Anaphes ( Anaphes ) maxim Triapitsyn, 2021

Anaphes ( Anaphes) maxim Triapitsyn , sp. n.

http://zoobank.org/NomenclaturalActs/ 5EC4CFD0-7160-4F11-8625-76742D6B985F

Figs 55–60 View Figs 55–60

TYPE MATERIAL. Holotype female [ ZIN] on slide ( Fig. 55 View Figs 55–60 ): Russia: Primorskii krai,

Ussuriyskiy rayon, Gornotayozhnoye, 43.66°N 132.25°E, 200 m, 15–17.VII 2000 (M. V. GoogleMaps

Michailovskaya), MT. Paratypes (all on slides): Russia: Primorskii krai, Ussuriyskiy rayon ,

Gornotayozhnoye, 43.66°N 132.25°E, 200 m (M. V. Michailovskaya): 26–31.VIII 2000, MT GoogleMaps

[ 1 ♀, UCRC]; 15–30.IX 2000, MT [ 1 ♀, IBPV] ; 5–20.X 2002, YPT [ 1 ♀, UCRC]. Sakhalinskaya oblast’, Sakhalin Island, 2–3 km E of Sokol, a tributary of Belaya River , 47°14.385’N

142°46.347’E, 18.VII 2001 ( N. Minakawa), MT , SK-01-DJB-128C [ 1 ♀, CAS] .

DESCRIPTION. FEMALE ( holotype). Body dark brown, appendages brown except scape and pedicel light brown and coxae dark brown. Vertex with mesh-like sculpture, mesoscutum and scutellum with faint sculpture. Antenna ( Fig. 56 View Figs 55–60 ) with scape (excluding radicle) 4.7× as long as wide, almost smooth; F1 longer than wide, F2–F6 much longer than pedicel, F2 4.5×

as long as wide, F3 the longest funicular, F2–F6 each with 2 mps; clava with 6 mps, 3.9× as long as wide, almost as long as combined length of F5 and F6 (just slightly shorter). Fore wing

( Fig. 57 View Figs 55–60 ) 7.2× as long as wide; longest marginal seta 1.3× maximum wing width; marginal space separated from medial space by 2 complete lines of setae. Hind wing ( Fig. 58 View Figs 55–60 ) almost

23× as long as wide; longest marginal seta 3.9× maximum wing width, disc with 1 short row of 3 or 4 setae apically. Metatarsomere 1 about as long as metatarsomere 2 ( Fig. 60 View Figs 55–60 ). Ovipositor

( Fig. 59 View Figs 55–60 ) occupying about 0.6× length of gaster (and thus not extending forward under mesosoma), not exserted beyond apex of gaster posteriorly, and 0.7× length of metatibia.

Measurements (µm) of the holotype. Mesosoma 303; metatibia: 297; gaster 327; ovipositor 206. Scape (minus radicle) 128; pedicel 54; F1 24; F2 82; F3 89; F4 85; F5 82; F6 76;

clava 154. Fore wing 867:121; longest marginal seta 160. Hind wing 824:36; longest marginal seta 139.

Variation ( paratypes). Body length 0.72–0.78 mm (slide-mounted specimens). Antenna with F2 3.5–3.6× as long as wide, F2–F4 often subequal in length and the longest funiculars,

clava about 3.6× as long as wide. Fore wing 0.79–0.92 mm long, 7.0–7.1× as long as wide,

longest marginal seta 1.3–1.4× maximum wing width, marginal space separated from medial space by 1 or 2 lines of setae. Hind wing 20–23× as long as wide, longest marginal seta 3.3–

3.8× maximum wing width, disc with 1 irregular row of 4 to 8 setae apically. Ovipositor

0.7–0.8× length of metatibia (usually 0.7×).

57) fore wing, 58) hind wing, 59) ovipositor, 60) metatibia and metatarsus.

MALE. Unknown.

DIAGNOSIS. Anaphes ( Anaphes) maxim is characterized by the combination of female antenna ( Fig. 56 View Figs 55–60 ) bearing 2 mps on F2, F2–F6 being notably longer than wide and longer than pedicel, and a relatively narrow female fore wing ( Fig. 57 View Figs 55–60 ) which is at least 7.0× as long as wide. This new species is similar to the Nearctic A. ( Anaphes) nigrellus Girault, 1911 , known from Canada and USA (Huber, 1992), from which it differs mainly by the relatively longer

F2–F6 of the female antenna ( Fig. 56 View Figs 55–60 ), a longer female fore wing (at least 0.79 mm long),

and a relatively shorter ovipositor (at most 0.8× as long as the metatibia, usually 0.7×). In the positively identified North American female specimens of A. ( Anaphes) nigrellus , the fore wing is at most 0.78 mm long and the ovipositor is about 0.9× length of the metatibia (Huber,

1992). By its relatively long female fore wing ( 0.79–0.92 mm), A. ( Anaphes) maxim is also similar to the Nearctic A. ( Anaphes) hercules Girault, 1911 , known from Illinois, USA,

whose female fore wing is, however, less than 5.7× as long as wide (at least 7.0× as long as wide in A. ( Anaphes) maxim ), F2 of the female antenna bears only 1 mps, and the ovipositor is a little more than 0.8× length of the metatibia (Huber, 1992).

ETYMOLOGY. The species name, which is treated as a noun in apposition, is the first name of Maxim Yur’evich Proshchalykin, a fellow hymenopterist who kindly helped me with information on the specimens of Anaphes ( Anaphes) nipponicus Kuwayama, 1932 deposited in IBPV.

HOSTS. Unknown.