Tovomita riedeliana Engl.

47. Tovomita riedeliana Engl. View in CoL in Martius & al., Fl. Bras. 12(1): 449. 1888. – Holotype: Brazil, “loco accuratius haud addicto”, L. Riedel [123] (LE photo!).

Description — Trees up to 20 m tall, prop roots conspicuous; exudate yellow, abundant. Petioles 2.2–3.5 cm long, green or reddish, smooth, lenticels absent. Leaf blades 7.5–15.5 × 3–7 cm, copper-coloured in sicco, black dots absent, coriaceous, oblong to obovate, base convex to decurrent, apex acuminate; papillae, lenticels and fungal spots absent; exudate canals inconspicuous. Venation: secondary veins 16–20 pairs, 3–4 mm apart from each other, forming angle 45°–65° to midvein, prominent abaxially, immersed adaxially, straight near margin; intersecondary veins present, one per intercostal area, similar to secondary veins, parallel to a major secondary,> 50 % of subjacent secondary length; tertiary veins inconspicuous; intramarginal vein present. Inflorescences: ♂ lax cyme with 3 basal branches and up to 15 flowers, lacking terminal flower, ♀ dichasium, lenticels absent. Pedicels 13–55 mm long, green, distal and proximal portion with same gauge, proximally or in middle articulated in all flowers of dichasia; calyptrae and lenticels absent. Floral buds 5.5–8 mm long, spheroid, apex rounded, lenticels absent. Sepals 2–4, 5.5–8.5 × 5–6.5 mm, circular to oblong, apex rounded, light green; petals 4, 6.5–9 × 6–7 mm, circular to oblong, reflexed, apex rounded, pale yellow. Staminate flowers: stamens 45–50, 4–5 mm long, isodynamous; filaments terete, yellow; anthers c. 1 mm long, yellow, connective not exceeding thecae; pistillode inconspicuous, yellow. Pistillate flowers: staminodes 45–50, 3–3.5 mm long, yellow; ovary c. 4 mm long, not costate or lobed, 4-locular, yellow, styles fused 1.5–2 mm long, stigmas 4, 1.5–2 mm in diam. Capsules fleshy, 3.5– 4 × 1.5–2.5 cm, 4-septate, pyriform when closed, rostrum present, 3–8 mm long, epicarp smooth, green when immature and mature, mesocarp red to purplish-red; sepals, petals and staminodes caducous, stigmas persistent. Aril orange. Fig. 69.

Iconography — Illustration available in Engler (1888: 97).

Distribution — Brazil (Alagoas, Bahia, Espírito Santo, Minas Gerais, Pernambuco). Fig. 70.

Conservation status — Least Concern (LC, Marinho & Beech 2019).

Recognition and discussion — Tovomita riedeliana is similar to T. glazioviana , mainly by the number of secondary veins (16–20 vs 10–17 pairs in T. glazioviana ). They are sympatric in the state of Bahia, Brazil, and can be differentiated by the generally longer leaves in T. riedeliana (to c. 15 cm long vs rarely reaching 13 cm long in young individuals of T. glazioviana ), and pedicel length (13–55 mm vs 6–20 mm long in T. glazioviana ). The relatively large anthers in T. riedeliana (c. 1 mm vs ≤ 0.8 mm long mm long) were reported by Engler (1888) as a striking feature of this species.

Selected specimens examined — BRAZIL: ALAGOAS: Ibateguara, Coimbra / Grota do Dudé , 24 Oct 2001, fr., M . Oliveira & A . A . Grilo 649 ( UEC!, UFP!) . BAHIA: Ilhéus, Estação Hidromineral de Olivença , 8 Feb 1982, fl. ♂, L .

A. Mattos-Silva & al. 1470 ( CEPEC!, UFP!). ESPÍRITO SANTO: Santa Teresa , 25 Feb 2014, fl. ♀ , L. C. Marinho & J. Molino 800 ( CEPEC!). MINAS GERAIS: Rio Novo, Sep 1895, bud, Schwacke 11805 ( RB!). PERNAMBUCO: Gravatá, Fazenda Harmonia , 6 Sep 1970, bud, fl. ♂, D. Andrade-Lima 70/6005 ( CEPEC!, IPA!, UFP!) .