Sufetula choreutalis (Snellen, 1880)

Sufetula choreutalis (Snellen, 1880) View in CoL

( Figures 5 View Figures 3-8 , 21 View Figures 19-26 , 34 View Figures 32-37 , 45 View Figures 42-45 )

Pseudochoreutes choreutalis Snellen, 1880: 202 - 203 View in CoL . Type locality: Indonesia, Sulawesi [¼ Celebes]: Makassar, Saleyer

¼ Sufetula chloreutalis Whalley, 1962: 108

Material examined. 7 Ƌ (specimen identifiers coll.mfn-berlin.- de_u_de00bb, b4b0fe, 2a6f59, 2a6f5a, 2a6f5b, 2a6f5c, e80bee), 3 ♀ (specimen identifiers coll.mfn-berlin.de_u_2a6f58, 6d7f3a, 430abf), 2 unsexed (specimen identifiers coll.mfn-berlin.de_u_87a5c0, 73609d). Detailed information in Table S1.

Similar species. Sufetula sunidesalis Walker, 1859 , Sufetula gigantea sp. nov.

Diagnosis. This species ( Figure 5 View Figures 3-8 ) is superficially similar to Sufetula gigantea sp. nov., but its wing length of 5 - 6 mm is conspicuously smaller than in S. gigantea sp. nov. (wing length¼ 9.5 - 12 mm). The brown nuclei of the costal spots of the forewing as well as the continuous submarginal lines on the fore- and hindwing further separates S. choreutalis from S. gigantea sp. nov. In the male genitalia, the short, tapering uncus and the quadrangular shape of the valva with a pointed apex separates this species from other Sufetula species. In the female genitalia ( Figure 45 View Figures 42-45 ), the bulge at the posterior end of the ductus bursae with irregular sclerotisation is unique to this species.

Description. Wings ( Figure 5 View Figures 3-8 ). Forewing length: 9.5 - 12.3 mm; forewing ground colour brown. Antemedian area with two oblique cream stripes. Antemedian line cream with distal edge dark brown, slightly arched outwards, forming three spikes pointing distally. Costa with three conspicuous cream spots with innerside dark brown, the third one fused with postmedian line. Discoidal spot rather large, round, black. Postmedian line cream, curved outwards on basal 1/3, abruptly angled at 1/3, running inwardly; third 1/4 running downwards towards dorsum, abruptly angled at 3/4, running outwardly, then down to dorsum. Subterminal area with dark brown patch at apex, with thin, dark brown submarginal line. Wing margin deeply notched at M 1 -M 2. Fringes basally cream, distally brown. Hindwing basally cream speckled with dark brown, with antemedian patch of dark brown. Median line jagged, cream-coloured. Distal part light to dark brown, with thin, dark brown submarginal line. Forewing CuP vein present. Tympanal organs ( Figure 21 View Figures 19-26 ). Ƌ ♀. Bullae tympani with lateral fourth recessed lateral of venula prima. Saccus tympani small, wider than long, deeper and more demarcated in male, opened ventrad. Venulae secundae short, kinked twice, extending to midlength of sternum II. Male genitalia ( Figure 34 View Figures 32-37 ). Uncus ca 2/3 of tegumen arm length, slender, slightly enlarging toward middle, with scattered hairs on lateral edges, tip pointed. Valva quadrangular, innerside of ventral margin with tuft of dense hairs; ventral part protruding distally into rounded projection. Phallus with one small cornutus, phallus apex sclerotised, notched. Female genitalia ( Figure 45 View Figures 42-45 ). Posterior apophyses ca 2.25 × tergite VIII length. Anterior apophyses ca 2 × tergite VIII length, at basal 1/5 with both ventral and dorsal short bump. Colliculum ca 1/4 of tergite VIII length, ring-shaped. Ductus bursae with small, round, irregular sclerite anterior of colliculum and ductus seminalis, posterior 3/4 of ductus bursae membranous, anterior 1/4 lined with spicules near corpus opening. Corpus bursae subequal to length of ductus, oval-shaped, with posterior area covered with short erected papillae; anterior area covered with erected acanthae. Signum ca half the corpus length, forming an oblong patch of short, sclerotised, dent-shaped thorns.

Distribution. INDONESIA: Sulawesi, Selayer [Salayer], 1,600 ft.; PAPUA NEW GUINEA: Rossel I.; Astrolabe Bay; Alor; Talaut; Sariba I.; Witu ¼ French Is.; Hydrographer Mts, 2000 ft.; St Matthias I.; Fergusson Is.; Milne Bay; Neu Pommern [New Britain]; Feni I, E. of New Ireland; Key I.; Little Kei; Squally Is.; Rossel Is; Sangir; INDONESIA: Tenimber [Tanimbar}; SOLOMONS: Rennel Is.; Bellona I, Matahenua, Hutuna; Vella Lavella; Nissan I.; S. side of Choiseul Is.; Bougainville; PHILIPPINES. Leyte, Luzon (Quezon), Mindanao (90 - 216m); Los Banos; Kolambugan, Lano, plains, Mindanao ( Figure 55 View Figure 55 ); SAMOA: Apia, Upolu.

DNA barcoding. An intraspecific variation of 0.3 % was found between the specimen MFNLEP970 from Leyte and the BOLD record LNAUW 3158-17 from Luzon (Laguna, Mt Makiling).

Remarks. This species is sexually dimorphic, with females exhibiting wing length 1 - 2 mm longer than males.