Ephydra pseudomurina Krivosheina, 1983

Ephydra pseudomurina Krivosheina, 1983 View in CoL

Figs 6 View Figs 5–6 , 12 View Figs 7–15 , 21 View Figs 16–24 , 30 View Figs 25–30 , 39 View Figs 34–39 .

pseudomurina Krivosheina 1983: 368 View in CoL ( Ephydra View in CoL ). Type-locality: Mekhnatabadskiy Distr.   GoogleMaps , Yangier [ 40.276ºN 68.816ºE] ( Uzbekistan).

MATERIAL. Astrakhan’ Oblast: salt lake Baskunchak ( 48.193°N 46.813°E), 17.06.2003, A. Ovchinnikov ( 9 ♂♂, ZISP); same place , 2–4. V .2010 GoogleMaps , K. Tomkovich ( 3 ♂♂, ZMUM); Kharabali   GoogleMaps ( 47.393ºN, 47.248ºE), salt lake, 4. VI .2009 GoogleMaps , P. Mel’nik ( 1 ♂, ZMUM); Crimea: Kerch   GoogleMaps env. ( 45.2ºN 36.1ºE), 26.IV.2014, N. Vikhrev ( 2 ♂♂, 5 ♀, ZMUM); Dzhankoy ( 45.71ºN 34.4ºE) GoogleMaps , 19. V .1963 GoogleMaps , K. Gorodkov ( 1 ♂, ZISP); Koktybel’   GoogleMaps ( 44.964ºN 35.241ºE), 1–8.IX.2007, K. Tomkovich ( 1 ♂, ZMUM); Lake Koyanskoe ( 45.025ºN 36.111ºE), reared, 2008, A. Przhiboro ( 1 ♂, ZISP); Krasnodar Krai: Ubinskaya ( 44.74ºN 38.54ºE) GoogleMaps , 8. VI.1971, V. Kovalev ( 1 ♂, ZMUM); Rostov Oblast: Gigant vill. ( 46.507ºN 41.34ºE), on the light GoogleMaps , 17–22. VI .2009 , A. Koval ( 1 ♂, ZMUM).

ADDITIONAL MATERIAL. Azerbaijan: Avrora [=Hirkan, 38.67ºN 48.799ºE], 19. V .1981 GoogleMaps , N. Krivosheina ( 1 ♂, ZMUM); Georgia: Tiflis [= Tbilisi, 41.71ºN 44.791ºE], 1903, K. Satunin ( 2 ♂♂, ZISP); Kazakhstan: Golodnaya   GoogleMaps step’, 7.VII.1959, P. Lehr ( 1 ♂, ZISP); Tadjikistan: Nature Reserve Tigrovaya balka , 15.IV. 1986, 15–17.V.1987, N. Krivosheina ( 2 ♂♂, ZMUM); Turkmenistan: Ashgabat ( 37.934ºN 58.387ºE) GoogleMaps , 15.XI.1983, N. Krivosheina ( 2 ♂♂, ZMUM); Repetek ( 38.563ºN 63.178ºE) GoogleMaps , 27.XII.1965, K. Gorodkov ( 2 ♂♂, ZISP); same place , 25.I.1966, V . Zaitzev ( 1 ♂, ZISP); Lake Sarygamysh (ca. 42.052º9N 57.697ºE) , 19–26.IV.1984, M. Krivosheina ( 1 ♂, ZMUM); Ukraine: Odessa, Kuyal’nitskiy liman (ca. 46.565ºN 30.717ºE) GoogleMaps , 16.IX.1938, B. Rohdendorf ( 7 ♂♂, ZMUM); same place , 21. VI .1926 GoogleMaps , L. Zimin ( 2 ♂♂, ZISP); Odessa env. (ca. 46.466ºN 30.719ºE), salt puddle near oil tank, 8. V .1926 GoogleMaps , L. Zimin ( 2 ♂♂, ZMUM); Aleshki   GoogleMaps ( 46.62ºN 32.719ºE), 18. VI .1926 GoogleMaps , L. Zimin ( 2 ♂♂, ZISP); Chernomorskiy Nature Reserve (ca. 46.314ºN 31.890ºE), cordon Geroyskiy   GoogleMaps , 6–7.VII.1977, E. Klechkovskiy ( 16 ♂♂, ZMUM); Uzbekistan: Mekhnatabadskiy Distr., Yangier ( 40.276ºN 68.816ºE) GoogleMaps , 12. V.–10. VI .1980 GoogleMaps , M. Krivosheina ( 17 ♂♂, 20 ♀♀, ZMUM); Khiva   GoogleMaps ( 41.394ºN 60.381ºE), 22.III.1927, L. Zimin; Aral Sea, Syr-Dar’inskaya Distr ., 30. V .1928 , N. Olenev, V. Popov ( 1 ♂, ZISP) .

DESCRIPTION. Head golden-grey pollinose except metallic shining frontal vitta and upper face, with bluish-greenish tinge; fronto-orbital plate golden pollinose. 3–4 long fronto-orbital setae, anterior interfrontal thinner and 1/3 as long as ocellar. Postocular setae weak and short. Palpus yellow. Antenna blackish, greyish dusted; arista pectinate on basal half of dorsal surface ( Fig. 6 View Figs 5–6 ). Gena and parafacial greyish dusted.

Thorax black, grey pollinose. Scutum metallic subshining with greenish tinge. Acrostichals setulose in many irregular rows, intra-alars absent.

Legs. Femora yellow, greyish dusted or yellow; tibiae and tarsi of all legs yellow, darkened dorsally.

Abdomen black, grey pollinose, metallic subshining with greenish tinge. Tergite 5 about 1.5–2.0 times as long as tergite 4. Male genitalia large, as long as tergites 4+5. Epandrium elongate in dorsal (posterior) view ( Fig. 30 View Figs 25–30 ); surstylus broad, rounded apically. Epandrium parallel-sided in lateral view ( Fig. 39 View Figs 34–39 ); surstylus with oval median protuberance. Aedeagus with round- ed tip in lateral view ( Fig. 12 View Figs 7–15 ). Gonite stout at base in lateral view, bifurcate apically, of complex shape, giving different images if slightly rotated ( Fig. 21 View Figs 16–24 ).

NOTE. Wirth [1975] described E. murina from the territories of Iraq ( holotype), Bulgaria, Iran, Romania, Turkey and Ukraine. The figures of surstyli dorsally were closer to E. flavip es, relatively broad basally and evenly tapering apically. Krivosheina [1983] described the species E. pseudomurina from Uzbekistan; surstyli of this species are broad almost to the tip and narrowed at the very apex. The same species was described by Hu & Yang [2001], who did not see the previous work, under the name of E. hejingensis Hu & Yang, 2001 . Beschovsky [2009] gave figures of surstyli the same as in E. pseudomurina and determined this species as E. murina . Przhiboro & Shadrin [2012] registered the same species in Russia under the name of E. murina . Krivosheina [2010] obviously had two morphological forms of E. pseudomurina . However we discovered specimens of Ephydra from Tajikistan, which surstyli are closer to the figures given for E. murina by Wirth [1975]. This prevented us from synonymizing pseudomurina , hejingensis and murina in the present work. We leave the solution to this question for the future, when the holotype of E. murina becomes available for study.

DISRTIBUTION. Russia: Astrakhan’ Oblast (incl. Krivosheina [2010]), Crimea (incl. Przhiboro & Shadrin, [2012]; Krivosheina [2014]), Krasnodar Krai, Orenburg Oblast [ Shayhutdinova, Krivosheina, 2020], Rostov Oblast. — Palearctic: Turkmenistan, Uzbekistan [ Krivosheina, 1983, 1986, 2010; Mathis, Zatwarnicki, 1995]; Azerbaijan ( first record), Georgia ( first record), Kazakhstan ( first record), Tadjikistan ( first record), Ukraine ( first record).