Rhytomus sodalis Saxton & Génier, 2025

Génier, François, Gunter, Nicole L. & Saxton, Natalie A., 2025, Rhytomus Génier & Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae), Zootaxa 5575 (1), pp. 87-110 : 104-106

publication ID

https://doi.org/10.11646/zootaxa.5575.1.3

publication LSID

lsid:zoobank.org:pub:521D404D-EB9D-4110-989E-A8A273C10152

DOI

https://doi.org/10.5281/zenodo.14808927

persistent identifier

https://treatment.plazi.org/id/038987E8-FFE6-6E08-FF46-15C0FDEAFD7D

treatment provided by

Plazi

scientific name

Rhytomus sodalis Saxton & Génier
status

sp. nov.

Rhytomus sodalis Saxton & Génier , new species

urn:lsid:zoobank.org:act:7168813D-24AF-4CCA-8B46-B5168014A9E3

Figs. 7H View FIGURE 7 , 8H View FIGURE 8 , 10C View FIGURE 10 , 12C View FIGURE 12

Name-bearing type data. Holotype ♂ (CMNC): || PAPUA NEW GUINEA: Western | Prov., P’nvang , CI RAP Survey | Camp 1. 05º29.5S 141º32.6E | 573 m. May 2013. T. Larsen ||; || Canadian Museum of | Musée canadien de la | NATURE | CMNEN 00049583 || [QR code label]; || HOLOTYPE ♂ | Rhytomus | sodalis | Des. N. Saxton & F. Génier, 2024 || [red card] GoogleMaps .

Material examined (69 ♂♂, 24 ♀♀). PAPUA NEW GUINEA: WESTERN PROVINCE, CI RAP Survey Camp 1, P’nvang, 575 m, 05°29’S, 141°32’E, v.2013, T. Larsen — 2 ♂♂, 1 ♀ paratypes ( ANIC), GoogleMaps 2 ♂♂, 1 ♀ paratypes ( NHMUK), GoogleMaps ♂ holotype, ♀ allotype GoogleMaps , 23 ♂♂, 12 ♀♀ paratypes ( CMNC), GoogleMaps 2 ♂♂, 1 ♀ paratypes ( MNHN), GoogleMaps 2 ♂♂, 1 ♀ paratypes ( NMPC), GoogleMaps 1 ♂, 1 ♀ paratypes ( USNM); GoogleMaps CI RAP Survey Camp 2 , P’nvang , 800 m, 05°30’S, 141°34’E, v.2013, T. Larsen — 15 ♂♂, 3 ♀♀ paratypes ( CMNC); GoogleMaps CI RAP Survey Camp 3 , P’nvang , 1110 m, 05°32’S, 141°33’E, v.2013, T. Larsen — 19 ♂♂, 1 ♀ paratypes ( CMNC), GoogleMaps 2 ♂♂, 2 ♀♀ paratypes ( QM) GoogleMaps .

Description. Measurements. Body length 3.0– 4.6 mm, maximum body width 2.1–2.7 mm. Holotype: body length 3.8 mm, maximum body width 2.5 mm. Body. Overall body shape in dorsal view elongate-oval, elytral edge slightly subangular at apical third. Head. Clypeal edge lateral to clypeal teeth lacking emargination, head margin at clypeogenal suture of females slightly protruding. Dorsal parts of eyes semi-ovoid, separated by an interocular space of about 3.2 times eye width. Pronotum. Pronotal width to length along midline ratio 1.7, lateral pronotal edge sharply defined throughout, pronotal pubescence short and fine, pronotal basal portion along midline lacking fine longitudinal depression. Elytra. Elytral maximum width to length along suture ratio 1.0, elytral outline moderately convex in lateral view, elytral pubescence long and conspicuous, elytral surface between punctures appearing glossy, at most with ill-defined microsculptures, basal elytral pit wide and deep, basal surface of pseudepipleuron with irregularly distributed more or less defined punctures, interstriae 3–5 on basal fifth with two rows of punctures along striae, with at most a few scattered punctures in some individuals. Legs. Profemoral anterior surface flat or might appear furrowed in some individuals, male metatibial apicointernal tooth approximately in line with posterior tibial edge ( Fig. 12C View FIGURE 12 ). Pygidium. Pygidial surface along midline with punctures similar in density to rest of discs, pygidial surface along basal groove with coarse transverse microsculptures ( Fig. 8H View FIGURE 8 ), pygidium of female depressed basally. Aedeagus. Left paramere apex truncated apically, distinctly angular externally ( Fig. 10C View FIGURE 10 ). Female genitalia. Spermatheca with two apodemes basally and one apodeme apically.

Etymology. The specific epithet of this species is a masculine noun in apposition “sŏdālis ” (companion, mate, fellow) relating to the fact that it was collected along with R. larseni and R. latidentatus .

Remarks. One female specimen (excluded from type series) was discovered from the same collecting event (i.e., Camp 1, 575 m) that had coarser pronotal punctation and a barely indicated meso-metaventrite suture. This specimen likely represents a fourth species. However, because no male specimens could be found at the time of this study, we refrain from describing it.

ANIC

Australia, Australian Capital Territory, Canberra City, CSIRO, Australian National Insect Collection

CMNC

Canada, Ottawa, Canadian Museum of Nature

MNHN

France, Paris, Museum National d'Histoire Naturelle

NMPC

Czech Republic, Prague, National Museum (Natural History)

USNM

USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]

QM

Australia, Queensland, South Brisbane, Queensland Museum

ANIC

Australian National Insect Collection

NHMUK

Natural History Museum, London

MNHN

Museum National d'Histoire Naturelle

NMPC

National Museum Prague

USNM

Smithsonian Institution, National Museum of Natural History

QM

Queensland Museum

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Scarabaeidae

SubFamily

Scarabaeinae

Genus

Rhytomus

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