Fissidens ferrugineus Müll.Hal.

Fissidens ferrugineus Müll.Hal.

( Fig. 5 View FIG )

Botanische Zeitung (Berlin) 22: 341 ( Müller 1864).

— Lectotype (designated here): Madagascar • Pervillé 51A; BM [ BM000871895 ] mixed with the lectotype of F. madecassus Schimp. ex Müll.Hal. ; isolectotype: BM [ BM000871896 ]; two stems mixed between six stems of the isolectotype of F. madecassus .

Fissidens ugandae Dixon & P. de la Varde in P. de la Varde , Annales de Cryptogamie Exotique 3: 45, 1, f. 3. (P. de la Varde 1930).

— Lectotype (designated here): Uganda • Musandama; Budd s.n.; 1926; BM [ BM000871873 ]; isolectotype: MNHN [ PC0095552 ], syn. nov.

Fissidens gibbonii Bizot , Revue bryologique et lichénologique 40 (2): 131 ( Bizot 1973).

— Lectotype (designated byPócs 1977 as holotype): Tanzania • Morogoro District, Uluguru Mts, on top of Bondwa Peak ; on the ground of subalpine heath; 2100 m alt.; 12.X.1969; Pócs & Gibbon 6052 /CR ( EGR; isolectotypes: MNHN [ PC0132125 ] and DSM), syn. nov.

DIAGNOSIS (distinctive features in bold). — Plants often rusty brown, leaves narrow, 3.5-5 (-6) times as long as wide, lanceolate with narrow acute to acute-acuminate tips; limbidium thick yet unistratose ( Fig. 5J View FIG ), smooth to denticulate, marginal, present on the vaginant laminae of all leaves, typically extending the whole length of the vaginant lamina, often confluent at the apex and frequently extending onto the apical lamina; vaginant lamina 2/3-3/4 the leaf length; costa percurrent to excurrent, or ending 1-2 cells below the apex; laminal cells can have 1-2 or 2-4 papillae per cell (rarely 1(-2), or 2-6), punctate, less often irregular ( Fig. 5I View FIG ) papillae; antheridia terminal on short stems (rhizautoicous), less often terminal on longer stems or on short perigonial branches in the axils of lower leaves; setae 3-10 mm long, theca cylindric to strumulose, with 32-58 columns of exothecial cells. Some collections have thecae with ± 32 as well as with 40 or more exothecial cells. Since ±32 columns are typical for subg. Polypodiopsis 32 is not conclusive. When, however, there are 40 or more exothecial cells, this is diagnostic for F. ferrugineus .

OTHER EXAMINED MATERIAL (Specimens used for Fig. 5 View FIG ). — Madagascar • Pervillé 51A; BM [ BM000871895 ], lectotype F. ferrugineus . Tanzania • Meru Mts; in the crater ground near Njeku; swampy moorland patches, rupicolous; 2600 m alt.; Pócs et al. 8670 /D; EGR, L • Kilimanjaro Mts, Karanga River Gorge N of Kibosho; Montane evergreen forest on the steep; rocky E facing slope; with Albizia gummifera (J.F.Gmel.) C.A.Sm. ; on soil covered rock; 1880 m alt.; Pócs, Mialla & Margesi 90068 / G; EGR • Southern Highlands , Kipengere Range, above Salala Falls below Mt Ishinga ; in deep ravine of streamlet, on half shady rock; 2380-2400 m alt.; 26.V.1987; Pócs, Iversen, Persson & Petterson 87146 / O; EGR, private herbarium Bruggeman-Nannenga • Ukaguru Mts in Kilos District, on Mamiwa ridge; mossy elfin forest, rupicolous; 2150 m alt.; Pócs, Chamshama & Sylvander 86100 / T; EGR, L • Ukaguru Mts, NE slope of Mamwira W of Masenge village; on shady rocks in bamboo ( Arundinaria alpina K.Schum. ), thicket; 2050 m alt.; Pócs 88015 / C; EGR .

La Réunion • G. de L’ Isle s.n. ( PC) • Forêt de Bélouve on the Plateau des Cochons, Trou de Fer trail; 21°40’S, 55°32.3’E; 21°40’S, 55°32.4’E; Bruggeman-Nannenga 10995; L GoogleMaps .