Spinolochus laevifrons (Holmgren, 1860)

56. Spinolochus laevifrons (Holmgren, 1860) View in CoL

Material examined. Russia: Tyumen Prov ., Khanty- Mansi Autonomous Region, Lyamin River, 61°43′48′′N, 71°01′48′′E, 75 m a.s.l., 21.VII.2010, coll. K. Tomkovich, 1 female (ZMUM); Sakhalin Prov ., Kuril Is. : Matua I., inland from Dvoinaya Bay, 48°04′N, 153°15′E, 3.VIII.1999, coll. D.J. Bennett & B.K. Urbain, 3 females, 3 males (TAMU); Onekotan I., 49°17′N, 154°45′E, 9.VIII.1996, coll. B.K. Urbain, 1 female (TAMU).

Distribution. North America, Europe, Middle Asia, Russian Far East, Japan.

57. Tersilochus (Gonolochus) caudatus (Holmgren, 1860)

Material examined. France, “Touraine Desbrochers”, coll. Desbrochers, N . Kokujev collection, 1 female ( ZIN) . Germany, Bavaria: Allgäu, Kempten, Kempter Wald Blauseemoos, 47°40′21′′N, 10°28′56′′E, 880 m a.s.l., Malaise trap, 28. V GoogleMaps .–10. VI GoogleMaps .2015, coll. D. Doczkal & J. Voith, 1 female ( ZSM); N of Bopfingen, Ipf Mt., 48°52′12′′N, 10°21′25′′E, 580– 560 m a.s.l., 3. V GoogleMaps .2016, coll. A.I. Khalaim, 7 females, 2 males ( ZIN); S of Oggelshausen, Wildes Ried, 580 m a.s.l., 48°02.2′N, 09°38.7′E, 5. V GoogleMaps .2016, coll. A.I. Khalaim, 1 female, 1 male ( ZIN); SW of Heubach, Scheuelberg, 600–720 m a.s.l., meadows, forest, 48°46.8′N, 9°54.9′E, 6. V GoogleMaps .2016, coll. A.I. Khalaim, 8 females, 3 males ( ZIN) .

Remarks. General distribution of this species is given after Khalaim & Tereshkin (2019).

Distribution. Widespread in Palaearctic Region. Europe, Caucasus, Turkey, Kazakhstan, Middle Asia, Mongolia, Siberia, Russian Far East, Republic of Korea.

58. Tersilochus (Pectinolochus) acutangulus Khalaim, 2007

Material examined. Russia, Tuva, Turan, floodlands, Turanchik River , 3. VI .1975, coll. D.R. Kasparyan , 1 female ( ZIN) .

Remarks. Record of T. (P.) kerzhneri Khalaim, 2007 from Finland and Russian Karelia ( Khalaim & Várkonyi, 2018: 182) actually belongs to T. acutangulus . Thus, T. kerzhneri is excluded here from the fauna of Europe.

Distribution. Europe (north), Mongolia, Siberia, Russian Far East.

59. Tersilochus (Pectinolochus) asper Horstmann, 1981

Material examined. * Kazakhstan: Akmola Prov ., Kokshetau Mts. , 21. V .1957, coll. V. I . Tobias, 1 female ( ZIN); Karaganda [Qaraghandy] Prov ., 40 km S of Zhanaarka Station, Mt. Koksengir , 3. V .1959, coll. V. I . Tobias, 1 female ( ZIN); South Kazakhstan [ Turkistan ] Prov .: 15 km NE of Achisay, 1000–1300 m a.s.l., 11–13. V .1994, coll. D.A. Milko, 1 female ( ZIN); Pskem Mt. Range, near Nanai [north of Nanai?], 3000 m a.s.l., 23. V .1963, coll. V. I . Tobias, 4 females, 1 male ( ZIN); East Kazakhstan Prov ., 45 km SSW of Semipalatinsk [Semey], Semeytau Mt. , on blooming snowdrops, 1. V .1961, coll. V. I . Tobias, 5 females, 4 males ( ZIN) . * Uzbekistan, Samarkand Prov ., Kara-Tepa Mt. Range , Takhtakaracha Pass, 7–8. V .1982, coll. S. A . Belokobylskij, 1 female ( ZIN) .

Distribution. Europe, Kazakhstan, Middle Asia.

60. Tersilochus (Pectinolochus) coeliodicola (Silvestri, 1917)

Material examined. Mongolia, Dornod Aimag: 32 km SE of Salkhit Mt., Numregin River [46°58′N, 119°18′E], forest-steppe, 14–16. VI GoogleMaps .1976, coll. M. A GoogleMaps . Kozlov, 4 females ( ZIN); 30 km ENE of Tsagan-Ula [Tsagaan Ulaa] Mt., Modon-Obo Mt. [46°30′N, 116°26′E], 20. VI GoogleMaps .1976, coll. I. M GoogleMaps . Kerzhner , 1 female ( ZIN) . Russia: Leningrad Prov ., 8 km N of Pushkin , 16th km, 17. V .1987, coll. D.R. Kasparyan , 1 female ( ZIN) ; Yaroslavl Prov ., Yaroslavl, 7. V .1899, N. Kokujev collection, 1 female ( ZIN) ; Ul’yanovsk Prov ., Ul’yanovsk, left bank of Volga River, Verkhnyaya Terrace Station , 80 m a.s.l., 14.VIII [year illegible], coll. Z . A . Yefremova , 1 female ( ZIN) ; Rostov Prov ., 20 km W of Zavetnoe, Sal River , 17. V .1986, coll. D.R. Kasparyan , 1 female ( ZIN) ; Jewish Autonomous Prov ., Amurzet , oak forest, 18. VI .1985, coll. D.R. Kasparyan , 1 female ( ZIN) ; Khabarovsk Terr., Khekhtsyr Range , 18th km, 13. VI .1985, coll. D.R. Kasparyan , 1 female ( ZIN) . * Ukraine, Mykolaiv Prov ., Black Sea Nature Biosphere Reserve, Volyzhin Forest [ W of Vasilevka Vill., 46°32′N, 31°43′E], 21. V GoogleMaps .1974, coll. V. V GoogleMaps . Kostyukov , 1 female ( ZIN) .

Distribution. Europe, Mongolia, Russian Far East.

61. Tersilochus (Pectinolochus) ensifer (Brischke, 1880)

Material examined. * Lithuania, “Vil... Kalu...” [handwritten, illegible text], 1. V .1973, coll. A. Jakimavičius, 1 female ( ZIN) . Russia: Leningrad Prov ., 8 km N of Pushkin , 16th km , 17.V.1987, coll. D.R. Kasparyan, 1 female ( ZIN) ; Stavropol Terr., Kislovodsk , Kurortnyy Park, meadow, forest , 27.IV.2018, coll. S. A . Belokobylskij, 1 female ( ZIN) . * Ukraine: Kyiv Prov ., Kyiv, 21.IV.1900, coll. Yu. Vagner, 1 female ( ZIN) ; Poltava Prov ., Poltava, 20.IV.1914, coll. A. A . Ogloblin, 1 female ( ZIN) .

Remarks. The species was recently record- ed from southern Finland and Russian Karelia ( Khalaim & Várkonyi, 2018: 181).

Distribution. Europe.

62. Tersilochus (Pectinolochus) hungaricus Horstmann, 1981

Material examined. Ukraine, Donets’k Prov ., NNE of Amvrosievka, Yasenevoe , forest, 15. V .1974, coll. D.R. Kasparyan , 1 female [det. K. Horstmann] ( ZIN) .

Remarks. In the original description, Horstmann (1981: 43–44) recorded this species from Ukraine, based on the Leningrad collection, but without any details. One specimen of T. hungaricus from the Donets’k Province of Ukraine, which is stored in the ZIN collection and is provided with the identification (non-type) label of Horstmann, was examined.

Distribution. Hungary, Ukraine.

63. Tersilochus (Pectinolochus) intermedius Horstmann, 1981

Material examined. Russia, Yaroslavl Prov ., “Gedenowo”, “Dan. [former Danilov Uezd]”, 27. V .1918, coll. A. Schestakow , 1 female ( ZIN) .

Remarks. The species was recently record- ed from Russian Karelia ( Khalaim & Várkonyi, 2018: 182).

Distribution. Ukraine (east), Russia (Republic of Karelia, Yaroslavl Province).

64. Tersilochus (Pectinolochus) lapponicus Hellén, 1958

Material examined. * Georgia, Georgian Military Road, Krestovyy Pass , 2150 m a.s.l., 23. V .1967, coll. D.R. Kasparyan , 1 female ( ZIN) . * Lithuania, N of Vilnius, Visoriai , 28. V .1999, coll. V. Jonaitis , 1 female ( ZIN) . Russia: Leningrad Prov ., 52 km S of St Petersburg, Susanino. , 13. V .1990, coll. D.R. Kasparyan, 1 female ( ZIN); Novgorod Prov ., 20 km NW of Pestovo, Tychkino Vill. , 1. VI .1993, coll. V.I. Tobias, 1 female ( ZIN); Stavropol Terr., Kislovodsk, road to Bol’shoe Sedlo Mt. , meadow, forest, 1. V .2018, coll. S.A. Belokobylskij , 1 female ( ZIN) .

Remarks. The species was recently recorded from Finland and the Murmansk Province of Russia ( Khalaim & Várkonyi, 2018: 182).

Distribution. Europe, Caucasus.

65. Tersilochus (Pectinolochus) luteicornis (Hellén, 1958)

Material examined. Russia, Leningrad Prov ., 55 km S of St Petersburg, 4 km W of Krasnitsy , aspen forest, Malaise trap, 24–31.V.2008, coll. D. R. Kasparyan, 2 females ( ZIN) .

Remarks. The species was recently recorded from Finland and Russian Karelia ( Khalaim & Várkonyi, 2018: 182).

Distribution. Europe.

66. Tersilochus (Pectinolochus) rossicus Horstmann, 1981

Material examined. * Kazakhstan, West Kazakhstan Prov ., Khar’kin [Khar’kino], lower Ural River , 12. V .1951, coll. Rudolf, 1 female ( ZIN) . Ukraine, Lugans’k Prov ., 3 km NW of Antratsyt , forest, 10. V .1974, coll. D.R. Kasparyan, 3 females ( ZIN) .

Distribution. Ukraine, Kazakhstan (west).

67. Tersilochus (Pectinolochus) spiracularis Horstmann, 1971

Material examined. * Kazakhstan, Akmola Prov ., [without locality], 15. V .1942, coll. B. Kuzin, 2 females ( ZMUM) . * Latvia, Bausk, on willow, 4. V .1954, coll. V. Negrobov, 2 females ( ZIN) . Russia: Leningrad Prov .: 43 km S of St Petersburg , Semrino, 8. V .1983, coll. D.R. Kasparyan, 1 female ( ZIN); 8 km N of Pushkin , 16th km, 17. V .1987, coll. D.R. Kasparyan, 2 females ( ZIN); “ St Petersburg Governorate ”, on willow, 20.IV.1897, coll. Pomerantsev, N . Kokujev collection, 2 females ( ZIN); south of Leningrad Prov . [without locality], on willow flowers, 7. V .1972, coll. V.A. Trjapitzin, 3 females, 1 male ( ZIN); Moscow Prov .: SW of Moscow, Moskovsky , 55°34′44′′N, 37°20′30′′E, 25.IV.2016, coll. K GoogleMaps . Tomkovich, 1 female ( ZIN); Mikhnevo , 1. V .1962, coll. E. Antonova, 1 female [det. K. Horstmann; examined in ZMUM, deposited in ZIN]; Mytishchi , 2. V .1933, coll. G. Kostylev, 1 female ( ZMUM); Yaroslavl Prov ., Yaroslavl, N . Kokujev collection, 1 female ( ZIN) .

Remarks. Horstmann (1981: 21) recorded this species from “Mittelrussland”, based on the collection of Viktorov, but without any details. One specimen of T. spiracularis from the Moscow Province of Russia, which is stored in the ZMUM collection and is provided with the identification label of Horstmann (see Material examined section), was examined.

The species was recently recorded from Finland, Russian Karelia and the Leningrad Province of Russia. ( Khalaim & Várkonyi, 2018: 183).

Distribution. Europe, Kazakhstan.

68. Tersilochus (Pectinolochus) striola (Thomson, 1889)

Material examined. * Lithuania: N of Vilnius, Visoriai , 28. V .1999, coll. V. Jonaitis, 1 female ( ZIN); Avižieniai , 27.IV.1977, coll. V . Jonaitis, 1 female, 2 males ( ZIN) . Russia: Leningrad Prov . : St Petersburg, 3 females, 1 male ( ZIN); 43 km S of St Petersburg, Semrino , 6. VI .1981, coll. D.R. Kasparyan, 1 female ( ZIN); same data, but 8. V .1983, 3 females, 8 males ( ZIN); N of St Petersburg, Sosnovo , 13. V .1984, coll. D.R. Kasparyan, 2 females ( ZIN); Vyborg Distr., Privetninskoe , on birch, 2. V .1990, coll. A. Matov, 1 female ( ZIN); SE of St Petersburg, Nevsky Forest Park , 8. V .1983, coll. S.A. Belokobylskij, 2 females ( ZIN) ; Yaroslavl Prov ., “ Yarosl. ”, 24.IV.1944, coll. “Kom…”, N . Kokujev collection, 1 female ( ZIN) ; Moscow Prov .: Pushchino, near Tulchino , 54°49′44′′N 37°41′28′′E, yellow pan trap, 22–24.IV.2016, coll. K GoogleMaps . Tomkovich, 1 female ( ZMUM); Orekhovo-Zuevo Distr., Station of 73rd km, on fir-trees, 8.IV.2004, coll. K . Tomkovich, 2 females ( ZMUM); Podol’sk, Grivno Station , 16. V .2004, coll. K. Tomkovich, 1 female ( ZIN) ; Voronezh Prov ., Voronezh Nature Reserve , 19.IV.1949, coll. D. Dovnar [- Zapol’skiy], “ Diaparsis rufipes Holmg. det. D. Dovnar [-Zapol’skiy]”, 1 female ( ZMUM) ; Stavropol Terr., Kislovodsk, road to Bol’shoe Sedlo Mt. , meadow, forest, 1. V .2018, coll. S.A. Belokobylskij, 1 female ( ZIN); Yakutia : 40 km SSW of Yakutsk, Tekhtyur [Tyokhtyur], on willow, V .1976, coll. Vinokurov, 2 females ( ZIN); same locality, steppe, 25. V .1976, coll. Vinogradov, 1 female ( ZIN) ; Sakhalin Prov ., Sakhalin I ., Yuzhnosakhalinsk , 27. V .1973, coll. I.M. Kerzhner, 2 females ( ZIN) ; Magadan Prov ., 50 km N of Magadan , on willow, 24. VI .1975, coll. V. Marshakov, 2 females ( ZIN) ; Kamchatka Terr.: Klyuchevskoe Vill, Kamchatka River , 5. VI .1909, coll. A. Derzhavin, 2 females, 1 male ( ZIN); Avachinsky Volcano , 1000 m a.s.l., tundra, 26–27.VII.1985, coll. D. R . Kasparyan, 20 females ( ZIN) . * Ukraine, Kyiv Prov .: “Spartak” [? Station NW of Kyiv], 16. V .2003, coll. A.G. Kotenko, 1 female ( ZIN) ; Kyiv, “ O. K. Kiril. Kladbishche [Kirillovskoe Cemetery]”, on Salix caprea L ., 31.III.1917, coll. “ V. Alek.”, 5 females, 2 males ( ZIN) .

Distribution. North America, Europe,? Iran, Siberia, Russian Far East.

69. Tersilochus (Pectinolochus) terebrator ( Horstmann, 1971)

Material examined. Russia: Leningrad Prov ., [St Petersburg], Smolenskoe Cemetery , 1. V .1903, coll. Jacobson & Zubov, 1 female ( ZIN) ; Primorsky Terr., Kedrovaya Pad’ Nature Reserve , 1. V .1974, coll. Zhiltsova, 1 female ( ZIN) ; Magadan Prov ., 50 km N of Magadan, on Chosenia , 25. VI .1975, coll. V. Marshakov , 1 female ( ZIN) .

Distribution. Europe, Russian Far East.

70. Tersilochus (Tersilochus) cognatus (Holmgren, 1860)

Material examined. Armenia, Syunik Prov ., Meghri Municipality., Gudemnis, 1500–2000 m a.s.l., oak forest, 20.VI.1981, coll. A.G. Kotenko, 1 female (ZIN). * Azerbaijan, Abscheron [Abşeron] Distr., 1200–1400 m a.s.l., 30.V.1981, coll. A.G. Kotenko, 1 female (ZIN). Crimea: Crimean Nature Reserve, Al’ma River, 27.VI.1976, coll. A.G. Kotenko, 1 female (ZIN); S of Dobroe, Krasnolesye, 13.VI.1990, coll. D.R. Kasparyan, 1 female (ZIN). Czech Republic, South Bohemia, near České Budějovice, Černiš wetland, 1.V.2002, coll. A. Lozan, 3 females (ZIN). Germany, Bavaria, S of Oggelshausen, Wildes Ried, 580 m a.s.l., 48°02.2′N, 09°38.7′E, 5.V.2016, coll. A.I. Khalaim, 6 females, 5 males (ZIN). Hungary: Kétvölgy, 18.V.2001, coll. A.G. Kotenko, 1 female (ZIN); Kőszeg, forest, 19.V.2001, coll. A.G. Kotenko, 3 females (ZIN). * Kazakhstan, West Kazakhstan Prov ., 10 km SE of Mirgorodka, Aktau, 30.V.1986, coll. D.R. Kasparyan, 3 females (ZIN). Lithuania, “W. Russland Jurburg Winogradoff-Nikitin.” [Jurbarkas], 1 female (ZIN). Russia: Kaliningrad Prov .: Curonian Spit, 34th km, Biological Station “Rybachy”, bay, 23–28.V.2001, coll. A.I. Khalaim, 3 females, 3 males (ZIN); Curonian Spit, 7th km [between Zelenogradsk and Lesnoy], 11.V.1991, coll. A.R. Manukyan, 1 male (ZIN); Karelia: Kon, Malyy Lelikovskiy I., 61°98′N, 35°16′E, meadows, 1. VII.2017, coll. A.E. Humala, 1 male (HUM); Kol, Petrozavodsk, 21.VI.2006, coll. A.E. Humala, 1 female (ZIN); Leningrad Prov .: 70 km N of St Petersburg, Sosnovo, coll. D.R. Kasparyan (all in ZIN): 9.V.1990 (2 females, 3 males), 2–4.VI.1973 (5 females, 1 male [of them 3 females, det. K. Horstmann]), 8.VII.1987 (1 female), 11–17.VII.2009 (1 female); 19 km S of St Petersburg, near Pushkin, 29.V.1984, coll. D.R. Kasparyan, 6 females, 1 male (ZIN); same data, but 20.VI.1973, 1 female [det. K. Horstmann] (ZIN); NE of St Petersburg, Murino, 12.VI.1983, coll. S.A. Belokobylskij, 2 females (ZIN); same data, but 12.VI.1991, 1 female (ZIN); NW of St Petersburg, Roshchino, mixed forest, 15.VI.1991, coll. S.A. Belokobylskij, 1 female (ZIN); 40 km S of St Petersburg, Kobralovo, 17.VI.1984, coll. D.R. Kaspary- an, 1 female (ZIN); 50 km S of St Petersburg, 7. VII.1969, coll. D.R. Kasparyan, 1 female (ZIN); 55 km S of St Petersburg, 4 km W of Krasnitsy, aspen forest, Malaise trap, 9–23.V.2008, coll. D.R. Kasparyan, 3 females (ZIN); same data, but 24–31.V.2008, 9 females, 2 males (ZIN); St Petersburg, Vasilyevsky I., Smolenskoe Cemestry, 26.V.1991, coll. A. Matov, 1 male (ZIN); Novgorod Prov ., 20 km NW of Pestovo, Tychkino Vill., coll. V.I. Tobias (all in ZIN): 8–10.VII.1990 (1 female), 27.V.–18.VI.1991 (6 females, 1 male), 1.VI.–2.VII.1993 (26 females), 7.VI.2000 (1 female), 7.VI.2001 (1 female); Smolensk Prov ., Smolenskoye Poozer’e National Park, 10–15.VI.1993, coll. D.R. Kasparyan, 20 females (ZIN); Yaroslavl Prov .: Yaroslavl, V–VII.1995 –1996, N. Kokujev collection, 23 females, 4 males [6 females, 2 males, det. K. Horstmann] (ZIN); Berditsyno [20 km SE of Yaroslavl], 25.V.1896, N. Kokujev collection, 1 female (ZIN); “Gedenowo. Jarossl. Dan. [Yaroslavl, former Danilov Uezd] 9.VII.916. A. Schestakow”, 2 females (ZIN); “Zhedenovo, Dan. u. [former Danilov Uezd]”, 10.V.1914, coll. A. Schestakow, 1 female (ZIN); Nizhny Novgorod, Volchikha, “Gorbatovsk. u. [former Gorbatovskiy Uezd]”, 7–14.VI.1894, coll. Jacobson, 1 female (ZIN); Moscow Prov .: SW of Moscow, Moskovsky, 55°35′17′′N, 37°19′59′′E, yellow pan trap, 17– 21.V.2014, coll. K. Tomkovich, 1 female (ZMUM); same data, but 55°34′51′′N, 37°20′15′′E, 10–15.VI.2016, 1 female (ZMUM); SE of Moscow, Tsaritsyno Park, 19.V.2005, coll. K. Tomkovich, 1 male (ZMUM); Chekhovo Distr., Petruhovo, garden, 18–21.VI.2003, coll. K. Tomkovich, 1 male (ZIN); SE of Moscow, 27. VI.2005, coll. K. Tomkovich, 1 female (ZIN); E of Moscow, Fryazevo, 20.VII.–30.VIII.2001, coll. M.E. Tretyakov, 1 female (ZIN); Orekhovo-Zuevo, 23.V.1989, coll. V. Barták, 1 male (OLML); Losiny I., 26.V.1989, coll. V. Barták, 2 females, 1 male, 1 specimen [metasoma absent, sex unknown] (OLML); Abramcevo, 28.V.1989, coll. V. Barták, 1 female (OLML); W of Moscow, Skorotovo, 30.V.1989, coll. V. Barták, 9 males (OLML); Kursk Prov ., Central Chernozemnyy [Black Earth] Nature Reserve, Streletskaya Step’ [steppe], 5–10.V.2008, coll. K. Tomkovich, 1 male (ZIN); Voronezh Prov .: Voronezh Nature Reserve, 27.IV.1950, coll. D. Dovnar[-Zapol’skiy], “ Thersilochus triangularis Gr. det. D. Dovnar[-Zapol’skiy]”, 1 female (ZMUM); same locality and collector, 7.V.1950, “ Thersilochus jocator Gr. det. D. Dovnar[-Zapol’skiy]”, 1 male (ZMUM); same locality and collector, 14.[month illegible].1950, “ Thersilochus truncorum Holmgr. det. D. Dovnar[-Zapol’skiy]”, 1 female (ZMUM); Astrakhan Prov ., Baskunchak Salt Lake, 48°10′01′′N, 46°49′48′′E, semidesert, 3–6.V.2010, coll. K. Tomkovich, 1 male (ZMUM); Krasnodar Terr.:, Lago-Naki Plateau, 44°05′56′′N, 39°59′38′′E, 1700 m a.s.l., 6–8.V.2013, coll. N.E. Vikhrev, 1 male (ZMUM); Gelendzhik, 3.V.1957, coll. G. Viktorov, 1 female (ZMUM); Stavropol Terr.: Shpakovsky Distr., Sengileevskoe reservoir, 3.V.2011, coll. A.S. Ukrainsky, 1 female (ZMUM); 10– 15 km NW of Shpakovskoe [Mikhaylovsk], 4.VI.1987, coll. S.A. Belokobylskij, 1 female (ZIN); 20 km SW of Kislovodsk, Medovye Waterfalls, 20.V.2009, coll. D.R. Kasparyan, 5 females, 7 males (ZIN); Karachay-Cherkessia: Arkhyz, valley of Kizgych River, 1.VII.1976, coll. D.R. Kasparyan, 1 female (ZIN); Teberda, 1300 m a.s.l., broadlived forest, 25–30.V.2009, coll. D.R. Kasparyan, 2 females, 1 male (ZIN); Ingushetia, Nesterovskaya Stanitsa, 9.VI.1972, coll. D.R. Kaspary- an, 1 female [“ T. jocator ”, det. K. Horstmann] (ZIN); Sverdlovsk Prov .: W Ekaterinburg, Verkh-Isetsky Distr., 12.V.2005, coll. T.S. Kostromina, 2 males (ZIN); Ekaterinburg, 20.IV.2005, coll. P. Rudoiskatel’, 1 female (ZIN); Chelyabinsk Prov ., Ilmen’ Nature Reserve, Bol’shoe Miassovo Lake, 21.VI.1983, coll. A.G. Kotenko, 1 female (ZIN); Omsk Prov ., Omsk, 54°51′36′′N, 73°21′36′′E, 23.V.2012, coll. O. Kosterin, 1 female (ZMUM); Altay Terr., 8 km SE of Biysk, Ust’-Katun’, pine forest, 7–8.VII.2007, coll. A.I. Khalaim, 1 female (ZIN); Altai Republic, Turochak Distr., 13–15 km along Lebed’ River, 18.VI.1999, coll. A.G. Kirejtsuk, 1 female (ZIN). * Slovakia, 12 km NE of Bratislava, Šúr Nature Reserve, 30.VI.1991, coll. D.R. Kasparyan, 1 female (ZIN). Sweden, Öland I., Skogsby, forest, meadows, 26.V.2011, coll. D.R. Kasparyan, 3 females (ZIN). Ukraine: Zakarpats’ka Prov ., Beregove Distr., meadow in oak forest, 23.V.1983, coll. Boganich, 1 female (ZIN); Ternopil’ Prov ., Medobori Nature Reserve, Gorodnitskoe Forestry, right bank of Zbruch River, forest, 25.V.1984, coll. V.I. Tolkanitz, 1 female (ZIN); Odessa Prov ., Izmail Distr., W of Yalpuh Lake, Plavni Vill., 11.V.1979, coll. A.G. Kotenko, 1 female (ZIN); Kyiv Prov .: NNE of Kyiv, Novosilky Vill., 4.V.1984, coll. A.G. Kotenko, 1 female (ZIN); SW of Kyiv, Fastiv, 11.V.2003, coll. A.G. Kotenko, 1 female (ZIN); S of Kyiv, Feofaniya Park, 3.VI.1992, coll. A.G. Kotenko, 1 female (ZIN); S of Kyiv, Teremky, 8.VI.2009, coll. D.R. Kasparyan, 1 female (ZIN); Poltava Prov ., Myrgorod, 19.V.1984, coll. A.G. Kotenko, 1 female (ZIN); Kherson Prov ., Black Sea Biosphere Reserve, WSW of Golaya Pristan’, 2.VI.1974, coll. D.R. Kasparyan, 1 female [“ T. jocator ”, det. K. Horstmann] (ZIN); Donets’k Prov .: SE of Debal’tseve, Ol’khovatka Vill., forest, 11.V.1974, coll. D.R. Kasparyan, 1 female [“ T. jocator ”, det. K. Horstmann] (ZIN); 45 km N of Mariupol’, 20.V.1974, coll. D.R. Kasparyan, 1 female (ZIN); NW of Amvrosievka, forest, 15.V.1974, coll. D.R. Kasparyan, 1 female [“ T. jocator ”, det. K. Horstmann] (ZIN); NNE of Amvrosievka, Blagodatnoe Vill., forest, 15.V.1974, coll. D.R. Kasparyan, 1 male (ZIN); Novoazovsk Distr., Khomutovskaya Step’ Nature Reserve, 27.IV.1978, coll. A.G. Kotenko, 1 male (ZIN); Lugans’k Prov .: 15 km E of Sverdlovsk, Provalska Step’ [steppe] Nature Reserve, 5–6.V.1974, coll. D.R. Kasparyan, 7 females, 1 male [5 females, 1 male, “ T. jocator ”, det. K. Horstmann] (ZIN); 3 km NW of Antratsyt, forest, 1–10.V.1974, coll. D.R. Kasparyan, 4 females, 1 male [3 females, 1 male, “ T. jocator ”, det. K. Horstmann] (ZIN); “Nizhnyaya Ilyenka”, right bank of Derkul Riv- er, 2.V.1978, coll. A.G. Kotenko. United Kingdom, East Midlands, Derbyshire, Clough Wood, 10.VII.1996, coll. A. Tereshkin, 1 male (ZIN).

Remarks. Horstmann (1981: 22) record- ed this species (as T. jocator ) from “Mittel- und Südrussland”, Ukraine and Caucasus, based on the Leningrad collection, but without any details. Twenty six specimens of T. cognatus from the Republic of Ingushetia, Leningrad and Yaroslavl provinces of Russia, and the Donets’k, Kherson and Lugans’k provinces of Ukraine, which are stored in the ZIN collection and is provided with the identification labels of Horstmann (see Material examined section), were examined.

Distribution. Europe, Caucasus, Turkey, Kazakhstan, Western Siberia.

71. Tersilochus (Tersilochus) curvator Horstmann, 1981

Material examined. Russia, Moscow Prov ., “Maykovo”, 24. V .1936, coll. Y. Kostylev , 1 female ( ZMUM) .

Distribution. Europe, China (Qinghai), Russian Far East.

72. Tersilochus (Tersilochus) filicornis (Thomson, 1889)

Material examined. Ukraine, Lugans’k Prov ., 15km E of Sverdlovsk, Provalska Step’ [steppe] Nature Reserve, 5. V .1974, coll. D.R. Kasparyan , 1 female [det. K. Horstmann] ( ZIN) .

Remarks. Horstmann (1981: 21) recorded this species from Ukraine, based on the Leningrad collection, but without any details. One specimen of T. filicornis the Lugans’k Province of Ukraine, which is stored in the ZIN collection and have the identification label of Horstmann (see Material examined section), was examined.

Distribution. Europe.

73. Tersilochus (Tersilochus) fulvipes (Gravenhorst, 1829)

Material examined. * Armenia, Syunik Prov ., 8 km E of Ordubad, 20.IV.1982, coll. D.R. Kasparyan, 1 female (ZIN). * Russia: Rostov Prov ., left bank of Don River, in front of Konstantinovsk, 16.V.1986, coll. D.R. Kasparyan, 1 female (ZIN); Stavropol Terr., 10 km SE of Georgievsk, 14.V.1974, coll. D.R. Kasparyan, 1 female [“ Tersilochus? fulvipes ”, det. K. Horstmann] (ZIN); Karachay-Cherkessia, Teberda, 1300 m a.s.l., broadleaf forest, 30.V.2009, coll. D.R. Kasparyan, 2 females (ZIN); Sakhalin Prov ., Kuril Is., Kunashir I. , Yuzhno-Kuril’sk, 11.VII.1973, coll. D.R. Kasparyan, 1 female (ZIN); same island, “ 4 km above river”, 12.VI.1976, coll. Anufriev, 1 female (ZIN).

Distribution. Europe, Caucasus, Russian Far East.

74. Tersilochus (Tersilochus) grandiculus Khalaim, 2012

Material examined. Russia: Jewish Autonomous Prov .: Amurzet , airport, 16. VI .1985, coll. D.R. Kasparyan, 1 female, 1 male ( ZIN); 20 km NW of Amurzet , steppe slopes, 17. VI .1985, coll. D.R. Kasparyan, 1 male ( ZIN); Amurzet , oak forest, 18. VI .1985, coll. D.R. Kasparyan, 6 females, 2 males ( ZIN) ; Primorsky Terr.: Kedrovaya Pad’ Nature Reserve , broadleaf forest, meadows, 4.IX.1982, coll. I . M . Kerzhner, 1 female ( ZIN); 30 km SW of Slavyanka, Andreevka Vill. , forest, meadows, 26.VIII.2003, coll. S . A . Belokobylskij, 1 female ( ZIN); Chernigovka Distr., 15 km E of Dmitrievka, Merkushevka Vill. , forest, meadows, 21.VII.1991, coll. S . A . Belokobylskij, 1 female ( ZIN); Lazovskiy Nature Reserve , cordon Amerika, forest, meadows, 24–29. VI .2006, coll. S. A . Belokobylskij, 1 female ( ZIN); Novokachalinsk, Khanka Lake , mixed forest, meadows, 10.VIII.2001, coll. S . A . Belokobylskij, 1 female ( ZIN); 20 km SE of Spassk-Dal’niy, mixed forest, meadows, 31.VII.1998, coll. S . A . Belokobylskij, 1 female ( ZIN); 20 km ESE of Spassk-Dal’niy, Siniy Mt. Range , forest, meadows, 6.IX.2001, coll. S . A . Belokobylskij, 1 female ( ZIN); Spassk-Dal’niy, forest, meadows, 16. VI .1995, coll. S. A . Belokobylskij, 1 female ( ZIN); NE of Spassk-Dal’niy, Vasil’kovka Vill., forest, meadows, 13.VII.1993, coll. S . A . Belokobylskij, 1 female ( ZIN); Anisimovka , forest, meadows, 29.VIII.2001, coll. S . A . Belokobylskij, 1 female ( ZIN) .

Variation. The species is found to be rath- er variable in body size and morphology. Malar space sometimes short, 0.6 times as long as basal mandibular width. Foveate groove of mesopleuron thin, short to long, S-curved, sometimes extending almost entire length of mesopleuron. Gena and mesopleuron sometimes with fine punctures, mesopleuron often granulate and impunctate below the foveate groove. Ovipositor moderately slender to robust, always strongly upcurved at apex, with two distinct dorsal teeth, and lower valve subapically swollen.

Distribution. Southern Siberia, Russian Far East.

75. Tersilochus (Tersilochus) heterocerus (Thomson, 1889)

Material examined. * Belarus, S Minsk, Lošyca Distr. , at Sinapis , 6–7.VII.1963, coll. T . Bezdenko, 2 females [“ Isurgus heterocerus Thoms. ”, det. K. Horstmann] ( ZMUM) . Germany, Bavaria: NW of Lauterstein, Heldenberg, 560–760 m a.s.l., 48°43.5′N, 9°53.15′E, 4. V GoogleMaps .2016, coll. A. I GoogleMaps . Khalaim, 2 females ( ZIN); N of Bopfingen, Ipf Mt., 48°52′12′′N, 10°21′25′′E, 580– 560 m a.s.l., 3. V GoogleMaps .2016, coll. A. I GoogleMaps . Khalaim, 2 males ( ZIN) .

Remarks. Horstmann (1981: 22) recorded this species from “ Weissrussland ”, based on the collection of Viktorov, but without any details. Two specimens of T. heterocerus from the Minsk Province of Belarus, which are stored in the ZMUM collection and are provided with the identification labels of Horstmann (see Material examined section), were examined.

Distribution. Europe,?Caucasus, Turkey.

76. Tersilochus (Tersilochus) hokkaidoensis Khalaim, 2012

Material examined. * Russia, Sakhalin Prov ., Kuril Is., Kunashir I., Dubovoe Vill. , 31.VIII.1973, coll. D.R. Kasparyan, 1 female (ZIN).

Variation. Female from Kunashir Island is somewhat paler than specimens from Japan; its pterostigma is pale brown and antennal flagellum comprises 20 flagellomeres.

Distribution. Russian Far East, Japan.

77. Tersilochus (Tersilochus) impunctator Khalaim, 2012

Material examined. Russia, Primorsky Terr.: Anisimovka, 43°10′01′′N, 132°45′32′′E, 250 m a.s.l., forest, 5. VI GoogleMaps .1993, coll. S.A. Belokobylskij, 2 females ( ZIN); Vol’no-Nadezhdenskoe ,forest, 4.VII.1996,coll. S. A . Belokobylskij, 1 female ( ZIN); 9 km E of Rettikhovka Vill. , 44°10′40′′N, 132°52′37′′E, 170 m a.s.l., 29. V GoogleMaps .2016, coll. S.A. Belokobylskij, 2 females ( ZIN); Arsen’ev , 44°07′19′′N, 133°16′12′′E, 200 m a.s.l., 30. V GoogleMaps .2016, coll. S.A. Belokobylskij, 1 female ( ZIN); Chuguevka Distr. , N Samarka River , forest, 29. V .1993, coll. S.A. Belokobylskij, 1 female ( ZIN); Gornotaezhnoe , 43°41′24′′N, 132°09′36′′E, 21–22. V GoogleMaps .2016, coll. S.A. Belokobylskij, 3 females, 3 males ( ZIN); Spassk-Dal’niy, forest, meadows, 7,8. VI .1990, coll. S.A. Belokobylskij, 2 females ( ZIN); Ussuriyskiy Nature Reserve , 43°38′38′′N, 132°20′46′′E, 150 m a.s.l., 20–23. V GoogleMaps .2016, coll. S.A. Belokobylskij, 4 females, 1 male ( ZIN) .

Remarks. The male of T. impunctator is record- ed here for the first time. It is similar to the female, with antennal flagellum somewhat slenderer than in the female and comprising 18 flagellomeres.

Distribution. Russian Far East.

78. Tersilochus (Tersilochus) liopleuris (Thomson, 1889)

Material examined. Russia, Primorsky Terr.: Gornotaezhnoe, 43°41′56′′N, 132°09′58′′E, 21–22. V GoogleMaps .2016, coll. S.A. Belokobylskij, 4 females ( ZIN); 20 km SE of Nakhodka , 42°41′17′′N, 133°05′35′′E, 12. VI GoogleMaps .2016, coll. S.A. Belokobylskij, 3 females ( ZIN) .

Distribution. Europe, Russian Far East.

79. Tersilochus (Tersilochus) longicornis (Thomson, 1889)

Material examined. * Lithuania: Kaunas Distr., Ringovė [?Vill.], 8. VI .1990, coll. A. Jakimavičus, 1 female ( ZIN); Varėna Distr. , V .1975, coll. A. Jakimavičus , 1 female ( ZIN) . Russia: Leningrad Prov .: N of St Petersburg, Sosnovo , 2. VI .1984, coll. D.R. Kasparyan , 1 female ( ZIN); 40–43 km S of St Petersburg, between Kobralovo and Semrino, 8.VII.1984, coll. D. R . Kasparyan, 1 female ( ZIN); S of St Petersburg, Pushkin , 29. V .1984, coll. D.R. Kasparyan , 2 females ( ZIN) ; Novgorod Prov ., 20 km NW of Pestovo , Tychkino Vill., coll. V . I . Tobias (all in ZIN): 27. V .–22. VI .1991 (7 females), 1–20. VI .1993 (3 females), 30. V .–4. VI .2000 (2 females).

Distribution. Europe.

80. Tersilochus (Tersilochus) microgaster (Szépligeti, 1899)

Material examined. * Russia, Leningrad Prov .,

55 km S of St Petersburg, Krasnitsy , 6.V.1984, coll .

D. R. Kasparyan, 1 female ( ZIN) . Distribution. Europe.

81. Tersilochus (Tersilochus) nitidipleuris Horstmann, 1971

Material examined. * Russia, Leningrad Prov ., 55 km S of St Petersburg, 4 km W of Krasnitsy , 1–23. V .2011, coll. D.R. Kasparyan , 9 females ( ZIN) . Ukraine, Lugans’k Prov ., NW of Antratsyt, 1. V .1974, coll. D.R. Kasparyan , 1 female [det. K. Horstmann] ( ZIN) .

Remarks. Horstmann (1981: 22) recorded this species from Ukraine, based on the Leningrad collection, but without any details. One specimen of T. nitidipleuris from the Lugans’k Province of Ukraine, which is stored in the ZIN collection and has the identification label of Horstmann (see Material examined section), was examined.

Distribution. Europe.

82. Tersilochus (Tersilochus) obliquus (Thomson, 1889)

Material examined. * Russia, Yaroslavl Prov ., Yaroslavl, N. Kokujev collection, 1 female ( ZIN) . Distribution. Europe.

83. Tersilochus (Tersilochus) obscurator (Aubert, 1959)

Material examined. * Russia, Moscow Prov .: [without locality], ex weevil in stem, 10.X.1968, coll. Voskresenskaya, 2 females [1 female, det. K. Horstmann] (currently deposited in ZIN); Mikhnevo Town , 1. V .1962, coll. E. Antonova, 1 female [det. K. Horstmann ] ( ZMUM) . Ukraine: Donets’k Prov .: NW of Slavyansk , 30.IV.1974, coll. D. R . Kasparyan, 1 female [det. K. Horstmann] ( ZIN); 20 km N of Novoazovsk, Khomutovskaya Step’ [steppe] Nature Reserve, 16–19. V .1974, coll. D.R. Kasparyan, 3 females [det. K. Horstmann] ( ZIN); Lugans’k Prov .: 15 km E of Sverdlovsk, Provalska Step’ [steppe] Nature Reserve, 5. V .1974, coll. D.R. Kasparyan, 1 female [det. K. Horstmann ] ( ZIN); N of Antratsyt, “Lby”, forest, 5–9. V .1974, coll. D.R. Kasparyan, 3 females [det. K. Horstmann ] ( ZIN) .

Remarks. Horstmann (1971: 122–123) record- ed this species from “Zentral- und Südrussland”, based on the Viktorov’s and Prague collections, and from Ukraine ( Horstmann, 1981: 22), based on the Leningrad collection, but without any details. Two specimens of T. obscurator , which are stored in the ZMUM collection, and eight specimens from the ZIN collection, all with the identification labels of Horstmann (see Material examined section), were examined; all of them are from the Moscow Province of Russia and the Lugans’k and Donets’k provinces of Ukraine.

Distribution. Europe, Turkey.

84. Tersilochus (Tersilochus) rufovarius Horstmann, 1981

Material examined. Russia, Moscow Prov ., S of Moscow, N of Vidnoe Town , 26.V.2008, coll. K. Tomkovich, 1 female ( ZIN). Distribution. Europe .

85. Tersilochus (Tersilochus) spasskensis Khalaim, 2012

Material examined. Russia, Primorsky Terr.: Vladivostok, Morskoe Cemetery , forest, 22. VI .2003, coll. S. A . Belokobylskij , 2 females ( ZIN); 25 km SSE of Spassk-Dal’niy, Evseevka Vill., forest, 27. VI .2016, coll. S. A . Belokobylskij , 1 female ( ZIN) .

Distribution. Russian Far East, China (northeast), Japan.

86. Tersilochus (Tersilochus) subdepressus (Thomson, 1889)

Material examined. * Russia: Leningrad Prov ., N of St Petersburg, Sosnovo , 2. VI .1984, coll. D.R. Kasparyan , 1 female ( ZIN) ; Stavropol Terr., Kislovodsk, Kurortny Park , meadow, forest, 27.IV.2018, coll. S . A . Belokobylskij , 1 female ( ZIN) .

Distribution. Europe.

87. Tersilochus (Tersilochus) triangularis (Gravenhorst, 1807)

Material examined. * Lithuania, W of Alytus, Łukśniany Vill. , 20. V.1967, coll. A. Stanionyte , 1 female ( ZIN) . * Russia: Yaroslavl Prov ., Yaroslavl, N. Kokujev collection, 1 female ( ZIN); same data, 9. V.1996, 1 female ( ZIN) ; Kirov Prov ., Urzhum , 25. V.– 10.VI.1900, coll. Krulikovskiy, N. Kokujev collection, 1 female ( ZIN) . Ukraine: Donets’k Prov ., 10 km NE of Amvrosievka , forest, 15. V.1974, coll. D. R. Kasparyan, 1 female ( ZIN); Lugans’k Prov .: 15 km E of Sverdlovsk, Provalska Step’ [steppe] Nature Reserve, 5–6. V.1974, coll. D. R. Kasparyan , 4 females, 1 male [2 females, det. K. Horstmann] ( ZIN); N of Antratsyt, “Lby”, forest, 9. V.1974, coll. D. R. Kasparyan , 1 female ( ZIN) .

Remarks. Horstmann (1981: 22) recorded this species from Ukraine, based on the Leningrad collection, but without any details. Two specimens of T. triangularis from the Lugans’k Province of Ukraine, which are stored in the ZIN collection and have the identification labels of Horstmann (see Material examined section), were examined.

Distribution. Europe, Turkey.

88. Tersilochus (Tersilochus) tripartitus (Brischke, 1880)

Material examined. * Ukraine: Donets’k Prov .: NE of Amvrosievka, “Yasenevoe”, forest, 15.V.1974, coll. D.R. Kasparyan, 2 females [“ Tersilochus? tripartitus ”, det. K. Horstmann] (ZIN); NW of Slavyansk, Slavyanogorsk Town, 30.IV.1974, coll. D.R. Kasparyan, 1 female [“ Tersilochus? tripartitus ”, det. K. Horstmann] (ZIN); Lugans’k Prov ., N of Antratsyt, “Lby”, forest, 9.V.1974, coll. D.R. Kasparyan, 1 female [“ Tersilochus? tripartitus ”, det. K. Horstmann] (1 female).

Distribution. Europe.