Hypotrabala maryannae, Friend & Sulak & Beavogui & Müller & Revay & Yakovlev & Saldaitis & Volkova & Prozorova & Prozorov, 2025

Diagnosis. Hypotrabala maryannae sp. n. differs from:

1) H. neavei by lighter background color of forewing (compare Figs 19–24 View Figures 16–24 and 2–9 View Figures 2–9 ), wider socia and narrower valva, smaller processes of the eighth sternite, lack of apical plate on vesica (compare Figs 43–46 View Figures 43–46 and 27–30 View Figures 27–32 );

2) H. lunda by stable background color of male forewing (compare Figs 19–20, 22–23 View Figures 16–24 and 10, 13– 15 View Figures 10–15 ) and less contrastingly colored female forewing (compare Figs 21, 24 View Figures 16–24 and 11–12 View Figures 10–15 ), overall smaller clasping apparatus, smaller socius and valva, about twice shorter processes of eighth sternite (compare Figs 43–46 View Figures 43–46 and 31–34 View Figures 27–32 View Figures 33–35 ), larger sterigma with wider lateral arms, smaller signum (compare Figs 38–39 and 36–37 View Figures 36–39 );

3) H. liviae sp. n. by lighter background color of male forewing (compare Figs 19–20, 22–23 and 16–17 View Figures 16–24 ), wider socia and narrower valva, smaller processes of the eighth sternite, lack of apical plate on vesica (compare Figs 43–46 View Figures 43–46 and 41–42 View Figures 40–42 );

4) H. pearlae sp. n. by less vivid coloration of forewings with more intense dark speckles (compare Figs 19–20, 22–23 and 18 View Figures 16–24 ), overall smaller clasping apparatus, shorter socia and valva, lack of apical plate on vesica, smaller processes of eighth sternite (compare Figs 43–46 View Figures 43–46 and 40 View Figures 40–42 ).

Description. Male ( Figs 19–20, 22–23 View Figures 16–24 ). The flagellum is straw. The head and thorax are mottled, with brown, orangish-straw, and straw scales. The forewing is 16–18 mm long, elongate, somewhat triangular, with a slightly concave costal margin, a blunt apex, and an outer margin with the slightest waviness. The background color is yellow with more or less intense brown or reddish-brown speckles, and straw in the anal area. The pattern consists of a barely pronounced blurred brown basal line, a faint double crenulated brown antemedial and dark-brown postmedial lines, a slightly wavy dashed dark brown subterminal line, an oval silverish medial spot with brown contour, a brown area expanding from under the discal spot towards subterminal line, and between Rs4 and 1A, and dark brown terminal field with mottled area along the subterminal line and mottled veins, covered with yellow and orange scales. The fringe is mottled, with brown and orange scales. The hindwing is somewhat ovoid with a slightly wavy outer margin. The background color is straw or orangish-straw, and mottled, with brown and straw scales, along the costal margin. The fringe is straw or orangish-straw along the anal and outer margins, and brown with sparse orange scales along the costal margin. The abdomen dorsally is orangish-straw. Genitalia ( Figs 43–46 View Figures 43–46 ). The tegumen is a somewhat W-shaped ribbon; it medially bears a pair of elongate digitiform socia with semioval ventral outgrowth. The valva is claw-like, with a pointed apex, about twice as long as a socius. The vinculum is split with juxta into a pair of elongate, somewhat rectangular lobes. The juxta is a ribbon surrounding the phallus. The phallus is claw-like, upwardly bent, with a rounded apex, and dorsally has a wide opening for the vesica. The vesica is bag-like with a pair of longitudinal serrated plates going from the center of the opening towards the apex of the vesica, but not reaching it; two smaller serrated plates are more or less pronounced after a small break separating them from the larger plates. The eighth sternite is a narrow plate with a posterior medial concavity, a pair of heavily sclerotized, nearly straight posterior processes with pointed apices, and lateral elongate apodemes. Female ( Figs 21, 24 View Figures 16–24 ). Reminiscent of the male but overall larger, with a longer forewing of straw background color instead of yellow with more intense brown speckles, a dark brown discal spot instead of the silverish one, and a lack of dark brown area in the subterminal field. The forewing is 24– 25 mm long. The hindwing has a blurred transversal band of brown scales. Genitalia ( Figs 38–39 View Figures 36–39 ). The papilla analis is somewhat oval, densely covered with chaetae, and ventrobasally, near the base of the posterior apophysis, has a wrinkled, sclerotized area. The posterior apophysis is almost twice as long as the anterior one. The sterigma is heavily sclerotized and shield-like, with an anterior cup-like extension near the ostium, which is hidden with a beak-like outgrowth, and lateral arms about half of the sterigma’s length. The ductus bursae is heavily sclerotized, semioval, and flat, about two times smaller than the bursa copulatrix. The bursa copulatrix is membranous, somewhat pear-shaped, and dorsomedially bears a small transversal plate-like signum with lateral minute dents.

Distribution and biology. The type series was collected in November–December and February–March during the wet season within the dry miombo woodlands ecoregion in Zimbabwe ( Olson et al. 2001, Dinerstein et al. 2017).

Etymology. The species is named in honor and in loving memory of Mary Ann Boncelet (Highland Park, NJ, USA). She passed away on May 31, 2025, at the age of 97. Mary Ann was born in McCook, Nebraska. Mrs. Boncelet received her bachelor’s degree from the University of Colorado and a master’s degree from Fairleigh Dickinson University. She taught first grade in the Woodbridge School System for 25 years and retired in 1991. She loved to knit and would make family members beautiful blankets and scarves. Her loving son, Peter, flew back and forth from Naples, Florida, to ensure that she received the best care. She is survived by four children, seven grandchildren, and two great-grandchildren.

Acknowledgments

We thank †John Rawlins (CMNH), †Martin Krüger (DMNH), Bernard Landry and Bernhard Merz (MHNG), †Martin Honey and Geoff Martin (NHMUK), Moira FitzPatrick (NHMZ), †Ugo Dall’Asta (RMCA), and Bert Gustaffson (SNHM) for providing access to the material and helping to process it. We thank †Raymond Murphy (Mzuzu, Malawi), Petr Ustjuzhanin (Novosibirsk, Russia), and Vasily Kovtunovich (Moscow, Russia) for the donation of specimens of the new species, and Thierry Bouyer (Chênée, Belgium) for helping with locating one collection locality in the DRC.

†Vadim Zolotuhin and Alexey Prozorov did a portion of the presented work with the financial support of the Thomas-Witt-Stiftung.

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