Cycas

KEY TO THE SPECIES OF PHILIPPINE CYCAS View in CoL

1. Seeds ribbed.................................. 2

1. Seeds not ribbed............................... 4

2. Seeds with 5 – 6 shallow ribs................ C. wadei View in CoL

2. Seeds with more than 6 prominent ribs.............. 3

3. Leaves 180–260 cm long, petiole 35– 55 cm long, 15– 30 % of entire leaf length; leaflets 150–360; seeds ≥ 45 mm long...................................... C. curranii View in CoL

1 Biology Department, De La Salle University-Manila, Taft Avenue, Manila, The Philippines;

corresponding author e-mail: esperanza.agoo@dlsu.edu.ph.

3. Leaves 160–190 cm long, petiole 40–60 cm long, 25– 35 % of entire leaf length; leaflets 100–380; seeds, ≤ 40 mm long.................................... C. saxatilis View in CoL

4. Megasporophyll lamina entire, without teeth........ 5

4. Megasporophyll lamina with teeth................. 6

5. Megasporophyll lamina not winged shaped; seeds flattened ovoid, 45 –66 by 35–50 mm............. C. edentata View in CoL

5. Megasporophyll lamina winged shaped; seeds obovoid, 36 by 27 mm........................ C. vespertilio View in CoL

6. Leaflets undulating; lateral spines of megasporophyll along triangular top of lamina............. C. sancti-lasallei View in CoL

6. Leaflets not undulating; lateral spines of megasporophyll on the margins of the lamina..................... 7

7. Petiole and rachis very tomentose, leaflets rigid and stiff................................. C. zambalensis View in CoL

7. Petiole and rachis not tomentose, leaflets not rigid and stiff......................................... 8

8. Leaves long, 220– 300 cm; megasporophyll lamina lanceolate.............................. C. aenigma View in CoL

8. Leaves short, less than 200 cm long; megasporophyll lamina not lanceolate.......................... 9

9. Leaves drooping; seeds without spongy endotesta........................................ C. lacrimans View in CoL

9. Leaves not drooping; seeds with spongy endotesta.. 10

10. Apical spine of microsporophyll prominent, 13 –23 mm long; seeds flattened ovoid, spongy endotesta on upper half or at the apical portion of the endosperm, 10–15 mm thick..................................... C. nitida View in CoL

10. Apical spine of microsporophyll short, up to 10 mm; seeds ovoid, spongy endotesta very thin, less than 1 mm, surrounding the endosperm.............. C. riuminiana View in CoL