the internal genitalia of Hypoxys Amyot & Serville, 1843 (Hemiptera: Pentatomidae: Edessinae). Zootaxa 5168 (5): 501-520, DOI: 10.11646/zootaxa.5168.5.1
03C787C3FFDEFFF5C2ABFDC2FC77FD1C.taxon	distribution	Distribution (Fig. 12). PERU: Loreto; BRAZIL: Pará, Amazonas, Rondônia; SURINAME: Sipaliwini; GUIANA: Georgetown; VENEZUELA: Monagas; COLOMBIA: Meta.	en	Cunha, Eduardo Victor De Paiva, Fernandes, José Antônio Marin (2022): New species of the Hypoxys triangularis group and the first contribution on the morphology of the internal genitalia of Hypoxys Amyot & Serville, 1843 (Hemiptera: Pentatomidae: Edessinae). Zootaxa 5168 (5): 501-520, DOI: 10.11646/zootaxa.5168.5.1
03C787C3FFD8FFF5C2ABFD7BFAE2F954.taxon	etymology	Etymology. The specific epithet refers to the paramere posterior face striated.	en	Cunha, Eduardo Victor De Paiva, Fernandes, José Antônio Marin (2022): New species of the Hypoxys triangularis group and the first contribution on the morphology of the internal genitalia of Hypoxys Amyot & Serville, 1843 (Hemiptera: Pentatomidae: Edessinae). Zootaxa 5168 (5): 501-520, DOI: 10.11646/zootaxa.5168.5.1
03C787C3FFD8FFF5C2ABFD7BFAE2F954.taxon	materials_examined	Holotype. BRAZIL. Pará: 1 ♂ Óbidos, VIII. 1978, A. C. Domingos Coll. (MNRJ). Paratypes (n = 3). BRAZIL. Pará: 1 ♀ Óbidos, XII. 1963, Dirings Coll. (MZUSP); 1 ♀ e 1 ♂ Santarém, VI. 1919, S. M. Klages Coll. (CMNH).	en	Cunha, Eduardo Victor De Paiva, Fernandes, José Antônio Marin (2022): New species of the Hypoxys triangularis group and the first contribution on the morphology of the internal genitalia of Hypoxys Amyot & Serville, 1843 (Hemiptera: Pentatomidae: Edessinae). Zootaxa 5168 (5): 501-520, DOI: 10.11646/zootaxa.5168.5.1
03C787C3FFD8FFF5C2ABFD7BFAE2F954.taxon	description	Measurements. Total length 14.3 – 14.8; total width 7.7 – 8.0; head length 1.4 – 1.5; head width 3.2 – 3,4; interocular distance: 1.6 – 1.8; length of antenomeres I: 0.9 – 1.0; II: 1.0 – 1.1; III: 1.5; IV: 3.1 – 3.2; V: 3.6; pronotum length 2.7 – 3.0; pronotum width 8.9 – 9.2; scutellum length 6.5 – 6.7; scutellum width 5.0 – 5.3. Diagnose. Antennomeres with increasing lengths progressively (I <II <III <IV <V). Anterolateral margin of pronotum not punctured (Fig. 10 C). Corium reddish brown (Fig. 10 C). Metasternal process with arms of anterior bifurcation receiving 2 / 3 of third rostral segment (Fig. 10 D). Pseudosutures of abdomen with adjacent dark green spot (Fig. 10 D). Male genitalia. Dorsal surface with spot restricted to coarse area of the pygophore (Fig. 2 A). Dorsal rim sinuous (Fig. 2 A); lateral expansions slightly globose and almost level with posterolateral angles (Fig. 2 C). Superior process of the genital cup subtriangular, small and swollen (Fig. 2 B). Paramere with head slightly concave; anterior and posterior lobe subtriangular; anterior lobe large with external face striated and reaching superior process of the genital cup; posterior lobe smaller than the anterior lobe and laterally directed (Fig. 2 D). Proctiger shallowly excavated laterally, excavation covered by long and dense setae; posterior face elliptical (Fig. 2 B). Inner wall with small swollen area contiguous to posterolateral angles (Fig. 8 F, black arrow). Ventral rim in widely opened V-shaped with expansions hardly visible (Fig. 2 C). Female genitalia (Fig. 2 E). Valvifers VIII with distal margin slightly arched; outer and inner angles of the posterior margin rounded; mediolateral crest slightly swollen (Fig. 9 F, black arrow); mesial borders with excavation narrow, deep and U-shaped. Valvulae IX clearly exposed.	en	Cunha, Eduardo Victor De Paiva, Fernandes, José Antônio Marin (2022): New species of the Hypoxys triangularis group and the first contribution on the morphology of the internal genitalia of Hypoxys Amyot & Serville, 1843 (Hemiptera: Pentatomidae: Edessinae). Zootaxa 5168 (5): 501-520, DOI: 10.11646/zootaxa.5168.5.1
03C787C3FFD8FFF5C2ABFD7BFAE2F954.taxon	distribution	Distribution (Fig. 12). BRAZIL: Pará.	en	Cunha, Eduardo Victor De Paiva, Fernandes, José Antônio Marin (2022): New species of the Hypoxys triangularis group and the first contribution on the morphology of the internal genitalia of Hypoxys Amyot & Serville, 1843 (Hemiptera: Pentatomidae: Edessinae). Zootaxa 5168 (5): 501-520, DOI: 10.11646/zootaxa.5168.5.1
03C787C3FFD8FFF5C2ABFD7BFAE2F954.taxon	discussion	Comments. The superior process of the genital cup of H. triangularis and H. striatus sp. n. are the smallest processes in the group, but in H. triangularis it is rounded and concave and in H. striatus sp. n. it is subtriangular and swollen. This species is the only one with expansions of the ventral rim hardly visible. The mesial borders of the valvifers VIII of H. curvatus sp. n., H. capito and H. subrastratus have the excavation forming a “ U ”. However, H. striatus sp. n. can be separated from the other two species by the more deeply excavated mesial borders.	en	Cunha, Eduardo Victor De Paiva, Fernandes, José Antônio Marin (2022): New species of the Hypoxys triangularis group and the first contribution on the morphology of the internal genitalia of Hypoxys Amyot & Serville, 1843 (Hemiptera: Pentatomidae: Edessinae). Zootaxa 5168 (5): 501-520, DOI: 10.11646/zootaxa.5168.5.1
03C787C3FFD8FFF8C2ABF8AEFF14FED8.taxon	etymology	Etymology. The specific epithet refers to Province of Salta (Argentina) where the specimens collected.	en	Cunha, Eduardo Victor De Paiva, Fernandes, José Antônio Marin (2022): New species of the Hypoxys triangularis group and the first contribution on the morphology of the internal genitalia of Hypoxys Amyot & Serville, 1843 (Hemiptera: Pentatomidae: Edessinae). Zootaxa 5168 (5): 501-520, DOI: 10.11646/zootaxa.5168.5.1
03C787C3FFD8FFF8C2ABF8AEFF14FED8.taxon	materials_examined	Holotype. ARGENTINA. Province of Salta: 1 ♂ Quebrada of San Lorenzo, [Quebrada of Catayete, Province of Salta?], 10. II. 1953, R. Foerster Coll. (MMB).	en	Cunha, Eduardo Victor De Paiva, Fernandes, José Antônio Marin (2022): New species of the Hypoxys triangularis group and the first contribution on the morphology of the internal genitalia of Hypoxys Amyot & Serville, 1843 (Hemiptera: Pentatomidae: Edessinae). Zootaxa 5168 (5): 501-520, DOI: 10.11646/zootaxa.5168.5.1
03C787C3FFD8FFF8C2ABF8AEFF14FED8.taxon	description	Measurements. Total length 13.6; total width 6.6; head length 1.4; head width 3.0; interocular distance: 1.6; length of antennomeres I: 0.9; II: 0.7; III: 1.4; IV: 2.5; V: lost; pronotum length 2.6; pronotum width 7.7; scutellum length 5.7; scutellum width 4.4.	en	Cunha, Eduardo Victor De Paiva, Fernandes, José Antônio Marin (2022): New species of the Hypoxys triangularis group and the first contribution on the morphology of the internal genitalia of Hypoxys Amyot & Serville, 1843 (Hemiptera: Pentatomidae: Edessinae). Zootaxa 5168 (5): 501-520, DOI: 10.11646/zootaxa.5168.5.1
03C787C3FFD8FFF8C2ABF8AEFF14FED8.taxon	diagnosis	Diagnosis. Second antennomer smaller than the others (II <I <III <IV). Anterolateral margin of pronotum punctured (Fig. 10 E). Scutellum with punctures of the same size (Fig. 10 E). Hemelytrom membrane transparent (Fig. 10 E). Metasternal process with arms of anterior bifurcation receiving 2 / 3 of third rostral segment (Fig. 10 F). Pseudosutures of abdomen without adjacent dark green spot (Fig. 10 F). Male genitalia. Dorsal surface with spot extending about ½ of the length of the pygophore (Fig. 3 A). Dorsal rim with lateral expansions level with posterolateral angles and rounded (Figs. 3 A, 3 C). Superior process of the genital cup elliptic, face rough in posterior view (Fig. 3 B). Paramere spearhead-shaped, lobes slightly developed, apex truncated (Figs. 3 B, 3 D). Proctiger strongly excavated laterally, excavation covered by long and dense setae; dorsal face tapering posteriorly; posterior face subpentagonal, basal third constricted (Figs. 3 B, 3 D). Inner wall with swollen area large and close to posterolateral angles (Fig. 3 D, black arrow). Ventral rim strongly excavated, U-shaped; expansions ending laterally in a rounded projection (Fig. 4 C). Female genitalia. Unknown.	en	Cunha, Eduardo Victor De Paiva, Fernandes, José Antônio Marin (2022): New species of the Hypoxys triangularis group and the first contribution on the morphology of the internal genitalia of Hypoxys Amyot & Serville, 1843 (Hemiptera: Pentatomidae: Edessinae). Zootaxa 5168 (5): 501-520, DOI: 10.11646/zootaxa.5168.5.1
03C787C3FFD8FFF8C2ABF8AEFF14FED8.taxon	distribution	Distribution (Fig. 12). ARGENTINA: Salta.	en	Cunha, Eduardo Victor De Paiva, Fernandes, José Antônio Marin (2022): New species of the Hypoxys triangularis group and the first contribution on the morphology of the internal genitalia of Hypoxys Amyot & Serville, 1843 (Hemiptera: Pentatomidae: Edessinae). Zootaxa 5168 (5): 501-520, DOI: 10.11646/zootaxa.5168.5.1
03C787C3FFD8FFF8C2ABF8AEFF14FED8.taxon	discussion	Comments. The species H. subrastratus, H. myrtacivorus sp. n. and H. saltensis sp. n. share the proctiger strongly laterally excavated and tapered posteriorly. The species H. saltensis sp. n. differs from the other species of the group H. triangularis by anterior and posterior lobes of the paramere barely developed, giving a spearhead shape.	en	Cunha, Eduardo Victor De Paiva, Fernandes, José Antônio Marin (2022): New species of the Hypoxys triangularis group and the first contribution on the morphology of the internal genitalia of Hypoxys Amyot & Serville, 1843 (Hemiptera: Pentatomidae: Edessinae). Zootaxa 5168 (5): 501-520, DOI: 10.11646/zootaxa.5168.5.1
03C787C3FFD5FFF8C2ABFE22FBEFF9EC.taxon	etymology	Etymology. The specific epithet refers to the humeral angle curved backwards.	en	Cunha, Eduardo Victor De Paiva, Fernandes, José Antônio Marin (2022): New species of the Hypoxys triangularis group and the first contribution on the morphology of the internal genitalia of Hypoxys Amyot & Serville, 1843 (Hemiptera: Pentatomidae: Edessinae). Zootaxa 5168 (5): 501-520, DOI: 10.11646/zootaxa.5168.5.1
03C787C3FFD5FFF8C2ABFE22FBEFF9EC.taxon	materials_examined	Holotype. BOLIVIA. Cercado: 1 ♂ Rio Cristal Mavu 50 mi NE Cochabamba, VIII – II. 1944, L. Pena Collector (USNM). Paratypes (n = 5). PERU. Loreto: 1 ♂ Iquitos, Amazon River, 4. II. 1927, H. Bassler Coll. (USNM); Huanuco Department: 1 ♂ Puente Cinchavito, 25 km S Tíngo Maria, 3400 ’, 11 – 17. IV. 1987, J. E. Eger Coll. (JEE); BOLIVIA. Santa Cruz: 1 ♀ Prov. Ichilo 3 – 5 km SSE of Buena Vista, ± 400 m; 5 – 12. II. 2000, 17 ° 29 ’ 96 ” S 63 ° 39 ’ 13 ” W, W. B. Warner (JEE); BRAZIL. Acre: 1 ♂ Triunfo, X. 9.1977, D. Dominick Coll. (INPA).	en	Cunha, Eduardo Victor De Paiva, Fernandes, José Antônio Marin (2022): New species of the Hypoxys triangularis group and the first contribution on the morphology of the internal genitalia of Hypoxys Amyot & Serville, 1843 (Hemiptera: Pentatomidae: Edessinae). Zootaxa 5168 (5): 501-520, DOI: 10.11646/zootaxa.5168.5.1
03C787C3FFD5FFF8C2ABFE22FBEFF9EC.taxon	description	Measurements. Total length 13.1 – 14.5; total width 6.9 – 7.9; head length 2.9 – 3.2; head width 2.9 – 3.2; interocular distance: 1.6 – 1.7; length of antennal articles I: 0.6 – 0,9; II: 0.9 – 1.0; III: 1.2 – 1.5; IV: 2.5 – 2.9; V: 2.8 – 3.0; pronotum length 2.5 – 3.0; pronotum width 9.2 – 10.1; scutellum length 6.0 – 6.7; scutellum width 4.5 – 5,3.	en	Cunha, Eduardo Victor De Paiva, Fernandes, José Antônio Marin (2022): New species of the Hypoxys triangularis group and the first contribution on the morphology of the internal genitalia of Hypoxys Amyot & Serville, 1843 (Hemiptera: Pentatomidae: Edessinae). Zootaxa 5168 (5): 501-520, DOI: 10.11646/zootaxa.5168.5.1
03C787C3FFD5FFF8C2ABFE22FBEFF9EC.taxon	diagnosis	Diagnosis. Antennomeres with increasing lengths progressively (I <II <III <IV ≤ V). Anterolateral margin of pronotum unpunctured (Fig. 11 A). Humeral angles posteriorly curved, longer than eyes width (Fig. 11 A). Corium reddish brown (Fig. 11 A). Metasternal process with arms of anterior bifurcation receiving 2 / 3 of third rostral segment (Fig. 11 B). Pseudosutures of abdomen with adjacent dark green spot (Fig. 11 B). Male genitalia. Dorsal surface with spot restricted to coarse area of the pygophore (Fig. 4 A). Dorsal rim less excavated than other species (Fig. 4 A); lateral expansion slightly surpassing level of posterolateral angles (Fig. 4 C). Superior process of the genital cup elliptic, long and narrow (Fig. 4 B). Paramere head with distal margin strongly concave; anterior and posterior lobes subrectangular, apices rounded; anterior lobe large, reaching the superior process of the genital cup; posterior lobe short and laterally curved (Fig. 4 D). Proctiger moderately excavated laterally, excavation covered by short sparse setae; posterior face triangular (Fig. 4 B). Inner wall with slight swollen, narrow area contiguous to posterolateral angles (Fig. 8 E, black arrow). Ventral rim excavated, V-shaped; expansions little developed, rounded and curved dorsally (Fig. 4 C). Female genitalia (Fig. 4 E). Valvifers VIII distal and outer lateral margins almost straight and sloping; outer angle of the posterior margin forming a spine overlapping valvifers IX base (Fig. 9 E, red arrow); mediolateral crest swollen globose (Fig. 9 E, black arrow); mesial borders with deep U-shaped excavation. Valvulae IX exposed, with median keel, and somewhat sclerotized.	en	Cunha, Eduardo Victor De Paiva, Fernandes, José Antônio Marin (2022): New species of the Hypoxys triangularis group and the first contribution on the morphology of the internal genitalia of Hypoxys Amyot & Serville, 1843 (Hemiptera: Pentatomidae: Edessinae). Zootaxa 5168 (5): 501-520, DOI: 10.11646/zootaxa.5168.5.1
03C787C3FFD5FFF8C2ABFE22FBEFF9EC.taxon	distribution	Distribution (Fig. 12). PERU: Loreto, Huanuco Department; BOLIVIA: Cercado, Santa Cruz; Brazil: Acre.	en	Cunha, Eduardo Victor De Paiva, Fernandes, José Antônio Marin (2022): New species of the Hypoxys triangularis group and the first contribution on the morphology of the internal genitalia of Hypoxys Amyot & Serville, 1843 (Hemiptera: Pentatomidae: Edessinae). Zootaxa 5168 (5): 501-520, DOI: 10.11646/zootaxa.5168.5.1
03C787C3FFD5FFF8C2ABFE22FBEFF9EC.taxon	discussion	Comments. The female genitalia of H. curvatus sp. n., H. myrtacivorus sp. n., H. ocanensis sp. n., and H. triangularis show the outer angle forming a spine, but in H. curvatus sp. n. and H. ocanensis sp. n. this spine is dislocated to the middle of the valvifers IX base (in H. myrtacivorus sp. n. (Fig. 1 E) and H. triangularis the outer angle spine is lateral). Hypoxys curvatus sp. n. can be separated from H. ocanensis sp. n. by mesial borders of valvifers VIII deeply excavated and U-shaped (less excavated and V-shaped in H. ocanensis sp. n.). The spinose outer angle separates these four species from the others of H. triangularis group.	en	Cunha, Eduardo Victor De Paiva, Fernandes, José Antônio Marin (2022): New species of the Hypoxys triangularis group and the first contribution on the morphology of the internal genitalia of Hypoxys Amyot & Serville, 1843 (Hemiptera: Pentatomidae: Edessinae). Zootaxa 5168 (5): 501-520, DOI: 10.11646/zootaxa.5168.5.1
03C787C3FFD5FFFAC2ABF90DFA50FADA.taxon	etymology	Etymology. The specific epithet refers to the municipality of Ocaña (Colombia, Department Santander) where the type material was collected.	en	Cunha, Eduardo Victor De Paiva, Fernandes, José Antônio Marin (2022): New species of the Hypoxys triangularis group and the first contribution on the morphology of the internal genitalia of Hypoxys Amyot & Serville, 1843 (Hemiptera: Pentatomidae: Edessinae). Zootaxa 5168 (5): 501-520, DOI: 10.11646/zootaxa.5168.5.1
03C787C3FFD5FFFAC2ABF90DFA50FADA.taxon	materials_examined	Holotype. COLOMBIA. Department Santander: 1 ♀ Ocaña, PL VII. 2014 MNHN (EH) 25135 (MNHN).	en	Cunha, Eduardo Victor De Paiva, Fernandes, José Antônio Marin (2022): New species of the Hypoxys triangularis group and the first contribution on the morphology of the internal genitalia of Hypoxys Amyot & Serville, 1843 (Hemiptera: Pentatomidae: Edessinae). Zootaxa 5168 (5): 501-520, DOI: 10.11646/zootaxa.5168.5.1
03C787C3FFD5FFFAC2ABF90DFA50FADA.taxon	description	Measurements. Total length 15.6; total width 8.3; head length 2.1; head width 3.1; interocular distance: 1.8; length of antennomeres I: 0.9; II: 1.4; III: 1.0; IV: 2.6; V: 3.4; pronotum length 2.6; pronotum width 9.0; scutellum length 7.2; scutellum width 5.5.	en	Cunha, Eduardo Victor De Paiva, Fernandes, José Antônio Marin (2022): New species of the Hypoxys triangularis group and the first contribution on the morphology of the internal genitalia of Hypoxys Amyot & Serville, 1843 (Hemiptera: Pentatomidae: Edessinae). Zootaxa 5168 (5): 501-520, DOI: 10.11646/zootaxa.5168.5.1
03C787C3FFD5FFFAC2ABF90DFA50FADA.taxon	diagnosis	Diagnosis. Third antennomer smaller than the second (I <III <II <IV <V). Anterolateral margin of pronotum not punctured (Fig. 11 C). Posterior part of pronotum with black punctures (Fig. 11 C). Scutellum with black punctures anteriorly (Fig. 11 C). Metasternal process with arms of anterior bifurcation robust and receiving 1 / 2 of third rostral segment (Fig. 11 D). Connexivum flat (Fig. 11 C). Pseudosutures without adjacent dark green spot (Fig. 11 D). Female genitalia (Fig. 5). Valvifers VIII with distal margin sloping (red arrow) and slightly arched; outer angle acuminated and carinated, this carina also swollen along mediolateral crest (black arrow); mesial borders shallowly excavated. Valvulae IX not exposed. Male genitalia. Unknown.	en	Cunha, Eduardo Victor De Paiva, Fernandes, José Antônio Marin (2022): New species of the Hypoxys triangularis group and the first contribution on the morphology of the internal genitalia of Hypoxys Amyot & Serville, 1843 (Hemiptera: Pentatomidae: Edessinae). Zootaxa 5168 (5): 501-520, DOI: 10.11646/zootaxa.5168.5.1
03C787C3FFD5FFFAC2ABF90DFA50FADA.taxon	distribution	Distribution (Fig. 12). COLOMBIA: Department Santander.	en	Cunha, Eduardo Victor De Paiva, Fernandes, José Antônio Marin (2022): New species of the Hypoxys triangularis group and the first contribution on the morphology of the internal genitalia of Hypoxys Amyot & Serville, 1843 (Hemiptera: Pentatomidae: Edessinae). Zootaxa 5168 (5): 501-520, DOI: 10.11646/zootaxa.5168.5.1
03C787C3FFD5FFFAC2ABF90DFA50FADA.taxon	discussion	Comments. This species is the only one in H. triangularis group that has black punctures on pronotum. Hypoxys ocanensis sp. n. and H. curvatus sp. n. share the distal margin of each valvifer VIII with a distal spine. The H. ocanensis sp. n. can be separated by valvifers VIII carinate (in H. curvatus sp. n. valvifers VIII never carinate).	en	Cunha, Eduardo Victor De Paiva, Fernandes, José Antônio Marin (2022): New species of the Hypoxys triangularis group and the first contribution on the morphology of the internal genitalia of Hypoxys Amyot & Serville, 1843 (Hemiptera: Pentatomidae: Edessinae). Zootaxa 5168 (5): 501-520, DOI: 10.11646/zootaxa.5168.5.1
03C787C3FFD7FFFCC2ABFA24FB7BFAD2.taxon	description	(Fig. 9, 19 E – F, 22 — Nunes et al. (2020); 6 D, 7 A, 8 A, 9 A, 12)	en	Cunha, Eduardo Victor De Paiva, Fernandes, José Antônio Marin (2022): New species of the Hypoxys triangularis group and the first contribution on the morphology of the internal genitalia of Hypoxys Amyot & Serville, 1843 (Hemiptera: Pentatomidae: Edessinae). Zootaxa 5168 (5): 501-520, DOI: 10.11646/zootaxa.5168.5.1
03C787C3FFD7FFFCC2ABFA24FB7BFAD2.taxon	description	Edessa scabriventris Stål, 1859: 231; Stål, 1860: 28; Walker, 1868: 429; Stål, 1872: 56; Lethierry & Severin, 1893: 194 (catalog); Kirkaldy, 1909: 164 (catalog); Silva et al. 2018: 424 (checklist)	en	Cunha, Eduardo Victor De Paiva, Fernandes, José Antônio Marin (2022): New species of the Hypoxys triangularis group and the first contribution on the morphology of the internal genitalia of Hypoxys Amyot & Serville, 1843 (Hemiptera: Pentatomidae: Edessinae). Zootaxa 5168 (5): 501-520, DOI: 10.11646/zootaxa.5168.5.1
03C787C3FFD7FFFCC2ABFA24FB7BFAD2.taxon	diagnosis	Diagnosis. See Nunes et al. (2020). Characteristic added here: Male genitalia. Dorsal surface with spot extending about ¼ of the length of the pygophore (9 A — Nunes et al. (2020 )). Inner wall with well-swollen area attached to dorsal rim (Fig. 8 A, black arrow). Phallus (Fig. 7 A). Vesica tapering dorsally in lateral view. Phallotheca curved. Female genitalia. Pars intermedialis subcylindrical medially (Fig. 6 D). Capsula seminalis with three thick projections: one projection half the length of the others, the other two projections with half of the length of pars intermedialis (Fig. 6 D).	en	Cunha, Eduardo Victor De Paiva, Fernandes, José Antônio Marin (2022): New species of the Hypoxys triangularis group and the first contribution on the morphology of the internal genitalia of Hypoxys Amyot & Serville, 1843 (Hemiptera: Pentatomidae: Edessinae). Zootaxa 5168 (5): 501-520, DOI: 10.11646/zootaxa.5168.5.1
03C787C3FFD7FFFCC2ABFA24FB7BFAD2.taxon	distribution	Distribution (Fig. 12): FRENCH GUIANA; BRAZIL: Pernambuco, Bahia, Alagoas, Espírito Santo, Rio de Janeiro, Paraná, Santa Catarina, Rio Grande do Sul.	en	Cunha, Eduardo Victor De Paiva, Fernandes, José Antônio Marin (2022): New species of the Hypoxys triangularis group and the first contribution on the morphology of the internal genitalia of Hypoxys Amyot & Serville, 1843 (Hemiptera: Pentatomidae: Edessinae). Zootaxa 5168 (5): 501-520, DOI: 10.11646/zootaxa.5168.5.1
03C787C3FFD7FFFCC2ABFA24FB7BFAD2.taxon	discussion	Comments. The H. triangularis can be separated from the other species of the H. triangularis group by posterior stripe of pronotum reddish brown and body ventrally with large number of small yellow calli.	en	Cunha, Eduardo Victor De Paiva, Fernandes, José Antônio Marin (2022): New species of the Hypoxys triangularis group and the first contribution on the morphology of the internal genitalia of Hypoxys Amyot & Serville, 1843 (Hemiptera: Pentatomidae: Edessinae). Zootaxa 5168 (5): 501-520, DOI: 10.11646/zootaxa.5168.5.1
03C787C3FFD1FFFCC2ABFA2CFCC3F8C4.taxon	diagnosis	Diagnosis. See Nunes et al. (2020). Characteristic added here: Male genitalia. Dorsal surface of the pygophore with spot restricted to coarse area (Fig. 10 A — Nunes et al. (2020 )). Inner wall with swollen area globose close to superior process of genital cup (Fig. 8 B, black arrow). Female genitalia. Valvifers VIII with mediolateral crest slightly swollen (Fig. 9 B, black arrow).	en	Cunha, Eduardo Victor De Paiva, Fernandes, José Antônio Marin (2022): New species of the Hypoxys triangularis group and the first contribution on the morphology of the internal genitalia of Hypoxys Amyot & Serville, 1843 (Hemiptera: Pentatomidae: Edessinae). Zootaxa 5168 (5): 501-520, DOI: 10.11646/zootaxa.5168.5.1
03C787C3FFD1FFFCC2ABFA2CFCC3F8C4.taxon	distribution	Distribution (Fig. 12): PARAGUAY: Cordillera, Central, Paraguarí.	en	Cunha, Eduardo Victor De Paiva, Fernandes, José Antônio Marin (2022): New species of the Hypoxys triangularis group and the first contribution on the morphology of the internal genitalia of Hypoxys Amyot & Serville, 1843 (Hemiptera: Pentatomidae: Edessinae). Zootaxa 5168 (5): 501-520, DOI: 10.11646/zootaxa.5168.5.1
03C787C3FFD1FFFCC2ABFA2CFCC3F8C4.taxon	discussion	Comments. The swollen area on inner wall of the pygophore is the innermost one compared to other species of H. triangularis group. See H. striatus sp. n. comments.	en	Cunha, Eduardo Victor De Paiva, Fernandes, José Antônio Marin (2022): New species of the Hypoxys triangularis group and the first contribution on the morphology of the internal genitalia of Hypoxys Amyot & Serville, 1843 (Hemiptera: Pentatomidae: Edessinae). Zootaxa 5168 (5): 501-520, DOI: 10.11646/zootaxa.5168.5.1
03C787C3FFCDFFE1C2ABFF26FC28FE84.taxon	description	(Figs. 11, 19 I – J, 22 — Nunes et al. (2020); 6 E, 7 B, 8 C, 9 C, 12)	en	Cunha, Eduardo Victor De Paiva, Fernandes, José Antônio Marin (2022): New species of the Hypoxys triangularis group and the first contribution on the morphology of the internal genitalia of Hypoxys Amyot & Serville, 1843 (Hemiptera: Pentatomidae: Edessinae). Zootaxa 5168 (5): 501-520, DOI: 10.11646/zootaxa.5168.5.1
03C787C3FFCDFFE1C2ABFF26FC28FE84.taxon	diagnosis	Diagnosis. See Nunes et al. (2020). Characteristic added here: Male genitalia. Dorsal surface of the pygophore with spot restricted to coarse area (Fig. 11 A — Nunes et al. (2020 )). Phallus (Fig. 7 B). Vesica widening ventrally in lateral view. Phallotheca curved. Female genitalia. Valvifers VIII with mediolateral crest well swollen (Fig. 9 C, black arrow). Pars intermedialis slightly swollen medially (Fig. 6 E). Capsula seminalis with three thick and long projections: two slightly surpassing anterior annular crest and the other almost reaching base of the pars intermedialis (Fig. 6 E).	en	Cunha, Eduardo Victor De Paiva, Fernandes, José Antônio Marin (2022): New species of the Hypoxys triangularis group and the first contribution on the morphology of the internal genitalia of Hypoxys Amyot & Serville, 1843 (Hemiptera: Pentatomidae: Edessinae). Zootaxa 5168 (5): 501-520, DOI: 10.11646/zootaxa.5168.5.1
03C787C3FFCDFFE1C2ABFF26FC28FE84.taxon	distribution	Distribution (Fig. 12): BRAZIL: Pará, Maranhão, Goiás, Minas Gerais, São Paulo, Santa Catarina, Rio Grande do Sul; ARGENTINA: Candelaria; BOLIVIA: La Paz; PARAGUAY: Paraguarí.	en	Cunha, Eduardo Victor De Paiva, Fernandes, José Antônio Marin (2022): New species of the Hypoxys triangularis group and the first contribution on the morphology of the internal genitalia of Hypoxys Amyot & Serville, 1843 (Hemiptera: Pentatomidae: Edessinae). Zootaxa 5168 (5): 501-520, DOI: 10.11646/zootaxa.5168.5.1
03C787C3FFCDFFE1C2ABFF26FC28FE84.taxon	discussion	Comments. This species can be separated by dorsal rim of pygophore with rectangular median notch and inner wall with the larger swollen area between species of H. triangularis group.	en	Cunha, Eduardo Victor De Paiva, Fernandes, José Antônio Marin (2022): New species of the Hypoxys triangularis group and the first contribution on the morphology of the internal genitalia of Hypoxys Amyot & Serville, 1843 (Hemiptera: Pentatomidae: Edessinae). Zootaxa 5168 (5): 501-520, DOI: 10.11646/zootaxa.5168.5.1
