Sicarius crustosus, Nicolet, 1849

Sicarius crustosus View in CoL : Simon , 1893: 271.

Note: This species has been recognized by the strongly widened macrosetae in the femora.

Diagnosis: Differs from S. thomisoides and S. yurensis stat. nov. in the small size and by possessing a posteriorly truncate sternum, and white macrosetae and black prolateral setae in the femora. Differs from S. fumosus by the widened macrosetae in the femora ( Fig. 40C, D View Figure 40 ; not widened in S. fumosus , Fig. 40B View Figure 40 ), and by having brown markings at the border of the sternum ( Fig. 30D View Figure 30 ). The most similar species is S. lanuginosus : males differ by the longer, slightly sinuous embolus of the male ( Fig. 41A – C View Figure 41 ) (shorter and with a proximal hump in S. lanuginosus ); females differ by lacking rounded reservatories at the apex of the spermathecae branches, and by the gradual reduction in length of the spermathecae branches ( Fig. 41D – I View Figure 41 ) (rounded reservatories present and the two median pairs of branches clearly longer than the laterals in S. lanuginosus ).

Description: Male (Paposo, Antofagasta, Chile, AMNH). Coloration: carapace and chelicerae reddish brown, labium and endites light brown, sternum light brown with dark brown markings on margin, legs light brown, abdomen brownish yellow. Total length 6.14. Carapace length 3.12, anterior width 1.39, thoracic region width 2.92. Eye diameters and interdistances: PME 0.15, ALE 0.18, PLE 0.15, PME – PLE 0.25, ALE – PLE 0.25. Clypeus height 0.56. Eye tubercles present. Macrosetae in lateral border of the carapace in multiple rows. Femora with prolateral black setae in a zig-zag pattern, white macrosetae present, widened at the base. Sternum length 1.35, width 1.63, cordiform. Femoral thorns: 5. Leg I: femur 3.88 long, patella 1.14, tibia 4.36, metatarsus 3.2, tarsus 1.63. II: fe 4.16, ti 4.2. III: fe 3.65, ti 3.63. IV: fe 3.96, pa 1.09, ti 3.8, mt 3.14, ta 1.6. Abdomen: length 3.18, width 3.02. Palp as in Figure 41A and B View Figure 41 .

Female (Paposo, Antofagasta, Chile, AMNH). As in male, except for the following. Total length 7.58. Carapace length 3.1, anterior width 1.27, thoracic region width 2.98. Eye diameters and interdistances: PME 0.13, ALE 0.13, PLE 0.14, PME – PLE 0.25, ALE – PLE 0.2. Clypeus height 0.56. Femora macrosetae widened at the apex. Sternum length 1.3, width 1.5, cordiform. Femoral thorns: 4. Leg I: femur 3.08 long, patella 1.14, tibia 3.22, metatarsus 2.17, tarsus 1.19. II: fe 3.12, ti 3.15. III: fe 2.78, ti 2.75. IV: fe 3.12, pa 1.12, ti 2.21, mt 2.32, ta 1.37. Abdomen: length 4.67, width 4.36. Spermathecae branches: 8. Ventral branches present, one or two pairs, straight. Spermathecae as in Figure 41D and E View Figure 41 .

Variation: Males (N = 3): carapace length 3.1 – 3.27 (3.16), femoral thorns 4 – 5, femur II length 4.08 – 4.87 (4.37). Females (N = 5): carapace length 2.85 – 3.76 (3.15), femoral thorns 4 – 5, femur II length 3.08 – 4.11 (3.39), spermathecae branches 8 – 12. The number of ventral branches in the spermathecae of females varies (one pair, one pair and one unpaired, two pairs). The femoral macrosetae morphology is different in males and females, the first having them widened at the base ( Fig. 40C View Figure 40 ) and the latter having them widened at the apex ( Fig. 40D View Figure 40 ).

Natural history: Specimens have been collected under rocks in deserts, dry coastal vegetation and xeric scrublands. This is a very cryptic species: individuals remain motionless when disturbed.

Distribution: Dry coast of Chile, from Antofagasta to Coquimbo ( Fig. 26D).

Additional material examined: CHILE. Regi on � II de Antofagasta: Antofagasta, Antofagasta, 4 km N Paposo [70.38333, 23.51667, 20 m], N.I. Platnick et al., 11/X/1992, 1 $ ( AMNH); 6 km E Paposo GoogleMaps [70.33333, 23.56667, 894 m], 12/X/1992, 1 # 1 imm. ( AMNH); Taltal , 24 km S Paposo GoogleMaps [70.41667,

25.2, 50 m], 11/X/1992, 1 # (AMNH). Regi on � III de Atacama: Cha naral ~, Las Lomitas, Parque Nacional Pan de Az ucar � (70.60601, 26.00979, 614 m), M.J. Ram � ırez et al., 26/X/2011, 1 # 1 $ (MACN-Ar 32330); Copiap o �, Caldera, Rodillo [70.83333, 27.08333, 26 m], XI/1991, 1 $ (LEULS); Copiapo , 35 km S Copiapo [70.31667, 27.66667, 1326 m], L.E. Pe na ~, 16 – 17/X/1992, 1 $ (AMNH); 40 km S Copiapo [70.31667, 27.73333, 700 m], 23 – 25/X/1983, 2 $ (AMNH); Ruta 5, km. 758 (70.48717, 27.76693, 610 m), A.D. Brescovit et al., 9/XI/2013, 1 $ (IBSP 165572); Huasco, Huasco, Caleta Angosta (71.14792, 28.22308, 97 m), C.J. Grismado et al., 18/VIII/2009, 1 # (MACN-Ar 32331), 1 # (MACN-Ar 34224); Freirina (71.16243, 28.48288, 118 m), A.D. Brescovit et al., 9/XI/2013, 1 # (IBSP 165577), 1 $ (IBSP 165578); Parque Nacional Llanos de Challe (71.11356, 28.18403, 304 m), C.J. Grismado et al., 17 – 18/VIII/2009, 1 $ (MACN-Ar 32328); same locality (71.0885, 28.19222, 304 m), 1 $ 1 imm. (MACN-Ar 32329). Regi on � IV de Coquimbo: Elqui, La Higuera, El Totoralillo (71.2615, 29.59264, 154 m), M. Izquierdo et al., 12/VII/2009, 1 $ (MACNAr 30189), 1 $ 1 imm. (MACN-Ar 30205), 1 $ (MACNAr 30206); Los Choros [71.46047, 29.24025, 40 m], A. Gonzalez, 2007, 1 $ (LEULS); La Serena, Cuesta Porotitos (71.29217, 29.79447, 48 m), C.J. Grismado et al., 16/VIII/2009, 1 $ (MACN-Ar 32327); 1 # (MACN-Ar 30212); Limar � ı, Ovalle, Lorenzo Peralta (71.58207, 30.5741, 230 m), A.D. Brescovit & I.L.F. Magalh ~ aes, 12/XI/2013, 1 $ (IBSP 165584).