identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03C787C3FFDCFFF3C2ABFC14FA57FDB3.text	03C787C3FFDCFFF3C2ABFC14FA57FDB3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypoxys triangularis (Dallas 1851) Nunes et al. 2020	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Hypoxys triangularis Group </p>
            <p>(Figs. 1–12)</p>
            <p> Diagnosis. Body uniformly green with corium reddish-brown (Fig. 10) (green with yellow spots in  H. triangularis — see Nunes et al. 2020: Fig. 19E–F), and a median large yellow spot on the ventral surface. Species of the  H. triangularis group also lack any transversal black stripe on pronotum and ventral surface, like in the other species groups of  Hypoxys , except  H. triangularis (see Nunes et al. 2020: Fig. 19E). Trichobothria with base yellowish (Fig. 10F). Male genitalia. Paramere (Pa) usually ax-shaped (Fig. 1B). Inner wall of pygophore laterally swollen (Figs. 3B–C, 8 —black arrow). Ventral rim (Vr) with expansions slightly developed and rounded (Fig. 4C). Female genitalia (Fig. 9). Valvifers VIII (Vf8) with mediolateral crest swollen (black arrow); outer lateral angle of the posterior margin (red arrow) rounded or ending in an acute projection. </p>
            <p>Redescription. Head. Mandibular plates without spot (Figs. 10, 11). Antennae yellow with setae more densely distributed on antennomeres III, IV and V.</p>
            <p> Thorax.Anterolateral margin of pronotum usually not punctured, except  H. subrastratus (see Nunes et al. 2020: Fig. 19I) and  H. saltensis sp. n. (Fig. 10E). Apex of humeral angle with margin brown (Figs. 10, 11). Pronotum and scutellum usually with brown punctures (Fig. 10A). Anterior margin of scutellum without yellow stripe, like the one found in some species of  H. quadridens and  H. oxyacanthus groups. Costal margin of corium with a black spot reaching approximately 1/3 of corial extension (Figs. 10, 11). Propleura with concolorous punctures concentrated on posterior half. Evaporatorium slightly whitish (Figs. 10F, 11A). Metasternal process with arms of anterior bifurcation receiving fourth and most of third rostral segment (Fig. 10A). </p>
            <p> Abdomen. Dorsal surface with brown spots. Connexivum with two barely excavated areas (Fig. 10C) and usually not associated with dark spot (except  H. triangularis ). Pseudosutures with or without adjacent dark green spot (Figs. 10B, 11D). </p>
            <p>Male. Pygophore dorsally with brown to reddish brown spot occupying about 1/2 of the length of the pygophore; distal part of dorsal surface coarse (Fig. 1A). Superior process (Pc) of genital cup brown and positioned near to dorsal rim (Fig. 2C). Paramere directed posterolaterally (Fig. 9); base with a swollen and rounded protuberance (Fig. 9, blue arrow). Proctiger (Proc) with posterior face triangular, elliptical or pentagonal, and bent ventrally (Fig. 8).</p>
            <p>Female. Valvifers VIII and Valvifers IX (Vf9) convex (Fig. 2E). Laterotergites VIII (La8) with margin black (Fig. 9). Laterotergites IX (La9) with outer lateral margin sinuous, well projected beyond sclerite uniting laterotergites VIII; almost reaching level of laterotergites VIII (Fig. 4E). Capsula seminalis (Cs) with two thick projections of equal lengths (Fig. 6D); projections of capsula seminalis of the same width as pars intermedialis (Pi) (Fig. 6F).</p>
            <p> Comments. Five new species are described into  H. triangularis group. Therefore, the new composition of the group is:  Hypoxys triangularis ;  H. capito ;  H. subrastratus ;  H. myrtacivorus sp. n. ;  H. saltensis sp. n. ;  H. striatus sp. n. ;  H. curvatus sp. n. and  H. ocanensis sp. n. The  H. triangularis group has species associated with cultivated plants of the family  Myrtaceae Juss. (Angiosperma;  Myrtales ):  H. triangularis associated the  Eugenia uniflora L. (brazilian-cherry) and  Psidium guajava (guava) (Lima et al. 2010);  H. subrastratus associated the  Eugenia uniflora (brazilian-cherry) (data base Inaturalist; Fernandes pers. Comm.);  H. myrtacivorus sp. n. associated the  Myrciaria dubia (Kunth) McVaugh (camu-camu) (Panduro et al. 2001),  Eugenia malaccensis L. (red jambo) and  Eugenia uniflora L. (see comments of  H. myrtacivorus sp. n. ). Parameres of this species group are also almost straight in dorsal view and never strongly curved laterally, characteristic shared with  Hypoxys balteatus group (see in Mendonça et al. 2021). The internal genitalia of some Edessinae genera were described, but never compared or discussed in a comprehensive way (  Pantochlora vivida Stål, 1870 , ilustrated— Kumar (1969);  Brachystethus Laporte, 1832 — Grazia &amp; Barcellos (2003);  Paraedessa Silva &amp; Fernandes, 2013 — Silva et al. (2013);  Ascra Say, 1832 — Santos et al. (2015);  Plagaedessa Almeida &amp; Fernandes, 2018 — Almeida et al. (2018);  Pygoda Amyot &amp; Serville, 1843 — Fernandes et al. (2018);  Graziaedessa anastrephae Eger, 2021 , only the phallus described— Eger (2021);  Edessa ovina group— Fernandes &amp; Silva (2021)). The internal genitalia of species of  Hypoxys have not been describe yet. This is the first attempt to compare and use internal genitalia to characterize the groups of species. Studying the internal genitalia of species belonging to the  Hypoxys , we observed some characteristics that support  Hypoxys and its groups of species. The female internal genitalia present the following characteristics for  Hypoxys : distal part of vesicular area of the ductus receptaculi sclerotized (Fig. 6E, yellow arrow); base of the capsula seminalis spherical and sclerotized (Fig. 6D, green arrow); capsula seminalis dilated with thick projections well-developed (Fig. 6B); posterior annular crest more developed than anterior annular crest (Fig. 6A). The phallus of  Hypoxys species have the phallotheca cylindrical and curved dorsoventrally (Fig. 7). The differences observed between the species belonging to different groups of  Hypoxys were: the  H. quadridens (Fig. 6A) and  H. oxyacanthus (Fig. 6B) groups share the projections of capsula seminalis wider and more dilated than the pars intermedialis. In  H. triangularis (Fig. 6D, E, F) and  H. balteatus (Fig. 6C) groups the projections of capsula seminalis are almost as wide as pars intermedialis. Groups  H. quadridens and  H. oxyacanthus share median portion of pars intermedialis thickened and spherical, while in  H. triangularis group this structure is spherical or cylindrical, and in  H. balteatus group the same structure is cylindrical. The  H. quadridens group shows the base of capsula seminalis conical (Fig. 6A, red arrow).  Hypoxys oxyacanthus group has the globose base of capsula seminalis as wide as the segment X. A comparative study of the internal genitalia in all taxa of Edessinae can help to understand the evolution of the genitalia in the subfamily and add diagnostic characteristics to the genera. </p>
            <p>Distribution (Fig. 12). BRAZIL; FRENCH GUIANA; VENEZUELA; PARAGUAY; SURINAME; GUIANA; BOLIVIA; PERU; COLOMBIA, and ARGENTINA.</p>
            <p> Key to species of the  Hypoxys triangularis group </p>
            <p> 1. Pronotum with posterior smoky brown stripe; ventral surface of the body with small yellow calli (See Nunes et al. 2020: Fig. 19E–F)..................................................................................  H. triangularis</p>
            <p>1’. Pronotum without dark stripe; ventral surface of the body without yellow calli (Figs. 10A, 11A)...................... 2</p>
            <p>2. Male............................................................................................... 3</p>
            <p>2’. Female..............................................................................................8</p>
            <p>3. Proctiger strongly (Figs. 1B, 3B, See Nunes et al. 2020: Fig. 11A–B) or moderately excavated laterally (Fig. 4D).........4</p>
            <p>3’. Proctiger shallowly excavated laterally (Figs. 2D, 8B)........................................................ 7</p>
            <p> 4. Dorsal rim of pygophore medially notched (See Nunes et al. 2020 – Fig. 11A). Superior process of genital cup subtriangular, almost the same size on swollen area on inner wall of pygophore (Fig. 8C)............................  H. subrastratus</p>
            <p>4’. Dorsal rim entire. Superior process of genital cup elliptic, bigger than swollen area on inner wall of pygophore (Figs. 1B, 3B, 4B)................................................................................................ 5</p>
            <p> 5. Superior process of genital cup coarse in posterior view (Fig. 3B). Paramere spearhead-shaped (Fig. 3D)..  H. saltensis sp. n.</p>
            <p>5’. Superior process of genital cup smooth in posterior view (Figs. 1B, 4B). Paramere ax-shaped (Fig. 1D, 4D)............. 6</p>
            <p> 6. Swollen area on inner wall globose, developed, close to posterolateral angles (Fig. 8D). Ventral rim with bottom of excavation convex (Fig. 1C).....................................................................  H. myrtacivorus sp. n.</p>
            <p> 6”. Swollen area on inner wall narrow, slightly developed, contiguous to posterolateral angles (Fig. 8E). Ventral rim with bottom of excavation flat (Fig. 4C)...............................................................  H. curvatus sp. n.</p>
            <p> 7. Superior process of genital cup subtriangular, small, swollen in posterior view (Fig. 2B). Expansions of ventral rim hardly visible (Fig. 2C)..........................................................................  H. striatus sp. n.</p>
            <p> 7’. Superior process of genital cup elliptic, narrow, smooth in posterior view (Fig. 8B). Expansions of ventral rim visible (See Nunes et al. 2020: Fig. 10C)......................................................................  H. capito</p>
            <p>8. Valvifers VIII with outer lateral angle rounded (Figs. 2E, See Nunes et al. 2020: Figs. 10D, 11D)..................... 9</p>
            <p>8’. Valvifers VIII with outer lateral angle acute (Figs. 1E, 4E, 5A)................................................ 11</p>
            <p> 9. Valvifers VIII with excavation in “V” (See Nunes et al. 2020: Fig. 10D).................................. ...  H. capito</p>
            <p>9’. Valvifers VIII with excavation in “U” (Figs. 2E, See Nunes et al. 2020: Fig. 11D)................................. 10</p>
            <p> 10. Valvifers VIII with mediolateral crest slightly swollen (Fig. 9F); median excavation narrow, clearly narrower than base of valvifers IX (Fig. 2E)....................................................................  H. striatus sp. n.</p>
            <p> 10’. Valvifers VIII with mediolateral crest swollen (Fig. 9C); median excavation wide, almost as wide as base of valvifers IX (See Nunes et al. 2020: Fig. 11D)................................................................  H. subrastratus</p>
            <p> 11. Mediolateral crest of valvifers VIII carinated (Fig. 5B).........................................  H. ocanensis sp. n.</p>
            <p>11’. Mediolateral crest of valvifers VIII not carinated (Fig. 8D, 8E)................................................ 12</p>
            <p> 12. Valvifers VIII with mesial borders forming a spiniform median projection (Fig. 1E)...............  H. myrtacivorus sp. n.</p>
            <p> 12’. Valvifers VIII with mesial borders not forming a spiniform projection. (Fig. 4E).....................  H. curvatus sp. n.</p>
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	https://treatment.plazi.org/id/03C787C3FFDCFFF3C2ABFC14FA57FDB3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cunha, Eduardo Victor De Paiva;Fernandes, José Antônio Marin	Cunha, Eduardo Victor De Paiva, Fernandes, José Antônio Marin (2022): New species of the Hypoxys triangularis group and the first contribution on the morphology of the internal genitalia of Hypoxys Amyot & Serville, 1843 (Hemiptera: Pentatomidae: Edessinae). Zootaxa 5168 (5): 501-520, DOI: 10.11646/zootaxa.5168.5.1
03C787C3FFDEFFF5C2ABFDC2FC77FD1C.text	03C787C3FFDEFFF5C2ABFDC2FC77FD1C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypoxys myrtacivorus Cunha & Fernandes 2022	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Hypoxys myrtacivorus sp. n.</p>
            <p>(Figures: 1, 6F, 7C, 8D, 9D, 10A–B, 12)</p>
            <p> Etymology. The specific epithet refers to the feeding plants of this species (family  Myrtaceae ). </p>
            <p>  Holotype. PERU. Loreto: 1 ♂ Iquitos, 14.V.1992, G. Couturier Col./ Host Plant  Myrciaria dubia (Myrtaceae) / Isla Mayuy INIAA/IVITA MNHN(EH) 25126 (MNHN). </p>
            <p> Paratypes (n=23). No data, 1 ♀ 3093, 72304 (MZUSP); PERU. Loreto: 2 ♂ and 3 ♀ Carretera Iquitos–Nauta, Estacion El Dorado INIAA Unamurú, IX–2006, J. Vasquez B., sobre  Myrciaria dubia (Myrtaceae) “camu camu” MNHN (EH) 25130-25134 (MNHN); same data 1 ♂ and 1 ♀ (UFPA); 1 ♂ Cocha Supay, Jenaro Herrero, 26.X.1988, 4° 55’S 73° 40’W, G. Couturier &amp; H. Inga col., Plante–hote:  Myrciaria dubia MNHN (EH) 25127 (MNHN); 1 ♂ Iquitos, 27. V.1991, G. Couturier Col., Host Plant  Eugenia malaccensis L. (  Myrtaceae ) MNHN (EH) 25128 (MNHN); BRAZIL. Pará: 1 ♂ and 1 ♀ Tucuruí, Rio Tocantins, Acamp. Bagagem, 03.VII.1984, coleta Noturna (MPEG); 1 ♂ Belém, Bairro Umarizal, XI. 2014 (UFPA); 1 ♀ Belém, ICB— UFPA Campus Guamá, 17.XI.2021, E. V. P. Cunha col., sobre  Eugenia uniflora L., Obs.: indivíduo se alimentando no ramo (UFPA); 1 ♀ Belém, UFPA —Campus Guamá, 09.XII.2016, E. V. P. Cunha col.; Host Plant  Eugenia malaccensis L. (  Myrtaceae ) (UFPA); 1 ♀ Santarém, E. le Moult. (RMNH); Amazonas: 1 ♂ Igarapé Tacana, Solimões, II.1962, Cândido &amp; Dyrce (MNRJ); 1 ♀ Benjamin Constant, Rio Javari, II.1942, Pko. (MNRJ); Rondônia: 2 ♀ 62 km Ariquemes, nr Fazenda Rancho Grande, 8– 20.XI.1994, J. E. Eger &amp; L. B. &amp; C. W. O’Brien (JEE); SURINAME. Sipaliwini: 1 ♀ Kayser Gebergte, Exped. H. A. Beatty, 25.XI–14.XII.1960, Sergiman Moentari leg. (FMNH); GUIANA. Georgetown: 1 ♀ Belair, Y. VII.1931, R. T. Potts (USNM); VENEZUELA. Monagas: 1 ♀ Josepín, 50m, 27.IX.1955, F. Fernandez y C. J. Rosales (MBUC); COLOMBIA. Meta: 1 ♀ Peralonso, Caño Pachiaquiari, 4.IX.1965, J. A. Ramos Collector (USNM). </p>
            <p>Measurements. Total length 11.1–15.9; total width 6.2–8.5; head length 1.9–2.3; head width 2.9–3.4; interocular distance: 1.5–1.8; length of antennomeres I: 0.7–1.0; II: 0.9–1.5; III: 1.3–1.6; IV: 2,5–3,1; V: 2.5–3.4; pronotum length 2.4–3.8; pronotum width 7.7–9.3; scutellum length 5.9–7.8; scutellum width 4.4–5.4.</p>
            <p> Diagnosis. Antennomeres with increasing lengths progressively (I&lt;II&lt;III&lt;IV&lt;V). Anterolateral margin of pronotum unpunctured (Fig. 10A). Metasternal process with arms of anterior bifurcation robust and receiving all third rostral segment (Fig. 10B). Pseudosutures of abdomen with adjacent dark green spot (Fig. 10B). Trichobothria with yellow base (Fig. 10B). Male genitalia. Dorsal surface with dark spot occupying 1/3 of the length of the pygophore (Fig. 1A). Superior process of the genital cup long and narrow (Fig. 1B). Paramere with dorsal margin strongly concave; anterior lobe large, subrectangular, reaching dorsal rim; posterior lobe short, rectangular laterally directed (Figs. 1B, 1D). Proctiger strongly excavated laterally, excavation and posterior face covered by long and sparse setae; dorsal face tapering posteriorly; posterior face triangular, small, below the level of dorsal face and ventrally bent, lateral areas slightly swollen (Fig. 1D, 8D). Inner wall swollen area globose and close to posterolateral angles (Fig. 8D, black arrow). Ventral rim strongly excavated; bottom of median excavation convex (Fig. 1C).  Phallus (Fig. 7C). Vesica (vs) half-moon-shaped in lateral view. Phallotheca (ph) slightly curved. Female genitalia (Figs. 1E). Valvifers VIII with rounded excavation; booth distal angles dentiform (Fig. 9, red arrow); inner angles reaching valvifers IX; mediolateral crest swollen along outer lateral margin (Fig. 9D, black arrow); mesial borders contiguous forming a spiniform projection (Fig. 9D). Valvulae IX (Va9) exposed and sclerotized. Pars intermedialis (pi) spherical medially (Fig. 6F). Capsula seminalis with three thick projections almost as long as capsula seminalis (Fig. 6F). </p>
            <p> Comments. This species was collected only on  Myrtaceae plants,  E. malaccensis (red jambo),  E. uniflora (brazilian-cherry) and  M. dubia (camu-camu). This species was observed feeding on branches from  E. uniflora L., but in  M. dubia it was observed feeding on fruits.  Hypoxys myrtacivorus sp. n. was considered pest of camu-camu,  M. dubia , in Peru, causing a transparent spot with a central red dot (Panduro et al. 2001).  Hypoxys myrtacivorus sp. n. had nymphal stages and biological aspects described by Iannacone et al. (2007), identified as  Edessa aff. aulacosterna Stål, 1872 , but author mentioned that the species probably was new to science. Nevertheless,  H. myrtacivorus sp. n. differs from  Edessa aulacosterna by several aspects, but two are very conspicuous: humeral angles acute and slightly developed with apex brown to black (not developed, rounded and concolorous in  E. aulacosterna ); anterior arms of metasternal process swollen and tapering anteriorly (laterally compressed in  E. aulacosterna ). The shape of the paramere of this species resembles  H. curvatus sp. n. , but differ because the apex of the posterior lobe is acuminate in  H. myrtacivorus sp. n. (in  H. curvatus sp. n. the apices of the anterior and posterior lobes are rounded). </p>
            <p>Distribution (Fig. 12). PERU: Loreto; BRAZIL: Pará, Amazonas, Rondônia; SURINAME: Sipaliwini; GUIANA: Georgetown; VENEZUELA: Monagas; COLOMBIA: Meta.</p>
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	https://treatment.plazi.org/id/03C787C3FFDEFFF5C2ABFDC2FC77FD1C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cunha, Eduardo Victor De Paiva;Fernandes, José Antônio Marin	Cunha, Eduardo Victor De Paiva, Fernandes, José Antônio Marin (2022): New species of the Hypoxys triangularis group and the first contribution on the morphology of the internal genitalia of Hypoxys Amyot & Serville, 1843 (Hemiptera: Pentatomidae: Edessinae). Zootaxa 5168 (5): 501-520, DOI: 10.11646/zootaxa.5168.5.1
03C787C3FFD8FFF5C2ABFD7BFAE2F954.text	03C787C3FFD8FFF5C2ABFD7BFAE2F954.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypoxys striatus Cunha & Fernandes 2022	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Hypoxys striatus sp. n.</p>
            <p>(Figures: 2, 8F, 9F, 10C–D, 12)</p>
            <p>Etymology. The specific epithet refers to the paramere posterior face striated.</p>
            <p>  Holotype. BRAZIL.  Pará : 1 ♂ Óbidos, VIII.1978, A. C. Domingos Coll. (MNRJ). </p>
            <p>  Paratypes (n=3). BRAZIL. Pará: 1 ♀  Óbidos , XII.1963, Dirings Coll. (MZUSP)  ; 1 ♀ e  1 ♂ Santarém, VI  .1919, S. M.  Klages Coll. (CMNH)  . </p>
            <p>Measurements. Total length 14.3–14.8; total width 7.7–8.0; head length 1.4–1.5; head width 3.2–3,4; interocular distance: 1.6–1.8; length of antenomeres I: 0.9–1.0; II: 1.0–1.1; III: 1.5; IV: 3.1–3.2; V: 3.6; pronotum length 2.7– 3.0; pronotum width 8.9–9.2; scutellum length 6.5–6.7; scutellum width 5.0–5.3.</p>
            <p>Diagnose. Antennomeres with increasing lengths progressively (I&lt;II&lt;III&lt;IV&lt;V). Anterolateral margin of pronotum not punctured (Fig. 10C). Corium reddish brown (Fig. 10C). Metasternal process with arms of anterior bifurcation receiving 2/3 of third rostral segment (Fig. 10D). Pseudosutures of abdomen with adjacent dark green spot (Fig. 10D). Male genitalia. Dorsal surface with spot restricted to coarse area of the pygophore (Fig. 2A). Dorsal rim sinuous (Fig. 2A); lateral expansions slightly globose and almost level with posterolateral angles (Fig. 2C). Superior process of the genital cup subtriangular, small and swollen (Fig. 2B). Paramere with head slightly concave; anterior and posterior lobe subtriangular; anterior lobe large with external face striated and reaching superior process of the genital cup; posterior lobe smaller than the anterior lobe and laterally directed (Fig. 2D). Proctiger shallowly excavated laterally, excavation covered by long and dense setae; posterior face elliptical (Fig. 2B). Inner wall with small swollen area contiguous to posterolateral angles (Fig. 8F, black arrow). Ventral rim in widely opened V-shaped with expansions hardly visible (Fig. 2C). Female genitalia (Fig. 2E). Valvifers VIII with distal margin slightly arched; outer and inner angles of the posterior margin rounded; mediolateral crest slightly swollen (Fig. 9F, black arrow); mesial borders with excavation narrow, deep and U-shaped. Valvulae IX clearly exposed.</p>
            <p>Distribution (Fig. 12). BRAZIL: Pará.</p>
            <p> Comments. The superior process of the genital cup of  H. triangularis and  H. striatus sp. n. are the smallest processes in the group, but in  H. triangularis it is rounded and concave and in  H. striatus sp. n. it is subtriangular and swollen. This species is the only one with expansions of the ventral rim hardly visible. The mesial borders of the valvifers VIII of  H. curvatus sp. n. ,  H. capito and  H. subrastratus have the excavation forming a “U”. However,  H. striatus sp. n. can be separated from the other two species by the more deeply excavated mesial borders. </p>
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	https://treatment.plazi.org/id/03C787C3FFD8FFF5C2ABFD7BFAE2F954	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cunha, Eduardo Victor De Paiva;Fernandes, José Antônio Marin	Cunha, Eduardo Victor De Paiva, Fernandes, José Antônio Marin (2022): New species of the Hypoxys triangularis group and the first contribution on the morphology of the internal genitalia of Hypoxys Amyot & Serville, 1843 (Hemiptera: Pentatomidae: Edessinae). Zootaxa 5168 (5): 501-520, DOI: 10.11646/zootaxa.5168.5.1
03C787C3FFD8FFF8C2ABF8AEFF14FED8.text	03C787C3FFD8FFF8C2ABF8AEFF14FED8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypoxys saltensis Cunha & Fernandes 2022	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Hypoxys saltensis sp. n.</p>
            <p>(Figures: 3, 10E –F, 12)</p>
            <p>Etymology. The specific epithet refers to Province of Salta (Argentina) where the specimens collected.</p>
            <p>  Holotype. ARGENTINA. Province of Salta: 1 ♂  Quebrada of San Lorenzo , [Quebrada of Catayete, Province of Salta?], 10.II.1953, R. Foerster Coll. (MMB). </p>
            <p>Measurements. Total length 13.6; total width 6.6; head length 1.4; head width 3.0; interocular distance: 1.6; length of antennomeres I: 0.9; II: 0.7; III: 1.4; IV: 2.5; V: lost; pronotum length 2.6; pronotum width 7.7; scutellum length 5.7; scutellum width 4.4.</p>
            <p>Diagnosis. Second antennomer smaller than the others (II&lt;I&lt;III&lt;IV).Anterolateral margin of pronotum punctured (Fig. 10E). Scutellum with punctures of the same size (Fig. 10E). Hemelytrom membrane transparent (Fig. 10E). Metasternal process with arms of anterior bifurcation receiving 2/3 of third rostral segment (Fig. 10F). Pseudosutures of abdomen without adjacent dark green spot (Fig. 10F). Male genitalia. Dorsal surface with spot extending about ½ of the length of the pygophore (Fig. 3A). Dorsal rim with lateral expansions level with posterolateral angles and rounded (Figs. 3A, 3C). Superior process of the genital cup elliptic, face rough in posterior view (Fig. 3B). Paramere spearhead-shaped, lobes slightly developed, apex truncated (Figs. 3B, 3D). Proctiger strongly excavated laterally, excavation covered by long and dense setae; dorsal face tapering posteriorly; posterior face subpentagonal, basal third constricted (Figs. 3B, 3D). Inner wall with swollen area large and close to posterolateral angles (Fig. 3D, black arrow). Ventral rim strongly excavated, U-shaped; expansions ending laterally in a rounded projection (Fig. 4C). Female genitalia. Unknown.</p>
            <p>Distribution (Fig. 12). ARGENTINA: Salta.</p>
            <p> Comments. The species  H. subrastratus ,  H. myrtacivorus sp. n. and  H. saltensis sp. n. share the proctiger strongly laterally excavated and tapered posteriorly. The species  H. saltensis sp. n. differs from the other species of the group  H. triangularis by anterior and posterior lobes of the paramere barely developed, giving a spearhead shape. </p>
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	https://treatment.plazi.org/id/03C787C3FFD8FFF8C2ABF8AEFF14FED8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cunha, Eduardo Victor De Paiva;Fernandes, José Antônio Marin	Cunha, Eduardo Victor De Paiva, Fernandes, José Antônio Marin (2022): New species of the Hypoxys triangularis group and the first contribution on the morphology of the internal genitalia of Hypoxys Amyot & Serville, 1843 (Hemiptera: Pentatomidae: Edessinae). Zootaxa 5168 (5): 501-520, DOI: 10.11646/zootaxa.5168.5.1
03C787C3FFD5FFF8C2ABFE22FBEFF9EC.text	03C787C3FFD5FFF8C2ABFE22FBEFF9EC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypoxys curvatus Cunha & Fernandes 2022	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Hypoxys curvatus sp. n.</p>
            <p>(Figures: 4, 8E, 9E, 11A–B, 12)</p>
            <p>Etymology. The specific epithet refers to the humeral angle curved backwards.</p>
            <p>  Holotype. BOLIVIA. Cercado: 1 ♂  Rio Cristal Mavu 50 mi NE Cochabamba, VIII– II.1944, L. Pena Collector (USNM). </p>
            <p>  Paratypes (n=5). PERU. Loreto: 1 ♂ Iquitos,  Amazon River , 4.II.1927, H. Bassler Coll. (USNM)  ;   Huanuco  Department :   1 ♂  Puente Cinchavito , 25 km S Tíngo Maria, 3400’, 11–17. IV  .1987, J. E. Eger Coll. (JEE) ;   BOLIVIA. Santa Cruz: 1 ♀  Prov. Ichilo 3–5 km SSE of Buena Vista, ± 400m; 5–12.II.2000, 17°29’96”S 63°39’13”W, W. B. Warner (JEE)  ;   BRAZIL. Acre: 1 ♂  Triunfo , X. 9.1977, D. Dominick Coll. (INPA)  . </p>
            <p>Measurements. Total length 13.1–14.5; total width 6.9–7.9; head length 2.9–3.2; head width 2.9–3.2; interocular distance: 1.6–1.7; length of antennal articles I: 0.6–0,9; II: 0.9–1.0; III: 1.2–1.5; IV: 2.5–2.9; V: 2.8–3.0; pronotum length 2.5–3.0; pronotum width 9.2–10.1; scutellum length 6.0–6.7; scutellum width 4.5–5,3.</p>
            <p>Diagnosis. Antennomeres with increasing lengths progressively (I&lt;II&lt;III&lt;IV≤V). Anterolateral margin of pronotum unpunctured (Fig. 11A). Humeral angles posteriorly curved, longer than eyes width (Fig. 11A). Corium reddish brown (Fig. 11A). Metasternal process with arms of anterior bifurcation receiving 2/3 of third rostral segment (Fig. 11B). Pseudosutures of abdomen with adjacent dark green spot (Fig. 11B). Male genitalia. Dorsal surface with spot restricted to coarse area of the pygophore (Fig. 4A). Dorsal rim less excavated than other species (Fig.4A); lateral expansion slightly surpassing level of posterolateral angles (Fig. 4C). Superior process of the genital cup elliptic, long and narrow (Fig. 4B). Paramere head with distal margin strongly concave; anterior and posterior lobes subrectangular, apices rounded; anterior lobe large, reaching the superior process of the genital cup; posterior lobe short and laterally curved (Fig. 4D). Proctiger moderately excavated laterally, excavation covered by short sparse setae; posterior face triangular (Fig. 4B). Inner wall with slight swollen, narrow area contiguous to posterolateral angles (Fig. 8E, black arrow). Ventral rim excavated, V-shaped; expansions little developed, rounded and curved dorsally (Fig. 4C). Female genitalia (Fig. 4E). Valvifers VIII distal and outer lateral margins almost straight and sloping; outer angle of the posterior margin forming a spine overlapping valvifers IX base (Fig. 9E, red arrow); mediolateral crest swollen globose (Fig. 9E, black arrow); mesial borders with deep U-shaped excavation. Valvulae IX exposed, with median keel, and somewhat sclerotized.</p>
            <p>Distribution (Fig. 12). PERU: Loreto, Huanuco Department; BOLIVIA: Cercado, Santa Cruz; Brazil: Acre.</p>
            <p> Comments. The female genitalia of  H. curvatus sp. n. ,  H. myrtacivorus sp. n. ,  H. ocanensis sp. n. , and  H. triangularis show the outer angle forming a spine, but in  H. curvatus sp. n. and  H. ocanensis sp. n. this spine is dislocated to the middle of the valvifers IX base (in  H. myrtacivorus sp. n. (Fig. 1E) and  H. triangularis the outer angle spine is lateral).  Hypoxys curvatus sp. n. can be separated from  H. ocanensis sp. n. by mesial borders of valvifers VIII deeply excavated and U-shaped (less excavated and V-shaped in  H. ocanensis sp. n. ). The spinose outer angle separates these four species from the others of  H. triangularis group. </p>
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	https://treatment.plazi.org/id/03C787C3FFD5FFF8C2ABFE22FBEFF9EC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cunha, Eduardo Victor De Paiva;Fernandes, José Antônio Marin	Cunha, Eduardo Victor De Paiva, Fernandes, José Antônio Marin (2022): New species of the Hypoxys triangularis group and the first contribution on the morphology of the internal genitalia of Hypoxys Amyot & Serville, 1843 (Hemiptera: Pentatomidae: Edessinae). Zootaxa 5168 (5): 501-520, DOI: 10.11646/zootaxa.5168.5.1
03C787C3FFD5FFFAC2ABF90DFA50FADA.text	03C787C3FFD5FFFAC2ABF90DFA50FADA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypoxys ocanensis Cunha & Fernandes 2022	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Hypoxys ocanensis sp. n.</p>
            <p>(Figures: 5, 11C–D, 12)</p>
            <p>Etymology. The specific epithet refers to the municipality of Ocaña (Colombia, Department Santander) where the type material was collected.</p>
            <p>  Holotype. COLOMBIA. Department Santander: 1 ♀  Ocaña , PL VII.2014 MNHN (EH) 25135 (MNHN). </p>
            <p>Measurements. Total length 15.6; total width 8.3; head length 2.1; head width 3.1; interocular distance: 1.8; length of antennomeres I: 0.9; II: 1.4; III: 1.0; IV: 2.6; V: 3.4; pronotum length 2.6; pronotum width 9.0; scutellum length 7.2; scutellum width 5.5.</p>
            <p>Diagnosis. Third antennomer smaller than the second (I&lt;III&lt;II&lt;IV&lt;V). Anterolateral margin of pronotum not punctured (Fig. 11C). Posterior part of pronotum with black punctures (Fig. 11C). Scutellum with black punctures anteriorly (Fig. 11C). Metasternal process with arms of anterior bifurcation robust and receiving 1/2 of third rostral segment (Fig. 11D). Connexivum flat (Fig. 11C). Pseudosutures without adjacent dark green spot (Fig. 11D). Female genitalia (Fig. 5). Valvifers VIII with distal margin sloping (red arrow) and slightly arched; outer angle acuminated and carinated, this carina also swollen along mediolateral crest (black arrow); mesial borders shallowly excavated. Valvulae IX not exposed. Male genitalia. Unknown.</p>
            <p>Distribution (Fig. 12). COLOMBIA: Department Santander.</p>
            <p> Comments. This species is the only one in  H. triangularis group that has black punctures on pronotum.  Hypoxys ocanensis sp. n. and  H. curvatus sp. n. share the distal margin of each valvifer VIII with a distal spine. The  H. ocanensis sp. n. can be separated by valvifers VIII carinate (in  H. curvatus sp. n. valvifers VIII never carinate). </p>
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	https://treatment.plazi.org/id/03C787C3FFD5FFFAC2ABF90DFA50FADA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cunha, Eduardo Victor De Paiva;Fernandes, José Antônio Marin	Cunha, Eduardo Victor De Paiva, Fernandes, José Antônio Marin (2022): New species of the Hypoxys triangularis group and the first contribution on the morphology of the internal genitalia of Hypoxys Amyot & Serville, 1843 (Hemiptera: Pentatomidae: Edessinae). Zootaxa 5168 (5): 501-520, DOI: 10.11646/zootaxa.5168.5.1
03C787C3FFD7FFFCC2ABFA24FB7BFAD2.text	03C787C3FFD7FFFCC2ABFA24FB7BFAD2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypoxys triangularis (Dallas 1851) Nunes et al. 2020	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Hypoxys triangularis (Dallas, 1851)</p>
            <p>(Fig. 9, 19E–F, 22— Nunes et al. (2020); 6D, 7A, 8A, 9A, 12)</p>
            <p> Edessa triangularis Dallas, 1851: 320 (original description). </p>
            <p> Edessa triangularis Walker, 1868: 425 ; Stål, 1872: 60; Lethierry &amp; Severin, 1893: 195 (catalog); Kirkaldy, 1909: 165 (catalog) </p>
            <p> Edessa scabriventris Stål, 1859: 231 ; Stål, 1860: 28; Walker, 1868: 429; Stål, 1872: 56; Lethierry &amp; Severin, 1893: 194 (catalog); Kirkaldy, 1909: 164 (catalog); Silva et al. 2018: 424 (checklist) </p>
            <p> Edessa leprosula Breddin, 1904a: 144 ; Bergroth, 1908: 179; Kirkaldy, 1909: 159 (catalog) </p>
            <p> Hypoxys triangularis (Dallas, 1851) Nunes et al. (2020) : 420 (diagnosis and distribution). </p>
            <p> Diagnosis. See Nunes et al. (2020). Characteristic added here: Male genitalia. Dorsal surface with spot extending about ¼ of the length of the pygophore (9A— Nunes et al. (2020)). Inner wall with well-swollen area attached to dorsal rim (Fig. 8A, black arrow).  Phallus (Fig. 7A). Vesica tapering dorsally in lateral view. Phallotheca curved. Female genitalia. Pars intermedialis subcylindrical medially (Fig. 6D). Capsula seminalis with three thick projections: one projection half the length of the others, the other two projections with half of the length of pars intermedialis (Fig. 6D). </p>
            <p>Distribution (Fig. 12): FRENCH GUIANA; BRAZIL: Pernambuco, Bahia, Alagoas, Espírito Santo, Rio de Janeiro, Paraná, Santa Catarina, Rio Grande do Sul.</p>
            <p> Comments. The  H. triangularis can be separated from the other species of the  H. triangularis group by posterior stripe of pronotum reddish brown and body ventrally with large number of small yellow calli. </p>
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	https://treatment.plazi.org/id/03C787C3FFD7FFFCC2ABFA24FB7BFAD2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cunha, Eduardo Victor De Paiva;Fernandes, José Antônio Marin	Cunha, Eduardo Victor De Paiva, Fernandes, José Antônio Marin (2022): New species of the Hypoxys triangularis group and the first contribution on the morphology of the internal genitalia of Hypoxys Amyot & Serville, 1843 (Hemiptera: Pentatomidae: Edessinae). Zootaxa 5168 (5): 501-520, DOI: 10.11646/zootaxa.5168.5.1
03C787C3FFD1FFFCC2ABFA2CFCC3F8C4.text	03C787C3FFD1FFFCC2ABFA2CFCC3F8C4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypoxys capito (Breddim 1904)	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Hypoxys capito (Breddim, 1904)</p>
            <p>(Figure: 10, 19G–H, 22— Nunes et al. (2020); 7B, 8B, 11)</p>
            <p> Edessa capito Breddin, 1904: 151 (original description); Bergroth, 1908: 178; Kirkaldy, 1909: 155 (catalog) </p>
            <p> Hypoxys capito (Breddin, 1904) Nunes et al. (2020) : 422 (diagnosis and distribution). </p>
            <p>Diagnosis. See Nunes et al. (2020). Characteristic added here: Male genitalia. Dorsal surface of the pygophore with spot restricted to coarse area (Fig. 10A — Nunes et al. (2020)). Inner wall with swollen area globose close to superior process of genital cup (Fig. 8B, black arrow). Female genitalia. Valvifers VIII with mediolateral crest slightly swollen (Fig. 9B, black arrow).</p>
            <p>Distribution (Fig. 12): PARAGUAY: Cordillera, Central, Paraguarí.</p>
            <p> Comments. The swollen area on inner wall of the pygophore is the innermost one compared to other species of  H. triangularis group. See  H. striatus sp. n. comments. </p>
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	https://treatment.plazi.org/id/03C787C3FFD1FFFCC2ABFA2CFCC3F8C4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cunha, Eduardo Victor De Paiva;Fernandes, José Antônio Marin	Cunha, Eduardo Victor De Paiva, Fernandes, José Antônio Marin (2022): New species of the Hypoxys triangularis group and the first contribution on the morphology of the internal genitalia of Hypoxys Amyot & Serville, 1843 (Hemiptera: Pentatomidae: Edessinae). Zootaxa 5168 (5): 501-520, DOI: 10.11646/zootaxa.5168.5.1
03C787C3FFCDFFE1C2ABFF26FC28FE84.text	03C787C3FFCDFFE1C2ABFF26FC28FE84.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hypoxys subrastratus (Bergroth 1891) Nunes et al. 2020	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Hypoxys subrastratus (Bergroth, 1891)</p>
            <p>(Figs. 11, 19I–J, 22— Nunes et al. (2020); 6E, 7B, 8C, 9C, 12)</p>
            <p> Edessa subrastrata Bergroth, 1891: 233 ; Lethierry &amp; Severin, 1893: 195 (catalog); Kirkaldy, 1909: 165 (catalog) </p>
            <p> Hypoxys subrastratus (Bergroth, 1891) Nunes et al. (2020) : 423 (diagnosis and distribution). </p>
            <p> Diagnosis. See Nunes et al. (2020). Characteristic added here: Male genitalia. Dorsal surface of the pygophore with spot restricted to coarse area (Fig. 11A — Nunes et al. (2020)).  Phallus (Fig. 7B). Vesica widening ventrally in lateral view. Phallotheca curved. Female genitalia. Valvifers VIII with mediolateral crest well swollen (Fig. 9C, black arrow). Pars intermedialis slightly swollen medially (Fig. 6E). Capsula seminalis with three thick and long projections: two slightly surpassing anterior annular crest and the other almost reaching base of the pars intermedialis (Fig. 6E). </p>
            <p>Distribution (Fig. 12): BRAZIL: Pará, Maranhão, Goiás, Minas Gerais, São Paulo, Santa Catarina, Rio Grande do Sul; ARGENTINA: Candelaria; BOLIVIA: La Paz; PARAGUAY: Paraguarí.</p>
            <p> Comments. This species can be separated by dorsal rim of pygophore with rectangular median notch and inner wall with the larger swollen area between species of  H. triangularis group. </p>
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	https://treatment.plazi.org/id/03C787C3FFCDFFE1C2ABFF26FC28FE84	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Cunha, Eduardo Victor De Paiva;Fernandes, José Antônio Marin	Cunha, Eduardo Victor De Paiva, Fernandes, José Antônio Marin (2022): New species of the Hypoxys triangularis group and the first contribution on the morphology of the internal genitalia of Hypoxys Amyot & Serville, 1843 (Hemiptera: Pentatomidae: Edessinae). Zootaxa 5168 (5): 501-520, DOI: 10.11646/zootaxa.5168.5.1
