Paramannophorus skillmani (Chemsak and Hovore, 2010) Eya, 2024

Paramannophorus skillmani (Chemsak and Hovore in: Eya, 2010), new combination

( Fig. 235–244 View Figures 235–244 )

Ischnocnemis skillmani Chemsak and Hovore View in CoL : in Eya 2010: 7; MacRae et al. 2012: 180 (bio.)

Materials examined. MEXICO: Oaxaca: MX 190 vic Metatlan ( Matatlán ), microwave tower, 19 Oct. 2001, F. Skillman and J. Davidson (paratypes: 6 males, 4 females; 1 male, EMEC) ; road to Micro. Ocotepec, 6 km S Matatlan (Matatlán), Hwy 190, 2000–2100 m, 16°49.37′, 96°21.73′, 7 Oct. 2003, N. Schiff (paratype: 1 male, EMEC) ; 6 km SE Santiago Matatlán, HWY 190, R.M.O. Ocotepec , 16 Oct. 2005, B.K. Eya (paratypes: 1 male, 1 female, BKEC) ; 21 Oct. 2005 B.K. Eya (paratype: 1 female, BKEC) ; Microondas, Ocotepec road, 6 km S Matalán ( Matatlán ), 6800 ft, gps N 16°49.32′, W 96°21.42′, 16 Oct. 2004, C.L. Belamy (paratypes: 1 male, 1 female, EMEC) GoogleMaps .

Discussion. Paramannophorus skillmani (Chemsak and Hovore) new combination is small (8–13 mm), and subparallel. The “integument is shining black with vague, dark, greenish overtones,” and the antennae are “pale, brownish from the apices of third or fourth” antennomere (Chemsak an Hovore in Eya: 2010: 7). The pubescence is moderately dense, pale, and erect on the head, pronotum and prosternum, and long and depressed on the remainder of underside, and shorter and suberect on the elytra. There are two eburneous vittae on each elytron, a broader subsutural vitta extending from base almost to apices, and a narrower submarginal vitta starting behind humerus extending to about apical one-tenth of elytra. Front is declivous with mid-cranial sulcus narrow, deeply canaliculate, and “punctures moderately coarse and subconfluent.” Vertex is coarsely, confluently punctate throughout, and the midline vaguely elevated. The antennal tubercles are divergent and the integument between tubercles is impressed forming a shallow V-shaped valley sloping down to the mid-cranial sulcus. The antennae are slender, extending about three antennomeres beyond the elytra in males, and attaining or about an antennomere longer than the elytra in females. The pronotum is “broader than long (L/W: 0.88), sides obtusely angulate” in middle with apical margin not elevated on the dorsum with lateral collar-like projection absent. The pronotal disc is coarsely and subcontiguously punctate with a shallow, longitudinally elongate, glabrous callus in the middle. The proepisternal areas are confluently punctate. Prosternum is transversely rugulose and confluently punctate. The mesosternum is coarsely, sparsely punctate in middle, and confluently punctate on sides. The metasternum is nitid, coarsely, discretely punctate, and glabrous along midline. The elytra are “about two and half times as long as broad” (L/W: 2.5). The punctures on elytra between vittae are “dense and coarse becoming finer” apically, those between subsutural vittae and suture are shallower and sparser, and those between submarginal vittae and the epipleural margins are coarse and contiguous. The elytral apices are sinuate truncate with exterior angles dentate.

According to Chemsak and Hovore in Eya (2010: 8), “this is another Ischnocnemis species with longitudinal yellow vittate elytra. It differs generally from the other described vittate Ischnocnemis by its smaller size, pale brown distal antennal segments, and vaguely angulated sides of the pronotum.” Although this species superficially resembles Ischnocnemis , the morphology of the head capsule with declivous front, divergent antennal tubercles, and prognathous mouthparts ( Fig. 17a, 17b View Figures 16–23 ) differs from the later with convex, subvertical or vertical front and retracted mouthparts ( Fig. 23a, 23b View Figures 16–23 ). This species also lacks the elevated collar-like lateral projection and raised anterior margin of pronotum, which are the key characters for Mannophorus as designated by LeConte. This species is therefore, moved to Paramannophorus Eya new genus. Paramannophorus skillmani (Chemsak and Hovore) resembles Mannophorus laetus LeConte in overall form with broader pronotum and elytra, declivous front, coarser texture of integument, and shorter and explanate metatarsomeres ( Fig. 36a View Figures 36–39 ). This species has pronotum that is coarsely punctate and densely clothed with long, erect pubescence, and the exterior angle of elytral apices are obtusely dentate. The genae of P. skillmani are narrow as in Ischnocnemis and Lophaliamorpha Eya new genus; however, the mandibles are not retracted with sides arcuate from base to apex. The pale brown distal coloration of the antennae is variable, starting from the apex of third or from the fourth, fifth or sixth antennomere.