Aname warialda Raven, 1985

Aname warialda Raven, 1985 View in CoL

Figs 1 View Fig , 11 View Fig , 81–82 View Fig View Fig

Aname warialda Raven, 1985: 407 View in CoL , figs 7, 18, 22, 35, 46, 56, 58.

Aname View in CoL “MYG687” – Rix et al. 2021: figs 3, 6–7.

Diagnosis

Males of A. warialda can be distinguished from all species for which males are known except A. bifaceta sp. nov., A. occivillosa sp. nov., A. scutitheca sp. nov., and A. villosa by a moderate to large body size (carapace length>4.0 mm), the presence of a long embolus (embolus length / bulb length>1.5) that is relatively straight, a short tibial megaspine (megaspine length/ tibia length <0.2), and three or more spines on the prolateral patellae of the pedipalp and leg I ( Fig. 81A–Q View Fig ). Males of A. warialda can be distinguished from those of A. occivillosa and A. villosa by the presence of a longer embolus (embolus length / bulb length>2.5) and a longer asetose depression on the palp tibia (depression length / palp tibia length> 0.6) ( Fig. 81J–M View Fig ; cf. Figs 74 View Fig , 78 View Fig ). Males of A. warialda can be distinguished from those of A. scutitheca by the presence of a tibia I that stays about the same width from the proximal end to the base of the tibial spur when in lateral view and a less spiny palp tibia ( Fig. 81J–K, N–P View Fig ; cf. Fig. 76 View Fig ). Males of A. warialda can be distinguished from those of A. bifaceta by the presence of a longer asetose depression on palp tibia (depression length / palp tibia length ~0.71), and a knuckle at the base of the tibial spur ( Fig. 81J–K, N–P View Fig ; cf. Fig. 71 View Fig ).

Females of A. warialda can be distinguished from all species for which females are known except A. bifaceta sp. nov. and A. scutitheca sp. nov. by the presence of spermathecae with a single elongate, undulating vesicle (lateral vesicle length/ genitalia width>0.5) ( Fig. 82L View Fig ). Females of A. warialda can be distinguished from those of A. scutitheca by the absence of a large rounded extension of the posterior genital plate (“scute”) over the epigastric furrow ( Fig. 82D, L View Fig ; cf. Fig. 77 View Fig ). Females of A. warialda can be distinguished from those of A. bifaceta by the presence of larger, longer spermathecae (lateral vesicle length / genitalia width ~ 0.8; cf. ~ 0.5 in A. bifaceta ) ( Fig. 82L View Fig ; cf. Fig. 72 View Fig ).

Type material

Holotype

AUSTRALIA – New South Wales • ♂; Warialda ; 29°32′ S, 150°35′ E; 7 Jan. 1980; R. Hobbs leg.; QMB S1291 View Materials .

GoogleMaps

Paratypes

AUSTRALIA – Queensland • 1 ♂; Girraween National Park ; 28°50′ S, 151°56′ E; 5 Feb. 1979; H. Todd leg.; QMB S1296 View Materials . – New South Wales • 1 ♀; Toowoomba ; 27°34′ S, 151°57′ E; 10 Feb. 1965; T. Passlow leg.; QMB S1299 View Materials GoogleMaps • 2 ♂♂; Tenterfield ; 29°06′ S, 151°54′ E; Apr. 1983; G. Colvin leg.; QMB S1294 View Materials GoogleMaps • 1 ♂; Texas Caves [ Ashford Caves ]; 29°12′ S, 150°59′ E; 4 Apr. 1979; L.R. Jeffrey leg.; QMB S1298 View Materials GoogleMaps • 1 ♀; Inverell ; 29°46′ S, 151°07′ E; May 1979; QMB S1314 View Materials GoogleMaps • 1 ♂; Bingara ; 29°49′ S, 150°25′ E; Jan. 1980; R. Hobbs leg.; QMB S1292 View Materials GoogleMaps .

GoogleMaps

Other material examined

AUSTRALIA – Queensland • 1 ♂; Carnarvon National Park, Mount Moffatt Section, near Marlong Arch ; 24°59′ S, 147°54′ E; 760 m a.s.l.; 12 Dec. 2012 – 16 Jan. 2013; G.B. Monteith and G. Keith leg.; gutter trap, eucalypt forest; QMB S52397 GoogleMaps • 2 ♂♂; Carnarvon National Park, Mount Moffatt Section ; 25°04′ S, 148°02′ E; 13–15 Dec. 1987; G.B. Monteith, J. Thompson and D. Yeates leg.; QMB S11261 GoogleMaps • 1 ♀; Mount Tabor Station , ca 147.2 km NNE of Morven; 25°10′ S, 147°36′ E; 8 Dec. 2020; C. Eddie and E. Amsters leg.; excavated, eucalypt forest; QMB S118236 View Materials GoogleMaps • 1 ♀; Mount Tabor Station , ca 145.4 km NNE of Morven; 25°11′ S, 147°37′ E; 22 Oct. 2021; E. Amsters and R. Aisthorpe leg.; excavated, eucalypt forest, near watercourse; QMB S118245 View Materials GoogleMaps • 1 ♀; Mount Tabor Station , ca 145.0 km NNE of Morven; 25°11′ S, 147°36′ E; 21 Oct. 2021; E. Amsters and R. Aisthorpe leg.; excavated, eucalypt forest; QMB S118242 View Materials GoogleMaps • 1 ♀; 51.5 km NNW of Injune ; 25°25′ S, 148°21′ E; 21 Sep. 2021; E. Amsters leg.; excavated, Callitris forest; QMB S118241 View Materials GoogleMaps • 1 ♀; 51.3 km NNW of Injune ; 25°26′ S, 148°21′ E; 21 Sep. 2021; E. Amsters leg.; excavated, Callitris forest; QMB S118239 View Materials GoogleMaps • 1 ♀; 51.3 km NNW of Injune ; 25°26′ S, 148°21′ E; 21 Sep. 2021; E. Amsters leg.; excavated, Callitris forest; QMB S118240 View Materials GoogleMaps • 1 ♂; Hutton Creek , “Oak Wells”; 25°46′ S, 148°17′ E; 600 m a.s.l.; 15 Dec. 2001 – 7 Mar. 2002; G.B. Monteith and D.J. Cook leg.; pitfall trap, vine scrub; QMB S59308 View Materials GoogleMaps • 1 ♂; Kingaroy ; 26°31′ S, 151°48′ E; 1 Dec. 1995; D. Kamholtz leg.; QMB S29591 GoogleMaps • 1 ♂; Roma ; 26°34′ S, 148°47′ E; 24 Feb. 1986; R. Kunde leg.; QMB S100534 View Materials GoogleMaps • 1 ♂; 40 km S of Charleville, “ Wallal ”; 26°44′ S, 146°07′ E; 23 Jan. 1996; P. McRae leg.; QMB S29635 GoogleMaps • 1 ♂; Tara ; 27°16′ S, 150°27′ E; 20 Nov. 2001; Queensland Department of Primary Industries leg.; QMB S118360 View Materials GoogleMaps • 1 juv.; Lake Broadwater Conservation Park , N of lake; 27°20′ S, 150°33′ E; 335 m a.s.l.; 7 Dec. 2019; M.G. Rix and M.S. Harvey leg.; excavated, open burrow on ground, open woodland; QMB S111487 View Materials GoogleMaps • 1 ♀; Lake Broadwater Conservation Park, Wilga Bush Camping Area ; 27°20′ S, 151°06′ E; 345 m a.s.l.; 28 Oct. 2022; J.D. Wilson, E. Briggs, M.G. Rix and A.G. Rix leg.; excavated; QMB S118219 View Materials GoogleMaps • 1 ♀; Lake Broadwater Conservation Park, Wilga Bush Camping Area ; 27°20′ S, 151°06′ E; 345 m a.s.l.; 28 Oct. 2022; J.D. Wilson, E. Briggs, M.G. Rix and A.G. Rix leg.; excavated; QMB S118220 View Materials GoogleMaps • 1 ♂; Lake Broadwater Conservation Park ; 27°21′ S, 151°05′ E; 3 Jan.–25 Feb. 1986; Queensland Museum Party leg.; pitfall trap; QMB S3153 View Materials GoogleMaps • 1 ♀; Moonbah, campsite [ Alton National Park ]; 27°59′ S, 149°19′ E; 9–11 Jan. 1979; R.J. Raven leg.; QMB S1295 View Materials GoogleMaps • 1 ♂; Ula Ula National Park ; 27°59′ S, 149°28′ E; 11 Nov. 2016; R.C. Santana leg.; QMB S110191 View Materials GoogleMaps • 1 ♀; Durikai State Forest, Durikai Road ; 28°12′ S, 151°37′ E; 531 m a.s.l.; 19 Feb. 2023; M.G. Rix, J.D. Wilson and M.S. Harvey leg.; excavated, open burrow on ground; QMB S118230 View Materials GoogleMaps • 1 ♂; 18 km N of Inglewood, Robert Wicks Research Centre ; 28°18′ S, 151°09′ E; 10 Jan. 1999; G. Rettke leg.; hand collected, on path at night, Callitris forest; QMB S42819 GoogleMaps • 1 juv.; Oman Ama , E of Inglewood, Donovans Road; 28°23′ S, 151°20′ E; 342 m a.s.l.; 19 Feb. 2023; M.G. Rix, J.D. Wilson and M.S. Harvey leg.; excavated, open burrow on ground next to road; QMB S118229 View Materials GoogleMaps • 1 juv.; Warroo , SE of Inglewood, Cement Mills Road; 28°31′ S, 151°23′ E; 409 m a.s.l.; 14 Aug. 2022; E.J. Briggs and M. S. Woolley leg.; excavated, y-shaped burrow, Callitris forest; QMB S118218 View Materials GoogleMaps • 1 ♀; Warroo , SE of Inglewood, Cement Mills Road; 28°32′ S, 151°21′ E; 374 m a.s.l.; 19 Feb. 2023; M.G. Rix, J.D. Wilson and M.S. Harvey leg.; excavated, open burrow on ground next to road; QMB S118219 View Materials GoogleMaps • 1 juv.; Warroo , SE of Inglewood, Cement Mills Road; 28°32′ S, 151°21′ E; 374 m a.s.l.; 19 Feb. 2023; M.G. Rix, J.D. Wilson and M.S. Harvey leg.; excavated, open burrow on ground next to road; QMB S118228 View Materials GoogleMaps • 1 ♂; The Summit ; 28°34′ S, 151°57′ E; 16 Jan. 1984; Queensland Department of Primary Industries leg.; QMB S100536 View Materials GoogleMaps • 1 ♂; Stanthorpe ; 28°37′ S, 151°30′ E; 21 Dec. 1994; Mr and Mrs Kuenstner leg.; QMB S26087 GoogleMaps • 1 ♂; Stanthorpe ; 28°37′ S, 151°30′ E; Apr. 2016; M. Hodgetts leg.; QMB S118359 View Materials GoogleMaps • 1 ♀; Nundubbermere Falls , 25 km SW of Stanthorpe; 28°47′ S, 151°41′ E; 1–4 Apr. 1988; G.B. Monteith leg.; QMB S4192 View Materials GoogleMaps • 1 ♂; Tenterfield, Wallangarra ; 28°55′ S, 151°56′ E; Apr. 1983; M. Wanstall leg.; QMB S10000 GoogleMaps . – New South Wales • 1 ♂; Texas ; 28°51′ S, 151°09′ E; 4 Feb. 1992; E.J. Greentree leg.; QMB S19135 GoogleMaps • 1 ♂; Texas Caves [Ashford Caves]; 29°12′ S, 150°59′ E; 1–2 Feb. 1975; G.V. Czechura leg.; QMB S1297 View Materials GoogleMaps • 1 ♂; Deepwater ; 29°27′ S, 151°49′ E; 10 Dec. 1985; R. Hobbs leg.; QMB S9804 View Materials GoogleMaps • 1 ♂; Emmaville ; 29°30′ S, 150°19′ E; 13 Jun. 1996; F. Ginns leg.; QMB S34590 GoogleMaps • 1 ♂; The Brigalows , 25 km N of Inverell; 29°37′ S, 151°06′ E; 1 Jan. 1991; F. Ginns leg.; QMB S34586 GoogleMaps • 1 ♂; 15 km W of Armidale ; 30°31′ S, 151°31′ E; 7 Dec. 1980; R. Hobbs leg.; QMB S9747 View Materials GoogleMaps • 1 ♀; Armidale , near cemetary; 30°31′ S, 151°40′ E; AMS KS31709 GoogleMaps • 1 ♂; Moore Creek , 8–10 km N of Tamworth; 31°02′ S, 150°58′ E; Jan. 1984; D. McCalee leg.; AMS KS13803 GoogleMaps .

Description

Male (holotype, QMB S1291)

GENERAL ( Fig. 81A–Q View Fig ). Body length 22.77, in good condition, colour presumably faded due to preservation.

DORSAL PROSOMA ( Fig. 81A, E–F View Fig ). Carapace length 8.81, width 7.71, length/width 1.14, clypeus to fovea length/carapace length 0.71, caput width/carapace width 0.66, carapace red, caput slightly darker than thorax, reflective setae present, heavy on caput, moderate on thorax, fovea procurved, fovea width/ carapace length 0.16 ( Fig. 81A, F View Fig ); chelicerae dark red-brown, rastellum absent or inconspicuous, chelicerae length/carapace length 0.46 ( Fig. 81A View Fig ); eye group rectangular, width/length 1.84, eye tubercle present ( Fig. 81E View Fig ).

ABDOMEN ( Fig. 81B, D View Fig ). Abdomen length 9.63, brown, dorsal pattern absent, with consistent cover of short setae.

VENTRAL PROSOMA ( Fig. 81C, G–I View Fig ). Labium cuspules absent ( Fig. 81H View Fig ); maxillae heel distinct, cuspules present, count =about 150, extending posteriorly onto heel, extending laterally about 40% of maxillae length ( Fig. 81C, I View Fig ); coxae cuspules absent, thorn-like setae on prolateral face present ( Fig. 81C, I View Fig ); sternum length/width 1.09, central sternum with consistent covering of short setae, row of longer setae around posterior edges ( Fig. 81G–H View Fig ); posterior sigilla elongate, central sternum to posterior sigilla length/sternum length 0.25, posterior sigilla length/sternum length 0.18 ( Fig. 81G–H View Fig ); other sigilla small and lateral, medial sigilla semi-elongate, anterior sigilla round ( Fig. 81G–H View Fig ).

LEG I ( Fig. 81N–Q View Fig ). Leg I orange-brown, femur length 6.88, patella length 4.41, tibia length 4.86, metatarsus length 4.63, tarsus length 3.24, total length 24.02, leg I length/carapace length 2.73 ( Fig. 81N– O View Fig ); scopulae on distal metatarsus and tarsus ( Fig. 81N–O View Fig ); spine count Fe D 4, Fe PL 1, Pa PL 3, Ti PL 0, Ti RL 0, Me PL 0, Me RL 0, Ta 0 ( Fig. 81N–O View Fig ); tibia length/width [TIL/TID] 3.21, even width along length, spur present, digitiform, knuckle present, megaspine angled at 3 degrees, length to distal face of spur/tibia length [TIS/TIL] 0.51, spur height/tibia width [TISH/TID] 0.61, megaspine length/tibia length 0.18 ( Fig. 81N–P View Fig ); metatarsus slightly sinuous, proximal excavation present, excavation concave with pronounced heel, heel rounded, excavation length/metatarsus length [MIPEL/MIL] 0.39, metatarsus length/width [MIL/MID] 3.78 ( Fig. 81N–O, Q View Fig ).

PEDIPALP ( Fig. 81J–M View Fig ). Tibia length 3.62, width 1.48, length/width [PTL/PTD] 2.45, asetose depression present, depression length/palp tibia length [PDL/PTL] 0.71, retrolateral face with consistent covering of light setae, ventral face with two elongate bristle-like setae below depression, prolateral face with patch of spines on distal half, disto-medial spine present ( Fig. 81J–K View Fig ); patella prolateral face with 3 spines ( Fig. 81J–K View Fig ); cymbium with scopulae present distally ( Fig. 81J–K View Fig ); copulatory organ total length 2.78, length/palp tibia length 0.77 ( Fig. 81L–M View Fig ); bulb length/width 0.94 ( Fig. 81L–M View Fig ); embolus demarcated and roughly perpendicular to bulb, attenuate, long and straight with slightly swollen base, width at base/ bulb width 0.24, embolus length/bulb length 2.56 ( Fig. 81L–M View Fig ).

Female (QMB S118220)

GENERAL ( Fig. 82A–L View Fig ). Body length 25.16, in good condition.

DORSAL PROSOMA ( Fig. 82A, E–F View Fig ). Carapace length 8.96, width 8.65, length/width 1.04, clypeus to fovea length/carapace length 0.78, caput width/carapace width 0.74, carapace red-brown, reflective setae present, light on caput, light on thorax, fovea procurved, fovea width/carapace length 0.21 ( Fig. 82A, F View Fig ); chelicerae dark red-brown, rastellum absent or inconspicuous, chelicerae length/carapace length 0.62 ( Fig. 82A View Fig ); eye group rectangular, width/length 1.82, eye tubercle present ( Fig. 82E View Fig ).

ABDOMEN ( Fig. 82B, D View Fig ). Abdomen length 10.77, dark brown, dorsal pattern absent, with consistent cover of short setae.

VENTRAL PROSOMA ( Fig. 82C, G–I View Fig ). Labium cuspules absent ( Fig. 82H View Fig ); maxillae heel distinct, cuspules present, count =about 216, extending posteriorly onto heel, extending laterally about 40% of maxillae length ( Fig. 82C, I View Fig ); coxae cuspules absent, thorn-like setae on prolateral face present ( Fig. 82C, I View Fig ); sternum length/width 1.06, central sternum with consistent covering of short setae, row of longer setae around posterior edges, setae at higher density around anterior edges ( Fig. 82G–H View Fig ); posterior sigilla elongate, central sternum to posterior sigilla length/sternum length 0.23, posterior sigilla length/sternum length 0.20 ( Fig. 82G–H View Fig ); other sigilla small, round and lateral ( Fig. 82G–H View Fig ).

LEG I ( Fig. 82J–K View Fig ). Leg I dark coffee-brown, darker on patella and tibia, femur length 6.85, patella length 4.64, tibia length 4.58, metatarsus length 4.29, tarsus length 2.76, total length 23.12, leg I length/ carapace length 2.58; scopulae on distal metatarsus and tarsus; spine count Fe D 0, Fe PL 1, Pa PL 1, Ti PL 0, Ti RL 4, Me PL 1, Me RL 4, Ta 0; tibia length/width [TIL/TID] 3.01.

GENITALIA ( Fig. 82D, L View Fig ). Epigastric furrow extending slightly, posterior edge with rounded shape ( Fig. 82D View Fig ); spermathecae with one vesicle each ( Fig. 82L View Fig ); lateral vesicle undulating, with several distinct bends, length 1.47, lateral vesicle length/genitalia width 0.81, length/width at base 4.04, crown slightly wider than stem ( Fig. 82L View Fig ).

Distribution and natural history

Aname warialda has a wide distribution in northern New South Wales and south-eastern Queensland, in the Brigalow Belt South, Nandewar and New England Tablelands bioregions, from Tamworth north to Carnarvon National Park, and east to Toowoomba ( Fig. 11 View Fig ). It constructs an open, silk-lined burrow with silk spilling out from the entrance, sometimes with a built-up mound of soil around the entrance. The burrow has a short, hidden ‘wishbone’ entrance, which, when excavated, is revealed to terminate in a distinctly thick, white sheath of silk ( Fig. 11 View Fig ).