Coproica vagans (Haliday, 1833)

Coproica vagans (Haliday, 1833) View in CoL

Figs. 8 View FIGURES 6–8 , 289 View FIGURES 289–291 –298

Borborus vagans Haliday, 1833: 178 View in CoL [both sexes?]. Type locality: Ireland, Downshire , Holywood. STs (NMID).

Limosina opacula Stenhammar, 1855: 389 View in CoL [both sexes]. Type localities: Sweden, " Ostrogothia " and "Gottlandia". LT male, designated by Roháček , 1983e: 159 (UZIU) [no specimen of type series is labelled and, therefore, no definite type locality can be selected on the basis of lectotype].

Limosina albipennis Rondani, 1880: 41 View in CoL [both sexes?]. Type locality: Italy, nr. Parma. STs (MZUF).— Duda, 1918: 210 [synonymy].

Limosina (Coprophila) vagans var. flava Duda, 1918: 210 [male, illustr.]. Type locality: Tunisia, Gafsa. STs males (ZMHB).— Papp, 1979 a: 104 [synonymy].

Limosina vagans var. rufa View in CoL .—Collin, 1949: 201 [lapsus].— Richards, 1960 c: 205 [synonymy].

Description. Body length 1.3–2.3 mm. Body brown. Face light brown, frons brown. Interfrontal bristles in 4–5 subequal pairs, postocellar bristles convergent, 1 pair of small postocellar setulae. Eye small, eye to gena ratio 1.3:1–1.5:1; 6 genal setae, anterior seta 2 times as long as posterior bristles.

Scutellum with 20–25 discal setae arranged in 3 sparse rows between basal 1/5 and basal 3/5, posterior row of setae approximately 1.5 times as long as other discal setae; 3 lateral setae between larger anterior and posterior marginal scutellar bristles. Katepisternum with 2 long, subequal dorsal bristles, anterior and posterior bristles each approximately 3/5 and 4/5 as long as distance between dorsal margin of katepisternum and wing base respectively. Legs and fore coxa brown to light brown. Mid tibia with small additional anterodorsal bristle at 1/10, and proximal posterodorsal bristles present at 1/4 and 1/2. Mid bastitarsus with strong anterior setae sub-basally and at 1/2, sub-basal ventral pair of bristles present, posteroventral sub-basal bristle approximately 1.3 times as long as anteroventral bristle present, anteroventral bristle at 2/5 and approximately 2 times longer than width of mid basitarsus, anteroventral bristle at 4/5, and small posteroventral bristles at 2/5 and 3/5. Second costal sector 1.1–1.7 times as long as third costal sector. Costal bypass short, terminating within 5 vein widths of R 4+5.

Male terminalia: Sternite 5 rectangular, discal bristles larger laterally. Sternite 6 straight medially. Epandrium with enlarged pair of dorsolateral bristles; subanal plate present, narrow; cercus with 1 outstanding bristle, ventral margin with broad rounded medial processes. Surstylus with 2 small dorsally curved processes separated by an emargination, 3 or more enlarged anteroventral bristles, and several small bristles along ventral margin. Basiphallus L-shaped, open posteriorly; epiphallus short. Distiphallus: lateral sclerites rounded, with blunt subapical dorsal process, an apicoventral posteromedially deflected process; central sclerite fused with lateral sclerites on apicoventral process; membranous sheets broad.

Female terminalia: Tergite 8 long laterally and short dorsally, not divided medially. Tergite 10 pollinose, narrowly fused to cerci laterally; cerci conical, pollinose, apical bristles sinuate. Sternite 7 broad, rounded. Sternite 8 broad, rounded, width approximately 3/4 width of sternite 7, with pair of lateral, elongate bristles. Sternite 10 with medial and lateral pairs of large bristles, posterior margin not curved dorsally. Spermathecae spherical; duct length approximately 2 times as long as spermathecal bulb.

Type material. Syntypes: Ireland: Downshire ( NMID, not examined) .

Material examined. See Appendix A.

Distribution: Afrotropical: Ethiopia, South Africa, Tanzania, Zaire; Australasian/Oceanian: Australia (QLD), Guam (USA), Hawaii (USA); Nearctic: Canada (AB, BC, NS, ON, SK) , USA (AR, AZ, CA, CO, CT, DC, FL, ID, MI, MO, NC, NJ, NM, OK, OR, PA, SD, TX, UT, WA, WY); Neotropical: Argentina, Bermuda, Bolivia, Chile, Costa Rica, Jamaica, Mexico (BCS, MEX); Oriental: Japan (Okinawa), Taiwan; Palaearctic: Afghanistan, Algeria, Andorra, Austria, Azerbaidjan, Belgium, Bulgaria, Canary Is. (Spain), Cyprus, Czech Republic, Denmark, Egypt, Finland, France (incl. Corsica), Georgia, Germany, Great Britain, Greece (incl. Crete, Thíra), Hungary, Iceland, Ireland, Israel, Italy (incl. Sardinia, Sicily), Japan, Latvia, Macedonia, Malta, Mongolia, Netherlands, Norway, Poland, Romania, Russia (CET, NET, SET), Saudia Arabia, Serbia, Slovakia, Spain (incl. Balearic Is.), Sweden, Switzerland, Tadjikistan, Tunisia, Turkey, United Arab Emirates, South Atlantic: Saint Helena.

Comments. We were unable to examine the type specimens of C. vagans but Richards’ (1960) concept of this species is clear and is followed here.

18192021222324252627282930313233 C. bispatha 0 0 0 0 1 0 1 1 0 1 0 0 0 0 2 1 C. brachystyla 0 0 0 0 1 0 1 1 0 1 1 0 0 0 2 1 C. cacti ???? 0?? 1 0 2 1 0 0 0 2 1 C. coreana 0 0 0 0 1 0 0 1 1 1 0 0 0 1 0 0 C. diabolica 1 0 0 1 1 0 2 1 0 1 1 0 0 0 1 0 C. digitata 1 0 1 1 1 0 2 1 1 1 1 0 1 1 0 0 C. emarginata 0 0 0 0 1 1 0 1 0 1 1 0 0 0 1 0 C. ferruginata 0 0 0 0 1 1 1 1 0 1 0 0 0 1 1 0 C. galapagosensis 0 0 0 0 0 1 0 1 0 2 0 0 0 1 0 0 C. hirticula 1 0 0 0 1 0 0 1 0 1 0 0 0 1 0 0 C. hirtula 0 0 0 0 1 1 0 1 0 3 0 0 0 1 1 0 C. hirtuloidea 0 0 0 0 1 1 0 1 0 0 1 0 0 0 2 0 C. lacteipennis 0 0 0 0 1 0 0 1 0 3 0 0 0?? 0 C. lugubris 0 1 1 0 1 1 4 1 1 0 0 1 0 1 2 0 C. mitchelli 1 0 1 1 1 1 4 1 1 1 0 1 0 1 0 0 C. novacula 1 0 0 1 1 1 1 1 0 1 0 0 0 1 2 0 C. pappi 0 0 1 1 1 1 3 1 1 2 0 0 0 1 2 0 C. pusio 0 0 1 1 0 1 4 1 1 0 0 1 0 1 0 0 C. rohaceki 0 1 0 0 1 0 1 1 0 3 0 0 0 1 0 0 C. rufifrons 0 0 0 0 1 0 0 1 0 3 0 0 0 1 0 0 C. serra ???? 1?? 1 1 0 0 0 0 1 0 0 C. setulosa 0 0 0 0 1 1 1 1 0 1 0 0 0 1 2 0 C. testudinea 0 0 0 0 1 1 1 1 0 1 0 0 0 1 2 0 C. urbana 0 0 0 0 1 0 1 1 0 1 1 0 0 0 2 1 C. vagans 0 0 0 0 1 1 0 1 0 0 0 0 0 1 0 0 3435363738394041424344 C. emarginata 0 0 0 0 1 0 0 2 2 0 0 C. ferruginata 1 1 0 1 0 0 0 2 0 0 0 C. galapagosensis 0 0 0 0 0 0 1 2 2 1 0 C. hirticula 0 0 2 0 0 0 0 1 0 1 0 C. hirtula 0 0 0 1 0 0 0 2 2 0 0 C. hirtuloidea 0 0 0 0 0 0 0 2 1 1 0 C. lacteipennis 0 0??? 0? 2 1 1 0 C. lugubris 0 0 0 0 0 0 0 1 0 0 0 C. mitchelli 0 0 0 0 0 1 0 1 0 0 0 C. novacula 0 0 0 1 0 0 0 1 0 1 0 C. pappi 0 0 0 0 0 0 0 1 0 0 0 C. pusio 0 0 0 0 0 1 0 2 1 0 0 C. rohaceki 1 1 0 0 1 0 0 1 0 0 0 C. rufifrons 0 0 0 1 0 0 1 1 0 1 0 C. serra 0 0??? 0????? C. setulosa 1 1 0 0 1 0 1 1 0 0 0 C. testudinea 0 0 0 0 1 0 0 2 1 1 0 C. urbana 0 0 3 0 1 0 1 1 0 1 1 C. vagans 0 0 0 1 0 0 0 1 0 0 0