Cochlis ukrainensis Pedriali, Sosso & Dell’Angelo, 2019

Cochlis ukrainensis Pedriali, Sosso & Dell’Angelo, 2019 View in CoL

Figs 2F View FIGURE 2 , 18–20 View FIGURE 18 View FIGURE 19 View FIGURE 20

Natica View in CoL ] millepunctata Lmk.— Hauer 1837: 421 View in CoL ( pars) [ non Natica millepunctata Lamarck, 1822 View in CoL = Naticarius stercusmuscarum ( Gmelin, 1791) View in CoL ].

Natica millepunctata Lam.— Hörnes 1848: 23 View in CoL ( pars) [ non Natica millepunctata Lamarck, 1822 View in CoL = Naticarius stercusmuscarum ( Gmelin, 1791) View in CoL ].

Natica millepunctata Lam.—Hörnes 1856: 518 View in CoL ( pars) [ non Natica millepunctata Lamarck, 1822 View in CoL = Naticarius stercusmuscarum ( Gmelin, 1791) View in CoL ].

Natica millepunctata View in CoL — Naumann 1852: pl. 69, fig. 12 [ non Natica millepunctata Lamarck, 1822 View in CoL = Naticarius stercusmuscarum ( Gmelin, 1791) View in CoL ].

Natica millepunctata Lam.— Naumann 1854: 1065 View in CoL [ non Natica millepunctata Lamarck, 1822 View in CoL = Naticarius stercusmuscarum ( Gmelin, 1791) View in CoL ].

Natica millepunctata tigrina (Defr.) View in CoL — Csepreghy-Meznerics 1950: 45, 91, pl. 2, fig. 12 [ non Cochlis tigrina ( Defrance, 1825) View in CoL ].

N [ atica]. ( N [ acca].) millepunctata View in CoL hörnesi Fisch. et Tourn.— Sieber 1958: 140 [ non Cochlis hoernesi ( Fischer & Tournouër, 1873) ].

N [ atica]. ( N [ acca].) millepunctata tigrina Defr.— Sieber 1958: 140 [ non Cochlis tigrina ( Defrance, 1825) ].

Natica millepunctata Lamarck, 1822 View in CoL — Nicorici 1972: 139, pl. 14, figs 1–2 [ non Natica millepunctata Lamarck, 1822 View in CoL = Naticarius stercusmuscarum ( Gmelin, 1791) View in CoL ].

Natica tigrina Defrance, 1825 View in CoL — Švagrovský 1982: 31, pl. 10, fig. 1 [ non Cochlis tigrina ( Defrance, 1825) View in CoL ].

Natica ( Naticarius) millepunctata hoernesi Fischer & Tournouer [ sic]— Schultz 1998: 198, pl. 22, fig. 12 [ non Cochlis hoernesi ( Fischer & Tournouër, 1873) ].

* Cochlis ukrainensis View in CoL sp. nov. — Pedriali et al. 2019: 166 View Cited Treatment , figs 24.1−6.

Type material. Holotype, MPUM 11735, SL: 35.4 mm, MD: 35.7 mm, Varivtsi ( Ukraine), illustrated in Pedriali et al. (2019: 24.1a–c), Figs 18A View FIGURE 18 1 –A View FIGURE 1 4 View FIGURE 4 . Paratypes: MPUM 11736, SL: 21.3 mm, MD: 20.7 mm, Varivtsi ( Ukraine), illustrated in Pedriali et al. (2019: 24.2a–b), Figs 18B View FIGURE 18 1 –B View FIGURE 1 4 View FIGURE 4 . For further paratypes and material see Pedriali et al. (2019: 166).

Illustrated material. NHMW 1855/0045/0226c, SL: 30.1 mm, MD: 29.9 mm, Steinebrunn ( Austria), Figs18C View FIGURE 18 1 – C View FIGURE 1 4 View FIGURE 4 . NHMW 1855/0045/0226a, SL: 32.6 mm, MD: 33.4 mm, Steinebrunn ( Austria), Figs 18D View FIGURE 18 1 –D View FIGURE 1 4 View FIGURE 4 . NHMW 2024/0238/0001, SL: 37.5 mm, MD: 35.1 mm, Steinebrunn ( Austria), Figs 19A View FIGURE 19 1 –A View FIGURE 1 4 View FIGURE 4 . NHMW 2024/0238/0002, SL: 31.4 mm, MD: 31.1 mm, Steinebrunn ( Austria), Figs 19B View FIGURE 19 1 –B View FIGURE 1 4 View FIGURE 4 . NHMW 2024/0235/0002, SL: 35.1 mm, MD: 37.5 mm, Gainfarn ( Austria), Figs 19C View FIGURE 19 1 –C View FIGURE 1 4 View FIGURE 4 . NHMW 2024/0235/0003, SL: 25.0 mm, MD: 24.1 mm, Gainfarn ( Austria), Figs 19D View FIGURE 19 1 –D View FIGURE 1 4 View FIGURE 4 . PIN 5904/121, SL: 32.5 mm, MD: 33.5 mm, Shushkivtsi, Ukraine, Middle Miocene, late Badenian, Figs 20A View FIGURE 20 1 –A View FIGURE 1 4 View FIGURE 4 . PIN 5904/122, SL: 20.4 mm, MD: 20.8 mm, Gorodok, Ukraine, Middle Miocene, late Badenian, Figs 20B View FIGURE 20 1 –B View FIGURE 1 4 View FIGURE 4 . PIN 5904/123, SL: 29.4 mm, MD: 28.8 mm, ravine Zhabyak, Ukraine, Middle Miocene, late Badenian, Figs 20C View FIGURE 20 1 –C View FIGURE 1 4 View FIGURE 4 . PIN 5904/124, SL: 20.1 mm, MD: 19.4 mm, Varivtsi, Ukraine, Middle Miocene, late Badenian, Figs 20D View FIGURE 20 1 –D View FIGURE 1 4 View FIGURE 4 .

Additional material. 8 spec., NHMW 2024 View Materials /0235/0004, Gainfarn ( Austria) ; 28 spec., NHMW 1855 View Materials /0045/0225, Steinebrunn ( Austria) ; 21 spec., NHMW 1997 View Materials z0178/1372, Gainfarn ( Austria) ; 12 spec., NHMW 2013 View Materials /0479/1005, Gainfarn ( Austria) ; 12 spec., NHMW 1846 View Materials /0037/0442, Gainfarn and Enzesfeld ( Austria) ; 6 spec., NHMW 1855 View Materials /0002/0053, Enzesfeld ( Austria) ; 11 spec., NHMW 1851 View Materials /0010/0061, Mikulov ( Czech Republic) .

Revised description. Medium sized to large, globose, solid shell of about four teleoconch whorls; often only slightly higher than wide ( SL /MD = ~1.0). Spire low-conical, slightly elevated to rather depressed, with convex whorls; apical angle 120–135°. Protoconch small, depressed turbiniform, of 1.75 convex whorls. Suture linear, strongly incised. Last teleoconch whorl with narrow, weakly convex subsutural shelf, delimited by faint to subobsolete shoulder angulation. Last whorl moderately expanding, slightly inflated, attaining ~92–95% of total height. Periphery regularly convex, placed mid-whorl (~49%). Growth lines distinct and weakly prosocyrt in apical view, weaker to subobsolete at periphery. Aperture moderately wide, D-shaped. Position of adapical tip of aperture low (~75%), distinctly above mid-whorl, far below shoulder. Aperture attaining ~80–85% of total height. Columellar lip thick, basal lip slightly expanded, outer lip thin. Columellar angle ~16–32°. Opercular ridge in outer lip distinct. Umbilicus moderately wide (~16%), deep, with prominent growth lines. Parietal callus long (~53%), weakly thickened not expanding over base, with straight to weakly concave margin. Anterior lobe small, rounded, flat. Funicle broad, convex, semicircular, not very raised, moderately delimited. Adapical and abapical sulci wide, weakly concave. Umbilical callus small, thick, semicircular, well-separated from parietal callus. Basal fasciole solid, wide, rounded with prominent growth lines. Color pattern of uniform light brown to brown background, with uneven whitish collabral stripes (according to Pedriali et al. 2019: 166). Operculum unknown.

Discussion. Specimens with depressed spire and spherical outline ( Fig. 18 View FIGURE 18 ) co-occur with shells with more elevated spire and more elongate outline ( Fig. 19 View FIGURE 19 ). These morphs are united by transitional morphs. There are also variations in the strength of the funicle. Neither form has color pattern preserved. Only opercula would help to decide if these are two cryptic species. Therefore, we provisionally consider these morphologies to reflect the intraspecific variability of Cochlis ukrainensis Pedriali, Sosso & Dell’Angelo, 2019 . Pedriali et al. (2019) introduced this species based on specimens from the upper Badenian of Ukraine but did not refer to any other Paratethyan occurrences. Nevertheless, it is probably one of the most frequent Naticidae in shallow marine deposits of the Central Paratethys Sea, although it has been confused by most authors with Cochlis tigrina ( Defrance, 1825) from the Early Miocene of France. They differ in the narrower umbilicus of the French species, which has a deep umbilical channel surrounding the funicle (see Robba et al. 2016: pl. 4, figs 1–5). Cochlis raropunctata (Sasso, 1827) , from the Tortonian of Italy, differs in in much more prominent, strongly raised funicle (see Robba et al. 2016: pl. 3, figs 2–4). See Pedriali et al. (2019: 168) for further discussions.

Paleoenvironment. Inner neritic, shallow marine; occurrences at Gainfarn and Steinebrunn suggest sea-grass environments (Zuschin et al. 2007).

Distribution in Central Paratethys. Badenian (Middle Miocene): Ukrainian Fore-Carpathian Basin: Gorodok, Stara Pisochna, Varivtsi, Velyka Levada (Khmel’nytskyi Region), Rydomyl’, Shushkivtsi, ravine Zhabyak near Dzvynyacha ( Ternopil Region, Ukraine) ( Pedriali et al. 2019, hoc opus). Vienna Basin: Enzesfeld, Gainfarn, Steinebrunn ( Austria), ( Austria), Mikulov ( Czech Republic) ( hoc opus); Borský Mikuláš ( Slovakia) ( Švagrovský 1982). Pannonian Basin: Hidas ( Hungary) ( Csepreghy-Meznerics 1950); Şimleu Basin: Tusa ( Romania) ( Nicorici 1972); Moldavian Platform: Bursuc ( Moldova) ( hoc opus).