Tovomita nebulosa L. Marinho & Luján

42. Tovomita nebulosa L. Marinho & Luján View in CoL in Acta Amazonica 50: 326. 2020. – Holotype: Venezuela, Amazonas, Departamento Río Negro, Cerro de La Neblina Camp IV, 15 km NE of Pico Phelps   GoogleMaps , north branch of river in canyon, 780 m, 00°51'N, 65°57'W, 15–18 Mar

1984, R. Liesner 16670 ( NY [ NY02859712 ]!; isotypes: BRIT [ BRIT554535 About BRIT ] photo!, MO, US photo!, VEN [no. 327548] photo!) .

Description — Trees up to 3 m tall, prop roots and exudate not seen. Petioles 0.9–1.3 cm long, coloration not seen, smooth, lenticels absent. Leaf blades 4–10 × 2.6–5 cm, dark brown adaxially and light brown abaxially in sicco, black dots absent, subcoriaceous, oblong to obovate, base decurrent, apex acute to rounded, sometimes with drip tip; papillae, lenticels and fungal spots absent; exudate canals immersed on both surfaces, visible in thin blackish lines abaxially, parallel to secondary veins. Venation: secondary veins 20–23 pairs, 2.5–3.2 mm apart from each other, forming angle 65°–75° to midvein, prominent on both surfaces, slightly arcuate and connecting near margin; intersecondary veins present, one to two per intercostal area, similar to secondary veins,> 50 % of subjacent secondary length; tertiary veins percurrent sinuous; intramarginal vein present. Inflorescences: ♂ lax cyme with 3 basal branches and up to 35 flowers, with terminal flower, lenticels absent, ♀ not seen. Pedicels 4–9 mm long, coloration not seen, distal and proximal portion with same gauge, articulated in middle in lateral flowers of dichasia; calyptrae and lenticels absent. Floral buds 5–7 mm long, narrowly oblong, apex cuspidate to slightly rostrate, lenticels absent. Sepals 2, 5–7 × 2.5–3 mm, oblong to narrowly ovate, apex cuspidate to acuminate, coloration not seen; petals 4, 3.5–6.2 × 1–1.5 mm, linear to narrowly oblong, apex acute to cuspidate, coloration not seen. Staminate flowers: stamens c. 20, 3.3–4.8 mm long, heterodynamous; filaments terete, coloration not seen; anthers 0.3–0.5 mm long, coloration not seen, connective not exceeding thecae; pistillode c. 3 mm long, conic, rudimentary stigmas 4, amorphous. Pistillate flowers and capsules not seen.

Iconography — Illustration available in Marinho & al. (2020: 151).

Distribution — Venezuela (Amazonas). Fig. 66.

Conservation status — Data Deficient (DD, Marinho & al. 2020).

Recognition and discussion — The floral buds of Tovomita nebulosa become blackened in sicco, similar to those of T. carinata and T. tenuiflora . Tovomita nebulosa can be distinguished from both species by the number of stamens (c. 20 in T. nebulosa vs> 45 in T. carinata and T. tenuiflora ); moreover, it differs from T. tenuiflora by the number of secondary veins (<23 pairs vs> 35 pairs) and from T. carinata by the number of carpels (4 vs 5).