Galeopsomyia epidius ( Walker )

Galeopsomyia epidius ( Walker) View in CoL

( Figs 23 View Figs 23–28 , 648, 649 View Figs 648–651 , 658, 659 View Figs 656–659 , 887 View Figs 885–896 , 998 View Figs 997–998 , 1107)

Tetrastichus epidius Walker 1847:28 View in CoL . Lectotype ♀ (primary type no. 5.141 3, NHMUK 013458201) designated by Burks (1975:142), examined. U.S.A. ( Florida).

Aprostocetus epidius (Walker) View in CoL , Burks (1975:142).

Galeopsomyia epidius (Walker) View in CoL , Graham & LaSalle (1991:91).

Diagnosis (female). Antenna with a weak constriction between C1 and C2 ( Fig. 998 View Figs 997–998 ); genal carina absent; eyes small 1.0× as long as malar space ( Fig. 887 View Figs 885–896 ); mesoscutellum ( Figs 648 View Figs 648–651 , 658 View Figs 656–659 ) with strong reticulation, meshes isodiametric in posterior half, elongate in anterior half, submedian grooves indistinct and partly absent, slightly curved and diverging towards posterior part, with a ±complete median groove; dorsellum without median carina; propodeum with strong reticulation ( Figs 648 View Figs 648–651 , 658 View Figs 656–659 ), callus with six setae; gaster 3.1× as long as wide, Gt 2 0.3× as long as Gt 4, tergites with strong reticulation ( Figs 648 View Figs 648–651 , 658 View Figs 656–659 ).

Description (female nontype). Length of body 2.6mm (specimen measured) (variation in material 2.3–3.1mm).

Scape yellowish-brown with dorsal part brown, pedicel and flagellum dark brown. Head with frons bluish-green, antennal scrobes black, clypeus dark brown. Vertex bluish-green. Thoracic dorsum golden-green. Legs with fore and mid coxae black with metallic tinges, hind coxa golden-green; trochanters dark brown; femora black with apex yellowish-brown; fore tibia brown, mid and hind tibiae with basal half brown and apical half yellowish-brown; T1–3 yellowish-brown, T4 dark brown. Petiole black. Gaster golden-green; gonoplac black.

Antenna with a weak constriction between C1 and C2. Frons with strong reticulation, clypeal area smooth, antennal scrobes with weak reticulation; without genal carina. Vertex with strong reticulation.

Mesoscutum with strong reticulation, meshes isodiametric on sidelobes, elongate on midlobe; midlobe without median groove, with seven scattered adnotaular setae. Mesoscutellum with strong reticulation, meshes isodiametric in posterior half, elongate in anterior half; submedian grooves indistinct and partly absent, slightly curved and diverging towards posterior part; with a ±complete median groove; frenal groove wide and subdivided by carinae; with an expanded and strongly reticulate part behind frenal groove. Dorsellum with strong reticulation. Propodeum with a strong and relatively wide median carina that expands posteriorly; callus with six setae. Fore and hind coxae with strong reticulation, mid coxa with weak reticulation; hind coxa with a carina along posterior margin. Fore wing with five setae on dorsal surface of submarginal vein; speculum open.

Petiole very short, just a narrow band. Gaster elongate; tergites with strong reticulation.

Relative measurements: head length, dorsal view 21; head length, frontal view 41; POL 13; OOL 7; lateral ocellus diameter 4; head width 50; mouth width 16; malar space 18; eye length 18; scape length 21; scape width 4; pedicel+flagellum length 50; pedicel length 7.5; pedicel width, dorsal view 4; F1 length 8; F1 width 5; F2 length 7; F2 width 6; F3 length 6.5; F3 width 6; clava length 17; clava width 7; C3 length 5; spicule length 1; mesosoma length 62; mesosoma width 44; midlobe of mesoscutum length 24; mesoscutellum length 22; mesoscutellum width 21; median part of mesoscutellum width (measured medially) nm; median part of mesoscutellum, width in anterior part nm; median part of mesoscutellum, width in posterior part nm; lateral part of mesoscutellum (measured medially) nm; dorsellum length 3.5; propodeum length 8; costal cell length 34; costal cell width (measured at widest part) 2.5; marginal vein length 36; stigmal vein length 10.5; gaster length 124; gaster width 40; Gt 2 length (measured medially) 6.5; Gt 4 length (measured medially) 20; Gt 7 length (measured medially) 23; Gt

7 width (measured at base) 15; longest cercal seta length 11; shortest cercal seta length 5. Male. Length of body 1.6–1.8mm.

Antenna ( Fig. 998 View Figs 997–998 ) with scape dark brown, elongate with dorsal and ventral edges ±parallel; plaque dark brown and situated in apical half of scape; dorso-basal whorls of long setae present on F1–F4 and C1. Gaster long ovate. Otherwise as in female.

Relative measurements: head length, dorsal view 17; head length, frontal view 32; head width 44; mouth width 14; malar space 14; eye length 16; scape length 22; scape width 4.5; plaque length 9; pedicel length 6.5; pedicel+flagellum length 70; F1 length 6; F1 width 4; F2 length 11; F2 width 5; F3 length 12; F3 width 4.5; F4 length 11.5; F4 width 4; clava length 23; clava width 4; mesosoma length 57; mesosoma width 39; gaster length 66; gaster width 30; longest subbasal seta on F1, length 28.

Hosts. Reared from Asphondylia sp. ( Diptera : Cecidomyiidae ) on Ambrosia psilostachya ( Asteraceae ), and an unidentified Cecidomyiidae on Xanthium sp. ( Asteraceae ).

Distribution. U.S.A. ( Florida ( Walker 1847), Oklahoma, Texas ( new records)).

Material examined.

Type material: lectotype ♀ ( NHMUK). Additional material ( 14♀ 7♂, MZLU, NHMUK): 13♀ 7♂ from U.S.A., Oklahoma, Payne Co., Mehan, 36.014339°N, - 96.996774°W, 10.viii.2016, ex Asphondylia ( Diptera : Cecidomyiidae ) on Ambrosia psilostachya ( Asteraceae ), M.W. Palmer ( MZLU, NHMUK); 1♀ U.S.A., Texas, Navasota, v.1935, ex Cecidomyiidae on Xanthium, L.H. Hitchcock.