Tovomita foldatsii Cuello

20. Tovomita foldatsii Cuello View in CoL in Novon 9: 151. 1999. –

Holotype: Venezuela, Amazonas, Atures, bosque alto de tierra firme en el sector “Campamento Remo”, alto Río

Sinapo, 04°34'N, 67°18'W,

80–150 m, Jun 1989, E .

Foldats & J. Velázco 9577

( PORT [ PORT 46935] photo!;

isotype: MO [ MO05090713 ]

photo!).

Description — Trees up to 16

m tall, prop roots inconspicuous; exudate yellow, abun- Fig. 33. Distribution map of Tovomita crassidactyla (black circles) and T. xanthochlora (black stars).

dant. Petioles 2–3.5 cm long,

coloration not seen, smooth, Conservation status — Least Concern ( LC, Marinho & lenticels absent. Leaf blades 15–31 × 9.5–14 cm, brown- Beech 2019).

orangish in both surfaces in sicco, black dots present abaxially, coriaceous, broadly elliptic to ovate, base con- Recognition and discussion — This species was devex to decurrent, apex acuminate, sometimes with drip scribed based on fruiting specimens, and the flowers are tip; papillae, lenticels and fungal spots absent; exudate described here for the first time. Tovomita foldatsii recanals inconspicuous. Venation: secondary veins 11–16 sembles T. macrophylla vegetatively, but it can be difpairs, 10–24 mm apart from each other, forming angle ferentiated by the 5-carpellate (vs 4-carpellate) flowers, 65°–85° to midvein, prominent abaxially, immersed smooth (vs asperous) epicarp and fruits bearing a long adaxially, arcuate near margin; intersecondary veins (2–3.8 cm long) rostrum ( Fig. 35A) (vs absent). The represent, one or two per intercostal area, much thinner semblance of these two species is so evident that specithan secondary veins, distal course reticulating, c. 50 % mens of T. foldatsii were erroneously associated with the of subjacent secondary length; tertiary veins percurrent type collection of T. macrophylla (see discussion under T. sinuous; intramarginal vein absent. Inflorescences: ♂ lax macrophylla ). Tovomita foldatsii is named was “ Tovomicyme with 5 basal branches and up to 39 flowers, with ta sp. B ” in Cuello (1998).

a terminal flower, ♀ 5-flowered cyme, lenticels absent.

Pedicels 18–25 mm long, green, distal and proximal por- Selected specimens examined — BRAZIL: AMAZOtion with same gauge, proximally articulated on lateral NAS: Rio Negro, Rio Cueiras, Igarapé da Cachoeira , flowers of dichasia; calyptrae and lenticels absent. Flo- 22 Jun 1992, fr., S . Mori & C. Gracie 22454 ( CAS!, ral buds 8–15 mm long, ovoid, apex apiculate, lenticels NY!); RONDÔNIA: Basin of Rio Madeira, 1 km north absent. Sepals 4, 8–15 × 4.5–6.5 mm, ovate to oblong, of Ribeirão, road Abuña-Guajará-Mirim, 25 Jul 1963, apex rounded to apiculate, yellowish; petals 4, 12–16 × fr., G. T . Prance & al. 6449 ( INPA!). — COLOMBIA: 4–11 mm, oblong to lanceolate, reflexed, apex rounded, CAUCA: Piamonte, Vereda La Consolata , 01°01'24"N, yellowish. Staminate flowers: stamens with 50–60 sta- 76°07'35"W, 294 m, 4 Dec 2019, fr., L GoogleMaps . Rivera & al. mens, 5–8 mm long, isodynamous; filaments terete, 806 ( COAH!). CAQUETÁ: Río Caquetá , 01°15'03"S, yellowish; anthers c. 0.6 mm long, connective not seen; 69°44'35"W, 542 m, 6 Jun 2014, fr., S. B GoogleMaps . Correa 394 pistillode not seen. Pistillate flowers: staminodes 50–60, ( COAH!). VAUPÉS: Taraira, Estación Biológica Mosiro 5–6.5 mm long, yellowish; ovary 6–7 mm long, not cos- Itajura , Caparú , 01°04'48"S, 69°30'15"W, 18 Feb 2004, tate or lobed, 5-locular, yellowish, styles free 4–5 mm L GoogleMaps . Clavijo & Z . Cordero 387 ( HUA!). — VENEZUELA: long, stigmas 5, 1.5–2 mm in diam. Capsules fleshy, 7.2– AMAZONAS: Autana, Río Sipapo , Caño Gato, 04°54'– 10.2 × 2–4 cm, 5-septate, oblong to ovoid when closed, 05°03'N, 67°34'– 67°46'W, 225 m, 18 Aug 1997, bud, A GoogleMaps . sometimes falcate, not costate or lobed, rostrum present, Castillo 5430 ( M!) .

2–3.8 cm long, usually falcate due to abortion of a seed,