Coptorhinini Scholtz, Davis & de Klerk, 2025

Coptorhinini Scholtz, Davis & de Klerk View in CoL , new tribe

Type genus: Coptorhina Hope, 1833 View in CoL , here designated ( Fig. 8 View FIGURES 6–8 )

Diagnosis

Differs from other tribes by a combination of characters, primarily unique wing venation combined with an elongate to ovoid habitus and clypeal dentition varying from comparatively simple to complex (see Description).

Description

Head: Surface of head punctate in males, anterior part of clypeus smooth and shiny in females of all species; anterior clypeal margin distinctly divided into two lateral protrusions in Coptorhina , Frankenbergerius and Sarophorus ; lateral protrusions sexually dimorphic, extended into prongs in some male Frankenbergerius and Sarophorus but absent in females; anterior clypeal margin quadridentate in Delopleurus comprising four short spines with central two more extended than lateral two in some species.

Thorax: Pronotal disc with protuberances in Coptorhina , with a pattern of shiny tubercles in certain Sarophorus species, but evenly convex lacking carinae or protuberances in Frankenbergerius , Delopleurus and some Sarophorus species.

Wings: Metathoracic wings well developed featuring synapomorphic characters that are unique to Coptorhina , Delopleurus , Frankenbergerius and Sarophorus ( Frolov et al. 2008; Fig. 7b View FIGURES 6–8 ): (1) wing membrane brown except for almost transparent basal part, (2) reduced anal area with veins J and AP 3+4 absent, (3) CuA vein widened apically along the wing margin, (4) RA 4 vein not reaching wing margin becoming wide and indistinct apically. Elytra evenly convex and smooth, punctate or tubercula. Epipleurae moderately to strongly sinuate with mid-lateral convexity and correspondingly widened metepisternae in Coptorhina and Delopleurus , less so in Frankenbergerius and Sarophorus ( Frolov et al. 2008; Tarasov & Génier 2015: p. 63).

Legs: Tarsi inserted at tip of tridentate fore tibiae adjacent to terminal spur.

Male genitalia: In Frankenbergeriu s, Coptorhina and Deloplerus ( Coptorhinini new tribe), the arrangement of accessory sclerites in the basal part of the aedeagal sacs (BSc + A with TS spur) is similar to that in Paraphytus ( Paraphytini new tribe) and some other genera of the “basal Scarabaeinae ” of Tarasov & Dimitrov (2016) now in the tribes Elassocanthonini or Endroedyolini ( Tarasov & Génier 2015: pp. 35–37) . However, in the Coptorhinini other sclerites ( AS and FLP) are also present ( Tarasov & Génier 2015: pp. 36–37).

Sexual dimorphism: In addition to characters of the head cited above, further sexually dimorphic characters are present in all genera but relatively weak; in Sarophorus , the anterior tibial spur of males is more strongly curved inwards and downwards and the setation of apices of the middle tibiae is denser in males; in Coptorhina , males differ from females in bearing a slightly convex final abdominal sternite; spur of the anterior tibiae bifurcated in male Frankenbergerius and more or less bifurcated in male Delopleurus ; spur simple and acute in female Frankenbergerius , apically acute to rounded and curved inwards in female Delopleurus .