Tripterotyphis tripterus ( Grateloup, 1833 )

Tripterotyphis tripterus ( Grateloup, 1833) View in CoL

Figs 21 View FIGURE 21 , 25A–D View FIGURE 25

* Typhis tripterus Nob. View in CoL — Grateloup 1833: 160.

M [urex]. tripterus Grat. View in CoL — Grateloup 1845: pl. 30, fig. 22.

Murex (Typhis) Wenzelidesi Hörnes —Hörnes 1853: 264, pl. 26, figs. 12a–d.

Typhis Wenzelidesi — Quenstedt 1884: 654, pl. 211, fig. 39.

Typhis Wenzelidesi M. Hoern. —Hoernes & Auinger 1885: 228.

Typhis wenzelidesi M. Hö. View in CoL — Boettger 1902: 32.

Typhis Wenzelidesi Hörnes View in CoL — Peyrot 1903: 70, pl. 3, fig. 2.

Typhis wenzelidesi M. Hö. View in CoL — Boettger 1906: 46.

Pterotyphis tripterus Grateloup — Cossmann & Peyrot 1924: 240, pl. 15, figs. 13–14. Typhis View in CoL ( Pterotyphis View in CoL ?) wenzelidesi Hörnes View in CoL — Peyrot 1938: 197.

Typhis (Pterotyphis) wenzelidesi Hörnes, 1856 View in CoL — Glibert 1952: 295.

T [yphis]. (T [yphinellus].) wenzelidesi Hörn. View in CoL — Sieber 1958: 145.

[ Tripterotyphis View in CoL ] wenzelidesi Hörnes, 1853 View in CoL — Vokes 1971: 117.

Typhis wenzelidesi (Hoernes, 1853) View in CoL — Stojaspal 1978: 339.

Tripterotyphis wenzelidesi (Hörnes) — Vicián et al. 2017: 270, pl. 2, fig. 19. Tripterotyphis tripterus ( Grateloup, 1833) View in CoL — Kovács et al. 2018: 118, figs. 5C–D. Tripterotyphis tripterus (Grateloup) View in CoL — Kovács 2020: 463, pl. 2, figs. 5–6.

Type material. The syntypes are stored in the University of Bordeaux ( France) .

Illustrated material. NHMW 2024/0161/0001, SL: 10.0 mm, MD: 4.8 mm, Steinebrunn ( Austria), figs. 25A 1 –A 3. NHMW 1871/0011/0020, SL: 15.0 mm, MD: 7.0 mm, Steinebrunn ( Austria), figs. 25B 1 –B 3. NHMW 1860/0001/0221, SL: 12.5 mm, MD: 5.5 mm, Steinebrunn ( Austria), illustrated in Hörnes (1853: pl. 26, fig. 12), holotype of Murex (Typhis) wenzelidesi Hörnes, 1853 , figs. 25C 1 – C 3. NHMW 2024/0161/0002, SL: 9.9 mm, MD: 5.1 mm, Steinebrunn ( Austria), figs. 25D 1 –D 2 .

Revised description. Small, relatively slender fusiform, trivaricate shell; apical angle ~42°. Protoconch incomplete, high conical of about three convex whorls. Teleoconch of up to five whorls. Suture deeply incised, shallowly undulating. Early teleoconch whorls weakly convex with three very prominent, raised varices with adapically directed shoulder spine. Spiral sculpture strongly reduced; P1, P2 forming subobsolete broad cords with concave pits in interspaces at base of varices; P1 forming long tubular shoulder spine at end of first teleoconch whorl. Growth lines forming very delicately scabrose sculpture over entire shell surface. Last whorl attaining ~70% of total height; weakly convex periphery moderately constricted below with high base. P1–P6 prominent on varices; P1 (long tubular shoulder spine); P2–P6 forming bead-like tubercles on apertural side of varices, separated by deep, concave pits. Base of shoulder spines and terminal tubercles of P2–P6 fringed by delicate, raised foliose expansion; ADP, MP and ADP weak. Fasciole swollen, narrow, delimiting broad, shallow pseudoumbilicus. Aperture narrowly ovate. Outer lip strongly thickened by terminal varix, with crenulate edge. Prominent D1–D5; D2, D3 bifid. Anal canal narrowly incised at P1, accentuated by weak to subobsolete parietal denticle. Siphonal canal long, fused in fully grown specimens, straight to slightly bent to the left. Columella weakly excavated, smooth, slightly twisted at siphonal canal. Columellar callus forming broad, sharply delimited rim, erect over siphonal canal.

Discussion. Murex wenzelidesi was introduced by Hörnes (1853) as new name for “1840 Murex tripterus Grateloup ” (referring to the plates in Grateloup 1845), which he considered to be preoccupied. This assumption, however, was incorrect, as Grateloup (1833) had established this species in Typhis and not in Murex , which indeed would have been preoccupied by Murex tripterus Born, 1778 . Nevertheless, Murex (Typhis) wenzelidesi was not explicitly declared to be a new name for Grateloup’s species but was based on specimens from Steinebrunn ( Austria). Herein, we follow Kovács et al. (2018) and treat Murex (Typhis) wenzelidesi as subjective junior synonym of Tripterotyphis tripterus ( Grateloup, 1833) because the specimen illustrated by Cossmann & Peyrot (1924, pl. 15, figs. 13–14) from the Early Miocene of France is indistinguishable from the Paratethyan specimens, including the bifid denticles.

Among the extant congeners, Tripterotyphis jamesmacleani Wiedrick & Houart, 2021 , from the eastern Pacific, is very similar to Tripterotyphis tripterus in shape and sculpture but differs in its higher spire.

Paleoenvironment. Shallow marine, inner neritic; the occurrence at Steinebrunn might point to seagrass environments (own data, M.H.).

Distribution in Central Paratethys. Badenian (Middle Miocene): Vienna Basin: Steinebrunn ( Austria) (Hoernes & Auinger 1885); Eisenstadt-Sopron Basin: Forchtenau ( Austria) (Hoernes & Auinger 1885); Pannonian Basin: Letkés ( Hungary) ( Kovács et al. 2018); Bakony Mountains: Bánd ( Hungary) ( Kovács 2020); Făget Basin: CoŞteiu de Sus ( Romania) ( Boettger 1902).

Northeastern Atlantic. Aquitanian-Burdigalian (Early Miocene): Aquitaine Basin: Léognan, Saint-Paul-lésDax ( France) ( Cossmann & Peyrot 1924); Langhian (Middle Miocene): Loire Basin: Mirebeau ( France) ( Peyrot 1938).