Tovomita duckei Huber

17. Tovomita duckei Huber View in CoL in Bull. Soc. Bot. Genève 6: 189. 1915. – Lectotype (designated here) or perhaps holotype: Brazil, [Pará, Óbidos], hab. in silvis ad fl. Cumina-mirim , 16 Dec 1906, A. Ducke 7977 (RB [ RB00539002 ]!).

= Tovomita micrantha A. C. Sm. in Phytologia   GoogleMaps 1: 123. 1935, syn. nov. – Lectotype (designated here): Brazil, [Amazonas], terra firme near mouth of Rio Embira   GoogleMaps (tributary Rio Tarauaca), basin of Rio Jurua, [07°30'S, 70°15'W], 28 Jun 1933, B. A. Krukoff 5071 (NY

[ NY00578991 ]!; isolectotypes : A [ A00067913 ] !, G [ G00355500 ] photo !, G photo!).

Description — Trees up to 15 m tall, prop roots not seen; exudate yellow, abundant. Petioles 0.8–2.7 cm long, coloration not seen, smooth, lenticels absent. Leaf blades 10–22 × 4.5–12.3 cm, brown-orangish in sicco, black dots absent, coriaceous, oblong to ovate, base decurrent, apex acuminate; papillae, lenticels and fungal spots absent; exudate canals inconspicuous. Venation: secondary veins 10–15 pairs, 6–15 mm apart from each other, forming angle 50°–55° to midvein, prominent abaxially, immersed adaxially, arcuate near margin; intersecondary veins present, one or two per intercostal area, much thinner than secondary veins, parallel to major secondary,> 50 % of subjacent secondary length; tertiary veins percurrent sinuous; intramarginal vein absent. Inflorescences: ♂ lax cyme with 5 basal branches and up to 97 flowers, with terminal flower, ♀ cyme with 3 basal branches and up to 9 flowers, lenticels absent. Pedicels 4–5 mm long, green, distal and proximal portion with same gauge, proximally articulated on lateral flowers of dichasia; calyptrae and lenticels absent. Floral buds 3–4 mm long, spheroid, apex rounded to apiculate, lenticels absent. Sepals 4, 3–4.5 × 3–4 mm, ovate to oblong, apex rounded, greenish to pale yellow; petals 4, 2.8–3 × 1.5–1.8 mm, oblong, apex rounded, position not seen, greenish. Staminate flowers: stamens 25– 30, 1.3–3.5 mm long, heterodynamous; filaments terete, white; anthers 0.3–0.5 mm long, connective not exceeding thecae; pistillode inconspicuous. Pistillate flowers not seen. Capsules fleshy, 3–4.3 × 0.9–2.6 cm, 4-septate, pyriform when closed, not costate or lobed, rostrum absent, free styles c. 3 mm long, epicarp smooth, slightly verruculose, coloration not seen; sepals, petals, staminodes and stigmas persistent. Aril coloration not seen.

Distribution — Bolivia (La Paz), Brazil (Acre, Amazonas, Pará), Colombia (Vaupés), Peru (Loreto). Fig. 30.

Conservation status — Data Deficient (DD, Marinho & Beech 2019).

Nomenclatural notes — To designate the lectotype, we followed the same reasoning adopted for T. albiflora and T. caloneura , both of which were described by A. C. Smith.

Recognition and discussion — Tovomita duckei has the arrangement of secondary veins similar to that of T. guianensis ; however, T. duckei has wider (4.5–12.3 cm), brown-orangish leaves in sicco (vs 1.8–7.5 cm wide, dark brown leaves in T. guianensis ). The floral buds of T. duckei are small (3–4 mm long), spheroid, and with rounded apex, comparable to those of T. gracilipes , T. guianensis , and T. schomburgkii , but it can be distinguished from all of those species by the brown-orangish leaves in sicco.

Specimens examined — BOLIVIA: LA PAZ: Province Larecaja, Tuiri, near Mapiri , on left bank of Río Mapiri , 490–750 m, 12–30 Sep 1939, fr., B . A . Krukoff 10950 ( A!, NY!, UC!); ibid., Copacabana (about 10 km south of Mapiri ), 850–950 m, 8 Oct–15 Nov 1939, fr., B . A . Krukoff 11043 ( K!, NY!, UC!) . — BRAZIL: ACRE: Brasileia, estrada para Assis Brasil km 20, 3 Nov 1980, bud, C . A. Cid & al. 3119 ( INPA!); Sena Madureira, estrada Manuel Urbano, Rio Caeté , 10 Sep 1978, bud, J . Lima & al. 162 ( INPA!); ibid., J . Ramos & al. 165 ( INPA!); ibid., Seringal Camaru, Rio Caeté , 18 Sep 1978, bud, J . Ramos & al. 629 (INPA!); AMAZONAS: regição do Rio Madeira, Rio Canumã , 5 Nov 1957, bud, R . L . Fróes 33765 ( IAN!) . — COLOMBIA: VAUPÉS: Taraira, Estación Biológica Caparú , 01°04'36"S, 69°30'54"W, 100 m, 17 Jan 2008, fr., A GoogleMaps . Cano 58 ( COAH!) . — PERU: LORETO: Prov. Mariscal Ramón Castilla , 02°51'53.5"S, 71°24'54.1"W, 140 m, M GoogleMaps . A GoogleMaps . Ríos & al. 523 ( F photo!) .