Aname eddieorum, Wilson & Harvey & Simmons & Rix, 2025

Aname eddieorum sp. nov.

urn:lsid:zoobank.org:act:49D83F5A-9ED3-4B4F-A49B-B07BEBC0F0C2

Figs 1 View Fig , 7 View Fig , 39–40 View Fig View Fig

Aname barrema Raven, 1985 – Raven 1985 (pars): figs 13, 33, 70–71, 73–74 (illustrated female allotype QMB S1239 [Braemar State Forest], and female paratypes QMB S1244 [Moombah], QMB S1247 [Yuleba], and QMB S1245 [Stanthorpe], all here identified as A. eddieorum sp. nov.).

Diagnosis

Males of A. eddieorum sp. nov. can be distinguished from all species for which males are known except A. aurensis sp. nov., A. briggsi sp. nov., A. hughenden sp. nov., A. longitheca , A. mulgana sp. nov., A. rupicola sp. nov., and A. warrego sp. nov. by a moderate to large body size (carapace length> 4.0 mm), the presence of a long embolus (embolus length / bulb length>1.5) that has a relatively wide basal section tapering into an attenuate, sinuous distal section after about 0.4 of length, and the absence of a prominent sharp heel on metatarsus I (as in the pallida -complex) ( Fig. 39L–M View Fig ). Males of A. eddieorum can be distinguished from those of A. aurensis , A. briggsi , A. hughenden , A. mulgana , and A. rupicola by the presence of a tibia I that stays about the same width from the proximal end to the base of the tibial spur when in lateral view ( Fig. 39P View Fig ; cf. Figs 34 View Fig , 36 View Fig , 41 View Fig , 44 View Fig , 46 View Fig ). Males of A. eddieorum can be distinguished from those of A. warrego by the presence of a thicker palp tibia (palp tibia length / width <3) ( Fig. 39J–K View Fig ; cf. Fig. 48 View Fig ). Males of A. eddieorum can be distinguished from those of A. longitheca by the presence of a shorter, straighter embolus (embolus length / bulb length ~2.2; cf. ~ 2.7 in A. longitheca ) ( Fig. 39L–M View Fig ; cf. Fig. 42 View Fig ).

Females of A. eddieorum sp. nov. can be distinguished from all species for which females are known except A. aurensis sp. nov., A. briggsi sp. nov., A. camara , A. dingo sp. nov., A. longitheca , A. mulgana sp. nov., and A. rupicola sp. nov. by the presence of spermathecae with two vesicles, with relatively long and straight lateral vesicles (lateral vesicle length / genitalia width>0.25) and long medial vesicles (medial vesicle length / lateral vesicle length>1) that project medially or posteromedially, before undulating anteriorly ( Fig. 40L View Fig ). Females of A. eddieorum can be distinguished from those of A. aurensis , A. briggsi , A. dingo , A. longitheca , and A. rupicola by the presence of spermathecae with lateral vesicles terminating in relatively wide ends ( Fig. 40L View Fig ; cf. Figs 35 View Fig , 37–38 View Fig View Fig , 43 View Fig , 47 View Fig ). Females of A. eddieorum can be distinguished from those of A. camara by the presence of a darker body colouration and spermathecae with longer vesicles (lateral vesicle length / genitalia width>0.35) with less widely-spaced crowns (distance between crowns less than length of lateral vesicles) ( Fig. 40A–L View Fig ; cf. Fig. 107 View Fig ). Females of A. eddieorum can be distinguished from those of A. mulgana by the presence of bald patches on the sternum, lateral of the sigilla, the absence of thorn-like setae around the anterior edges of the sternum, and more rounded medioventral corners of the coxae ( Fig. 40G–I View Fig ; cf. Fig. 45 View Fig ).

Etymology

The specific epithet ‘ eddieorum ’ is named in honour of Craig and Merryl Eddie, founders of Boobook Ecological Consulting. The company provided many valuable specimens for this project.

Type material

Holotype

AUSTRALIA – Queensland • ♂; Kumbarilla State Forest, Halliford Road ; 27°29′ S, 150°54′ E; 326m a.s.l.; 4 Nov.–15 Dec. 2019; G.B. Monteith leg.; gutter trap, sandy open forest; QMB S111185 View Materials .

GoogleMaps

Paratypes

AUSTRALIA – Queensland • 2 ♂♂; Tara ; 27°16′ S, 150°27′ E; 20 Nov. 2001; Queensland Department of Primary Industries leg.; QMB S116964 View Materials • 1 ♂; Lake Broadwater Conservation Park ; 27°21′ S, 151°06′ E; 24 Nov. 1985 – 3 Jan. 1986; Queensland Museum Party leg.; pitfall trap; QMB S3151 View Materials GoogleMaps • 1 ♂; Lake Broadwater Conservation Park , south-western loop track; 27°22′ S, 150°34′ E; 342 m a.s.l.; 27 Jul. 2020; M.G. Rix leg.; excavated, mixed bulloak/cypress pine woodland; QMB S124044 View Materials GoogleMaps • 1 ♂; Windemere Station ; 27°25′ S, 149°41′ E; 5 Dec. 1987; R.J. Raven leg.; hand collected, on ground in low vegetation, brigalow; QMB S2476 View Materials GoogleMaps .

GoogleMaps

Other material examined

AUSTRALIA – Queensland • 1 ♂; Expedition Range National Park ; 25°12′ S, 148°59′ E; 560 m a.s.l.; 25 Sep.–18 Dec. 1997; D.J. Cook and G.B. Monteith leg.; intercept trap, open forest; QMB S44341 GoogleMaps • 1 ♂; Expedition Range National Park, Ampitheatre camp; 25°12′ S, 148°59′ E; 560 m a.s.l.; 17–19 Dec. 1997; D.J. Cook, G.B. Monteith and G. Thompson leg.; open forest; QMB S59368 GoogleMaps • 1 ♀; Kentucky Station, tributary of Baffle Creek , 44.3 km NNE of Injune; 25°34′ S, 148°53′ E; 14 Mar. 2021; C. Eddie and E. Amsters leg.; excavated, short, freshly dug burrow on sloping bank of rocky gorge, dry sclerophyll; QMB S118232 View Materials GoogleMaps • 1 ♂; Munduberra, Delubra Station ; 25°35′ S, 151°17′ E; 20 Oct. 1982; B. Bloxsome leg.; QMB S9757 View Materials GoogleMaps • 1 ♀; Strathblane ; 25°53′ S, 149°08′ E; 7 Feb. 2018; C. Eddie leg.; QMB S118224 View Materials GoogleMaps • 1 ♂; Lorraine Station , 35.3 km NNW of Roma; 26°10′ S, 148°41′ E; 12 Oct. 2022; J. Groat leg.; hand collected, active around house after rain, property homestead, sandy soil; QMB S118246 View Materials GoogleMaps • 2 ♂♂, 1 juv.; Barakula State Forest , S of Condarra Tower; 26°17′ S, 150°38′ E; 1 Oct.–3 Dec. 2012; C. Moeseneder and S. Moeseneder leg.; flight intercept trap; QMB S109552 View Materials GoogleMaps • 1 ♂; Barrakeela forestry [Barakula State Forest], via Chinchilla; 26°19′ S, 150°30′ E; 15 Dec. 1981; B.E. Smith leg.; QMB S9394 View Materials GoogleMaps • 1 ♂; Combabula State Forest ; 26°22′ S, 149°27′ E; 350 m a.s.l.; 5 Nov.–16 Dec. 2019; G.B. Monteith leg.; gutter trap, sandy open forest; QMB S111195 View Materials GoogleMaps • 1 ♂; Barakula State Forest, Auburn Road ; 26°23′ S, 150°41′ E; 30 Sep.–3 Dec. 2012; C. Moeseneder and S. Moeseneder leg.; flight intercept trap; QMB S108715 View Materials GoogleMaps • 1 ♀; Gurulmundi State Forest , ca 39.9 km NW of Miles, SEQ; 26°24′ S, 149°54′ E; 12 Sep. 2021; E. Amsters leg.; excavated, Arbanitis burrows observed in vicinity, sclerophyll forest, pale grey/brown loamy clay soil; QMB S118238 View Materials GoogleMaps • 1 ♂; Barakula State Forest, off Auburn Road ; 26°25′ S, 150°41′ E; 3 Dec. 2012 – 19 Apr. 2013; C. Moeseneder and S. Moeseneder leg.; flight intercept trap; QMB S118361 View Materials GoogleMaps • 1 ♂; Kingaroy, Gordonbrook Dam ; 26°27′ S, 151°44′ E; Nov. 1980; T. McAleer leg.; QMB S9368 View Materials GoogleMaps • 1 ♂; Kingaroy ; 26°31′ S, 151°46′ E; 24 Dec. 1986; K.J.M. leg.; QMB S64335 GoogleMaps • 1 ♀; Yuleba ; 26°37′ S, 149°23′ E; 19 Nov. 1957; M. Green leg.; QMB S1247 View Materials (paratype of Aname barrema Raven, 1985 ) GoogleMaps • 1 ♂; Condamine ; 26°38′ S, 149°56′ E; 27 Jun. 1985; P.G. Allsopp leg.; QMB S100538 View Materials GoogleMaps • 1 ♂; Miles ; 26°40′ S, 150°11′ E; Nov. 1973; G. May leg.; QMB S208 View Materials (paratype of Aname barrema Raven, 1985 ) GoogleMaps • 1 ♂; Miles ; 26°43′ S, 150°05′ E; Nov. 1973; G. May leg.; hand collected, wandering in grassland, grassland; QMB S208 View Materials GoogleMaps • 1 ♂; Chinchilla , “ Rockwood ”; 26°45′ S, 150°36′ E; Dec. 1985; W.D. McKenzie leg.; QMB S9799 View Materials GoogleMaps • 1 ♂; Condamine Highway ; 26°53′ S, 149°28′ E; 248 m a.s.l.; 6 Nov.–17 Dec. 2019; G.B. Monteith leg.; gutter trap, sandy open forest; QMB S111184 View Materials GoogleMaps • 1 ♀; Braemar State Forest, off Kumbarilla Lane ; 27°10′ S, 150°55′ E; 354 m a.s.l.; 4 Oct. 2020; M.G. Rix, A.G. Rix, A. Wojcieszek and M. Brien leg.; excavated, open woodland with cypress pine; QMB S124056 View Materials GoogleMaps • 1 ♀; Braemar State Forest ; 27°13′ S, 150°50′ E; 15–19 Oct. 1979; R. J. Raven leg.; excavated; QMB S1239 View Materials (allotype of Aname barrema Raven, 1985 ) GoogleMaps • 1 ♂; Toowoomba ; 27°34′ S, 151°51′ E; Mar. 1969; T. Passlow leg.; QMB S96436 GoogleMaps • 1 ♀; Moombah [E of St George]; 27°59′ S, 149°18′ E; 11 Feb. 1979; T. Adams and G.V. Czechura leg.; QMB S1244 View Materials (paratype of Aname barrema Raven, 1985 ) GoogleMaps • 1 ♂; Goondiwindi , N of town; 28°25′ S, 150°13′ E; D. O’Donoghue leg.; QMB S103498 View Materials GoogleMaps • 1 ♂; Inglewood ; 28°25′ S, 151°05′ E; 1979; Q. A. T. B. leg.; QMB S9742 View Materials GoogleMaps • 1 ♂; Stanthorpe ; 28°37′ S, 151°30′ E; 6 Nov. 1982; Queensland Ambulance Transport Brigade leg.; QMB S9744 View Materials GoogleMaps • 1 ♀; Stanthorpe ; 28°39′ S, 151°56′ E; 10 Nov. 1971; G. Smith leg.; QMB S1245 View Materials (paratype of Aname barrema Raven, 1985 ) GoogleMaps . – New South Wales • 1 ♂; Inverell ; 29°46′ S, 151°07′ E; 29 Dec. 1980; C. Easton leg.; AMS KS6463 GoogleMaps • 1 ♂; 100 km E of Walgett, near Yallambee ; 30°02′ S, 148°52′ E; 7 Oct. 1996; R.W. Blanch leg.; AMS KS49115 GoogleMaps .

Description

Male (holotype, QMB S111185)

GENERAL ( Fig. 39A–Q View Fig ). Body length 23.28, in good condition, colour presumably faded due to preservation.

DORSAL PROSOMA ( Fig. 39A, E–F View Fig ). Carapace length 8.17, width 7.52, length/width 1.09, clypeus to fovea length/carapace length 0.70, caput width/carapace width 0.74, carapace dark red-brown, reflective setae present, heavy on caput, moderate on thorax, fovea procurved, fovea width/carapace length 0.16 ( Fig. 39A, F View Fig ); chelicerae dark red-brown, rastellum absent or inconspicuous, chelicerae length/carapace length 0.64 ( Fig. 39A View Fig ); eye group rectangular, width/length 2.08, eye tubercle present ( Fig. 39E View Fig ).

ABDOMEN ( Fig. 39B, D View Fig ). Abdomen length 9.79, dark grey, dorsal pattern absent, with consistent cover of short setae.

VENTRAL PROSOMA ( Fig. 39C, G–I View Fig ). Labium cuspules present, count =2 ( Fig. 39H View Fig ); maxillae heel distinct, cuspules present, count = about 130, extending posteriorly onto heel, extending laterally about 40% of maxillae length ( Fig. 39C, I View Fig ); coxae cuspules absent, thorn-like setae on prolateral face present ( Fig. 39C, I View Fig ); sternum length/width 1.27, central sternum with consistent covering of moderate setae, row of longer setae around posterior edges, distinct bald patches laterally of all sigilla ( Fig. 39G–H View Fig ); posterior sigilla ovoid, central sternum to posterior sigilla length/sternum length 0.19, posterior sigilla length/sternum length 0.18 ( Fig. 39G–H View Fig ); other sigilla small, round and lateral ( Fig. 39G–H View Fig ).

LEG I ( Fig. 39N–Q View Fig ). Leg I red-brown, lighter on distal metatarsus and tarsus, femur length 7.11, patella length 4.51, tibia length 4.84, metatarsus length 4.74, tarsus length 2.89, total length 24.10, leg I length/ carapace length 2.95 ( Fig. 39N–O View Fig ); scopulae on distal metatarsus and tarsus ( Fig. 39N–O View Fig ); spine count Fe D 0, Fe PL 1, Pa PL 2, Ti PL 0, Ti RL 0, Me PL 0, Me RL 0, Ta 0 ( Fig. 39N–O View Fig ); tibia length/width [TIL/TID] 3.23, even width along length, spur present, digitiform, knuckle present, megaspine angled at 13 degrees, length to distal face of spur/tibia length [TIS/TIL] 0.43, spur height/tibia width [TISH/TID] 0.65, megaspine length/tibia length 0.22 ( Fig. 39N–P View Fig ); metatarsus slightly sinuous, proximal excavation present, excavation concave with pronounced heel, heel broadly rounded, excavation length/metatarsus length [MIPEL/MIL] 0.46, metatarsus length/width [MIL/MID] 3.54 ( Fig. 39N–O, Q View Fig ).

PEDIPALP ( Fig. 39J–M View Fig ). Tibia length 3.52, width 1.44, length/width [PTL/PTD] 2.44, asetose depression present, depression length/palp tibia length [PDL/PTL] 0.56, retrolateral face with consistent covering of light setae, ventral face with two elongate bristle-like setae below depression, prolateral face with two disto-ventral spines, disto-medial spine absent ( Fig. 39J–K View Fig ); patella prolateral face with 2 spines ( Fig. 39J–K View Fig ); cymbium with scopulae present distally ( Fig. 39J–K View Fig ); copulatory organ total length 2.17, length/palp tibia length 0.62 ( Fig. 39L–M View Fig ); bulb length/width 0.87 ( Fig. 39L–M View Fig ); embolus tapering from bulb, attenuate, with wide base tapering about halfway along into attenuate apical section, one slight bend, at about 0.4 of length, slight bend before tip, width at base/bulb width 0.29, embolus length/ bulb length 2.22 ( Fig. 39L–M View Fig ).

Female (QMB S118238)

GENERAL ( Fig. 40A–L View Fig ). Body length 22.51, in good condition.

DORSAL PROSOMA ( Fig. 40A, E–F View Fig ). Carapace length 9.70, width 8.15, length/width 1.19, clypeus to fovea length/carapace length 0.72, caput width/carapace width 0.80, carapace orange-brown, caput slightly darker than thorax and clypeus darker again, reflective setae present, moderate on caput, light on thorax, fovea procurved, fovea width/carapace length 0.16 ( Fig. 40A, F View Fig ); chelicerae dark red-brown, rastellum absent or inconspicuous, chelicerae length/carapace length 0.53 ( Fig. 40A View Fig ); eye group rectangular, width/length 1.8, eye tubercle present ( Fig. 40E View Fig ).

ABDOMEN ( Fig. 40B, D View Fig ). Abdomen length 8.14, brown, dorsal pattern absent, with consistent cover of short setae.

VENTRAL PROSOMA ( Fig. 40C, G–I View Fig ). Labium cuspules absent ( Fig. 40H View Fig ); maxillae heel distinct, cuspules present, count =about 110, extending posteriorly onto heel, extending laterally about 40% of maxillae length ( Fig. 40C, I View Fig ); coxae cuspules absent, thorn-like setae on prolateral face present ( Fig. 40C, I View Fig ); sternum length/width 1.14, central sternum with consistent covering of moderate setae, row of longer setae around posterior edges, distinct bald patches laterally of all sigilla ( Fig. 40G–H View Fig ); posterior sigilla semi-elongate, central sternum to posterior sigilla length/sternum length 0.17, posterior sigilla length/ sternum length 0.20 ( Fig. 40G–H View Fig ); other sigilla small, round and lateral ( Fig. 40G–H View Fig ).

LEG I ( Fig. 40J–K View Fig ). Leg I orange-brown, femur length 5.13, patella length 3.18, tibia length 3.53, metatarsus length 3.43, tarsus length 2.18, total length 17.45, leg I length/carapace length 1.80; scopulae on distal metatarsus and tarsus; spine count Fe D 0, Fe PL 1, Pa PL 2 (both weak), Ti PL 1, Ti RL 4 (weak), Me PL 0, Me RL 2, Ta 0; tibia length/width [TIL/TID] 3.30.

GENITALIA ( Fig. 40D, L View Fig ). Epigastric furrow unmodified ( Fig. 40D View Fig ); spermathecae with two vesicles each ( Fig. 40L View Fig ); lateral vesicle relatively straight, length 0.73, lateral vesicle length/genitalia width 0.42, length/width at base 2.08, crown slightly wider than stem ( Fig. 40L View Fig ); medial vesicle with distinct basal section angled medially, before undulating anteriorly, medial vesicle length/genitalia width 0.57, length/ width 5.15, medial vesicle length/lateral vesicle length 1.36 ( Fig. 40L View Fig ).

Distribution and natural history

Aname eddieorum sp. nov. has a broad distribution in south-eastern Queensland and northern New South Wales, predominantly in the Brigalow Belt South bioregion. It extends from around Walgett and Inverell north to Eidsvold, and from around Roma in the west to Toowoomba in the east ( Fig. 7 View Fig ). It constructs an open, silk-lined burrow without silk outside of the entrance, often on an angle, and with a hidden secondary ‘wishbone’ entrance ( Fig. 7 View Fig ).

Remarks

The female allotype of A. barrema , assigned by Raven (1985), as well as several imaged specimens identified as A. barrema in that study, actually represent this species.