Tovomita nidiae L. Marinho

44. Tovomita nidiae L. Marinho View in CoL in Willdenowia 49: 349. 2019. – Holotype: Venezuela, Bolívar, selva de galería y sabana a lo largo del Río Kanarakuni, norte y noroeste de la Misión de Campamento Sanidad del Río Kanarakuni, 400 m, 17–29 Mar 1967, J. Steyermark 98189 (NY [ NY02859717 ]!; isotype: NY!).

Description — Trees up to 15 m tall, prop roots inconspicuous; exudate yellow, abundant. Petioles 1.2–3.3 cm long, coloration not seen, smooth, lenticels absent. Leaf blades 8.4–14.3 × 3.3–5.8 cm, rich green above, pale green below in vivo, greenish to yellowish in sicco, black dots absent, subcoriaceous, obovate, base decurrent, apex rounded to straight; papillae, lenticels and fungal spots absent; exudate canals inconspicuous. Venation: secondary veins 5–6 pairs, 15–28 mm apart from each other, forming angle 40°–45° to midvein, immersed adaxially, prominent abaxially, arcuate connecting near margin; intersecondary veins present, one to three per intercostal area, much thinner than secondary veins, distal course reticulating, c. 50 % of subjacent secondary length; tertiary veins reticulate, irregular; intramarginal vein absent. Inflorescences: ♂ lax cyme with 5 basal branches and up to 35 flowers, lacking terminal flower, ♀ not seen. Pedicels 8–16 mm long, green, distal and proximal portion with same gauge, proximally articulated in lateral flowers of dichasia, rarely in central flower; calyptrae and lenticels absent. Floral buds 7–9 mm long, ovoid, apex rounded, lenticels absent. Sepals 4, 7–9 × 6–7 mm, ovate to oblong, apex rounded, greenish to white; petals 4, 11–12 × 3.5–4 mm, oblong, deflexed, apex rounded, white. Staminate flowers: stamens 60–65, 4.5–7 mm long, heterodynamous; filaments terete, creamy-white; anthers 0.5– 0.9 mm long, connective not exceeding thecae; pistillode c. 0.5 mm long, styles 6, amorphous. Pistillate flowers: staminodes not counted, 4.5–7 mm long; filaments terete, coloration not seen; ovary c. 7 mm long, locules and stigmas not counted; stigmas c. 2.5 mm in diam., coloration not seen. Capsules not seen.

Iconography — Illustration available in Marinho & al. (2019b: 347).

Distribution — Colombia (Caquetá), Venezuela (Bolívar). Fig. 66.

Conservation status — Data Deficient (DD, Marinho & al. (2019b).

Recognition and discussion — Tovomita nidiae can be recognized by the low number of secondary veins (5–6 pairs), which form an angle 40°–45° to the midvein and connect to each other near the margin ( Fig. 14B). This number of secondary veins is also found in T. schomburgkii , from which it can be differentiated even when sterile by the angle of the secondary veins with midvein (60°–75° in T. schomburgkii ).

Specimens examined — COLOMBIA: CAQUETÁ: Solano, región Araracuara, sector Chiribiquete , 00°15'00"S, 72°22'41"W, 210 m, 4 Dec 2010, bud, F GoogleMaps . Castro 10612 ( COAH!); Morelia, vereda Sinaí , 01°24'03"N, 75°45'49"W, 283 m, 4 Oct 2015, fl. ♀, D. Cárdenas & al. 46299 ( COAH!) GoogleMaps . — VENEZUELA: BOLÍVAR: Sucre, pica desde Santa Maria de Erebato , 04°59'N, 64°49'W, 430 m, Jan 1989, bud, E GoogleMaps . Sanoja 2382 ( NY!); Distrito Cedeño, along tributary of Río Erebato , 05°09'N, 64°34'W, 170 m, 12 Mar 1992, B GoogleMaps . Boom & E . Marin 10438 ( NY 2 sheets!) .