Sicarius thomisoides, WALCKENAER, 1847

SICARIUS THOMISOIDES WALCKENAER, 1847 View in CoL

Figures 12A View Figure 12 , 15A, B, 17D View Figure 17 , 18A View Figure 18 , 19A View Figure 19 , 26A, 30A View Figure 30 , 31B View Figure 31 , 32A View Figure 32 , 33A – C, 34 View Figure 34 , 35 View Figure 35 , 36 View Figure 36 , 37A View Figure 37

Sicarius thomisoides Walckenaer, 1847: 379 View in CoL . Description was not based on examination of specimens, but rather on the then unpublished illustrations by Nicolet (1849).

Thomisoides View in CoL terrosus Nicolet, 1849: 352 View in CoL , plate 1, fig. 9. Three immature syntypes from Valdivia , Chile (probably a mislabeling), deposited in MNHN 4159 About MNHN , examined, not found after the Instituto Butantan fire in 2010. SYN. NOV.

Thomisoides View in CoL rubripes Nicolet, 1849: 353 View in CoL , plate 1, fig. 10. Female ( MNHN 5488 About MNHN ) and immature ( MNHN 5491 About MNHN ) syntypes from Chile, examined, not found after the Instituto Butantan fire in 2010. SYN. NOV.

Thomisoides View in CoL minoratus Nicolet, 1849: 354 View in CoL . Two immature syntypes from Chile deposited in MNHN 5490 About MNHN , examined, not found after the Instituto Butantan fire in 2010. SYN. NOV.

Thomisoides View in CoL terrosus Keyserling, 1880: 268 View in CoL , plate 8, fig. 148.

Thomisoides View in CoL nicoleti Keyserling, 1880: 270 View in CoL , fig. 149. Male holotype from Chile, deposited in MNHN AR5492 About MNHN , examined, not found after the Instituto Butantan fire in 2010. SYN. NOV.

Sicarius terrosus View in CoL : Simon , 1893: 271.

Sicarius nicoleti View in CoL : Petrunkevitch, 1911: 122.

Synonymy: The syntypes of S. terrosus View in CoL and S. minoratus View in CoL are immatures, but somatic morphology (namely the shape of the sternum and leg macrosetae) indicates they belong to this species. The genital morphology of the types of S. nicoleti View in CoL ( Fig. 34C View Figure 34 ) and S. rubripes View in CoL ( Fig. 36G View Figure 36 ) falls within the variation of S. thomisoides View in CoL (see Variation below). The bulb of S. nicoleti View in CoL ’s holotype is expanded from the cymbium ( Fig. 34C View Figure 34 ), as can be noted in Keyserling’s illustration as well (1880, fig. 149).

Diagnosis: Distinguished from other species, except S. yurensis stat. nov., by the oval, longer-than-wide sternum ( Fig. 30A View Figure 30 ) (cordiform in other species). It is easily distinguished from S. yurensis stat. nov. by the narrow cephalic region ( Fig. 37A View Figure 37 ). Males are further distinguished by the gently curved palpal bulb body, without clear distinction between the bulb body and the embolus ( Fig. 34 View Figure 34 ). Females are further distinguished (except from S. yurensis stat. nov.) by the spermathecae with 8 – 20 branches usually bent at the apex and without a rounded reservatory ( Figs 35 View Figure 35 , 36 View Figure 36 ). We have been unable to find a genital feature in females to distinguish this species from S. yurensis stat. nov.

Description: Male (Illapel, Coquimbo, Chile, IBSP 165453). Coloration: carapace light reddish brown, chelicerae, labium and endites reddish brown, sternum orange, legs orange, abdomen brownish grey. Total length 14.7. Carapace length 7.7, anterior width 2.52, thoracic region width 7.1. Eye diameters and interdistances: posterior median eye (PME) 0.2, anterior lateral eye (ALE) 0.25, posterior lateral eye (PLE) 0.23, PME – PLE 0.61, ALE – PLE 0.31. Clypeus height 1.22. Eye tubercles absent. Macrosetae in lateral border of the carapace in a single row. Femora without prolateral black setae. Sternum length 3.76, width 3.02, oval. Femoral thorns: 5. Leg I: femur 9.6 long, patella 3.7, tibia 10, metatarsus 8.1, tarsus 4.1. II: fe 11, ti 11.38. III: fe 9.6, ti 9.2. IV: fe 9, pa 2.7, ti 8.7, mt 7.1, ta 3.7. Abdomen: length 6.7, width 5.4. Palp as in Figures 34A and B View Figure 34 .

Female (Illapel, Coquimbo, Chile, IBSP 165454). As in male, except by the following. Total length 14.85. Carapace length 7.25, anterior width 2.64, thoracic region width 7.08. Eye diameters and interdistances: PME 0.18, ALE 0.25, PLE 0.23, PME – PLE 0.69, ALE – PLE 0.28. Clypeus height 1.12. Sternum length 3.63, width 3.18. Femoral thorns: 5. Leg I: femur 7.92 long, patella 3.1, tibia 8.3, metatarsus 6, tarsus 3.3. II: fe 9, ti 8.6. III: fe 8.08, ti 7.5. IV: fe 8, pa 2.75, ti 7.33, mt 5.5, ta 3.08. Abdomen: length 6.58, width 6.25. Spermathecae branches: 14. Ventral branches present, one pair, straight ( Fig. 35A, B View Figure 35 ).

Variation: Males (N = 5): carapace length 4.72 – 7.7 (6.37), femoral thorns 5 – 6, femur II length 7.42 – 11.13 (9.92). Females (N = 5): carapace length 5.58 – 8.17 (6.98), femoral thorns 5 – 6, femur II length 6.75 – 9.1 (8.2), spermathecae branches 9 – 19. Genitalia are variable in both males ( Fig. 34 View Figure 34 ) and females ( Figs 35 View Figure 35 , 36 View Figure 36 ). The regions of Coquimbo and Atacama show the highest morphological diversity, with males with very long bulbs ( Fig. 34F View Figure 34 ) and aberrant females ( Figs 35C, D View Figure 35 , 36C, D View Figure 36 ). Some females do not have the spermathecae branches bent at the apex. There is a possibility that S. thomisoides , as delimited here, represents a species complex. Its treatment, however, is not trivial, and we prefer to consider it as a single species until further data are available. Subadult females have pre-spermathecae with several thin, short, digitiform branches ( Fig. 36H View Figure 36 ).

Natural history: This species occurs in a range of dry habitats in the arid coast of Chile, such as xeric woodlands in the Santiago region, semi-arid transition vegetation in the Coquimbo region, beaches and the extremely dry deserts in the north of the country. It is usually found beneath boulders, especially large ones. The eggcases ( Fig. 17D View Figure 17 ) can be found attached under those boulders. One of those, opened in the field, contained 17 eggs; another, that ecloded in the laboratory, gave rise to 15 juveniles. Dispersing-instar juveniles average 2.2 mg in dry weight (N = 15). Individuals have been observed in the field feeding on scolopendromorph centipedes, Loxosceles and gnaphosid spiders, solpugids, and on co-especifics .

Distribution: So far known only from the Chilean dry coast, from the extreme north of the country to south of the Santiago region ( Fig. 26A). A record from Buenos Aires, Argentina (deposited in AMNH), is certainly a mislabelling. There is no evidence that this species occurs in Argentina, and previous records from this country have been misidentified (see Sicarius levii sp. nov. below). The species probably extends its distribution into southern Peru, but this region is very sparsely sampled and we have not had the chance to examine many Sicarius specimens from there.

Additional material examined: CHILE. Regi on � XV de Arica y Parinacota: Arica , Arica, Playa Corazones (70.33067, 18.54817, 421 m), A.D.

Brescovit et al., 12/VII/2012, 1 $ 14 imm. ( IBSP 162896 View Materials ) , 1 $ ( IBSP 165451 View Materials ) ; [70.31667, 18.48333, 44 m], R. Calderon, 4/X/1970, 1 # ( UCCC) GoogleMaps ; Camarones , Azapa, Chitita [69.66667, 18.8, 2681 m], L.E. Pe na ~, 10/ VI /1986, 1 # 5 $ 42 imm. ( AMNH) , 1 # 1 imm. ( AMNH) , 2 # (AMNH); Codpa [69.73333, 18.81667, 2157 m], M. Codoceor, VII/ 1955, 1 $ ( MHNS) GoogleMaps ; Huara , Caleta Camarones (70.2635, 19.19283, 160 m), A.D. Brescovit et al., 11/VII/2012, 1 $ 4 imm. ( IBSP 162885 View Materials ) GoogleMaps , 1 $ ( IBSP 165476 View Materials ) . Regi on � I de Tarapac a �: Iquique , Huara, Pisagua (70.21683, 19.59833), A.D. Brescovit et al., 11/VII/2012, 5 $ 9 imm. ( IBSP 162900 View Materials ) GoogleMaps , 1 $ ( IBSP 165490 View Materials ) , 1 $ ( IBSP 165491 View Materials ) ; Quebrada de Tarapac a � [69.28333, 19.8, 1300 m], L.E. Pe na ~, 25/I/1992, 1 # 3 $ 6 imm. ( AMNH) ; Quebrada Mi nimi ~ ni ~ (70.006, 19.2035, 798 m), A.D. Brescovit et al., 11/VII/2012, 2 # 1 $ 3 imm. ( IBSP 162898 View Materials ) GoogleMaps ; Playa Chanabaya (70.13677, 20.88652, 73 m), 13/VII/2012, 2 # 1 imm. ( IBSP 162893 View Materials ) GoogleMaps , 1 $ ( IBSP 165446 View Materials ) , 1 # ( UFMG 15340 View Materials ) ; Playa Puenta Gruesa (70.11331, 20.04721, 691 m), 1 $ ( IBSP 162892 View Materials ) GoogleMaps ; Playa Quinteros (70.12075, 20.6109, 504 m), 2 $ ( IBSP 162879 View Materials ) GoogleMaps ; Playa R � ıo Seco (70.16417, 21.0023, 514 m), 1 $ 3 imm. ( IBSP 162883 View Materials ) GoogleMaps , 1 # ( IBSP 165475 View Materials ) ; Playa San Marcos (70.12075, 21.10068, 424 m), 1 $ 2 imm. ( IBSP 162880 View Materials ) GoogleMaps ; Punta Patache (70.18725, 20.80107, 52 m), 1$ 5 imm. ( IBSP 162887 View Materials ) GoogleMaps , 1 # ( IBSP 165477 View Materials ) , 1 $ ( IBSP 165478 View Materials ) ; Pica , Altos de Pica [68.95,

20.36667, 4149 m], M. Ferru, 21/II/2012, 1 $ ( IBSP 162891 View Materials ); Pozo Almonte, Centro Experimental Canchones (69.53383, 20.445, 997 m), A.D. Brescovit et al., 10/VII/2012, 1 $ 15 imm. ( IBSP 162899 View Materials ); La Huayca (69.54533, 20.43567, 997 m), 1 $ 10 imm. ( IBSP 162881 View Materials ), 1 $ 10 imm. ( IBSP 165449 View Materials ); Pampa del Tamarugal, 3 imm. ( IBSP 162910 View Materials ), 1 $ ( IBSP 165502 View Materials ); unspecific locality, 1$ 1 imm. ( ZMB 7894 View Materials ). Regi on � II de Antofagasta: Antofagasta, Antofagasta, 57 km N Antofagasta [70.33333, 23.11667, 435 m], H. Larrain, 5/VI/ 1992, 1 # ( AMNH); Juan Lopez (70.53637, GoogleMaps

23.50848, 70 m), A.D. Brescovit et al., 14/VII/2012, 1 imm. ( IBSP 162905 View Materials ) , 1 $ ( IBSP 165494 View Materials ) ; La Chimba (70.37787, 23.52132, 640 m), 15/VII/ 2012, 1 $ 2 imm. ( IBSP 162884 View Materials ) GoogleMaps ; La Negra [70.31667, 23.76667, 455 m], H. Larrain, 11/VII/ 1992, 1 $ 1 imm. ( AMNH) GoogleMaps ; Paposo [70.45, 25, 769 m], L.E. Pe na ~, 17 – 20/IX/1992, 1 $ 2 imm. ( AMNH) ; Quebrada La Chimba [70.37787,

23.52132, 640 m], J.C. Ortiz, 22/VIII/1996, 1 # ( UCCC); Vertedero Municipal (70.38565,

23.53375, 126 m), A.D. Brescovit et al., 15/VII/ 2012, 1 # 1 $ 3 imm. ( IBSP 162909 View Materials ), 1 # ( IBSP 165450 View Materials ); Mejillones, Hornitos [70.28333,

22.91667, 344 m], J.C. Ortiz, 22/VIII/1996, 1 $ ( UCCC); VIII/1996, 1 $ 1 imm. ( UCCC); (70.46267,

23.105, 28 m), A.D. Brescovit et al., 14/VII/2012, 1 $ 14 imm. ( IBSP 162894 View Materials ), 1 # ( IBSP 165479 View Materials ), 1 $ ( IBSP 165480 View Materials ), 1 $ ( IBSP 165481 View Materials ), 1 $ ( IBSP 165482 View Materials ), 1 $ ( UFMG 15356 View Materials ), 1 $ ( UFMG 15367 View Materials ); Taltal, 24 km S Paposo [70.41667, 25.2, 899 m], N.I. Platnick et al., 11/X/1992, 3 # 2 $ ( AMNH), 1 # 1 $ ( AMNH); same locality [70.41667, 25.2, 50 m], 3 # 1 $ 1 imm. ( AMNH); Caleta Paposo (70.46602, 24.99828, 45 m), A.D. Brescovit et al., 16/VII/2012, 1 $ ( IBSP 162904 View Materials ); Cave near road to Taltal (70.44578, 25.34698, 415 m), 17/VII/2012, 1 $ ( IBSP 162888 View Materials ); Cifuncho (70.64722, 25.65442, 44 m), 16/VII/2012, 4 imm. ( IBSP 162903 View Materials ), 1 $ ( IBSP 165493 View Materials ); Paposo [70.45, 25, 769 m], J. Stuardo, 23/VII/1964, 1 # ( UCCC); M. Elgueta, 8 – 10/X/1983, 1 # 2 $ ( MHNS 767 ); M. Ferr u �, II/2012, 1 imm. ( IBSP 162907 View Materials ), 1 # ( IBSP 165499 View Materials ); Paposo , La Rinconada, G. Arriagada , 25/X/1985, 1 $ ( MHNS); Quebrada del Medano [69.75, 25.28333, 2207 m], H. Niemeyer, VII/1985, 1 $ ( MHNS 900 ); Reserva Nacional Paposo (70.46602, 24.96367, 45 m), A.D. Brescovit et al., 16/VII/2012, 1 $ 4 imm. ( IBSP 162906 View Materials ), 1 # ( IBSP 165495 View Materials ), 1 $ ( IBSP 165496 View Materials ), 1 $ ( IBSP 165497 View Materials ), 1 $ ( IBSP 165498 View Materials ), 1 $ ( UFMG 15360 View Materials ); Taltal [69.75, 25.28333, 2207 m], 1 # ( AMNH); (70.4367, 25.26709, 252 m), A.D. Brescovit et al., 16/VII/2012, 1 $ 7 imm. ( IBSP 162895 View Materials ), 1 # ( IBSP 165483 View Materials ), 1 $ ( IBSP 165484 View Materials ), 1 $ ( IBSP 165485 View Materials ), 1 # ( IBSP 165486 View Materials ), 1 $ ( IBSP 165487 View Materials ), 1 $ ( IBSP 165488 View Materials ), 1 # ( UFMG 15358 View Materials ), 1 $ ( UFMG 15359 View Materials ); same locality [70.4367, 25.26709, 252 m], D.A. Masarey, 1910, 1 $ ( NHMB 459 View Materials a); El Loa, Calama, La Cascada [68.91667, 22.45, 2321 m], G. Arriagada, 10/III/1984, 1 $ 6 imm. ( MHNS 906 ); X/1982, 1 # 2 imm. ( MHNS 596 ); III/ 1983, 1 $ 3 imm. ( MHNS 635 ); Iquique, Huara, Quebrada de Tarapaca � [69.28333, 19.8, 2371 m], H. Larrain, 1 # 1 $ 1 imm. ( AMNH); Iquique, R � ıo Seco [70.11667, 20.25, 432 m], 21/VI/1998, 1 $ ( AMNH); Tocopilla , Mar � ıa Elena, R � ıo Loa Mouth [69.75, 21.43333, 10 m], L.E. Pe na ~, 12/II/1992, 3 $ ( AMNH); Tocopilla , 20 km N Tocopilla [70.18333, 22.08333, 263 m], 12/XII/1992, 1 $ 4 imm. ( AMNH); Caleta Buena (70.2444, 22.28246, 442 m), A.D. Brescovit et al., 14/VII/2012, 4 $ 9 imm. ( IBSP 162878 View Materials ), 1 # ( IBSP 165467 View Materials ), 1 $ ( IBSP 165468 View Materials ), 1 # ( IBSP 165469 View Materials ), 1 # ( IBSP 165470 View Materials ), 1 # ( IBSP 165471 View Materials ), 1 $ ( IBSP 165472 View Materials ), 1 $ ( IBSP 165473 View Materials ), 1 # ( UFMG 15354 View Materials ), 1 $ ( UFMG 15355 View Materials ); Caleta Urco Sur (70.15312, 21.76992, 285 m), 18/ VII/2012, 2 imm. ( IBSP 162902 View Materials ), 1 # ( IBSP 165492 View Materials ); Playa Consuela (70.24829, 22.32645, 524 m), 14/ VII/2012, 1 $ 2 imm. ( IBSP 162882 View Materials ), 1 $ ( IBSP 165474 View Materials ); Playa Punta Arenas (70.14537, GoogleMaps

21.64552), 13/VII/2012, 1 $ 3 imm. ( IBSP 162908 View Materials ), 1 $ ( IBSP 165500 View Materials ); Unspecific province, Camino a Paso Jama , 171-178 km, V. Jerez , 11/II/1997, 1 $ ( UCCC); Las Vertientes, H. Larrain, 12/VI/1992, 1 $ ( AMNH); 1 $ ( MHNS). Regi on � III de Atacama: Cha naral ~, Cha naral ~, Cachinal de la Costa [70.63333, 26.05, 228 m], J. Stuardo, 13/VII/1964, 1 # ( UCCC); Parque Nacional Pan de Az ucar � (70.6618, 26.14267, 120 m), A.D. Brescovit et al., 16/VII/2012, 3 imm. ( IBSP 162911 View Materials ), 1 $ ( IBSP 165503 View Materials ), 1 $ ( IBSP 165504 View Materials ); same locality (70.62823, 26.30238, 242 m), 17/VII/2012, 1 $ 3 imm. ( IBSP 162886 View Materials ), 1 $ ( UFMG 15343 View Materials ); Portofino (70.70087, 26.52123, 101 m), A.D. Brescovit et al., 10/XI/2013, 1 # ( IBSP 165564 View Materials ), 1 $ ( IBSP 165565 View Materials ), 1 imm. ( IBSP 165566 View Materials ); (70.63795, GoogleMaps

26.35577, 88 m), A.D. Brescovit et al., 17/VII/2012, 8 $ 9 imm. ( IBSP 162897 View Materials ), 1 # ( IBSP 165489 View Materials ), 1 # ( UFMG 15338 View Materials ), 1 $ ( UFMG 15339 View Materials ); Diego de Almagro , Inca de Oro [69.93333, 26.81667, 1954 m], L.E. Pe na ~, 8/X/1980, 3 $ 3 imm. ( AMNH); Flamenco coastal dunes [70.6, 26.61667, 10 m], N.I. Platnick et al., 10/X/1992, 1 $ ( AMNH); 13/X/ 1992, 1 $ ( AMNH); N of Cha naral ~ [70.6, GoogleMaps

26.36667, 204 m], L.E. Pe na ~, 10/X/1992, 1 $ ( AMNH); Copiapo, Caldera , 25 km SE Caldera [70.63333, 27.25, 591 m], 27 – 28/X/1983, 1 # 1 $ ( AMNH); Calderilla (70.85217, 27.0936, 12 m), A.D. Brescovit et al., 10/XI/2013, 1 $ ( IBSP 165546 View Materials ), 1 $ ( IBSP 165547 View Materials ); Caleta Los Patos (70.80013, GoogleMaps

26.95912, 11 m), 1 imm. (IBSP 165545), 1 # (UFMG 15325), 1 $ (UFMG 15326); E of Puerto Viejo [70.78333, 27.35, 152 m], L.E. Pe na ~, 9 – 10/ X/1980, 1 # 1 imm. (AMNH); Puerto Viejo (70.93197, 27.33708, 63 m), A.D. Brescovit et al., 10/XI/2013, 1 # (IBSP 165542), 1 imm. (IBSP 165548); same locality [70.93333, 27.33333, 14 m], L.E. Pe na ~, 15 – 16/X/1992, 1 # 2 imm. (AMNH), 1 # 2 imm. (AMNH); Quebrada de Leon [70.66667, 26.98333, 799 m], 8 – 9/X/1990, 2 $ 8 imm. (AMNH); same locality [70.66667, 26.98333, 140 m], N.I. Platnick et al., 10/X/1992, 1 $ (AMNH); Rodillo [70.83333, 27.08333, 26 m], 1989 – 1991, 67 # (LEULS); [70.8, 27.06667, 26 m], J. Moreno, 13/X/1982, 1 $ (AMNH); 20/VII/1983, 1 # 3 $ 3 imm. (AMNH); Copiapo , 40 km S Copiapo [70.31667,

27.73333, 700 m], 23 – 25/X/1983, 2 # 3 imm. (AMNH); 56 km N Vallenar [70.58333, 28.06667, 440 m], N.I. Platnick et al., 9/X/1992, 1 $ (AMNH); 60 km S Copiapo [70.33333, 27.88333, 1449 m], L.E. Pe na ~, 22/X/1991, 1 $ (AMNH); [70.31667,

27.36667, 589 m], H. Niemeyer, III/1986, 2 $ ( MHNS); J. Moreno, 11/X/1982, 1 # 3 $ 4 imm. ( AMNH); 20/X/1982, 2 $ 3 imm. ( AMNH); Elqui , Coquimbo, Tongoy [71.26667, 28.4], L.E. Pe na ~, 15/X/1990, 1 $ ( AMNH); Huasco, Freirina , 3.5 km E Carrizalillo (71.44536, 29.10379, 66 m), M.J. Ram � ırez et al., 30/X/2011, 1 # 2 $ 6 imm. (MACNAr 32342); 5.5 km N Carrizalillo (71.40012, GoogleMaps

29.05364, 157 m), 1 # (MACN-Ar 32339); Caleta Cha naral ~ de Aceituno (71.48782, 29.0788, 41 m), A.D. Brescovit et al., 11/XI/2013, 1 imm. ( IBSP 165551 View Materials ), 1 # ( UFMG 15327 View Materials ), 1 $ ( UFMG 15328 View Materials ); same locality (71.48759, 29.07202, 41 m), M.J. Ram � ırez et al., 30/X/2011, 1 # (MACN-Ar 32335); Carrizalillo (71.40623, 29.10455, 152 m), A.D. Brescovit et al., 11/XI/2013, 1 # ( IBSP 165561 View Materials ), 1 $ ( IBSP 165562 View Materials ), 1 $ ( IBSP 165563 View Materials ); Sauce Perez [71.11667, 28.63333, 853 m], L.E. Pe na ~, 26/X/ 1980, 1 $ 8 imm. ( AMNH); Huasco, Agua de Luna (71.16181, 28.33556, 107 m), C.J. Grismado et al., 18/VIII/2009, 1 $ 2 imm. (MACN-Ar 32336); Caleta Angosta (71.14792, 28.22308, 97 m), 1 # 2 $ 6 imm. (MACN-Ar 32334); Carrizal Bajo [71.15, GoogleMaps

28.08333, 39 m], J. Cepeda, X/2000, 2 # 1 $ 3 imm. ( LEULS); Freirina (71.16243, 28.48288, 118 m), A.D. Brescovit et al., 9/XI/2013, 1 # ( IBSP 165579 View Materials ), 1 $ ( IBSP 165580 View Materials ), 1 $ ( IBSP 165581 View Materials ); Los Pozos (71.15447, 28.16075, 198 m), C.J. Grismado et al., 18/VIII/2009, 1 $ 1 imm. (MACN-Ar 32333); Quebrada Talinay [71.21667, 28.46667, 110 m], L.E. Pe na ~, 13/X/1980, 1 $ 2 imm. ( AMNH); 20/III/1972, 1 # ( UCCC); Vallenar , 8 km N Vallenar [70.75, 28.5, 568 m], L.E. Pe na ~, 6/X/1990, 1 $ ( AMNH); Domeyko, road to Carrizalillo (70.98895, GoogleMaps

28.97858, 710 m), A.D. Brescovit et al., 11/XI/2013, 1 # ( UFMG 15334 View Materials ), 1 $ ( UFMG 15335 View Materials ); S of Vallenar, km 630 [70.75, 28.56667, 549 m], L.E. Pe na ~, 1/X/1980, 1 $ 5 imm. ( AMNH); Unspecific province , 1 km E Domeyko [800 m], N.I. Platnick et al., 8/X/1992, 1 # 2 $ 1 imm. ( AMNH); Juntas, R � ıo Huasco [1600 m], L.E. Pe na ~, 3/X/1980, 1 # ( AMNH); N of Cuesta Pajonales, 8/VIII/1982, 1 $ 4 imm. ( AMNH); X/1980, 2 # 1 $ ( AMNH); 21 – 23/X/ 1983, 1 # 3 imm. ( AMNH). Regi on � IV de Coquimbo: Choapa, Illapel, Cuesta El Espino (71.305, 31.75668, 629 m), A.D. Brescovit & I.L.F. Magalh aes ~, 8/IV/2013, 2 $ 7 imm. ( IBSP 163724 View Materials ); Cuesta Los Cristales (71.1388, 31.70447, 659 m), 1 $ 8 imm. ( IBSP 163744 View Materials ), 1 # ( IBSP 165453 View Materials ), 1 $ ( IBSP 165454 View Materials ), 1 # ( UFMG 15350 View Materials ), 1 $ ( UFMG 15351 View Materials ); Cuz-Cuz (71.29342, 31.67, 252 m), 1 # 7 $ 8 imm. ( IBSP 163734 View Materials ), 1 # ( IBSP 165509 View Materials ), 1 # ( IBSP 165510 View Materials ); Salamanca, El Tambo (71.0685, 31.77175, 453 m), 5 $ 5 imm. ( IBSP 163747 View Materials ); Los Vilos, Caleta Chigualoco (71.51118, GoogleMaps

31.75522, 20 m), 7/IV/2013, 3 imm. ( IBSP 163757 View Materials ); Near Puerto Punta Chungo (71.49122, 31.8767, 79 m), 8/IV/2013, 13 $ 7 imm. ( IBSP 163731 View Materials ), 1 # ( IBSP 165507 View Materials ), 1 # ( IBSP 165508 View Materials ), 1 # ( UFMG 15361 View Materials ); [71.5, 31.91667, 189 m], J. Pizarro-Araya, 28/X/2009, 1 $ ( LEULS); Mincha, Caleta Oscuro [71.6, 31.41667, 38 m], Mojano, XI/1974, 1 # ( UCCC); R. C. Digue-Loja, 2/XI/1974, 1 $ ( UCCC); Canela Baja, Los Pozos (71.20947, 31.30627, 912 m), A.D. Brescovit et al., 7/XI/2013, 1 $ ( IBSP 165558 View Materials ), 1 $ ( IBSP 165559 View Materials ), 1 $ ( IBSP 165560 View Materials ); Ermita Padre Hurtado [71.55, 31.46667, 256 m], A. Levican, 24/IX/2004, 1 # 1 imm. ( LEULS); Huentelauquen [71.53333, 31.58333, 127 m], L.E. Pe na ~, 26/IX/1980, 10 $ 10 imm. ( AMNH); Salamanca, Entre San Agust � ın y Zapallar (70.79339, 31.70568, 1430 m), M.J. Ram � ırez et al., 1/XI/2011, 1 # 1 $ 22 imm. (MACN-Ar 32341), 1 # 1 $ (MACN-Ar 32343); (70.98692, 31.79583, 771 m), A.D. Brescovit & I.L.F. Magalh ~ aes, 8/IV/ 2013, 2 imm. ( IBSP 163760 View Materials ); same locality [70.96667, 31.78333, 563 m], J. Aros, 1/V/1961, 1 $ 10 imm. ( AMNH); El Temblador [71.31667, GoogleMaps

29.46667, 83 m], 1/V/2004, 2 $ 2 imm. (LEULS); Elqui , Coquimbo, 16 km S Cruz Grande [71.41667,

30.35, 140 m], N.I. Platnick et al., 7/X/1992, 2 $ 1 imm. ( AMNH); Andacollo [71.2804, 30.2355, 260 m], J. Pizarro-Araya, VI – VII/2007, 1 # 3 $ 25 imm. ( LEULS); P. Augusto, 14/IX/2003, 1 $ 1 imm. ( LEULS); Horcon [71.215, 29.9243, 162 m], J. Pizarro-Araya, 14/VIII/2004, 1 # 1 imm. ( LEULS); La Serena, Cerro Grande [71.21667, 29.91667, 162 m], L. Espinosa , 16/VIII/2003, 1 # 2 imm. ( LEULS); Las Cardas [71.2613, 30.291, 360 m], 13/XII/2003, 1 $ 2 imm. ( LEULS); M. Moreno, I/ 1979, 1 # ( LEULS); Pan de Az ucar � (71.2321, GoogleMaps

30.12235, 174 m), A.D. Brescovit et al., 8/XI/2013, 1 $ ( IBSP 165588 View Materials ), 1 imm. ( IBSP 165589 View Materials ); Ruta 5, km. 453 (71.37845, 30.02547, 212 m), A.D. Brescovit & I.L.F. Magalh aes ~, 12/XI/2013, 1 # ( IBSP 165573 View Materials ), 1 $ ( IBSP 165574 View Materials ); Tongoy [71.48333, GoogleMaps

30.25, 31 m], 21/IX/1996, 1 # 2 $ 4 imm. ( AMNH); La Higuera , 6 km E El Tofo [71.16667, 29.43333, 764 m], N.I. Platnick & O.F. Francke, 7/I/1985, 1 $ 1 imm. ( AMNH); 6 km S Cruz Grande [71.31667, GoogleMaps

29.48333, 20 m], N.I. Platnick et al., 6/X/1992, 1 $ ( AMNH); same locality (71.31667, 29.48333, 5 m), 11/XI/1993, 2 # 3 $ 1 imm. ( AMNH); 79 km N La Serena, Rt. 5, km 553 [71.31667, 29.35, 300 m], 15/X/1992, 1 # ( AMNH); Caleta Hornos (71.28072, 29.62743, 449 m), A.D. Brescovit et al., 8/XI/2013, 1 # ( IBSP 165582 View Materials ), 1 $ ( IBSP 165583 View Materials ); Choros Bajos [71.3, 29.28333, 300 m], L.E. Pe na ~, 10 – 12/XI/1981, 1 $ 2 imm. ( AMNH); same locality [71.3, 29.28333, 79 m], 19/X/1983, 1 $ 8 imm. ( AMNH); Chungungo (71.30283, GoogleMaps

29.4446, 36 m), A.D. Brescovit et al., 11/XI/2013, 1 $ ( IBSP 165569 View Materials ), 1 # ( UFMG 15332 View Materials ), 1 $ ( UFMG 15333 View Materials ); Chungungo, Playa Blanca [71.30283,

29.4446, 36 m], J. Pizarro-Araya, 27/II/2005, 2 # 1 $ 1 imm. ( LEULS); El Tofo (71.25, 29.43333, 465 m), IX/1916, 1 # 1 imm. ( AMNH); El Totoralillo (71.2615, 29.59264, 154 m), M. Izquierdo et al., 12/VII/2009, 1 $ 18 imm. (MACN-Ar 32337); Isla Choros [71.5394, 29.25539], J. Pizarro-Araya, X/ 2012, 2 $ 5 imm. ( LEULS); L.E. Pe na ~, 7/VIII/1980, 3 $ 1 imm. ( AMNH); Llano de la Higuera [71.26667, 29.5, 645 m], 6/X/1990, 1 $ 1 imm. ( AMNH); 17/X/1983, 1 $ 3 imm. ( AMNH); Los Choros [71.46047, 29.24025, 40 m], A. Gonzalez, 3/IX/2007, 1 # 3 $ 3 imm. ( LEULS);, 2007, 1 $ 9 imm. ( LEULS); J. Pizarro-Araya, 27/V/2006, 3 $ 4 imm. ( LEULS); R. Villalon, 4/VI/2005, 1 # ( LEULS); Punta Choros (71.46028, 29.249, 40 m), A.D. Brescovit et al., 11/XI/2013, 1 # ( IBSP 165590 View Materials ), 1 # ( UFMG 15336 View Materials ), 1 $ ( UFMG 15337 View Materials ); same locality [71.46028, 29.249, 40 m], V. Rodriguez, 4/VI/ 2005, 1 $ ( LEULS); Quebrada Los Choros [71.066, GoogleMaps

29.35, 476 m], 1 # ( LEULS); Totoralillo [71.31667, 29.48333, 83 m], D. Valdivia, 1/IX/ 2006, 1 # ( LEULS); La Serena , 19 km N La Serena [71.33333, 29.76667, 182 m], N.I. Platnick & R. T. Schuh , 4/XI/1981, 1 $ ( AMNH); 20 km N La Serena, Rt. 5, km 491 [71.33333, 29.76667, 120 m], N.I. Platnick et al., 6/X/1992, 1 # ( AMNH); same locality (71.33333, 29.76667, 120 m), 8/II/1994, 1 $ ( AMNH); Cuesta Porotitos [71.29217, 29.79447, 48 m], 5/VII/2004, 1 $ ( LEULS); Marquesa (70.95792, 29.94158, 626 m), 10/I/2006, 1 $ 2 imm. ( LEULS); Monte Grande [71.2513, 29.881, 64 m], L. Espinoza & J. Pizarro-Araya, 13/VIII/2004, 3 $ 2 imm. ( LEULS); Pelicana [71.0087, 29.9871, 414 m], J. Pizarro-Araya, 1/XI/2006, 1 # ( LEULS); Punta de Teatinos [71.28333, 29.81667, 48 m], L. Espinoza, 6/IX/2003, 1 $ 2 imm. ( LEULS); R � ıo Hurtado, El Panque, 20 km S Vicuna [70.68333, GoogleMaps

30.21667, 1500 m], N.I. Platnick et al., 4/X/1992, 1 # 1 $ 6 imm. ( AMNH); Morrillos [70.7092,

30.2894, 1507 m], 24 – 31/X/2008, 1 # ( LEULS) ; Vicu na ~, Bahia Mausa [70.7, 29.71667, 1866 m], L.E. Pe na ~, 2 – 3/X/1993, 2 $ ( AMNH); Diaguitas (70.62603, 30.00844, 1005 m), O. Alcagaya, 18/ IX/2006, 1 # ( LEULS); Mayac un � [70.7, 29.71667, 1866 m] GoogleMaps , R. Donoso?, 1/ V /1961 , 1 $ 6 imm. (AMNH); Huasco, Huasco, Llanos de Challe [71.06667,

28.21667, 291 m], 19 – 21/XII/1997, 1 # 2 imm. ( LEULS); Limarı, Combarbal a �, Illapel, Cuesta El Espino [71.06667, 31.31667, 2200 m], L.E. Pe na ~, 1/IX/1984, 4 $ 2 imm. ( AMNH), 1 # 3 $ 3 imm. ( AMNH), 3 # ( AMNH); IX/1984, 2 # 3 $ 2 imm. ( AMNH); La Ligua Baja (71.04758, 31.0251, 711 m), A.D. Brescovit et al., 7/XI/2013, 1 $ ( IBSP 165570 View Materials ), 1 $ ( IBSP 165571 View Materials ); Ovalle , 24 km S Socos, Rt. 5, km 347 (71.61667, 30.88333, 360 m), N.I. Platnick et al., 9/XI/1993, 1 # ( AMNH); Caleta El Limar � ı [71.68333, 30.71667, 148 m], D. Valdivia, 21/IX/2004, 1 $ 1 imm. ( LEULS); Caleta La Cebada (71.64278, 30.97275, 53 m), A.D. Brescovit & I.L.F. Magalh aes ~, 14/XI/2013, 1 # ( IBSP 165553 View Materials ), 1 $ ( IBSP 165554 View Materials ), 1 # ( UFMG 15329 View Materials ); Camino a Fray Jorge (71.65365, 30.59622, 420 m), 12/XI/ 2013, 1 imm. ( IBSP 165557 View Materials ), 1 # ( UFMG 15330 View Materials ), 1 $ ( UFMG 15331 View Materials ); Huampulla [71.2, 30.6, 272 m], L.E. Pe na ~, 30/IX/1993, 1 $ 11 imm. ( AMNH); Huilmo, 11 km SSW Ovalle, 4 km N Los Cristales [71.13333, 30.76667, 858 m], 22 – 25/IX/1982, 1 $ 5 imm. ( AMNH); Lorenzo Peralta (71.58207, GoogleMaps

30.5741, 230 m), A.D. Brescovit & I.L.F. Magalh aes ~, 12/XI/2013, 1 # (IBSP 165585), 1 $ (IBSP 165586), 1 imm. (IBSP 165587); Los Maitenes, Corral El Julio [71.68052, 30.35201], S. Zunino, 6/VI/1975, 1 $ 4 imm. (UCCC); Los Maitenes, N of Amolantas [70.91667, 30.36667, 1384 m], L.E. Pe na ~, 27/IX/1980, 1 # 1 $ 3 imm. (AMNH); Socos [71.49294, 30.72781, 125 m], L. Espinoza, 27/IX/2003, 2 $ 5 imm. (LEULS); same locality [71.48333, 30.7, 151 m], T. Cekalovic, 8/ IX/1968, 1 $ 2 imm. (UCCC); same locality (71.49294, 30.72781, 125 m), 20/VII/2009, 1 # 2 $ 8 imm. (MACN-Ar 32338); Petorca, La Ligua, Pichidangu � ı, Valle de Quillimar � ı (71.50652,

32.10803, 25 m), A.D. Brescovit & I.L.F. Magalh aes ~, 7/IV/2013, 4 $ 5 imm. ( IBSP 163728 View Materials ); Playa La Ballena (71.46095, 32.29212, 169 m), 3 $ 7 imm. ( IBSP 163750 View Materials ); Unspecific province , El Encanto , 9/I/1972, 1 # 1 $ ( UCCC). Regi on � V de Valpara � ıso: Los Andes , Calle Larga [70.58333, GoogleMaps

32.83333, 878 m], H. Heatwole, 30/IX/1965, 1 # ( AMNH); Los Andes , Guarda Vieja (70.2737,

32.90292, 2094 m), A.D. Brescovit & I.L.F. Magalh aes ~, 6/IV/2013, 1 # ( IBSP 165515 View Materials ); Hidroeletrica Guarda Vieja, 1 $ 11 imm. ( IBSP 163748 View Materials ); same locality (70.5626, 32.83133, 878 m), 1 imm. ( IBSP 163754 View Materials ); same locality (70.2737, 32.90292, 2094 m), 1 $ ( IBSP 165516 View Materials ); R � ıo Blanco (70.45452, 32.8555, 1226 m), 12 imm. ( IBSP 163706 View Materials ); Petorca, La Ligua , 7 km E La Ligua [71.3, 32.46667, 185 m], N.I. Platnick & O.F. Francke, 9/I/1985, 1 # ( AMNH); Pichicuy , Quebrada Huaquen [71.46667, 32.33333, 10 m], N.I. Platnick et al., 2/X/1992, 1 $ ( AMNH); (71.32983, GoogleMaps

32.44067, 53 m), A.D. Brescovit & I.L.F. Magalh aes ~, 7/IV/2013, 1 $ 4 imm. ( IBSP 163725 View Materials ); Zapallar, Papudo (71.46545, 32.50513, 76 m), 9/ IV/2013, 1 # 3 $ 10 imm. ( IBSP 163733 View Materials ); Quillota, Olmue, Palmas de Ocoa (71.08547, 32.93833, 697 m), A.D. Brescovit et al., 23/III/2013, 4 imm. ( IBSP 163742 View Materials ); same locality (71.09033, GoogleMaps

32.91633, 377 m), 1 # 2 $ 7 imm. ( IBSP 163743 View Materials ); San Felipe de Aconcagua, Catemu, El Guayac an � (70.93057, 32.49723, 415 m), A.D. Brescovit & I.L.F. Magalh aes ~, 6/IV/2013, 1 $ 8 imm. ( IBSP 163718 View Materials ); San Felipe, Putaendo (70.71002, GoogleMaps

32.56667, 1017 m), 1 $ 10 imm. ( IBSP 163723 View Materials ), 1 $ ( IBSP 165505 View Materials ), 1 $ ( IBSP 165506 View Materials ); (70.73738,

32.706, 739 m), 4 imm. ( IBSP 163737 View Materials ); Valparaıso, Puchuncav � ı, La Laguna (71.424, 32.64933, 30 m), 9/IV/2013, 2 # 8 $ ( IBSP 163735 View Materials ), 1 # 6 $ 9 imm. ( IBSP 163736 View Materials ); La Laguna , road to Catapilco, km 16 (71.3974, 32.6155, 105 m), 2 # 5 $ 5 imm. ( IBSP 163730 View Materials ), 2 # 5 $ 1 imm. ( IBSP 163732 View Materials ), 1 # 4 $ 4 imm. ( IBSP 163746 View Materials ), 1 # ( IBSP 165455 View Materials ), 1 # ( IBSP 165513 View Materials ), 1 # ( IBSP 165514 View Materials ). Regi on � Metropolitana: Cachapoal, Mostazal, Huelquen (70.67338, 33.8749, 385 m), A.D. Brescovit & I.L.F. Magalh aes ~, 25/III/2013, 6 imm. ( IBSP 163726 View Materials ); Chacabuco, La Dormida, Til-Til [70.91667, 33.08333, 2000 m], L. Irarrazaval, 16 – 18/XI/1984, 1 # ( AMNH); Cordillera, Pirque, Cerro Chena [70.75, 33.58333, 529 m], G. Arriagada, VIII/1985, 1 $ ( MHNS 922 ); San Jose de Maipo, Camino al Embalse de Yeso (70.20333, 33.79117, 1965 m), A.D. Brescovit et al., 22/III/2013, 2 $ 12 imm. ( IBSP 163749 View Materials ), 1 # ( IBSP 165517 View Materials ), 1 # ( IBSP 165518 View Materials ); Camino Central Hidreletrica Los Maitenes (70.34133, 33.5705, 1329 m), 1 $ 4 imm. ( IBSP 163710 View Materials ); El Canelo [70.38333, 33.61667, 1243 m], L.E. Pe na ~, XII/1984, 2 $ 10 imm. ( AMNH); Farellones, Plazoleta Los Pumas (70.29033, GoogleMaps

33.35983, 2821 m), A.D. Brescovit et al., 21/III/ 2013, 1 # 1 imm. ( IBSP 163727 View Materials ); Farellones, Puente Nilhue ~ [70.3, 33.35, 2207 m], M. Elgueta, 1982, 1 $ 1 imm. ( MHNS 600 ); Pe nanolen ~ (70.51667,

33.48333, 892 m), C. Veloso, IV/2013, 4 imm. ( IBSP 163751 View Materials ); road to Farellones (70.36967,

33.35483, 1738 m), A.D. Brescovit et al., 21/III/ 2013, 1 $ ( IBSP 163740 View Materials ); Maipo, Calera de Tango, Quebrada de la Plata [70.88333, 33.5, 612 m], C. Veloso, 24/X/2010, 1 $ ( IBSP 161103 View Materials ); San Felipe de Aconcagua, Llaillay, Puente Pedro Bustus M. (70.84783, 32.86867, 757 m), A.D. Brescovit et al., 23/III/2013, 1 imm. ( IBSP 163705 View Materials ), 5 imm. ( IBSP 163753 View Materials ); Santiago, Santiago, Cuesta Lo Prado (70.8986, 33.45818, 562 m), 24/III/2013, 6 $ 14 imm. ( IBSP 163745 View Materials ), 1 # ( IBSP 165511 View Materials ), 1 $ ( IBSP 165512 View Materials ); Quilicura [70.71667, 33.35, 489 m], L.E. Pe na ~, 25/V/1979, 1 $ 2 imm. ( AMNH); V/1979, 1 # 3 imm. ( AMNH); VIII – IX/1979, 1 # 3 imm. ( AMNH); Renca [70.75, 33.38333, 514 m], 11/X/1984, 1 # ( AMNH); Valle Lo Campino (70.72447, 33.38665, 514 m), A.D. Brescovit et al., 24/III/2013, 3 imm. ( IBSP 163707 View Materials ). Regi on � VI de O’Higgins: Cachapoal, Machal � ı, Coya (70.61452, 34.1963, 941 m), 5/IV/2013, 1 $ 4 imm. ( IBSP 163708 View Materials ); Requinao, Do nihue ~ (70.84495, 34.18863, 483 m), 25/III/2013, 3 imm. ( IBSP 163755 View Materials ). No further data: Quebrada de Talca , 7/VIII/1997, 1 # ( MHNS). ARGENTINA. Buenos Aires: [58.36667, 34.6, 24 m], 15/IX/1996, 1 # 1 $ 2 imm. ( AMNH) (certainly a mislabelling) GoogleMaps .