Scopesis rufolabris ( ZETTERSTEDT , 1838) Riedel & H & Schmidt, 2024

Scopesis rufolabris ( ZETTERSTEDT, 1838) View in CoL , stat. nov. (figs 10, 28, 44, 64, 84, 100)

Original: Bassus rufolabris ZETTERSTEDT, 1838: 380 , ♀.

Lectotype: (♀) no label (Lund), synonymized with Sc. frontator by ROMAN (1912: 256), type examined by Horstmann.

syn. nov. Mesoleius rufonotatus HOLMGREN, 1876: 31 , ♀ ♁.

Lectotype: (♀) sc./Bhn. (Stockholm), type examined. syn. nov. Scopesus thomsoni HABERMEHL, 1925: 180. ♀ ♁, synonymized with Sc. rufonotata by

TEUNISSEN (1953: 35), type not examined.

D e s c r i p t i o n: ♀. Body length (6.0)– 7.5–9.5 mm. Flagellum with 36–44 segments; 1 st flagellar segment 5.0–6.3× as long as wide and 1.7–2.0× as long as 2 nd segment; 2 nd segment 2.5–3.0×, 3 rd segment 2.2–2.8× and preapical segment 1.5–1.7× as long as wide. Temple moderately and roundly narrowed. OED 1.4–1.7× ocellar diameter. Face granulated and with fine superficial punctures, dull. Clypeus moderately convex, ± bulging medially, with sparse rather coarse punctures; apical margin slightly concave, with small flat extensions laterally. Mandibular teeth equal or ventral mandibular tooth slightly larger than dorsal tooth. MI 0.4–0.5.

Mesoscutum finely granulate, dull. Notaulus usually impressed in frontal 0.3 of mesoscutum. Mesopleuron granulate, without clear punctures ventrally and with ± distinct striae dorsally. Speculum with microsculpture, shining. Area superomedia sometimes with weak but incomplete lateral carina. Area petiolaris with complete carina, sometimes obsolete medially. Lateral longitudinal and pleural carinae of propodeum sometimes ± obsolete. Hind femur 4.5–5.3× as long as wide. Vein 2rs-m 0.6× as long as vein M between 2rs-m and 2m-cu. Vein 1cu-a interstitial. Nervellus of hind wing inclivous, intercepted in its apical 0.7–0.8. 1 st tergite 1.4–1.8× as long as wide, without latero-median carina. 2 nd tergite 1.3–1.6× as wide as long

Color: black. Flagellum red or ochreous, ± black basally. Palps brown. Mandible centrally and apical ½–3/4 of clypeus reddish-yellow. Hind edge of pronotum brown. Tegula brown or reddish, sometimes yellow. Often 1 st tergite ±, 2 nd and 3 rd tergites mainly or entirely and sometimes 4 th tergite red or reddish-brown. Coxae, trochanters and trochantelli black; legs otherwise red; hind femur often slightly infuscate basally and/or apically; hind tibia red or reddish-yellow, rarely yellowish, sometimes narrowly infuscate basally and usually infuscate in apical 0.2–0.3; spurs of hind tibia ochreous or yellowish; hind tarsus black. Pterostigma yellowish to ochreous. Var. ♀: 1 specimen from Mt. Canigou / France ( ZSM): Flagellum very slender ; 1st flagellar segment 6.0×, 2 nd segment 2.9× and 3 rd segment 2.8× as long as wide. Hind femur 5.3× as long as wide. Metasoma black, 2 nd and 3 rd tergites with dark reddish shine .

♁. Body length 8.0– 8.5 mm. Flagellum with 35–37 segments; 1 st flagellar segment 5.1– 6.2× as long as wide and 1.7–1.8× as long as 2 nd segment; 2 nd segment 2.6–3.0× and 3 rd segment 2.5× as long as wide. Temple distinctly and roundly narrowed. OED 1.5×. Clypeus distinctly convex and bulging forwards medially, with sparse punctures, laterally with flat extensions. Lower mandibular tooth slightly larger. MI 0.3. Mesopleuron granulate, without punctures and distinct striae; speculum almost smooth, shining. Area superomedia with ± distinct lateral carina. Area petiolaris with complete carina. Hind femur 4.6–5.5× as long as wide. 1 st tergite 1.8–2.0× as long as wide, without latero-median carina. 2 nd tergite 1.1–1.2× as wide as long.

Color: black. Flagellum red, black basally. Palps, mandible, clypeus, face except brown median stripe, sometimes ventral half of frontal orbit and ventral spot at scape, sometimes spot on propleuron, hind edge of pronotum and tegula yellow. 2 nd to 4 th tergites red, rarely metasoma ± infuscate with dark reddish suffusion. Coxae black, apically yellow or fore and mid coxae entirely yellow; hind coxa sometimes yellow apically. Trochanters and trochantelli yellow, hind trochanter ± black dorsally. Legs red; hind femur sometimes slightly infuscate basally and usually apically; hind tibia reddish or reddish-yellow, infuscate in apical 0.3, hind tarsomeres black, with pale bases. Pterostigma brownish.

Host: Tenthredo ferruginea SCHRANK, 1776 (Hym. Tenthredinidae ) (ZSM).

D i s t r i b u t i o n: Probably Transpalaearctic, but the report from Irkutsk environment ( MOCSARY & SZÉPLIGETI 1901) is questionable and unconfirmed, studied material from Bulgaria *, Finland, France, Georgia *, Germany, Italy, Sweden, Switzerland, Turkey *, United Kingdom.

Taxonomical remarks: Scopesis rufolabris was usually regarded as a synonym of Sc. frontator since ROMAN (1912: 256) had synonymized both species (see HORSTMANN 2007: 81), but the comparison of the type material of both species shows some differences. The lectotype of Sc. rufolabris has longer basal flagellar segment (e. g. 3 rd segment 2.5× as long as wide) than Sc. frontator (3 rd flagellar segment 2.0× as long as wide). In Sc. rufolabris , the structure of mesopleuron is weaker and the red hind femur is slightly infuscate apically, whereas the mesopleuron has a coarser structure and the hind femur is entirely red in Sc. frontator . Due to these differences, we separate both species here. Sc. bicolor differs from Sc. rufolabris mainly by its entirely black hind femur.