Aname longitheca Raven, 1985

Aname longitheca Raven, 1985 View in CoL

Figs 1 View Fig , 7 View Fig , 42–43 View Fig View Fig

Aname longitheca Raven, 1985: 402 View in CoL , figs 16, 28, 65.

Diagnosis

Males of A. longitheca can be distinguished from all species for which males are known except A. aurensis sp. nov., A. briggsi sp. nov., A. eddieorum sp. nov., A. hughenden sp. nov., A. mulgana sp. nov., A. rupicola sp. nov., and A. warrego sp. nov. by a moderate to large body size (carapace length>4.0 mm), the presence of a long embolus (embolus length/ bulb length>1.5) that has a relatively wide basal section tapering into an attenuate, sinuous distal section after about 0.4 of length, and the absence of a prominent sharp heel on metatarsus I (as in the pallida -complex) ( Fig. 42L–M View Fig ). Males of A. longitheca can be distinguished from those of A. aurensis , A. briggsi , A. hughenden , A. mulgana , and A. rupicola by the presence of a tibia I that stays about the same width from the proximal end to the base of the tibial spur when in lateral view ( Fig. 42P View Fig ; cf. Figs 34 View Fig , 36 View Fig , 41 View Fig , 44 View Fig , 46 View Fig ). Males of A. longitheca can be distinguished from those of A. warrego by the presence of a thicker palp tibia (palp tibia length /width <3) ( Fig. 42J–K View Fig ; cf. Fig. 48 View Fig ). Males of A. longitheca can be distinguished from those of A. eddieorum by the presence of a longer, more curved embolus (embolus length / bulb length ~2.7; cf. ~ 2.2 in A. eddieorum ) ( Fig. 42L–M View Fig ; cf. Fig. 39 View Fig ).

Females of A. longitheca can be distinguished from all species for which females are known except A. aurensis sp. nov., A. briggsi sp. nov., A. camara , A. dingo sp. nov., A. eddieorum sp. nov., A. mulgana sp. nov., and A. rupicola sp. nov. by the presence of spermathecae with two vesicles, with relatively long and straight lateral vesicles (lateral vesicle length / genitalia width>0.25) and long medial vesicles (medial vesicle length / lateral vesicle length>1) that project medially or posteromedially, before undulating anteriorly ( Fig. 43L View Fig ). Females of A. longitheca can be distinguished from those of A. eddieorum and A. mulgana by the presence of spermathecae with lateral vesicles terminating in narrower ends ( Fig. 43L View Fig ; cf. Figs 40 View Fig , 45 View Fig ). Females of A. longitheca can be distinguished from those of A. aurensis and A. dingo by the presence of spermathecae with long medial vesicles (medial vesicle length/ genitalia width> 0.6) and dark patches of cuticle on the sternum, surrounding and lateral of the sigilla ( Fig. 43G–I, L View Fig ; cf. Figs 35 View Fig , 38 View Fig ). Females of A. longitheca can be distinguished from those of A. camara by the presence of a darker body colouration and spermathecae with longer vesicles (lateral vesicle length / genitalia width> 0.35) with less widely-spaced crowns (distance between crowns less than length of lateral vesicles) ( Fig. 43A–L View Fig ; cf. Fig. 107 View Fig ). Females of A. longitheca can be distinguished from A. briggsi by the presence of spermathecae with less elongate lateral and medial vesicles (medial vesicle length /genitalia width <0.65) ( Fig. 43L View Fig ; cf. Fig. 37 View Fig ). Females of A. longitheca can be distinguished from those of A. rupicola by the presence of spermathecae with straighter lateral vesicles ( Fig. 43L View Fig ; cf. Fig. 47 View Fig ).

Type material

Holotype

AUSTRALIA – Queensland • ♀; Rubyvale ; 23°25′ S, 147°42′ E; 29 Jun. 1980; I.A. Manthey leg.; QMB S1283 View Materials .

GoogleMaps

Other material examined

AUSTRALIA – Queensland • 1 ♂; Rubyvale ; 23°25′ S, 147°41′ E; 18 Jun. 2002; S. Eldridge leg.; QMB S548246 View Materials GoogleMaps • 1 ♀; Rubyvale, Capella Road ; 23°25′ S, 147°42′ E; 259 m a.s.l.; 18 Apr. 2023; J.D. Wilson and M.G. Rix leg.; excavated, open burrow on ground; QMB S118281 View Materials GoogleMaps .

Description

Male (QMB S548246)

GENERAL ( Fig. 42A–Q View Fig ). Body length 18.88, in good condition, colour presumably faded due to preservation.

DORSAL PROSOMA ( Fig. 42A, E–F View Fig ). Carapace length 7.89, width 6.89, length/width 1.15, clypeus to fovea length/carapace length 0.72, caput width/carapace width 0.71, carapace red-brown, reflective setae present, heavy on caput, moderate on thorax, fovea procurved, fovea width/carapace length 0.17 ( Fig. 42A, F View Fig ); chelicerae dark red-brown, rastellum absent or inconspicuous, chelicerae length/carapace length 0.55 ( Fig. 42A View Fig ); eye group rectangular, width/length 2.23, eye tubercle present ( Fig. 42E View Fig ).

ABDOMEN ( Fig. 42B, D View Fig ). Abdomen length 6.67, brown, dorsal pattern absent, with consistent cover of short setae.

VENTRAL PROSOMA ( Fig. 42C, G–I View Fig ). Labium cuspules present, count =2 ( Fig. 42H View Fig ); maxillae heel distinct, cuspules present, count = about 131, extending posteriorly onto heel, extending laterally about 40% of maxillae length ( Fig. 42C, I View Fig ); coxae cuspules absent, thorn-like setae on prolateral face present ( Fig. 42C, I View Fig ); sternum length/width 1.27, central sternum with consistent covering of short setae, row of longer setae around posterior edges, some shorter thorn-like setae around anterior edges, darker cuticle around sigilla ( Fig. 42G–H View Fig ); posterior sigilla semi-elongate, central sternum to posterior sigilla length/sternum length 0.21, posterior sigilla length/sternum length 0.12 ( Fig. 42G–H View Fig ); other sigilla small, round and lateral ( Fig. 42G–H View Fig ).

LEG I ( Fig. 42N–Q View Fig ). Leg I dark red-brown, lighter on distal metatarsus and tarsus, femur length 6.57, patella length 4.08, tibia length 4.64, metatarsus length 4.44, tarsus length 2.79, total length 22.53, leg I length/carapace length 2.86 ( Fig. 42N–O View Fig ); scopulae on distal metatarsus and tarsus ( Fig. 42N–O View Fig ); spine count Fe D 1, Fe PL 1, Pa PL 2, Ti PL 0, Ti RL 0, Me PL 0, Me RL 0, Ta 0 ( Fig. 42N–O View Fig ); tibia length/width [TIL/TID] 2.95, widening from proximal end to spur before narrowing again towards distal end, spur present, digitiform, knuckle present, megaspine angled at 14 degrees, length to distal face of spur/tibia length [TIS/TIL] 0.46, spur height/tibia width [TISH/TID] 0.51, megaspine length/tibia length 0.19 ( Fig. 42N–P View Fig ); metatarsus slightly sinuous, proximal excavation present, excavation concave with pronounced heel, heel broadly rounded, excavation length/metatarsus length [MIPEL/MIL] 0.46, metatarsus length/width [MIL/MID] 3.42 ( Fig. 42N–O, Q View Fig ).

PEDIPALP ( Fig. 42J–M View Fig ). Tibia length 3.47, width 1.29, length/width [PTL/PTD] 2.69, asetose depression present, depression length/palp tibia length [PDL/PTL] 0.63, retrolateral face with consistent covering of light setae, ventral face with without any distinctive setae, prolateral face with two disto-ventral spines, disto-medial spine absent ( Fig. 42J–K View Fig ); patella prolateral face with 2 spines ( Fig. 42J–K View Fig ); cymbium with scopulae present distally ( Fig. 42J–K View Fig ); copulatory organ total length 1.88, length/palp tibia length 0.54 ( Fig. 42L–M View Fig ); bulb length/width 0.81 ( Fig. 42L–M View Fig ); embolus demarcated and roughly perpendicular to bulb, attenuate, with wide base tapering about halfway along into attenuate apical section, one slight bend, at about 0.6 of length, slight bend before tip, width at base/bulb width 0.31, embolus length/bulb length 2.74 ( Fig. 42L–M View Fig ).

Female (holotype, QMB S1283)

GENERAL ( Fig. 43A–L View Fig ). Body length 20.28, in moderate condition, colour faded significantly due to preservation.

DORSAL PROSOMA ( Fig. 43A, E–F View Fig ). Carapace length 7.38, width 6.19, length/width 1.19, clypeus to fovea length/carapace length 0.75, caput width/carapace width 0.80, carapace orange-brown, reflective setae present, moderate on caput, light on thorax, fovea procurved, fovea width/carapace length 0.16 ( Fig. 43A, F View Fig ); chelicerae red-brown, rastellum absent or inconspicuous, chelicerae length/carapace length 0.55 ( Fig. 43A View Fig ); eye group rectangular, width/length 1.9, eye tubercle present ( Fig. 43E View Fig ).

ABDOMEN ( Fig. 43B, D View Fig ). Abdomen length 9.13, light brown, dorsal pattern absent, with consistent cover of short setae.

VENTRAL PROSOMA ( Fig. 43C, G–I View Fig ). Labium cuspules present, count =1 ( Fig. 43H View Fig ); maxillae heel distinct, cuspules present, count = about 162, extending posteriorly onto heel, extending laterally about 50% of maxillae length ( Fig. 43C, I View Fig ); coxae cuspules absent, thorn-like setae on prolateral face present ( Fig. 43C, I View Fig ); sternum length/width 1.14, central sternum with consistent covering of moderate setae, row of longer setae around posterior edges, some shorter thorn-like setae around anterior edges ( Fig. 43G–H View Fig ); posterior sigilla semi-elongate, central sternum to posterior sigilla length/sternum length 0.22, posterior sigilla length/sternum length 0.18 ( Fig. 43G–H View Fig ); other sigilla small, round and lateral ( Fig. 43G–H View Fig ).

LEG I ( Fig. 43J–K View Fig ). Leg I orange-brown, darker on patella and tibia, femur length 5.62, patella length 3.50, tibia length 3.63, metatarsus length 3.36, tarsus length 2.26, total length 18.38, leg I length/carapace length 2.49; scopulae on distal metatarsus and tarsus; spine count Fe D 1, Fe PL 1, Pa PL 2, Ti PL 4, Ti RL 4, Me PL 2, Me RL 2, Ta 0; tibia length/width [TIL/TID] 2.95.

GENITALIA ( Fig. 43D, L View Fig ). Epigastric furrow unmodified ( Fig. 43D View Fig ); spermathecae with two vesicles each ( Fig. 43L View Fig ); lateral vesicle relatively straight, length 0.52, lateral vesicle length/genitalia width 0.40, length/width at base 1.64, crown un-demarcated ( Fig. 43L View Fig ); medial vesicle with distinct basal section angled medially, before undulating anteriorly, medial vesicle length/genitalia width 0.64, length/width 6.2, medial vesicle length/lateral vesicle length 1.59 ( Fig. 43L View Fig ).

Distribution and natural history

Aname longitheca occurs in the ‘Gemfields’ of central Queensland, in the Brigalow Belt North bioregion, where it is known only from near the town of Rubyvale ( Fig. 7 View Fig ). It constructs an open, silk-lined burrow without silk outside of the entrance, on an angle, and with a hidden secondary ‘wishbone’ entrance ( Fig. 7 View Fig ).