Tovomita grandis L. Marinho

25. Tovomita grandis L. Marinho View in CoL in Willdenowia 49: 346. 2019. – Holotype: Peru, Loreto, Província Requena , swampy forest along creeks, dominated by palms (aguajal) and forest on sandy soil, 140 m, 11 Aug 1988, H. van der Werff & al. 10104 (NY [ NY02859715 ]!; isotype: MO [no. 3705235] photo!).

Description — Trees up to 18 m tall, prop roots not seen; exudate yellowish, abundant. Petioles 4.6–5 cm long, coloration not seen, smooth, lenticels absent. Leaf blades 29.5–35.5 × 12.5–14.3 cm, dark brown in sicco, black dots absent, coriaceous, oblanceolate to obovate, base decurrent, apex rounded; papillae, lenticels and fungal spots absent; exudate canals inconspicuous. Venation: secondary veins 8–9 pairs, 20–45 mm apart from each other, forming angle 30°–45° to midvein, prominent in both surfaces, slightly arcuate near margin; intersecondary veins usually absent, rarely present, one per intercostal area, thinner than secondary veins, distal course reticulating, <50 % of subjacent secondary length; tertiary veins percurrent, perpendicular; intramarginal vein absent. Inflorescences: ♂ lax cyme with 7 basal branches and up to 94 flowers, lacking terminal flower, ♀ not seen, lenticels absent. Pedicels 8–12 mm long, coloration not seen, distal portion slightly dilated, proximally articulat- ed in all flowers of dichasia; calyptrae and lenticels absent. Floral buds 8–12 mm long, oblong, apex rounded, lenticels absent. Sepals 4, 8–12 × 4–5 mm, oblong, apex rounded, greenish to white; petals 4, c. 16 × 6 mm, oblong, reflexed, apex rounded, white. Staminate flowers: stamens 45–50, 10–12 mm long, heterodynamous; filaments terete, white; anthers 1–1.1 mm long, connective not exceeding thecae; pistillode 1.2 mm long, conic. Pistillate flowers and capsules not seen.

Iconography — Illustration available in Marinho & al. (2019b: 345).

Distribution — Peru (Loreto). Fig. 44.

Conservation status — Data Deficient (DD, Marinho & al. 2019b).

Recognition and discussion — Tovomita grandis differs from T. longifolia and T. megantha by the angle of the secondary veins (30°–45° in T. grandis vs> 45° in T. longifolia and T. megantha ), intersecondary veins usually absent (vs present in T. longifolia and T. megantha ), and number of stamens (45–50 in T. grandis vs> 55 in T. longifolia and T. megantha ). Tovomita grandis is also similar to T. speciosa , due to overall leaf morphology and robust inflorescences, but T. speciosa has more than 11 pairs of secondary veins (vs up to 9 pairs in T. grandis ), and spheroid floral buds (vs oblong in T. grandis ).