Reishia haemastomoides (Hoernes & Auinger, 1882)

Reishia haemastomoides (Hoernes & Auinger, 1882)

Figs 48 View FIGURE 48 , 52A–D View FIGURE 52

[ Purpura View in CoL ] haemastoma Lam. View in CoL — Hörnes 1848: 17.

Purpura haemastoma Lin. View in CoL —Hörnes 1853: 167, pl. 13, fig. 18.

* Purpura (Stramonita) haemastomoides nov. form.—Hoernes & Auinger 1882: 151.

Purpura (Stramonita) haemastomoides Hö. Au. View in CoL — Boettger 1902: 23.

Purpura haemastomoides Hoernes & Auinger View in CoL —Friedberg 1928: 593, pl. 38, fig. 6.

Thais (Stramonita) haemastomoides Hoernes et Auinger — Csepreghy-Meznerics 1954: 36, pl. 4, figs. 9–10.

Th [ais]. (S [tramonita].) haemastomoides (R. Hörn., et Au.)— Sieber 1958: 146.

Thais (Stramonita) haemastomoides (Hoernes & Auinger, 1882) —Kojumdgieva in Kojumdgieva & Strachimirov 1960: 153, pl. 41, fig. 1.

Thais (Stramonita) haemastomoides Hoernes & Auinger, 1882 — Strausz 1966: 282, fig. 129a, pl. 35, figs. 13–15, pl. 36, figs. 1–5, pl. 64, fig. 1.

Thais (Stramonita) haemastomoides Hoernes & Auinger — Krach 1981: 68, pl. 17, fig. 9, pl. 18, figs. 1–4, 6.

Stramonita haemastomoides (Hoernes & Auinger, 1882) View in CoL — Landau et al. 2013: 155, pl. 23, fig. 4.

Janssenia echinulata (Pusch, 1837) View in CoL — Kovács et al. 2018: 126, figs. 8O–Q [non Janssenia echinulata (Pusch, 1837) View in CoL ].

Janssenia echinulata (Pusch, 1837) View in CoL — Kovács 2020: 458, pl. 4, figs. 3–10 [non Janssenia echinulata (Pusch, 1837) View in CoL ].

non Thais (Stramonita) haemastomoides Hoernes & Auinger, 1882 — Atanacković 1985: pl. 31, figs. 9–10 [= Janssenia echinulata (Pusch, 1837) View in CoL ].

Type material. Lectotype (designated herein): NHMW 1846/0037/0178, SL: 49.6 mm, MD: 32.4 mm, Gainfarn ( Austria), illustrated in Hörnes (1853: pl. 13, fig. 18), figs. 52B 1 –B 3 . Paralectotypes: NHMW 1867/0019/0066, SL: 37.3 mm, MD: 24.1 mm, CoŞteiu de Sus ( Romania), figs. 52A 1 –A 3. NHMW 1854/0035/0139a, SL: 34.0 mm, MD: 21.6 mm, Lăpugiu des Sus ( Romania), figs. 52C 1 –C 3. NHMW 1854/0035/0139b, SL: 39.9 mm, MD: 26.5 mm, Lăpugiu des Sus ( Romania), figs. 52D 1 –D 3 .

Additional material. 21 spec., NHMW 1854/0035/0138, Lăpugiu des Sus ( Romania) ; 19 spec., NHMW 1854/0035/0137, Lăpugiu des Sus ( Romania) ; 3 spec. NHMW A1521, Lăpugiu des Sus ( Romania) .

Revised description. Large, ovate shell with conical spire; apical angle ~68–73°. Protoconch unknown. Teleoconch of up to five whorls. Suture weakly impressed; shallowly undulating. Early teleoconch whorls with concave subsutural ramp and angled shoulder placed slightly below mid-whorl. Sculpture of about 13 nodes along shoulder; SP, adis, IP, abis weak, close-set; P1 prominent, s1 and tertiary threads indistinct. Penultimate whorl with swollen subsutural collar and scabrose spiral cords. Last whorl ovate, attaining ~80% of total height; swollen subsutural collar, deeply concave subsutural ramp, rounded shoulder, convex periphery, slowly contracting below. On sutural ramp indistinct SP, adis, IP, abis; on convex part of whorl prominent P1, P2, P3 forming prominent convex cords with up to 15 pointed tubercles; P4 prominent, but weaker than posterior primary cords; P5 weak on siphonal canal; secondary cords s1 to s5, weak, indistinguishable from tertiary threads. Fasciole strongly swollen, strongly twisted, delimiting deep, broad pseudoumbilicus. Aperture wide, ovate; outer lip thin, crenulate at primary cords, with wide basal margin; three wide-spaced, dot-like denticles deep within aperture corresponding to ID (weak), D1 to D4 (simple) Anal canal deeply incised, V-shaped. Siphonal canal moderately short, wide, open, weakly bent to the left. Columella broadly excavated, strongly angled at siphonal canal with weak columellar fold at angulation. Columellar callus forming broad, thick, well delimited rim, erect at pseudoumbilicus, adherent above.

Discussion. As discussed above, Janssenia echinulata (Pusch, 1837) and Reishia haemastomoides (Hoernes & Auinger, 1885) are very similar and can only be distinguished based on their apertural features. Therefore, literature records of both species are hard to confirm from the illustrations, especially if apertures are not well documented. Therefore, the chresonymy above may be partly incorrect and the paleogeographic distribution will need confirmation in some cases.

Paleoenvironment. Unknown.

Distribution in Central Paratethys. Badenian (Middle Miocene).

Distribution in Central Paratethys. Badenian (Middle Miocene): Vienna Basin: Gainfarn ( Austria), Mikulov ( Czech Republic) (Hoernes & Auinger 1882); Eisenstadt-Sopron Basin: Forchtenau ( Austria) (Hoernes & Auinger 1882); Pannonian Basin: Letkés ( Hungary) ( Kovács et al. 2018); Southern Pannonian Basin: Hrvaćani ( Bosnia and Herzegovina) ( Atanacković 1985); Bükk Mountains: Borsodbóta ( Hungary) ( Csepreghy-Meznerics 1972); Bakony Mountains: Bánd ( Hungary) ( Kovács 2020); Mecsek Mountains: Mecsekpölöske, Hidas, Hosszúhetény ( Hungary) ( Kovács 2020); Cserhát Hills: Kelet-Cserhát ( Hungary) ( Csepreghy-Meznerics 1972); Făget Basin: CoŞteiu de Sus, Buituri, Lăpugiu des Sus ( Romania) (Hoernes & Auinger 1882; Kovács 2019); Dacian Basin: Bivolare, Bozuritsa, Yasen, Opanec, Radomirci, Staropatica, Trifonovo ( Bulgaria) (Kojumdgieva in Kojumdgieva & Strachimirov 1960).

Proto-Mediterranean Sea. Serravallian (Middle Miocene): Karaman Basin: Akpınar ( Turkey) ( Landau et al. 2013).

Genus Penethais Vermeij, 2024

Type species. Purpura triangularis Blainville, 1832 View in CoL ; original designation by Vermeij (2024: 9). Present-day, Eastern Pacific.

Discussion. Herein, we tentatively place a single Paratethyan species in the eastern Pacific–western Atlantic genus Penethais . This species was originally described by Kovács et al. (2018) as Thaisella ? sp. Thaisella Clench, 1947 , as currently listed by MolluscaBase Eds. (2024c), comprises morphologically very different species. The type species Thaisella trinitatensis ( Guppy, 1869) has a moderately high, coronate spire, a globose-ovate last whorl with tuberculate spiral cords and strongly scabrose subsutural cord, a wide and deep pseudoumbilicus and a wide aperture with wide outer lip bearing delicate lirae within. Thaisella chocolata ( Duclos, 1832) is globose-ovate with an almost smooth shell with subobsolete pseudoumbilicus, whereas Thaisella coronata ( Lamarck, 1816) , Thaisella callifera ( Lamarck, 1822) and Thaisella guatemalteca Simone, 2017 have prominent, broad, tuberculate spiral cords and a very wide, deep pseudoumbilicus. The Paratethyan species has little in common with T. trinitatensis but is superficially similar to T. callifera . The presence of a weak columellar fold and the narrow pseudoumbilicus of the Paratethyan species make a relationship with Thaisella unlikely. In contrast, the presence of a columellar fold is consistent with the placement in Acanthais and Penethais . The poor preservation does not allow to confirm the presence of a labral tooth in the Paratethyan shell, which is typical for Acanthais (type species: Buccinum brevidentatum Wood, 1828 by original designation and single member of the genus). Kovács & Vicián (2024) followed this view and placed this species in? Acanthais and described it formally as new species. However, Acanthais brevidentata displays rounded internal denticles, lacking in the Paratethyan species. In addition, Vermeij (2024) erected Penethais , chose Purpura triangularis Blainville, 1832 as type species, and included two other species: P. brevicula ( Vermeij, 2001) from the Burdigalian of Venezuela and P. callaoensis (J.E. Gray, 1828) from Peru and Panama. Strong rows of nodes ranging from P1 to P4, missing rounded internal denticles and reduced or missing labral spines are features of Penethais shared with the Paratethyan species. Nevertheless, the Paratethyan species bears very prominent primary cords which are not found in Penethais species.